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<document id="5261D5004BE741463A7DF57CFE082EEA" ID="10.11646/zootaxa.4242.3.2SLASH10462" ID-CLB-Dataset="36297" ID-DOI="10.11646/zootaxa.4242.3.2" ID-GBIF-Dataset="0abf37ce-8190-42a4-87bc-96f3c8e10a38" ID-ISSN="1175-5326" ID-Zenodo-Dep="376905" ID-ZooBank="C67937C9-844F-461E-AABB-121B9C3CE5FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489393351114" checkinUser="plazi" docAuthor="Willems, Wim R., Reygel, Patrick, Steenkiste, Niels Van, Tessens, Bart &amp; Artois, Tom J." docDate="2017" docId="03FB87EB522CE34A57BEA9DFFE6DDEAA" docLanguage="en" docName="zootaxa.4242.3.2SLASH10462.pdf" docOrigin="Zootaxa 4242 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paulodora drepanophora Artois &amp; Tessens 2008" docType="treatment" docVersion="9" lastPageNumber="455" masterDocId="FFC2FF935222E3445729AD38FFA0D959" masterDocTitle="Kalyptorhynchia (Platyhelminthes: Rhabdocoela) from KwaZulu-Natal (South Africa), with the description of six new species" masterLastPageNumber="466" masterPageNumber="441" pageNumber="455" updateTime="1698437864087" updateUser="ExternalLinkService">
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<mods:title id="03EB875776327AA05572583C2786607D">Kalyptorhynchia (Platyhelminthes: Rhabdocoela) from KwaZulu-Natal (South Africa), with the description of six new species</mods:title>
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<mods:namePart id="77425CFA7593D10CB64EE8DC39D39B5A">Willems, Wim R.</mods:namePart>
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<mods:namePart id="B8CF7AD264786CFEED0B3F3F9CC4B90F">Reygel, Patrick</mods:namePart>
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<mods:namePart id="B1AE60D83BF7A79D44A34A87C42734B0">Steenkiste, Niels Van</mods:namePart>
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<mods:namePart id="AA92B3096E6D379FE69D370D635F3D61">Tessens, Bart</mods:namePart>
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<mods:namePart id="14F0CFAB60F9B99AC55DDF47BD062A4F">Artois, Tom J.</mods:namePart>
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<treatment id="03FB87EB522CE34A57BEA9DFFE6DDEAA" ID-DOI="http://doi.org/10.5281/zenodo.5689676" ID-GBIF-Taxon="127843769" ID-Zenodo-Dep="5689676" LSID="urn:lsid:plazi:treatment:03FB87EB522CE34A57BEA9DFFE6DDEAA" httpUri="http://treatment.plazi.org/id/03FB87EB522CE34A57BEA9DFFE6DDEAA" lastPageNumber="455" pageId="14" pageNumber="455">
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<paragraph id="8BED36FD522CE34A57BEA9DFFD4CDC5B" blockId="14.[151,748,1255,1315]" box="[151,748,1255,1282]" pageId="14" pageNumber="455">
<heading id="D0A58191522CE34A57BEA9DFFD4CDC5B" bold="true" box="[151,748,1255,1282]" fontSize="11" level="1" pageId="14" pageNumber="455" reason="1">
<taxonomicName id="4C524D7E522CE34A57BEA9DFFD4CDC5B" ID-CoL="75VB9" authority="Artois &amp; Tessens, 2008" authorityName="Artois &amp; Tessens" authorityYear="2008" box="[151,748,1255,1282]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" kingdom="Animalia" order="Rhabdocoela" pageId="14" pageNumber="455" phylum="Platyhelminthes" rank="species" species="drepanophora">
<emphasis id="B926EAEF522CE34A57BEA9DFFD4CDC5B" bold="true" box="[151,748,1255,1282]" pageId="14" pageNumber="455">
<emphasis id="B926EAEF522CE34A57BEA9DFFE68DC58" bold="true" box="[151,456,1255,1281]" italics="true" pageId="14" pageNumber="455">Paulodora drepanophora</emphasis>
<bibRefCitation id="EFC34B0C522CE34A56E6A9D0FD4CDC5B" author="Artois" box="[463,748,1256,1282]" pageId="14" pageNumber="455" refString="Artois, T. J. &amp; Tessens, B. S. (2008) Polycystididae (Rhabditophora: Rhabdocoela: Kalyptorhynchia) from the Indian Ocean, with the description of twelve new species. Zootaxa, 1849, 1 - 27." type="journal article" year="2008">Artois &amp; Tessens, 2008</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BED36FD522CE34A57BEA832FEBEDC7A" blockId="14.[151,748,1255,1315]" box="[151,286,1290,1315]" pageId="14" pageNumber="455">
(
<figureCitation id="13692A78522CE34A57B6A832FF42DC7A" box="[159,226,1290,1315]" captionStart="FIGURE 6" captionStartId="15.[151,250,1078,1100]" captionTargetBox="[288,1305,187,1062]" captionTargetId="figure@15.[278,1308,181,1064]" captionTargetPageId="15" captionText="FIGURE 6. Paulodora contortoides Artois &amp; Tessens, 2008. A. Prostate stylet type I. Paulodora drepanophora Artois &amp; Tessens, 2008. B. Prostate stylet type I. Phonorhynchoides gondwanae n. sp. C. umbrella-shaped &quot; nozzle &quot; D. Habitus of freeswimming animal. E. Caudal body end with the atrial organs (from a live specimen). F. Accessory stylet type IV (from the holotype). G. Prostate stylet type IV (from the holotype)." httpUri="https://zenodo.org/record/376911/files/figure.png" pageId="14" pageNumber="455">Fig. 6</figureCitation>
BC)
</paragraph>
</subSubSection>
<subSubSection id="C3486576522CE34A57BEA86AFBB6DF1A" pageId="14" pageNumber="455" type="description">
<paragraph id="8BED36FD522CE34A57BEA86AFF76DCA2" blockId="14.[151,1436,1362,2035]" pageId="14" pageNumber="455">
<materialsCitation id="3B3A3CA0522CE34A57BEA86AFB4EDCEA" ID-GBIF-Occurrence="1439511194" collectingDate="2009-12-05" county="Mission Rocks" latitude="-28.280666" location="New" longLatPrecision="1" longitude="32.485138" municipality="Eastern Shores" pageId="14" pageNumber="455" specimenCount="1">
<emphasis id="B926EAEF522CE34A57BEA86AFEE4DC32" bold="true" box="[151,324,1362,1387]" pageId="14" pageNumber="455">
<location id="8E8D6026522CE34A57BEA86AFF6CDC32" LSID="urn:lsid:plazi:treatment:03FB87EB522CE34A57BEA9DFFE6DDEAA:8E8D6026522CE34A57BEA86AFF6CDC32" box="[151,204,1362,1387]" county="Mission Rocks" latitude="-28.280666" longLatPrecision="1" longitude="32.485138" municipality="Eastern Shores" name="New" pageId="14" pageNumber="455">New</location>
localities.
</emphasis>
iSimangaliso
<location id="8E8D6026522CE34A56CEA86AFD23DC32" LSID="urn:lsid:plazi:treatment:03FB87EB522CE34A57BEA9DFFE6DDEAA:8E8D6026522CE34A56CEA86AFD23DC32" box="[487,643,1362,1387]" county="Mission Rocks" latitude="-28.280666" longLatPrecision="1" longitude="32.485138" municipality="Eastern Shores" name="Wetland Park" pageId="14" pageNumber="455">Wetland Park</location>
,
<collectingMunicipality id="6B89AC87522CE34A55A6A86BFC9BDC32" box="[655,827,1362,1387]" pageId="14" pageNumber="455">Eastern Shores</collectingMunicipality>
,
<collectingCounty id="628C4E71522CE34A5461A86AFC51DC32" box="[840,1009,1362,1387]" pageId="14" pageNumber="455">Mission Rocks</collectingCounty>
(
<geoCoordinate id="EE66503A522CE34A532AA86AFB03DC32" box="[1027,1187,1362,1387]" direction="south" orientation="latitude" pageId="14" pageNumber="455" precision="1" value="-28.280666">28°1650.4S</geoCoordinate>
,
<geoCoordinate id="EE66503A522CE34A5399A86AFAF2DC32" box="[1200,1362,1362,1387]" direction="east" orientation="longitude" pageId="14" pageNumber="455" precision="1" value="32.485138">32°2906.5E</geoCoordinate>
), sand and shrubby red algae from rock pool in the mid-eulittoral of a highly-exposed tidal area with sandstone terrace covered with invertebrates (barnacles, mussels, limpets, corals, tunicates),
<date id="FFEC103D522CE34A5339A8A2FB4EDCEA" box="[1040,1262,1434,1459]" pageId="14" pageNumber="455" value="2009-12-05">
<collectingDate id="EFA8E9D5522CE34A5339A8A2FB4EDCEA" box="[1040,1262,1434,1459]" pageId="14" pageNumber="455" value="2009-12-05">December 5, 2009</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3A3CA0522CE34A5228A8A3FF73DCA2" ID-GBIF-Occurrence="1439511185" collectingDate="2009-12-12" county="Mission Rocks" latitude="-28.278194" location="New" longLatPrecision="1" longitude="32.48628" municipality="Eastern Shores" pageId="14" pageNumber="455" specimenCount="1">
same locality (
<geoCoordinate id="EE66503A522CE34A57B6A886FE9BDC8E" box="[159,315,1470,1495]" direction="south" orientation="latitude" pageId="14" pageNumber="455" precision="1" value="-28.278194">28°1641.5&quot;S</geoCoordinate>
,
<geoCoordinate id="EE66503A522CE34A566FA886FE43DC8E" box="[326,483,1470,1495]" direction="east" orientation="longitude" pageId="14" pageNumber="455" precision="1" value="32.48628">32°2910.6&quot;E</geoCoordinate>
), red algae from a swirl hole close to the footpath from the parking lot,
<date id="FFEC103D522CE34A53D6A886FF73DCA2" pageId="14" pageNumber="455" value="2009-12-12">
<collectingDate id="EFA8E9D5522CE34A53D6A886FF73DCA2" pageId="14" pageNumber="455" value="2009-12-12">December 12, 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BED36FD522CE34A57EEAB3EFBB6DF1A" blockId="14.[151,1436,1362,2035]" pageId="14" pageNumber="455">
<materialsCitation id="3B3A3CA0522CE34A57EEAB3EFF7BDF1A" ID-GBIF-Occurrence="1439511195" collectorName="De Vocht" country="Kenya" location="Mombasa" pageId="14" pageNumber="455" specimenCount="1" stateProvince="Mombasa">
<emphasis id="B926EAEF522CE34A57EEAB3EFE1BDF46" bold="true" box="[199,443,1542,1567]" pageId="14" pageNumber="455">Known distribution.</emphasis>
<collectingCountry id="F345766D522CE34A56EDAB3FFDB0DF46" box="[452,528,1543,1567]" name="Kenya" pageId="14" pageNumber="455">Kenya</collectingCountry>
,
<collectingRegion id="4996F81F522CE34A5534AB3EFD2EDF46" box="[541,654,1542,1567]" country="Kenya" name="Mombasa" pageId="14" pageNumber="455">Mombasa</collectingRegion>
, McKenzie Point &amp; Tiwi (
<bibRefCitation id="EFC34B0C522CE34A5496AB3EFB6CDF46" author="Jouk" box="[959,1228,1542,1567]" pageId="14" pageNumber="455" refString="Jouk, P. E. H. &amp; De Vocht, A. J. - P. (1989) Kalyptorhynchia (Plathelminthes Rhabdocoela) from the Kenyan Coast, with descriptions of four new species. Tropical Zoology, 2, 145 - 157." type="journal article" year="1989">
Jouk &amp;
<collectorName id="26A7532B522CE34A5334AB3FFB27DF46" box="[1053,1159,1542,1567]" pageId="14" pageNumber="455">De Vocht</collectorName>
1989
</bibRefCitation>
;
<bibRefCitation id="EFC34B0C522CE34A53F1AB3EFF74DF1A" author="Artois" pageId="14" pageNumber="455" refString="Artois, T. J. &amp; Tessens, B. S. (2008) Polycystididae (Rhabditophora: Rhabdocoela: Kalyptorhynchia) from the Indian Ocean, with the description of twelve new species. Zootaxa, 1849, 1 - 27." type="journal article" year="2008">Artois &amp; Tessens 2008</bibRefCitation>
)
</materialsCitation>
.
<materialsCitation id="3B3A3CA0522CE34A57CFAB12FBB6DF1A" ID-GBIF-Occurrence="1439511187" box="[230,1046,1578,1603]" country="Somalia" location="Mogadiscio" pageId="14" pageNumber="455" specimenCount="1">
<collectingCountry id="F345766D522CE34A57CFAB12FEE5DF1A" box="[230,325,1578,1603]" name="Somalia" pageId="14" pageNumber="455">Somalia</collectingCountry>
, north of
<location id="8E8D6026522CE34A5699AB12FD97DF1A" LSID="urn:lsid:plazi:treatment:03FB87EB522CE34A57BEA9DFFE6DDEAA:8E8D6026522CE34A5699AB12FD97DF1A" box="[432,567,1578,1603]" country="Somalia" name="Mogadiscio" pageId="14" pageNumber="455">Mogadiscio</location>
, Hawadli and Warshek (
<bibRefCitation id="EFC34B0C522CE34A5461AB12FBA9DF1A" author="Schockaert" box="[840,1033,1578,1603]" pageId="14" pageNumber="455" refString="Schockaert, E. R. (1982) Turbellaria from Somalia 2. Kalyptorhynchia (Part 2). Monitore Zoologico Italiano, New Series, 18 (Supplement), 81 - 96." type="journal article" year="1982">Schockaert 1982</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3486576522CE34A57EEAB76FE6DDEAA" pageId="14" pageNumber="455" type="materials_examined">
<paragraph id="8BED36FD522CE34A57EEAB76FD85DFD2" blockId="14.[151,1436,1362,2035]" pageId="14" pageNumber="455">
<materialsCitation id="3B3A3CA0522CE34A57EEAB76FD85DFD2" ID-GBIF-Occurrence="1439511197" country="South Africa" location="All" pageId="14" pageNumber="455" specimenCount="1" typeStatus="holotype">
<emphasis id="B926EAEF522CE34A57EEAB76FE96DF3E" bold="true" box="[199,310,1614,1639]" pageId="14" pageNumber="455">Material.</emphasis>
Two whole mounts from
<collectingCountry id="F345766D522CE34A5577AB76FD4FDF3E" box="[606,751,1614,1639]" name="South Africa" pageId="14" pageNumber="455">South Africa</collectingCountry>
(HU, nos VII.3.49VII.3.50).
<location id="8E8D6026522CE34A5365AB76FBD0DF3E" LSID="urn:lsid:plazi:treatment:03FB87EB522CE34A57BEA9DFFE6DDEAA:8E8D6026522CE34A5365AB76FBD0DF3E" box="[1100,1136,1614,1639]" country="South Africa" name="All" pageId="14" pageNumber="455">All</location>
material from the original description, including the
<typeStatus id="54E9885F522CE34A5692AB4AFD81DFD2" box="[443,545,1650,1675]" pageId="14" pageNumber="455" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
<paragraph id="8BED36FD522CE34A57EEABAEFE6DDEAA" blockId="14.[151,1436,1362,2035]" pageId="14" pageNumber="455">
<emphasis id="B926EAEF522CE34A57EEABAEFE1BDFF6" bold="true" box="[199,443,1686,1711]" pageId="14" pageNumber="455">Additional remarks.</emphasis>
Habitus and general organisation do not differ from that of other species of
<taxonomicName id="4C524D7E522CE34A520DABAEFE96DF8A" authority="Marcus, 1948" authorityName="Marcus" authorityYear="1948" class="Rhabditophora" family="Polycystididae" genus="Paulodora" kingdom="Animalia" order="Rhabdocoela" pageId="14" pageNumber="455" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAEF522CE34A520DABAEFA3CDFF6" box="[1316,1436,1686,1711]" italics="true" pageId="14" pageNumber="455">Paulodora</emphasis>
<bibRefCitation id="EFC34B0C522CE34A57BEAB83FE96DF8A" author="Marcus" box="[151,310,1723,1747]" pageId="14" pageNumber="455" refString="Marcus, E. (1948) Turbellaria do Brasil. Boletins da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo, 13, 111 - 243." type="journal article" year="1948">Marcus, 1948</bibRefCitation>
</taxonomicName>
, a monophyletic taxon to which it undoubtedly belongs (see
<bibRefCitation id="EFC34B0C522CE34A5328AB83FB44DF8A" author="Tessens" box="[1025,1252,1722,1747]" pageId="14" pageNumber="455" refString="Tessens, B., Janssen, T. &amp; Artois, T. (2014) Molecular phylogeny of Kalyptorhynchia (Rhabdocoela, Platyhelminthes) inferred from ribosomal sequence data. Zoologica Scripta, 43, 519 - 530." type="journal article" year="2014">
Tessens
<emphasis id="B926EAEF522CE34A5348AB84FB3CDF8A" box="[1121,1180,1722,1747]" italics="true" pageId="14" pageNumber="455">et al.</emphasis>
2014
</bibRefCitation>
). The umbrellashaped, sclerotic &quot;nozzles&quot; on each of the oviducts typical of all species of
<taxonomicName id="4C524D7E522CE34A54D2ABE6FBD3DFAE" box="[1019,1139,1758,1783]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" kingdom="Animalia" order="Rhabdocoela" pageId="14" pageNumber="455" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAEF522CE34A54D2ABE6FBD3DFAE" box="[1019,1139,1758,1783]" italics="true" pageId="14" pageNumber="455">Paulodora</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC34B0C522CE34A539BABE6FF70DE42" author="Artois" pageId="14" pageNumber="455" refString="Artois, T. J. &amp; Schockaert, E. R. (1998) A cladistic re-assessment of the Polycystis species complex (Polycystididae, Eukalyptorhynchia). Hydrobiologia, 383, 97 - 102." type="journal article" year="1998">Artois &amp; Schockaert 1998</bibRefCitation>
) are clearly visible on live specimens as well as on whole mounts (
<figureCitation id="13692A78522CE34A54E2AA3AFBB0DE42" box="[971,1040,1794,1819]" captionStart="FIGURE 6" captionStartId="15.[151,250,1078,1100]" captionTargetBox="[288,1305,187,1062]" captionTargetId="figure@15.[278,1308,181,1064]" captionTargetPageId="15" captionText="FIGURE 6. Paulodora contortoides Artois &amp; Tessens, 2008. A. Prostate stylet type I. Paulodora drepanophora Artois &amp; Tessens, 2008. B. Prostate stylet type I. Phonorhynchoides gondwanae n. sp. C. umbrella-shaped &quot; nozzle &quot; D. Habitus of freeswimming animal. E. Caudal body end with the atrial organs (from a live specimen). F. Accessory stylet type IV (from the holotype). G. Prostate stylet type IV (from the holotype)." httpUri="https://zenodo.org/record/376911/files/figure.png" pageId="14" pageNumber="455">Fig. 6</figureCitation>
C). The prostate stylet
<typeStatus id="54E9885F522CE34A523BAA3BFAE2DE42" box="[1298,1346,1795,1819]" pageId="14" pageNumber="455">type</typeStatus>
I of the
<collectingRegion id="4996F81F522CE34A57BEAA1EFF78DE66" box="[151,216,1830,1855]" country="Tanzania" name="Kusini Unguja" pageId="14" pageNumber="455">South</collectingRegion>
African specimens is 6467 µm long (n = 2), which is only marginally longer than the known range (4262 µm;
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). In contrast with the specimens from the original description, the distal part of the stylet is not folded downwards. Therefore, the sickle-shape is less obvious. This folding is probably an artefact caused by the degree of squeezing of the specimen and also occurs in species with similar stylet morphology, such as
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<emphasis id="B926EAEF522CE34A579FAA8EFEC4DE96" box="[182,356,1974,1999]" italics="true" pageId="14" pageNumber="455">Paulodora felis</emphasis>
(
<bibRefCitation id="EFC34B0C522CE34A565AAA8FFDB0DE96" author="Marcus" box="[371,528,1975,1999]" pageId="14" pageNumber="455" refString="Marcus, E. (1948) Turbellaria do Brasil. Boletins da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo, 13, 111 - 243." type="journal article" year="1948">Marcus, 1948</bibRefCitation>
)
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</taxonomicName>
and
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<emphasis id="B926EAEF522CE34A54AFAA8FFB91DE97" box="[902,1073,1975,1998]" italics="true" pageId="14" pageNumber="455">P. asymmetrica</emphasis>
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</taxonomicName>
(see
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).
</paragraph>
</subSubSection>
</treatment>
</document>