134 lines
11 KiB
XML
134 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.788.21222" ID-GBIF-Dataset="f062cb75-2d34-4171-a3f5-72fb9771df04" ID-PMC="PMC6189229" ID-Pensoft-Pub="1313-2970-788-87" ID-PubMed="30337826" ID-ZBK="FCC76C6C5445497F9C6717A4F1A51464" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-788-87" ModsDocOrigin="ZooKeys 788" ModsDocTitle="A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)" checkinTime="1539050140006" checkinUser="pensoft" docAuthor="Goldstein, Paul Z., Janzen, Daniel H., Proshek, Benjamin, Dapkey, Tanya & Hallwachs, Winnie" docDate="2018" docId="83093074290DD9D686F475B08F24DDFF" docLanguage="en" docName="ZooKeys 788: 87-133" docOrigin="ZooKeys 788" docSource="http://dx.doi.org/10.3897/zookeys.788.21222" docTitle="Leucosigma solisae Goldstein, sp. n." docType="treatment" docUuid="C50ACC24-E816-4019-8FF5-7690574D9D3B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="102" masterDocId="FFC1FD22FF872E407E7FFF8E7B5FFFF9" masterDocTitle="A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)" masterLastPageNumber="133" masterPageNumber="87" pageNumber="100" updateTime="1668166274258" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Goldstein, Paul Z.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Janzen, Daniel H.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Proshek, Benjamin</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Dapkey, Tanya</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Hallwachs, Winnie</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2018</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>788</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>87</mods:start>
|
|
<mods:end>133</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.788.21222</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.788.21222</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-788-87</mods:identifier>
|
|
<mods:identifier type="ZBK">FCC76C6C5445497F9C6717A4F1A51464</mods:identifier>
|
|
<mods:identifier type="ZooBank">FCC76C6C5445497F9C6717A4F1A51464</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="148657019" LSID="urn:lsid:zoobank.org:act:C50ACC24-E816-4019-8FF5-7690574D9D3B" httpUri="http://treatment.plazi.org/id/83093074290DD9D686F475B08F24DDFF" lastPageId="15" lastPageNumber="102" pageId="13" pageNumber="100">
|
|
<subSubSection pageId="13" pageNumber="100" type="nomenclature">
|
|
<paragraph pageId="13" pageNumber="100">
|
|
<taxonomicName LSID="http://zoobank.org/C50ACC24-E816-4019-8FF5-7690574D9D3B" authority="Goldstein" class="Insecta" family="Noctuidae" genus="Leucosigma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucosigma solisae" order="Lepidoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="solisae">Leucosigma solisae Goldstein</taxonomicName>
|
|
<taxonomicNameLabel pageId="13" pageNumber="100">sp. n.</taxonomicNameLabel>
|
|
Figs 15, 16, 25, 26; Male genitalia: Figs 57, 58, 81, 82
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="14" pageNumber="101" type="materials_examined">
|
|
<paragraph pageId="14" pageNumber="101">
|
|
<pageBreakToken pageId="14" pageNumber="101" start="start">Material</pageBreakToken>
|
|
examined.
|
|
</paragraph>
|
|
<paragraph pageId="14" pageNumber="101">Type material. HOLOTYPE ♂. Voucher: D.H. Janzen & W. Hallwachs DB: http://janzen.sas.upenn.edu Area de Conservacion Guanacaste, COSTA RICA, Sector Pitilla: Estacion Pitilla, 10.98931, -85.42581, el. 675m, 02/17/2007, S. Rios & F. Quesada, collector, 07-SRNP-101229, ♂ USNM Dissection 148077, USNMENT01370294.</paragraph>
|
|
<paragraph pageId="14" pageNumber="101">PARATYPES (2♂). COSTA RICA: (1♂): Alajuela: Sector Rincon Rain Forest: Estacion Caribe, 10.90082, -85.2764, el. 391m, 10/10/2007, S. Rios & H. Cambronero, collector, 07-SRNP-109201, ♂ USNM Dissection 148078, USNMENT01437370. PERU (1♂): La Oroya, R. Inambari, Peru, Sept. 1904 3100 ft, dry seas., (G. Ockenden), Rothschild Bequest B.M. 1939-1., NHMUK010606203. Types at USNM.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="15" lastPageNumber="102" pageId="14" pageNumber="101" type="diagnosis">
|
|
<paragraph pageId="14" pageNumber="101">Diagnosis.</paragraph>
|
|
<paragraph lastPageId="15" lastPageNumber="102" pageId="14" pageNumber="101">
|
|
Forewing terminal area and distal part of the medial area appearing more uniform purplish gray than in other species, giving the wing a smoother, less granular appearance overall. The reniform-orbicular complex forms a continuous
|
|
<pageBreakToken pageId="15" pageNumber="102" start="start">green</pageBreakToken>
|
|
<normalizedToken originalValue="“Ч">"Ch</normalizedToken>
|
|
"(left wing) or
|
|
<normalizedToken originalValue="“μ”">"μ"</normalizedToken>
|
|
(right wing), the outline of which is broader than in other species except
|
|
<taxonomicName lsidName="L. albimixta" pageId="15" pageNumber="102" rank="species" species="albimixta">L. albimixta</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="L. viridipicta" pageId="15" pageNumber="102" rank="species" species="viridipicta">L. viridipicta</taxonomicName>
|
|
; the green (not silvery-white) apical patch distinguishes this and the following species from other members of the genus; hind wing underside with discal spot present but unpronounced, unlike
|
|
<taxonomicName lsidName="L. poolei" pageId="15" pageNumber="102" rank="species" species="poolei">L. poolei</taxonomicName>
|
|
below. Male genitalia most similar to those of
|
|
<taxonomicName lsidName="L. poolei" pageId="15" pageNumber="102" rank="species" species="poolei">L. poolei</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="L. uncifera" pageId="15" pageNumber="102" rank="species" species="uncifera">L. uncifera</taxonomicName>
|
|
, distinguished by a shorter and more gently arced distal part of the cucullus; vesica without small basal lobe as in
|
|
<taxonomicName lsidName="L. poolei" pageId="15" pageNumber="102" rank="species" species="poolei">L. poolei</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="15" pageNumber="102" type="description">
|
|
<paragraph pageId="15" pageNumber="102">Description.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Head. Antennae setose-ciliate, bifasciculate in males, scaled above with alternating bands of gray (basal) and tan (apical). Frons, vertex, and labial palpi scaled with an admixture of white, brown and black. Eyes smooth.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">
|
|
Thorax. Excepting vestiges of green visible immediately behind head, thoracic vestiture uniformly purplish gray. Wings. Forewing length 13.7 mm (holotype, male), average 13.3 mm (males, n = 3). Forewing predominantly lilacine, russet subcostally in the postmedial; apical and anal patch present; green scaling concentrated at inner margin, apical and anal patch, and
|
|
<normalizedToken originalValue="Ч">Ch</normalizedToken>
|
|
/μ-shaped medial stigma. Medial area cupreous; medial, antemedial, and postmedial lines visibly darker brown; terminal area green in Costa Rican specimens, more closely matching medial coloration or slightly darker reddish brown in Peruvian specimen; apical patch primarily green; forewing underside lightly suffused with green scaling in terminal area; no antemedial or postmedial lines present on underside except at most as costal striae. Legs. Scales predominantly purplish gray; femora and tibia with an admixture of lime-green among the tan-lilacine scales; a single pair of striped mid-tibial spurs, two pairs on hind-tibiae; three rows of tibial spines on all legs.
|
|
</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Abdomen. Vestiture uniformly tannish gray, paler than on thorax.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">
|
|
Male genitalia. Uncus elongate, widest subapically, upwardly curved with a very small apical point, and bearing ventral setal crest. Tegumen roughly hemi-circular, excepting a deformation of the dorsal edge at the base of the uncus. Vinculum a wide V-shape; saccus blunt. Juxta pentagonal, dorsal edge horizontal; annellar arms fused, hoop-like. Sacculus (1) densely setose, tapering to a blunt extension. Cucullus (2) appears chelicerate, the sclerotized part occupying ~1/3 the overall length and bending sharply at the outer edge near the sharply pointed apex, with a subapical tuft of
|
|
<normalizedToken originalValue="reëntrant">reentrant</normalizedToken>
|
|
spine-like setae. Dorsal process (3) coequal in width to cucullus, setose apically. Clasper (4) finger-like, gently curved. Aedeagus with minutely but differentially spinulose patch confined to apex. Vesica without cornuti; subbasal and medial diverticula reduced, with a weakly sclerotized ridge.
|
|
</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Female genitalia. Unknown.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Immature stages. Unknown.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="15" pageNumber="102" type="etymology">
|
|
<paragraph pageId="15" pageNumber="102">Etymology.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">
|
|
The name
|
|
<taxonomicName lsidName="solisae" pageId="15" pageNumber="102" rank="species" species="solisae">solisae</taxonomicName>
|
|
is given in honor of Dr. Alma Solis, lepidopterist at USDA/ USNM who has contributed her expertise to the systematics of the Costa Rican lepidopteran fauna for three decades.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="15" pageNumber="102" type="biology_ecology">
|
|
<paragraph pageId="15" pageNumber="102">Biology.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Unknown, collected only in rain forest light traps.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="15" pageNumber="102" type="distribution">
|
|
<paragraph pageId="15" pageNumber="102">Distribution.</paragraph>
|
|
<paragraph pageId="15" pageNumber="102">Costa Rica, Peru.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |