305 lines
36 KiB
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305 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.73.e102845" ID-Pensoft-Pub="2511-6428-1-9" ID-Pensoft-UUID="88A9F45828C552B281F9DB387032DC7E" ID-ZooBank="118596DA8D9C4569B6B1B004FE2AB136" ModsDocID="2511-6428-73-1-9" checkinTime="1687815002572" checkinUser="pensoft" docAuthor="Liston, Andrew" docDate="2023" docId="83859A1E333D5CEBBC574EA09ED35257" docLanguage="en" docName="ContribEntomol 73(1): 9-30" docOrigin="Contributions to Entomology 73 (1)" docPubDate="2023-06-26" docSource="http://dx.doi.org/10.3897/contrib.entomol.73.e102845" docTitle="Pamphilius mediterraneus Liston 2023, sp. nov." docType="treatment" docUuid="0B4CEBA2-A868-4C5D-977D-8ED9528B4474" docUuidSource="ZooBank" docVersion="1" id="88A9F45828C552B281F9DB387032DC7E" lastPageNumber="9" masterDocId="88A9F45828C552B281F9DB387032DC7E" masterDocTitle="Taxonomy, distribution and host plants of some southern European and North African Sawflies (Hymenoptera, Symphyta)" masterLastPageNumber="30" masterPageNumber="9" pageNumber="9" updateTime="1687815002572" updateUser="pensoft">
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<mods:titleInfo id="4D4E029CD9653026A486E5DAD1E4BD08">
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<mods:title id="2F9500E0962D603EB90255A245A98057">Taxonomy, distribution and host plants of some southern European and North African Sawflies (Hymenoptera, Symphyta)</mods:title>
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<mods:namePart id="56A191E0F6A07183EC2C93514137C4E6">Liston, Andrew</mods:namePart>
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<mods:affiliation id="7601106A098CF2E26850F0377CA58E27">Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany</mods:affiliation>
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<mods:nameIdentifier id="01BE85850185C0CF1B3F88A9B5F05142" type="email">andrew.liston@senckenberg.de</mods:nameIdentifier>
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<mods:title id="099FAF84D4617E8F005D3E522A026BED">Contributions to Entomology</mods:title>
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<mods:part id="1C1D7B7F9637D9222D4E0396F5304289">
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<mods:date id="CBF620049A327AB570222A12A4A47E13">2023</mods:date>
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<mods:number id="BE958297008D1437A1E5A1EC66C77D28">2023-06-26</mods:number>
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<mods:identifier id="508704BC3947A4A59E7CB39ED9C6357C" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.73.e102845</mods:identifier>
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<treatment id="83859A1E333D5CEBBC574EA09ED35257" LSID="urn:lsid:zoobank.org:act:0B4CEBA2-A868-4C5D-977D-8ED9528B4474" httpUri="http://treatment.plazi.org/id/83859A1E333D5CEBBC574EA09ED35257" lastPageNumber="9" pageId="0" pageNumber="9">
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<subSubSection id="E7F08A99475EC0F7851817FB7AAF0DD1" pageId="0" pageNumber="9" type="nomenclature">
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<paragraph id="483CB7B498365366DF3724C95D96C7F2" pageId="0" pageNumber="9">
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<taxonomicName id="279E5163B2AFD25BFA9ECF70728AF3B7" LSID="https://zoobank.org/0B4CEBA2-A868-4C5D-977D-8ED9528B4474" authority="Liston, 2023" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius mediterraneus" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="mediterraneus" status="sp. nov.">Pamphilius mediterraneus</taxonomicName>
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<taxonomicNameLabel id="A34129EB1BBF9983112707A1A549D1CA" pageId="0" pageNumber="9">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5D6F80F58DD74EECB7654BAB431E935C" pageId="0" pageNumber="9" type="description">
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<paragraph id="53B1AF79BAAB3FF6B4F18626BF40EBDF" pageId="0" pageNumber="9">
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<figureCitation id="24ADD9E648E0D93FD8F63A7D57629743" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="28D340858EF8FE023C2256A15DB117F8" pageId="0" pageNumber="9" type="reference_group">
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<paragraph id="6ACCB8800A46DA04BBC4773AF5CBDC46" pageId="0" pageNumber="9">
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<taxonomicName id="3EB5C3D0F345ABE45CC6C8961AC50B8F" authority="Liston, 2023" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius ignymontiensis" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="ignymontiensis" status="sp. nov.">Pamphilius ignymontiensis</taxonomicName>
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:
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<bibRefCitation id="49E6735A2F98B996B930705406692FFE" author="Liston, AD" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="0" pageNumber="9" pagination="29 - 65" refId="B49" refString="Liston, AD, Jacobs, H-J, Turrisi, GF, 2013. New data on the sawfly fauna of Sicily (Hymenoptera, Symphyta: Xyeloidea, Tenthredinoidea, Pamphilioidea, Cephoidea, Orussoidea). The Entomologist's Monthly Magazine 149 (1): 29 - 65" title="New data on the sawfly fauna of Sicily (Hymenoptera, Symphyta: Xyeloidea, Tenthredinoidea, Pamphilioidea, Cephoidea, Orussoidea)." volume="149" year="2013">Liston et al. 2013</bibRefCitation>
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, misidentification.
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</paragraph>
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</subSubSection>
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<subSubSection id="6355052DBAD6A805A9809CB4FA7CE428" pageId="0" pageNumber="9" type="type locality">
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<paragraph id="12D8339E1277B8D20A04E9102DED5252" pageId="0" pageNumber="9">Type locality.</paragraph>
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<paragraph id="963D2633D8B8AFFB1D915DBF6AF564E5" pageId="0" pageNumber="9">
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Italy, Sicily, 3 km east of Gratteri, approximately 800 m above sea-level,
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<geoCoordinate id="B1D812B1B620AF8BB4D5953EEED5D16B" degrees="37.968" direction="north" orientation="latitude" precision="55" value="37.968">37.968°N</geoCoordinate>
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,
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<geoCoordinate id="F0D545B02A9E57B7E496C61550094266" degrees="14.001" direction="east" orientation="longitude" precision="55" value="14.001">14.001°E</geoCoordinate>
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[the coordinates on the data label are rounded, whereas these are more precise: uncertainty 150 m. The coordinates on the label locate a spot on the other side of the public road, in an area with significantly less tree cover]. Mixed broadleaved woodland. Collection data: 21.05.2010, by sweeping, A. Liston leg.
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</paragraph>
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</subSubSection>
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<subSubSection id="114DCDB4970255469D2BF14F71386B24" pageId="0" pageNumber="9" type="type specimen">
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<paragraph id="DDDDFEB3DFA36E0DFBD4DBC606CD7084" pageId="0" pageNumber="9">Type specimen.</paragraph>
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<paragraph id="16BD2258EF600F8873BEF6DD9B900470" pageId="0" pageNumber="9">
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<emphasis id="CE9951190CBC0676BD9C335B1ACFDC9F" bold="true" italics="true" pageId="0" pageNumber="9">Holotype</emphasis>
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♀ (DEI-GISHym11123). Labels: "SICILY: 21.05.2010 Gratteri ca. 3 km NE (ca. 800 m) (
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<geoCoordinate id="FA7883FC6ECE56163578BF767A28A55E" degrees="37,97" direction="north" orientation="latitude" precision="555" value="37.97">37,97°N</geoCoordinate>
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<geoCoordinate id="9B77557A3CB6F10011A7AAFCCC545997" degrees="14,00" direction="east" orientation="longitude" precision="555" value="14.0">14,00°E</geoCoordinate>
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) leg. A. Liston",
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<normalizedToken id="7461F9D54803AD07E25081E06D1D9A45" originalValue="“DEI-GISHym11123”">"DEI-GISHym11123"</normalizedToken>
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, "
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<taxonomicName id="E86DD17AFFACA3711DDB742BCFBFE02C" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius ignymontiensis" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="ignymontiensis">Pamphilius ignymontiensis</taxonomicName>
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Lacourt det. A. Liston 2010", "DEI-GISHym 11123 ii/2013
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<taxonomicName id="F18E3947CB6A0378ED8DB103E5FBDD8E" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius ignymontiensis" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="ignymontiensis">Pamphilius ignymontiensis</taxonomicName>
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BASYM466-11 COI: 658 [0n] BIN:ABA1652" [blue], "
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<taxonomicName id="FBE9A97FF8F20C68DF6A933345011B3E" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius mediterraneus" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="mediterraneus">Pamphilius mediterraneus</taxonomicName>
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sp. n. ♀ det. A. Liston 2023" [red], deposited in SDEI.
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</paragraph>
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</subSubSection>
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<subSubSection id="2A7D3554691909A2EF2F7F2735FA503D" pageId="0" pageNumber="9" type="description">
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<paragraph id="389C4C9418F9634D72A72918AB74D680" pageId="0" pageNumber="9">Description.</paragraph>
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<paragraph id="B68A255A05CB7B0AFB36FA20AC759D0B" pageId="0" pageNumber="9">
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Head black (Fig.
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<figureCitation id="9F8E8D24E58F0F935C61DB10C4963F64" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1C, D</figureCitation>
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): pale are palpi, mandibles, clypeus, gena (partly), most of frons (but divided by a dark vertical stripe medially), a small oblique fleck on rear of postocular area (Fig.
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<figureCitation id="CC5223F32CE933EE5CCD4C66B5243111" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1B, C, E</figureCitation>
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). Antenna (Fig.
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<figureCitation id="71992C57FBA1FC831C73A86C0A025C33" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1B-E</figureCitation>
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): scape pale, pedicel slightly darker, flagellum progressively darkened towards apex. Thorax black (Fig.
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<figureCitation id="17B6135CA879892BCEDA44C76C073162" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A, B</figureCitation>
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): pale (whitish) are tegula, mesoscutellum (without appendage), metascutellum, upper edges and lowermost part of pronotum (Fig.
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<figureCitation id="855DDA29334A740FFD2B2FCA2611943A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A, B</figureCitation>
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). Legs pale yellowish (Fig.
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<figureCitation id="FCD93A5802B90E7C6C28A94114164562" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A, B</figureCitation>
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) except for black bases of coxae (Fig.
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<figureCitation id="56CAB54EF8F41DF57272019DFDEB07F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1F</figureCitation>
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). Veins of forewing pale to about half length of wing from base, apically dark; pterostigma yellow brown, anterior slightly darker (Fig.
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<figureCitation id="F9E0BE231951FD2D03873DA7318BC5AB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A</figureCitation>
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). Abdomen (Fig.
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<figureCitation id="6B46D4DEAEB71DBAF477B82C0B1E61F2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A, F</figureCitation>
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): tergum 1 mainly black except for very narrow white antero-lateral margin. Terga 2-5 entirely reddish-yellow. Tergum 6 largely black; anteriorly and laterally partly reddish-yellow. Tergum 7 black with small, pale antero-lateral fleck. Tergum 8 black. Tergum 10 basally dark, apically pale. Sterna pale except for 7 with small triangular baso-medial black marking (Fig.
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<figureCitation id="37B4B9142129E89E073CFE3EFB03C218" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1F</figureCitation>
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). Cerci (Fig.
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<figureCitation id="414ED1326EE5E4FC1A1F4E078497CD26" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1F</figureCitation>
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) and sawsheath pale, except for dark setae and peg on valvula 3 (Fig.
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<figureCitation id="12CC82F98BE5DD9C70B27345A28CCBD5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1G</figureCitation>
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).
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</paragraph>
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<caption id="A4F7001D7FFF498E316D427B3793B08A" doi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9" start="Figure 1" startId="F1">
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<paragraph id="B4A080F9C0B7770F6C3437EBF7FC9E07" pageId="0" pageNumber="9">
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<emphasis id="FFEB5F7B10161AF9AA5E071F34737B0F" bold="true" pageId="0" pageNumber="9">Figure 1.</emphasis>
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<taxonomicName id="C4FF5115F998B8BA7ADF9946A7BF11EA" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius mediterraneus" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="mediterraneus">
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<emphasis id="992C0C0FD48D46780A46639122B8939B" italics="true" pageId="0" pageNumber="9">Pamphilius mediterraneus</emphasis>
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</taxonomicName>
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sp. nov. holotype ♀ (DEI-GISHym11123):
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<emphasis id="AE4F3D4AB6676C13F3752B4DC0096101" bold="true" pageId="0" pageNumber="9">A, B.</emphasis>
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Complete;
|
||
<emphasis id="2E2D22B55D5AADBBFE9D5FB69AB506B8" bold="true" pageId="0" pageNumber="9">C-E.</emphasis>
|
||
Head;
|
||
<emphasis id="ED76968E05AC5B706ECBC08E9A548B04" bold="true" pageId="0" pageNumber="9">F.</emphasis>
|
||
Abdomen ventral;
|
||
<emphasis id="B283CC025D9B241CB79B5BA3EAD7B674" bold="true" pageId="0" pageNumber="9">G.</emphasis>
|
||
Valvula 3 lateral, peg arrowed. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="88D54E065A5812E9628A327DD7E047C7" pageId="0" pageNumber="9">
|
||
Upper head (Fig.
|
||
<figureCitation id="BF180754F8E98C786944B7947DC755F3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1C</figureCitation>
|
||
) largely glabrous; length and density of setae increasing towards the outer orbits; punctures shallow and very widely spaced; frontal crest moderately protruding, in dorsal view subtriangular; facial crests weakly protruding to about half height of frontal crest. Head below transverse sutures (Fig.
|
||
<figureCitation id="FEDF5C652450B69009C28AC5B0D40CAD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1D</figureCitation>
|
||
) moderately setose with diffuse shallow punctation, except for glabrous and impunctate paraantennal field; longest setae slightly longer than diameter of an ocellus; face below lateral transverse sutures slightly transversely corrugated down to level of lower edge of ocellar basin; clypeus densely punctate; frontoclypeal crest scarcely developed. Antenna (Fig.
|
||
<figureCitation id="8E5828B7595E0596B58DD0EB794B9A65" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1E</figureCitation>
|
||
): 21 antennomeres, scape 3.8
|
||
<normalizedToken id="994F337AAC03EEC7AA6D623B329D1695" originalValue="×">x</normalizedToken>
|
||
as long as apical width, 2.55
|
||
<normalizedToken id="5C61C8481AF6C7E885867A915ABA4EF6" originalValue="×">x</normalizedToken>
|
||
as long as pedicel; flagellomere 1 about 1.1
|
||
<normalizedToken id="86B738C82FD81C1C958BD4C4251EEF0A" originalValue="×">x</normalizedToken>
|
||
as long as scape, 2.3
|
||
<normalizedToken id="049CBCA369DBBB6A4CC6A19BD9732F99" originalValue="×">x</normalizedToken>
|
||
as long as flagellomere 2. Fore wing cell C entirely setose. Medial mesoscutal lobe glabrous, without punctation; lateral lobes medially glabrous and impunctate; moderately setose and weakly punctate laterally (Fig.
|
||
<figureCitation id="E0638FE86F12FC7BD5BC55CCF52F4BE9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1A</figureCitation>
|
||
). Mesepisternum (Fig.
|
||
<figureCitation id="F3C85A6D406864DCFD75A2C3EADB5F3E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1B</figureCitation>
|
||
) sparsely and shallowly punctate with adpressed setae about as long as diameter of ocellus; narrow glabrous impunctate stripe on lowest third. Inner tooth of claw slightly shorter than outer. Valvula 3 (Fig.
|
||
<figureCitation id="A67004FBD2F66690CD1057CFD9D80C4A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Pamphilius mediterraneus sp. nov. holotype ♀ (DEI-GISHym 11123): A, B. Complete; C-E. Head; F. Abdomen ventral; G. Valvula 3 lateral, peg arrowed. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.73.e102845.figure1" httpUri="https://binary.pensoft.net/fig/868214" pageId="0" pageNumber="9">1G</figureCitation>
|
||
) with dense apical fringe of dark setae; valvular peg conical, dark like setae, markedly shorter than longest setae. Terga 1-5 sculptured; sculpture less strong towards posterior of abdomen; distal terga nearly unsculptured.
|
||
</paragraph>
|
||
<paragraph id="0DDDC0D6FBBEEEE4381F5FEEE1879F6B" pageId="0" pageNumber="9">Length: 8.5 mm.</paragraph>
|
||
<paragraph id="1A4F62A37210C92818FD0CF967924717" pageId="0" pageNumber="9">
|
||
<emphasis id="464AA486B66DA36CFCA9C25E02376316" bold="true" pageId="0" pageNumber="9">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D86E2F8731B3FB81DA05996CCCD2B902" pageId="0" pageNumber="9" type="etymology">
|
||
<paragraph id="49D3979A77169993F7CC5510F756EF07" pageId="0" pageNumber="9">Etymology.</paragraph>
|
||
<paragraph id="B75AD4C3A19FCF089CCF0E3E137A132D" pageId="0" pageNumber="9">the species name, an adjective, relates to the Mediterranean Sea and its lands.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E0A9CB355C45296846433D5D5CE0C28E" pageId="0" pageNumber="9" type="host">
|
||
<paragraph id="1BBD235E41CBAA780849286CFB77B945" pageId="0" pageNumber="9">Host plant.</paragraph>
|
||
<paragraph id="D04A0A3BE7F6DFD76CE748C909D2310A" pageId="0" pageNumber="9">
|
||
Unknown. However, one or more
|
||
<taxonomicName id="2FDD0E336B3679DFE7A64441D800A404" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" lsidName="Acer" order="Sapindales" pageId="0" pageNumber="9" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="825B717A17DA65C69E15EEBDFB4779B0" italics="true" pageId="0" pageNumber="9">Acer</emphasis>
|
||
</taxonomicName>
|
||
species are probably hosts of
|
||
<taxonomicName id="FFF4BB658EAF27799EE53FE801706847" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="D81D2371E2EB64238827E1CF4B4DD2D2" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
, as for its closest relatives, such as
|
||
<taxonomicName id="9F44D4584CE0A67EB05AA9ED63A2EA34" lsidName="P. ignymontiensis" pageId="0" pageNumber="9" rank="species" species="ignymontiensis">
|
||
<emphasis id="155C7C040B3A0DBE1C3CF441D22B7380" italics="true" pageId="0" pageNumber="9">P. ignymontiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2B99D22ABC0618540D67831C6EE366D1" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="AEC3285032AADA6678607DCEB2CCF537" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
. Both
|
||
<taxonomicName id="FF23407E2793757486A76457ECB1A504" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" lsidName="Acer campestre" order="Sapindales" pageId="0" pageNumber="9" phylum="Tracheophyta" rank="species" species="campestre">
|
||
<emphasis id="D42B32D3823F84E1B3EBFEE5B267B350" italics="true" pageId="0" pageNumber="9">Acer campestre</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="76A24F8A24D28E2A3F2F3A75C92B13C1" lsidName="A. monspessulanum" pageId="0" pageNumber="9" rank="species" species="monspessulanum">
|
||
<emphasis id="75F823B72C00809F2AE3B0D712F15D9A" italics="true" pageId="0" pageNumber="9">A. monspessulanum</emphasis>
|
||
</taxonomicName>
|
||
occur at the type locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D92155A25B7CA4BAB8C1909F7FCF891" pageId="0" pageNumber="9" type="genetic data">
|
||
<paragraph id="261E2FE623EB94743A6254B5722CD77D" pageId="0" pageNumber="9">Genetic data.</paragraph>
|
||
<paragraph id="5BB8491892B4948C336D26B721698552" pageId="0" pageNumber="9">
|
||
in BOLD Systems the COI barcode sequence of the
|
||
<taxonomicName id="E6D1D5D96CB60D6FD5027917E49C8297" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="3C4AF6E915CE52260A3D39B949E36E54" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
holotype is the only specimen in BIN BOLD:ABA1652. Three German specimens of
|
||
<taxonomicName id="9A72959952959F0726BF9C02D245BE31" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="01B468BC07F3D90AA79C512AE2C5704F" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
with identical barcodes comprise BOLD:ABA1650. One German male
|
||
<taxonomicName id="E989A2D552F6CD1D4139805C60AFA13D" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="61CD08E5B4EE73B7E78832465F991020" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
comprises BOLD:ABV8260, and one Swiss male BOLD:ABV8261. The minimum divergence between
|
||
<taxonomicName id="C9F0B221F2470362D76B5F6B9EBBE932" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="56740CD11BC9DFD187B741D51B7202DA" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3BD33FB258544CCD16D915BEC505452" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="5A52DC364655F8E0E6BD8ABF989BDD9A" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
(BINs BOLD:ABA1650 and BOLD:ABV8260) is about 2.4%. Sequences (not in BOLD) of two
|
||
<taxonomicName id="7FB33750C2FC7A2B2001CE60753FFFEB" lsidName="P. ignymontiensis" pageId="0" pageNumber="9" rank="species" species="ignymontiensis">
|
||
<emphasis id="B3039464ABAA75A87F5FEBF649360E80" italics="true" pageId="0" pageNumber="9">P. ignymontiensis</emphasis>
|
||
</taxonomicName>
|
||
specimens (DEI-GISHym81330, DEI-GISHym89949) diverge slightly more from
|
||
<taxonomicName id="A6B1590A7C0915C4C1294CE9C5256B8F" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="BC50F88AFBA95A8674E3835D50E2DAA9" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
, by respectively approximately 2.5-2.6%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B45D4F02D380A0C7B064D5B9BF5EE127" pageId="0" pageNumber="9" type="diagnosis">
|
||
<paragraph id="22284095DFDE8BEF75B83CC1F44EA14B" pageId="0" pageNumber="9">Diagnosis.</paragraph>
|
||
<paragraph id="DDA580B184684704E3912E65A0EF31E2" pageId="0" pageNumber="9">
|
||
<taxonomicName id="88DBAB98DB2793070F233393C426A2FE" authorityName="Liston" authorityYear="2023" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius mediterraneus" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="mediterraneus">
|
||
<emphasis id="D30A34C133C12BF1AA79F726BA4B2D44" italics="true" pageId="0" pageNumber="9">Pamphilius mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="253B26DBCBE85E8321D7DB295B18901F" baseAuthorityName="A.Costa" baseAuthorityYear="1860" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius alternans" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="alternans">
|
||
<emphasis id="F5B37505260792E0A32E49D360A55CF9" italics="true" pageId="0" pageNumber="9">Pamphilius alternans</emphasis>
|
||
</taxonomicName>
|
||
sub-group of the
|
||
<taxonomicName id="C1E0825FDFDF597BFD045751F215195D" baseAuthorityName="A.Costa" baseAuthorityYear="1860" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pamphilius alternans" order="Hymenoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="alternans">
|
||
<emphasis id="321710CA9860A33055DC0346F91B8B28" italics="true" pageId="0" pageNumber="9">Pamphilius alternans</emphasis>
|
||
</taxonomicName>
|
||
species group, as defined by
|
||
<bibRefCitation id="6293F3BE050B905CB17D6BB855BD3890" author="Shinohara, A" editor="Viitasaari, M" journalOrPublisher="Tremex Press, Helsinki" pageId="0" pageNumber="9" pagination="359 - 438" refId="B77" refString="Shinohara, A, 2002. Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M, Ed., Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki: 359 - 438" title="Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview." volumeTitle="Sawflies (Hymenoptera, Symphyta) I." year="2002">Shinohara (2002)</bibRefCitation>
|
||
. All members of this species sub-group are West Palaearctic. In the key to species (females) of the sub-group by
|
||
<bibRefCitation id="B452BA0A2A7A7E869A479B6987A6476A" author="Shinohara, A" journalOrPublisher="Bulletin of the National Science Museum, Series A, Zoology" pageId="0" pageNumber="9" pagination="25 - 63" publicationUrl="archive.org" refId="B76" refString="Shinohara, A, 1991. Pamphilius alternans (Hymenoptera, Pamphiliidae) and its Close Relatives. Bulletin of the National Science Museum, Series A, Zoology 17 (1): 25 - 63, archive.org" title="Pamphilius alternans (Hymenoptera, Pamphiliidae) and its Close Relatives." url="archive.org" volume="17" year="1991">Shinohara (1991)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="0E4F2CA265AB7C74C3D27B637CE34335" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="5A349FE72D5692F188D4B29890BD061B" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
runs to couplet 8, but does not fit either alternative, because whereas its head is largely black, abdominal sternum 6 is mostly pale. In the key to European species (females) by
|
||
<bibRefCitation id="8A8617E3CE063755512531A25842EA23" author="Viitasaari, M" editor="Viitasaari, M" journalOrPublisher="A review of the suborder, the Western Palaearctic taxa of Xyeloidea and Pamphilioidea. Tremex, Helsinki" pageId="0" pageNumber="9" pagination="235 - 358" refId="B86" refString="Viitasaari, M, 2002b. The Northern European taxa of Pamphiliidae (Hymenoptera). In: Viitasaari, M, Ed., Sawflies (Hymenoptera, Symphyta) I. A review of the suborder, the Western Palaearctic taxa of Xyeloidea and Pamphilioidea. Tremex, Helsinki: 235 - 358" title="The Northern European taxa of Pamphiliidae (Hymenoptera)." volumeTitle="Sawflies (Hymenoptera, Symphyta) I." year="2002 b">Viitasaari (2002b)</bibRefCitation>
|
||
it runs to couplet 18 containing
|
||
<taxonomicName id="BDE4D511E2A8B28C05D769941E514EA2" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="3C7FE96D4EFD1AE82BD3088B51499674" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="04EDAB0C2A307B280D13AB893510C39C" lsidName="P. ignymontiensis" pageId="0" pageNumber="9" rank="species" species="ignymontiensis">
|
||
<emphasis id="F7966CCF811DBC314CE32D7431C7A716" italics="true" pageId="0" pageNumber="9">P. ignymontiensis</emphasis>
|
||
</taxonomicName>
|
||
but does not fit the combination of characters for either of the species. It is closer to
|
||
<taxonomicName id="275BC686BCBD7F05136DA1E63D5BC6DF" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="6C127BA7200FB1B7F156C6D22654D57B" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
in lacking a pale stripe from compound eye to posterior border of head, but differs in its very smooth postocular area (densely and rather deeply punctate in
|
||
<taxonomicName id="73B300A9E132C3DCE3D9B5F8973AC759" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="D0D77FD202D0742EA8F2625DF2C6B726" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
). The nearly entirely pale abdominal sterna of
|
||
<taxonomicName id="43254CA66F58A28E3CE972DF20A7A9FB" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="ECDD60F97D5C7274B7B79A383EF4B938" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
separate it from both
|
||
<taxonomicName id="A480ECB96125EC3CF8DBC12F58F0E675" lsidName="P. aurantiacus" pageId="0" pageNumber="9" rank="species" species="aurantiacus">
|
||
<emphasis id="5DE78A5753B2556AD5BA6D5E77D587EA" italics="true" pageId="0" pageNumber="9">P. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6820F4392CF6A461A6AD3FA3DD1588EE" lsidName="P. ignymontiensis" pageId="0" pageNumber="9" rank="species" species="ignymontiensis">
|
||
<emphasis id="20319D8355D038CC3FF1150AE9A8BB06" italics="true" pageId="0" pageNumber="9">P. ignymontiensis</emphasis>
|
||
</taxonomicName>
|
||
, with several sterna extensively dark. The single
|
||
<taxonomicName id="0CE5969C68D6CFEF142E38C2BD3198C2" lsidName="P. mediterraneus" pageId="0" pageNumber="9" rank="species" species="mediterraneus">
|
||
<emphasis id="D724523E24BA590F9A08D72193114762" italics="true" pageId="0" pageNumber="9">P. mediterraneus</emphasis>
|
||
</taxonomicName>
|
||
specimen is similar in size to
|
||
<taxonomicName id="0E21C38736BFD9674912B58B2AD66438" lsidName="P. ignymontiensis" pageId="0" pageNumber="9" rank="species" species="ignymontiensis">
|
||
<emphasis id="570A033AA07AB6362130559E313D355C" italics="true" pageId="0" pageNumber="9">P. ignymontiensis</emphasis>
|
||
</taxonomicName>
|
||
females, i.e. smaller than
|
||
<taxonomicName id="12898FD2F9DD09F173002D1D0071313B" lsidName="P. aurantiacus." pageId="0" pageNumber="9" rank="species" species="aurantiacus.">
|
||
<emphasis id="EFBB67E37F232BAFAFF5DAFB05041C50" italics="true" pageId="0" pageNumber="9">P. aurantiacus.</emphasis>
|
||
</taxonomicName>
|
||
Differences are summarized in the following key.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |