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<document id="E5E4395D013EB561735220BC3CE6CD68" ID-CLB-Dataset="277691" ID-DOI="10.11646/phytotaxa.622.3.3" ID-GBIF-Dataset="822e51ae-aa89-46b6-a723-dc5eedae9a36" ID-ISSN="1179-3163" ID-Zenodo-Dep="10146545" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700187957960" checkinUser="plazi" docAuthor="Monteros, Marco F., Restrepo, Eugenio, Iturralde, Gabriel A., Jiménez, Marco M. &amp; Baquero, Luis E." docDate="2023" docId="03FB9024FF9EFFB8F4E9834771A94306" docLanguage="en" docName="phytotaxa.622.3.3.pdf" docOrigin="Phytotaxa 622 (3)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.622.3.3/51255" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Lepanthes quadricornis Luer &amp; R. Escobar MFM 1994" docType="treatment" docVersion="2" lastPageNumber="211" masterDocId="FFC2E85CFF98FFB0F461843272014206" masterDocTitle="Contributions to the Lepanthes (Orchidaceae) knowledge in Ecuador: A new species from Cordillera del Toisán and new records for the country" masterLastPageNumber="214" masterPageNumber="204" pageNumber="210" updateTime="1700188752213" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F68647AAB05E64DCE4D104BDA5A930CC">Contributions to the Lepanthes (Orchidaceae) knowledge in Ecuador: A new species from Cordillera del Toisán and new records for the country</mods:title>
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<mods:namePart id="50A422AEF3B743B6A85F52000C915135">Monteros, Marco F.</mods:namePart>
<mods:nameIdentifier id="75BF40BBB5105D795D5B7E9F4F6B0A15" type="ORCID">0000-0002-6670-3687</mods:nameIdentifier>
<mods:affiliation id="B6A5831C746574C2F8A758DABCC83649">Fundacion EcoMinga, Mariscal Foch 7 - 21 y Juan León Mera, Quito, Ecuador. &amp; Reserva: The Youth Land Trust, Washington, D. C., USA. &amp; Instituto Nacional de Biodiversidad (INABIO), Rumipamba 341 y Av. De los Shyris, Quito, Ecuador. &amp; Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Ecuador. &amp; marcomonteros 24 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6670 - 3687</mods:affiliation>
<mods:nameIdentifier id="39AC63133E8FE73E2C46BE4390F33D18" type="email">marcomonteros24@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D8FA9FB667D6546CFDF03186711ED47C">Restrepo, Eugenio</mods:namePart>
<mods:nameIdentifier id="21E075957849C87114911A911D41DFB0" type="ORCID">0000-0002-7037-1670</mods:nameIdentifier>
<mods:affiliation id="2FF706A1E38FB309DF26484723929369">Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Ecuador. &amp; Programa de Biología, Facultad de Ciencias Exactas y Naturales, Universidad de Caldas, Calle 65 # 26 - 10, Manizales, Colombia. Grupo de Investigación Schultes, Fundación Ecotonos, Carrera 72 # 13 A- 56, Cali, Colombia. &amp; estambul 40 @ gmail. com; https: // orcid. org / 0000 - 0002 - 7037 - 1670</mods:affiliation>
<mods:nameIdentifier id="9748C09A8883E9AB4DA353063479FB83" type="email">estambul40@gmail.com</mods:nameIdentifier>
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<mods:namePart id="46F6EB8079F32C54516FF0561E3536F8">Iturralde, Gabriel A.</mods:namePart>
<mods:nameIdentifier id="4E3D6891E8FDD7A16B822A6CEF57C946" type="ORCID">0000-0003-2456-0929</mods:nameIdentifier>
<mods:affiliation id="797B9D06AA413FFEF614550F08A30DF0">Grupo de Investigación en Medio Ambiente y Salud BIOMAS, Facultad de Ingeniería y Ciencias Aplicadas, Universidad de Las Américas, Vía a Nayón, Quito 170124, Ecuador. &amp; gabriel. iturralde @ udla. edu. ec; https: // orcid. org / 0000 - 0003 - 2456 - 0929</mods:affiliation>
<mods:nameIdentifier id="57EDFC3CA3958D80311EB3AB1B746835" type="email">gabriel.iturralde@udla.edu.ec</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="E6E3491996928DFF38CC8B3949B08252">Jiménez, Marco M.</mods:namePart>
<mods:nameIdentifier id="8D6A1454141F0E5AD9A9D20F4CAE8238" type="ORCID">0000-0002-9502-5651</mods:nameIdentifier>
<mods:affiliation id="7684A77494F9883857B341D932C6544E">Grupo de Investigación en Medio Ambiente y Salud BIOMAS, Facultad de Ingeniería y Ciencias Aplicadas, Universidad de Las Américas, Vía a Nayón, Quito 170124, Ecuador. &amp; marco. jimenez. leon @ udla. edu. ec; https: // orcid. org / 0000 - 0002 - 9502 - 5651</mods:affiliation>
<mods:nameIdentifier id="B08465353BB12E4166C693692BD95947" type="email">marco.jimenez.leon@udla.edu.ec</mods:nameIdentifier>
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<mods:namePart id="A5E5A326A36A15F4B4B518234BE5D8A3">Baquero, Luis E.</mods:namePart>
<mods:nameIdentifier id="B850D68E680596143BA607EFAA72A8BE" type="ORCID">0000-0002-1444-5727</mods:nameIdentifier>
<mods:affiliation id="697CACDBD8321B3AD31151A1AA23DBC4">Grupo de Investigación en Medio Ambiente y Salud BIOMAS, Facultad de Ingeniería y Ciencias Aplicadas, Universidad de Las Américas, Vía a Nayón, Quito 170124, Ecuador. &amp; luis. baquero @ udla. edu. ec; https: // orcid. org / 0000 - 0002 - 1444 - 5727</mods:affiliation>
<mods:nameIdentifier id="0B9CBFD92D0EAD9408F808C5AE23D3C2" type="email">luis.baquero@udla.edu.ec</mods:nameIdentifier>
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<mods:title id="57D463C7A3E79476B499243A359A6D71">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0271FEE0FC21F20E200F0570B2A81D2D">
<mods:date id="A35DE78F07C33E840D5C176446052001">2023</mods:date>
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<mods:number id="1876000339B0B8205B0B5645F61B8C4F">2023-10-27</mods:number>
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<mods:number id="0A5122F483958A109F8464C64916C9B2">622</mods:number>
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</mods:location>
<mods:classification id="D51B11CCA8FCE6693322D405A6D78173">journal article</mods:classification>
<mods:identifier id="67D52C468D6DA4C77807D6E497F98949" type="CLB-Dataset">277691</mods:identifier>
<mods:identifier id="367C2B761C4B217EBFDD858EBC9EBA28" type="DOI">10.11646/phytotaxa.622.3.3</mods:identifier>
<mods:identifier id="9114601014E51BE8D8D8E48DCFB123A7" type="GBIF-Dataset">822e51ae-aa89-46b6-a723-dc5eedae9a36</mods:identifier>
<mods:identifier id="E5CB959EB190024F6A141C0BFE7AA136" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="01BB851FF273B27E647428B7F26BDB3C" type="Zenodo-Dep">10146545</mods:identifier>
</mods:mods>
<treatment id="03FB9024FF9EFFB8F4E9834771A94306" LSID="urn:lsid:plazi:treatment:03FB9024FF9EFFB8F4E9834771A94306" httpUri="http://treatment.plazi.org/id/03FB9024FF9EFFB8F4E9834771A94306" lastPageId="8" lastPageNumber="211" pageId="6" pageNumber="210">
<subSubSection id="C34872B9FF9EFFB6F4E98347719A4596" box="[136,923,1909,1936]" pageId="6" pageNumber="210" type="nomenclature">
<paragraph id="8BED2132FF9EFFB6F4E98347719A4596" blockId="6.[136,923,1909,1936]" box="[136,923,1909,1936]" pageId="6" pageNumber="210">
<heading id="D0A5965EFF9EFFB6F4E98347719A4596" box="[136,923,1909,1936]" fontSize="11" level="2" pageId="6" pageNumber="210" reason="3">
<taxonomicName id="4C525AB1FF9EFFB6F4E9834771084596" ID-CoL="3T8HM" authority="Luer &amp; R. Escobar (1994: 108)" authorityName="Luer &amp; R. Escobar MFM" authorityPageNumber="108" authorityYear="1994" box="[136,777,1909,1936]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="210" phylum="Tracheophyta" rank="species" species="quadricornis">
<emphasis id="B926FD20FF9EFFB6F4E98347707F4596" bold="true" box="[136,638,1909,1936]" pageId="6" pageNumber="210">
<emphasis id="B926FD20FF9EFFB6F4E98347739A4596" bold="true" box="[136,411,1909,1936]" italics="true" pageId="6" pageNumber="210">Lepanthes quadricornis</emphasis>
Luer &amp; R.Escobar
</emphasis>
(1994: 108)
</taxonomicName>
(
<figureCitation id="13693DB7FF9EFFB6F7798344718F4596" box="[792,910,1910,1936]" captionStart="FIGURE 4" captionStartId="6.[136,229,1769,1791]" captionTargetBox="[151,1436,245,1745]" captionTargetId="figure-40@6.[151,1436,245,1745]" captionTargetPageId="6" captionText="FIGURE 4. Images of the specimens of the new Lepanthes species for Ecuador. A. L. acrogenia Luer &amp; R.Escobar. MFM216 (QCNE). B. L. elata Reichenbach.f. MFM219 (QCNE). C. L. quadricornis Luer &amp; R.Escobar MFM220 (QCNE). D. L. ricaurtensis Luer &amp; R.Escobar. MFM218 (QCNE). Photographs by Marco F. Monteros. Plate by Eugenio Restrepo." figureDoi="http://doi.org/10.5281/zenodo.10146553" httpUri="https://zenodo.org/record/10146553/files/figure.png" pageId="6" pageNumber="210">Figure 4C</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34872B9FF9EFFB7F4E983F0705144D0" lastPageId="7" lastPageNumber="211" pageId="6" pageNumber="210" type="materials_examined">
<paragraph id="8BED2132FF9EFFB6F4E983F0716745FD" blockId="6.[136,1451,1985,2043]" pageId="6" pageNumber="210">
<emphasis id="B926FD20FF9EFFB6F4E983F072DB45D1" bold="true" box="[136,218,1986,2007]" pageId="6" pageNumber="210">Type:—</emphasis>
<materialsCitation id="3B3A2B6FFF9EFFB6F4BB83F0732745FD" collectingDate="1989-07-21" collectionCode="C, H" collectorName="Apparently &amp; Ricaurte" country="Colombia" location="La Planada Science Station" pageId="6" pageNumber="210" specimenCount="1" stateProvince="Narino">
<collectingCountry id="F34561A2FF9EFFB6F4BB83F0735C45D1" box="[218,349,1986,2007]" name="Colombia" pageId="6" pageNumber="210">COLOMBIA</collectingCountry>
.
<collectingRegion id="4996EFD0FF9EFFB6F50683F373AC45D1" box="[359,429,1985,2007]" country="Colombia" name="Narino" pageId="6" pageNumber="210">Nariño</collectingRegion>
:
<collectorName id="26A744E4FF9EFFB6F5D783F3702345D1" box="[438,546,1985,2007]" pageId="6" pageNumber="210">Apparently</collectorName>
collected near
<collectorName id="26A744E4FF9EFFB6F6D483F3710845D1" box="[693,777,1985,2007]" pageId="6" pageNumber="210">Ricaurte</collectorName>
, flowered in cultivation at the
<location id="8E8D77E9FF9EFFB6F05383F0773845D1" LSID="urn:lsid:plazi:treatment:03FB9024FF9EFFB8F4E9834771A94306:8E8D77E9FF9EFFB6F05383F0773845D1" box="[1074,1337,1985,2007]" country="Colombia" name="La Planada Science Station" pageId="6" pageNumber="210" stateProvince="Narino">La Planada Science Station</location>
#48,
<date id="FFEC07F2FF9EFFB6F11083F072EE45FD" pageId="6" pageNumber="210" value="1989-07-21">
<collectingDate id="EFA8FE1AFF9EFFB6F11083F072EE45FD" pageId="6" pageNumber="210" value="1989-07-21">21 Jul 1989</collectingDate>
</date>
,
<emphasis id="B926FD20FF9EFFB6F49883D4732745FD" box="[249,294,2022,2043]" italics="true" pageId="6" pageNumber="210">
<collectionCode id="ED43B9F7FF9EFFB6F49883D4730D45FD" box="[249,268,2022,2043]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="210" type="Herbarium">C</collectionCode>
.
<collectionCode id="ED43B9F7FF9EFFB6F57483D4732745FD" box="[277,294,2022,2043]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="6" pageNumber="210" type="Herbarium">H</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B926FD20FF9EFFB6F54683D4702945FD" box="[295,552,2021,2043]" italics="true" pageId="6" pageNumber="210">
.
<materialsCitation id="3B3A2B6FFF9EFFB6F55283D7739B45FD" box="[307,410,2021,2043]" collectionCode="R" collectorName="Dodson" country="Colombia" pageId="6" pageNumber="210" specimenCount="1" stateProvince="Narino">
<collectorName id="26A744E4FF9EFFB6F55283D7738145FC" box="[307,384,2021,2042]" pageId="6" pageNumber="210">Dodson</collectorName>
,
<collectionCode id="ED43B9F7FF9EFFB6F5E883D4739B45FD" box="[393,410,2022,2043]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="210">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3A2B6FFF9EFFB6F5C583D7702445FD" box="[420,549,2021,2043]" collectionCode="E" collectorName="Escobar" country="Colombia" pageId="6" pageNumber="210" specimenCount="1" stateProvince="Narino">
<collectorName id="26A744E4FF9EFFB6F5C583D773F545FC" box="[420,500,2021,2042]" pageId="6" pageNumber="210">Escobar</collectorName>
&amp;
<collectionCode id="ED43B9F7FF9EFFB6F67283D4702445FD" box="[531,549,2022,2043]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="6" pageNumber="210" type="Herbarium">E</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="3B3A2B6FFF9EFFB6F64C83D7716345FD" box="[557,866,2021,2043]" collectionCode="MO" collectorName="Valencia" country="Colombia" pageId="6" pageNumber="210" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
<emphasis id="B926FD20FF9EFFB6F64C83D770C345FC" box="[557,706,2021,2042]" italics="true" pageId="6" pageNumber="210">
<collectorName id="26A744E4FF9EFFB6F64C83D7708145FC" box="[557,640,2021,2042]" pageId="6" pageNumber="210">Valencia</collectorName>
18060
</emphasis>
(
<typeStatus id="54E99F90FF9EFFB6F6AE83D7712945FD" box="[719,808,2021,2043]" pageId="6" pageNumber="210" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED43B9F7FF9EFFB6F75083D4715D45FD" box="[817,860,2022,2043]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="6" pageNumber="210" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF2D71BAFF9FFFB7F4E98239738E446F" ID-DOI="http://doi.org/10.5281/zenodo.10146555" ID-Zenodo-Dep="10146555" httpUri="https://zenodo.org/record/10146555/files/figure.png" pageId="7" pageNumber="211" startId="7.[136,229,1547,1569]" targetBox="[151,1436,190,1523]" targetPageId="7" targetType="figure">
<paragraph id="8BED2132FF9FFFB7F4E98239738E446F" blockId="7.[136,1452,1547,1641]" pageId="7" pageNumber="211">
<emphasis id="B926FD20FF9FFFB7F4E9823972FC4426" bold="true" box="[136,253,1547,1569]" pageId="7" pageNumber="211">FIGURE 5.</emphasis>
Comparison between different morphotypes of
<taxonomicName id="4C525AB1FF9FFFB7F6AB823E76534427" authority="Luer &amp; R. Escobar. A. Plant" box="[714,1106,1547,1569]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="211" phylum="Tracheophyta" rank="species" species="acrogenia">
<emphasis id="B926FD20FF9FFFB7F6AB823E71424427" box="[714,835,1548,1569]" italics="true" pageId="7" pageNumber="211">L. acrogenia</emphasis>
Luer &amp; R.Escobar.
<emphasis id="B926FD20FF9FFFB7F0658239761A4426" bold="true" box="[1028,1051,1547,1568]" pageId="7" pageNumber="211">A.</emphasis>
Plant
</taxonomicName>
from Ecuador found at 2000 m, not collected.
<emphasis id="B926FD20FF9FFFB7F48D821D73034442" bold="true" box="[236,258,1583,1604]" pageId="7" pageNumber="211">B.</emphasis>
Plant from close to the type locality in Colombia. Photographs by Marco F. Monteros (A, C) &amp; Sebastian Moreno (B, D). Plate by Eugenio Restrepo.
</paragraph>
</caption>
<paragraph id="8BED2132FF9FFFB7F4E982A5705144D0" blockId="7.[136,1452,1687,2074]" pageId="7" pageNumber="211">
<emphasis id="B926FD20FF9FFFB7F4E982A5738244B7" bold="true" box="[136,387,1687,1713]" pageId="7" pageNumber="211">Examined specimens</emphasis>
:
<materialsCitation id="3B3A2B6FFF9FFFB7F5F582AA72FC44D3" collectingDate="2021-08-04" collectionCode="M, F" country="Ecuador" location="Reserva Dracula" pageId="7" pageNumber="211" specimenCount="1" stateProvince="Carchi">
<collectingCountry id="F34561A2FF9FFFB7F5F582AA701E44B4" box="[404,543,1688,1714]" name="Ecuador" pageId="7" pageNumber="211">ECUADOR</collectingCountry>
.
<collectingRegion id="4996EFD0FF9FFFB7F64D82AA707A44B4" box="[556,635,1688,1714]" country="Ecuador" name="Carchi" pageId="7" pageNumber="211">Carchi</collectingRegion>
:
<location id="8E8D77E9FF9FFFB7F6E682AA714744B4" LSID="urn:lsid:plazi:treatment:03FB9024FF9EFFB8F4E9834771A94306:8E8D77E9FF9FFFB7F6E682AA714744B4" box="[647,838,1688,1714]" country="Ecuador" name="Reserva Dracula" pageId="7" pageNumber="211" stateProvince="Carchi">Reserva Dracula</location>
, coordinates omitted for conservation purpose,
<date id="FFEC07F2FF9FFFB7F10782AA72C244D0" pageId="7" pageNumber="211" value="2021-08-04">
<collectingDate id="EFA8FE1AFF9FFFB7F10782AA72C244D0" pageId="7" pageNumber="211" value="2021-08-04">4 Aug 2021</collectingDate>
</date>
,
<emphasis id="B926FD20FF9FFFB7F4AF828E72FC44D3" box="[206,253,1724,1749]" italics="true" pageId="7" pageNumber="211">
<collectionCode id="ED43B9F7FF9FFFB7F4AF828E72E944D3" box="[206,232,1724,1749]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="7" pageNumber="211" type="Herbarium">M</collectionCode>
.
<collectionCode id="ED43B9F7FF9FFFB7F48A828E72FC44D3" box="[235,253,1724,1749]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="7" pageNumber="211" type="Herbarium">F</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B926FD20FF9FFFB7F49F828E730344D3" box="[254,258,1724,1749]" italics="true" pageId="7" pageNumber="211">.</emphasis>
<materialsCitation id="3B3A2B6FFF9FFFB7F569828E704D44D0" box="[264,588,1724,1750]" collectionCode="QCNE" country="Ecuador" location="Monteros" pageId="7" pageNumber="211" specimenCount="1" stateProvince="Carchi">
<emphasis id="B926FD20FF9FFFB7F569828E73E244D0" box="[264,483,1724,1750]" italics="true" pageId="7" pageNumber="211">
<location id="8E8D77E9FF9FFFB7F569828E737344D3" LSID="urn:lsid:plazi:treatment:03FB9024FF9EFFB8F4E9834771A94306:8E8D77E9FF9FFFB7F569828E737344D3" box="[264,370,1724,1749]" country="Ecuador" name="Monteros" pageId="7" pageNumber="211" stateProvince="Carchi">Monteros</location>
MFM220
</emphasis>
(
<collectionCode id="ED43B9F7FF9FFFB7F593828E704444D0" box="[498,581,1724,1750]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="7" pageNumber="211" type="Herbarium">QCNE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34872B9FF9FFFB8F4DC82ED71A94306" lastPageId="8" lastPageNumber="212" pageId="7" pageNumber="211" type="distribution">
<paragraph id="8BED2132FF9FFFB7F4DC82ED779D458C" blockId="7.[136,1452,1687,2074]" pageId="7" pageNumber="211">
<emphasis id="B926FD20FF9FFFB7F4DC82ED73EB44FC" bold="true" box="[189,490,1759,1786]" pageId="7" pageNumber="211">Distribution and ecology</emphasis>
:
<taxonomicName id="4C525AB1FF9FFFB7F59D82D2710A44FC" authorityName="Luer &amp; R. Escobar MFM" authorityYear="1994" box="[508,779,1760,1786]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="211" phylum="Tracheophyta" rank="species" species="quadricornis">
<emphasis id="B926FD20FF9FFFB7F59D82D2710A44FC" box="[508,779,1760,1786]" italics="true" pageId="7" pageNumber="211">Lepanthes quadricornis</emphasis>
</taxonomicName>
was previously known from southwestern
<collectingCountry id="F34561A2FF9FFFB7F16082D2777244FC" box="[1281,1395,1760,1786]" name="Colombia" pageId="7" pageNumber="211">Colombia</collectingCountry>
. The
<typeStatus id="54E99F90FF9FFFB7F4E9833672B94518" box="[136,184,1796,1822]" pageId="7" pageNumber="211">type</typeStatus>
locality is located near the La Planada reserve between
<quantity id="4CAA8CD7FF9FFFB7F74E833671CB4518" box="[815,970,1796,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="7" pageNumber="211" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800 m</quantity>
in the
<collectingRegion id="4996EFD0FF9FFFB7F079833676EA4518" box="[1048,1259,1796,1822]" country="Colombia" name="Narino" pageId="7" pageNumber="211">Nariño department</collectingRegion>
(
<bibRefCitation id="EFC35CC3FF9FFFB7F09B833672C44544" author="Luer, C. A. &amp; Thoerle, L." pageId="7" pageNumber="211" pagination="1 - 296" refId="ref5097" refString="Luer, C. A. &amp; Thoerle, L. (2012) Icones Pleurothallidinarum XXXII. Lepanthes of Colombia (Orchidaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 123: 1 - 296." type="journal article" year="2012">Luer &amp; Thoerle 2012</bibRefCitation>
).
<taxonomicName id="4C525AB1FF9FFFB7F4BA831A73EB4544" authorityName="Luer &amp; R. Escobar MFM" authorityYear="1994" box="[219,490,1832,1858]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="211" phylum="Tracheophyta" rank="species" species="quadricornis">
<emphasis id="B926FD20FF9FFFB7F4BA831A73EB4544" box="[219,490,1832,1858]" italics="true" pageId="7" pageNumber="211">Lepanthes quadricornis</emphasis>
</taxonomicName>
has recently been recorded from the cloud forests of northwestern
<collectingCountry id="F34561A2FF9FFFB7F160831A775F4544" box="[1281,1374,1832,1858]" name="Ecuador" pageId="7" pageNumber="211">Ecuador</collectingCountry>
in the
<collectingRegion id="4996EFD0FF9FFFB7F4E9837E733A4560" box="[136,315,1868,1894]" country="Ecuador" name="Carchi" pageId="7" pageNumber="211">Carchi province</collectingRegion>
at
<quantity id="4CAA8CD7FF9FFFB7F53D837973B54560" box="[348,436,1867,1894]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="211" unit="m" value="2150.0">2150 m</quantity>
. The species extends its distribution
<quantity id="4CAA8CD7FF9FFFB7F72A837E71934560" box="[843,914,1868,1894]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="211" unit="km" value="40.0">40 km</quantity>
south of the
<typeStatus id="54E99F90FF9FFFB7F043837E76534560" box="[1058,1106,1868,1894]" pageId="7" pageNumber="211">type</typeStatus>
locality and rises
<quantity id="4CAA8CD7FF9FFFB7F17D837977634560" box="[1308,1378,1867,1894]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="211" unit="m" value="350.0">350 m</quantity>
above the previous records from
<collectingCountry id="F34561A2FF9FFFB7F5D183427023458C" box="[432,546,1904,1930]" name="Colombia" pageId="7" pageNumber="211">Colombia</collectingCountry>
. In northwestern
<collectingCountry id="F34561A2FF9FFFB7F68A83427149458C" box="[747,840,1904,1930]" name="Ecuador" pageId="7" pageNumber="211">Ecuador</collectingCountry>
has been flowering in situ between May and August.
</paragraph>
<paragraph id="8BED2132FF9FFFB8F4DC83A171A94306" blockId="7.[136,1452,1687,2074]" lastBlockId="8.[136,1451,158,257]" lastPageId="8" lastPageNumber="212" pageId="7" pageNumber="211">
<emphasis id="B926FD20FF9FFFB7F4DC83A173C945AB" bold="true" box="[189,456,1939,1965]" pageId="7" pageNumber="211">Taxonomic Discussion</emphasis>
:
<taxonomicName id="4C525AB1FF9FFFB7F5B883A670E745A8" authorityName="Luer &amp; R. Escobar MFM" authorityYear="1994" box="[473,742,1940,1966]" class="Liliopsida" family="Orchidaceae" genus="Lepanthes" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="211" phylum="Tracheophyta" rank="species" species="quadricornis">
<emphasis id="B926FD20FF9FFFB7F5B883A670E745A8" box="[473,742,1940,1966]" italics="true" pageId="7" pageNumber="211">Lepanthes quadricornis</emphasis>
</taxonomicName>
is characterized by the proportionally large flowers on top of a spreading, broadly elliptical leaf. The sepals are densely ciliate and form a broad, shallow cup; the dorsal sepal is short-tailed, and the lateral sepals are acute. The petals are forked with erect, oblong, and sharply carinated lobes. The lip is “U” shaped, glabrous and encircles the column (
<bibRefCitation id="EFC35CC3FF9FFFB7F68A8C3271E24A1C" author="Luer, C. A. &amp; Thoerle, L." box="[747,995,2048,2074]" pageId="7" pageNumber="211" pagination="1 - 296" refId="ref5097" refString="Luer, C. A. &amp; Thoerle, L. (2012) Icones Pleurothallidinarum XXXII. Lepanthes of Colombia (Orchidaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 123: 1 - 296." type="journal article" year="2012">Luer &amp; Thoerle 2012</bibRefCitation>
). The Ecuadorian specimens are slightly different when compared to the Colombian ones (
<figureCitation id="13693DB7FF90FFB8F6A184AC712242BE" box="[704,803,158,184]" captionStart="FIGURE 6" captionStartId="8.[136,229,1859,1881]" captionTargetBox="[151,1436,317,1834]" captionTargetId="figure-71@8.[151,1436,317,1834]" captionTargetPageId="8" captionText="FIGURE 6. Comparison between different morphotypes of L. quadricornis Luer &amp; R.Escobar A. Plant from Ecuador, from M. Monteros 220 (QCNE). B. Plant from close to the type locality in Colombia. Photographs by Marco F. Monteros (A, C) &amp; Sebastian Moreno (B, D). Plate by Eugenio Restrepo." figureDoi="http://doi.org/10.5281/zenodo.10146557" httpUri="https://zenodo.org/record/10146557/files/figure.png" pageId="8" pageNumber="212">Figure 6</figureCitation>
), especially in the proportion of the flower sepals, being the dorsal sepal more concave with a longer tail in the apex; the lateral sepals are vinaceous suffused with ivory in the center and are longer and wider in relation to the Ecuadorian specimen.
</paragraph>
</subSubSection>
</treatment>
</document>
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