180 lines
21 KiB
XML
180 lines
21 KiB
XML
<document id="7894C6EB7FD7D334BD69106BF654DE4E" ID-DOI="10.11646/zootaxa.3857.2.5" ID-GBIF-Dataset="4ad4cc3b-5758-4f51-beab-68b53c9d25fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="229668" ID-ZooBank="32D449D9-0E04-4848-A3F9-88FB0FC24025" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460897827241" checkinUser="plazi" docAuthor="García-Romera, Carlos & Barrientos, Jose Antonio" docDate="2014" docId="03FB5521FFD33D08FF09FE31EF7864F8" docLanguage="en" docName="zt03857p260.pdf" docOrigin="Zootaxa 3857 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia arbuciensis Garcia-Romera, sp.nov." docType="treatment" docVersion="7" lastPageNumber="246" masterDocId="FFC22D59FFD23D0AFF9EFFCEEB116008" masterDocTitle="Nine new species of Phoridae (Insecta: Diptera) from Spain" masterLastPageNumber="260" masterPageNumber="244" pageNumber="245" updateTime="1698360198051" updateUser="plazi">
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<mods:titleInfo id="356634F53E067DE3D387D598ED0C87D3">
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<mods:title id="33C0EC49937C61CC6AC16B9E1781ABAD">Nine new species of Phoridae (Insecta: Diptera) from Spain</mods:title>
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<mods:name id="FB50846E1A0AB3CA74DFC18BC5B27E2C" type="personal">
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<mods:namePart id="F8AD5B6649D33A6C36917174DBF3A9ED">García-Romera, Carlos</mods:namePart>
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<mods:roleTerm id="0F7F41EAD160030C3E8D615EC1ED4484">Author</mods:roleTerm>
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<mods:namePart id="526DC413872BE44092C1F04D69EF4ED4">Barrientos, Jose Antonio</mods:namePart>
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<mods:typeOfResource id="093E932A69A36D70C74DE359A3411454">text</mods:typeOfResource>
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<mods:title id="4E0941AB5E2F001621B1C1DCFD9D6214">Zootaxa</mods:title>
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<mods:part id="D49ECF434FAC56767D8A6AFF84657D8C">
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<mods:date id="8F46FB3ADE0E18A4ADEED3CDB3C48692">2014</mods:date>
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<mods:number id="6A7FC1ABB9A73FDCC0C17A667C907509">3857</mods:number>
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<mods:number id="677678C809A3247655BCAEA3D34E946F">2</mods:number>
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<mods:identifier id="FA4433D39E1C11878064E8B2334FE6F7" type="DOI">10.11646/zootaxa.3857.2.5</mods:identifier>
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<treatment id="03FB5521FFD33D08FF09FE31EF7864F8" ID-DOI="http://doi.org/10.5281/zenodo.6136521" ID-GBIF-Taxon="119587654" ID-Zenodo-Dep="6136521" LSID="urn:lsid:plazi:treatment:03FB5521FFD33D08FF09FE31EF7864F8" httpUri="http://treatment.plazi.org/id/03FB5521FFD33D08FF09FE31EF7864F8" lastPageId="2" lastPageNumber="246" pageId="1" pageNumber="245">
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<subSubSection id="C348B7BCFFD33D0BFF09FE31EA1A6233" pageId="1" pageNumber="245" type="nomenclature">
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<paragraph id="8BEDE437FFD33D0BFF09FE31E9DC6212" blockId="1.[151,717,511,571]" box="[151,717,511,538]" pageId="1" pageNumber="245">
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<heading id="D0A5535BFFD33D0BFF09FE31E9DC6212" bold="true" box="[151,717,511,538]" fontSize="11" level="1" pageId="1" pageNumber="245" reason="1">
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<emphasis id="B9263825FFD33D0BFF09FE31E9DC6212" bold="true" box="[151,717,511,538]" pageId="1" pageNumber="245">
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<taxonomicName id="4C529FB4FFD33D0BFF09FE31E9606212" ID-CoL="8CS4W" authority="Garcia-Romera" authorityName="Garcia-Romera" box="[151,625,511,538]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="245" phylum="Arthropoda" rank="species" species="arbuciensis" status="sp. nov.">
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<emphasis id="B9263825FFD33D0BFF09FE31EAB56211" bold="true" box="[151,420,511,537]" italics="true" pageId="1" pageNumber="245">Megaselia arbuciensis</emphasis>
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García-Romera
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</taxonomicName>
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<taxonomicNameLabel id="A215855EFFD33D0BFDE9FDCEE9DC6212" box="[631,717,512,538]" pageId="1" pageNumber="245" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF09FDECEA1A6233" blockId="1.[151,717,511,571]" box="[151,267,546,571]" pageId="1" pageNumber="245">
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(
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<figureCitation id="1369F8B2FFD33D0BFF01FDECEA106233" box="[159,257,546,571]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,751,774]" captionTargetBox="[158,1428,203,723]" captionTargetId="figure@2.[151,1436,193,730]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Megaselia arbuciensis García-Romera sp. nov. male. 1, left face of hypopygium. 2, right face of hypopygium. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/229669/files/figure.png" pageId="1" pageNumber="245">Figs 1–2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C348B7BCFFD33D0BFF09FDA4E816628B" box="[151,775,618,643]" pageId="1" pageNumber="245" type="etymology">
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<paragraph id="8BEDE437FFD33D0BFF09FDA4E816628B" blockId="1.[151,1437,618,2014]" box="[151,775,618,643]" pageId="1" pageNumber="245">
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<emphasis id="B9263825FFD33D0BFF09FDA4EA0F628B" bold="true" box="[151,286,618,643]" pageId="1" pageNumber="245">Etymology.</emphasis>
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The species is named after its
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<typeStatus id="54E95A95FFD33D0BFDE8FDA5E9B7628B" box="[630,678,619,643]" pageId="1" pageNumber="245">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348B7BCFFD33D0BFF59FD40EA5063CF" pageId="1" pageNumber="245" type="diagnosis">
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<paragraph id="8BEDE437FFD33D0BFF59FD40EA5063CF" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">
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<emphasis id="B9263825FFD33D0BFF59FD40EA5062AF" bold="true" box="[199,321,654,679]" pageId="1" pageNumber="245">Diagnosis.</emphasis>
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Longest bristle on palp clearly longer than maximum width of palp; mesopleuron bare; scutellum with a posterior pair of bristles and an anterior pair of hairs; notopleuron with three bristles; abdominal venter with hairs on segments 3–6, with those on sixth segment weak; left side of epandrium with hairs which are more robust than hairs of cerci; posterior left lobe of hypandrium developed; anal tube shorter than length of dorsal face of epandrium; bristles at rear margin of sixth abdominal tergite subequal to most robust hairs on left side of epandrium; all femora brown; fore tarsus uniformly brown, with fifth segment being a little narrower than fourth one; longest hairs below basal half of hind femur about as long as those of anteroventral row of outer third; knob of haltere brown; with five axillary bristles; costal index less than 0.44; costal cilia (section 3) about as long as longest axillary bristle.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348B7BCFFD33D08FF59FC1CE95E648C" lastPageId="2" lastPageNumber="246" pageId="1" pageNumber="245" type="description">
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<paragraph id="8BEDE437FFD33D0BFF59FC1CEA0F6527" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">
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<emphasis id="B9263825FFD33D0BFF59FC1CEABD63E3" bold="true" box="[199,428,978,1003]" pageId="1" pageNumber="245">Description (male).</emphasis>
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Head. Frons brown, about as broad as long, with 113 hairs. Upper supra-antennal bristles about as long and strong as lower pair, and they are about as closer together as pre-ocellars, which are slightly lower on frons than mediolaterals and clearly closer together than either is from a mediolateral bristle. Upper supraantennal bristles at about the same level on frons as antials, which are lower on frons than anterolaterales, and about as far apart from these as from upper supra-antennal bristles. Gena with three long bristles and five shorter ones. Antennal postpedicel subglobose, brown and relatively small (diameter
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<quantity id="4CAA49D2FFD33D0BFB91FB49EF686497" box="[1039,1145,1159,1183]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="245" unit="mm" value="0.1">0.10 mm</quantity>
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), with SPS vesicles, each about diameter of basal socket of antial bristles. Palps dusky yellow,
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<quantity id="4CAA49D2FFD33D0BFC39FB65EF5D64CA" box="[935,1100,1195,1219]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="245" unit="mm" value="0.135" valueMax="0.17" valueMin="0.1">0.10–0.17 mm</quantity>
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long and
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<quantity id="4CAA49D2FFD33D0BFB20FB64EE3264CA" box="[1214,1315,1194,1219]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="245" unit="mm" value="0.05">0.05 mm</quantity>
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maximum width, with five bristles, the longest bristle being longer than greatest width of palp. Labella enlarged, yellow, with only a few short spinules below and a few long hairs. Labrum brown, about 0.85 × as wide as an antennal postpedicel.
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</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF59FAF4E8E46593" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">Thorax. Brown. Mesopleuron bare. Three notopleural bristles and no cleft in front of these. Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Propleura with 3–4 hairs on posterodorsal edge, 3–4 on posteroventral one and a few in middle, situated well forward of hindmargin.</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF59FA68EA26660F" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">Abdomen. Tergites light brown, with scattered short hairs with those of the rear margin of T6 only a little longer than the rest and about as long and strong as those on epandrium. Venter grey, with a few short hairs on segments 3–6.</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF59F9DCEF8E667B" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">
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<emphasis id="B9263825FFD33D0BFF59F9DCEA726623" bold="true" box="[199,355,1554,1579]" pageId="1" pageNumber="245">Hypopygium</emphasis>
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(
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<figureCitation id="1369F8B2FFD33D0BFEECF9DCEAC76623" box="[370,470,1554,1579]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,751,774]" captionTargetBox="[158,1428,203,723]" captionTargetId="figure@2.[151,1436,193,730]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Megaselia arbuciensis García-Romera sp. nov. male. 1, left face of hypopygium. 2, right face of hypopygium. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/229669/files/figure.png" pageId="1" pageNumber="245">Figs 1–2</figureCitation>
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). Left side of epandrium light brown, without bristles, with hairs stronger than those on cerci, which are weaker than those on proctiger. Anal tube yellow, a little shorter than dorsal face of epandrium. Hypandrium, light brown, with both posterior lobes developed and with minute hairs below.
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</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF59F9B1EA286743" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">
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Legs. Light brown, slightly clearing from the posterior to anterior legs, but without being yellowish. Front legs with a posterodorsal hair palisade on tarsal segments 1–4 and metatarsus a little enlarged. Length ratios of the fore tarsal segments 2.3:1.3:1.2:0.9–1:1, with fifth segment narrower than fourth one. Mid legs with a dorsal hair palisade on tibia extends about 2/3 of its length. Longest hairs below basal half of hind femur about as long as those of anteroventral row of outer third.
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<collectingCountry id="F345A4A7FFD33D0BFDB4F8C0E973672F" box="[554,610,1806,1831]" name="India" pageId="1" pageNumber="245">Hind</collectingCountry>
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tibia with 15–18 differentiated posterodorsal hairs and spinules of apical combs simple.
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</paragraph>
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<paragraph id="8BEDE437FFD33D0BFF59F898EA8C67D6" blockId="1.[151,1437,618,2014]" pageId="1" pageNumber="245">
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Wing. Length
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<quantity id="4CAA49D2FFD33D0BFEF4F898E9016766" box="[362,528,1878,1903]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" metricValueMax="1.76" metricValueMin="1.7" pageId="1" pageNumber="245" unit="mm" value="1.73" valueMax="1.76" valueMin="1.7">1.70–1.76 mm</quantity>
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. Membrane lightly tinged grey and veins light brown; with five axillary bristles. Subcosta (Sc) not reaching R1. One hair present at base of vein Rs,
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<quantity id="4CAA49D2FFD33D0BFC14F8B4EF5B679A" box="[906,1098,1914,1939]" metricMagnitude="-5" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="245" unit="mm" value="0.0325" valueMax="0.04" valueMin="0.025">0.025–0.040 mm</quantity>
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long. Costal index 0.35–0.40; costal ratios 3.5–3.8:1–1.2:1. Costal cilia (section 3)
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<quantity id="4CAA49D2FFD33D0BFD70F86CE88367B1" box="[750,914,1954,1978]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="1" pageNumber="245" unit="mm" value="0.125" valueMax="0.13" valueMin="0.12">0.12–0.13 mm</quantity>
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long, about as long as longest axillary bristle. Knob of haltere brown.
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</paragraph>
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<caption id="DF2DB4BFFFD03D08FF09FD21EA7F632C" httpUri="https://zenodo.org/record/229669/files/figure.png" pageId="2" pageNumber="246" targetBox="[158,1428,203,723]" targetPageId="2">
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<paragraph id="8BEDE437FFD03D08FF09FD21EA7F632C" blockId="2.[151,1437,751,804]" pageId="2" pageNumber="246">
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<emphasis id="B9263825FFD03D08FF09FD21EA2B630C" bold="true" box="[151,314,751,774]" pageId="2" pageNumber="246">FIGURES 1–2.</emphasis>
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<taxonomicName id="4C529FB4FFD03D08FEDEFD21E9D6630D" authority="Garcia-Romera" authorityName="Garcia-Romera" box="[320,711,751,773]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="246" phylum="Arthropoda" rank="species" species="arbuciensis" status="sp. nov.">
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<emphasis id="B9263825FFD03D08FEDEFD21E934630D" box="[320,549,751,773]" italics="true" pageId="2" pageNumber="246">Megaselia arbuciensis</emphasis>
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García-Romera
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</taxonomicName>
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<emphasis id="B9263825FFD03D08FD53FD3EE80C630D" bold="true" box="[717,797,752,773]" pageId="2" pageNumber="246">
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<taxonomicNameLabel id="A215855EFFD03D08FD53FD3EE80C630D" box="[717,797,752,773]" pageId="2" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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male.
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<emphasis id="B9263825FFD03D08FCFFFD21E87F630C" bold="true" box="[865,878,751,772]" pageId="2" pageNumber="246">1</emphasis>
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, left face of hypopygium.
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<emphasis id="B9263825FFD03D08FBE5FD21EF99630C" bold="true" box="[1147,1160,751,772]" pageId="2" pageNumber="246">2</emphasis>
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, right face of hypopygium. Scale bars = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BEDE437FFD03D08FF59FC85E95E648C" blockId="2.[151,1436,843,1264]" pageId="2" pageNumber="246">
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<emphasis id="B9263825FFD03D08FF59FC85EA4C636C" bold="true" box="[199,349,843,868]" pageId="2" pageNumber="246">Recognition.</emphasis>
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In the keys to the males of the British species of
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<taxonomicName id="4C529FB4FFD03D08FC08FC85EFAA636C" authority="Disney, 1989" authorityName="Disney" authorityYear="1989" box="[918,1211,843,868]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="246" phylum="Arthropoda" rank="genus">
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<emphasis id="B9263825FFD03D08FC08FC85EF1A636C" box="[918,1035,843,868]" italics="true" pageId="2" pageNumber="246">Megaselia</emphasis>
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(
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<bibRefCitation id="EFC399C6FFD03D08FB85FC85EFA2636C" author="Disney" box="[1051,1203,843,868]" pageId="2" pageNumber="246" refString="Disney, R. H. L. (1989) Scuttle flies. Diptera, Phoridae, genus Megaselia. Handbooks for the Identification of British Insects, 10 (8), 1 - 155." type="journal article" year="1989">Disney, 1989</bibRefCitation>
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)
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</taxonomicName>
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this species runs to couplet 87, after return of 182, to
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<taxonomicName id="4C529FB4FFD03D08FD90FCBEE8106380" authority="Wood" authorityName="Wood" box="[526,769,879,904]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="246" phylum="Arthropoda" rank="species" species="conformis">
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<emphasis id="B9263825FFD03D08FD90FCBEE9B26380" box="[526,675,880,904]" italics="true" pageId="2" pageNumber="246">M. conformis</emphasis>
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(Wood)
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</taxonomicName>
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, but its hypopygium clearly differs from the new species, in the shape of epandrium and development of posterior lobe of hypandrium; the subsequently added
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<taxonomicName id="4C529FB4FFD03D08FB65FC5BEA1B63D8" authority="Lundbeck" authorityName="Lundbeck" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="246" phylum="Arthropoda" rank="species" species="lactipennis">
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<emphasis id="B9263825FFD03D08FB65FC5BEE8D63A4" box="[1275,1436,915,940]" italics="true" pageId="2" pageNumber="246">M. lactipennis</emphasis>
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Lundbeck
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</taxonomicName>
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, runs to the same point; it differs from the new species clearly in the hypopygium (development of posterior lobes of hypandrium, shape of epandrium and thickness of hairs of epandrium and proctiger), in length of hairs of the rear margin of tergite 6, length of the hairs below basal half of hind femur and length of costal cilia (Disney, 2000). The subsequently described
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<taxonomicName id="4C529FB4FFD03D08FD19FBEBE87A6434" authority="Disney" authorityName="Disney" box="[647,875,1059,1084]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="246" phylum="Arthropoda" rank="species" species="putescavi">
|
||
<emphasis id="B9263825FFD03D08FD19FBEBE8076434" box="[647,790,1061,1084]" italics="true" pageId="2" pageNumber="246">M. putescavi</emphasis>
|
||
Disney
|
||
</taxonomicName>
|
||
runs to this couplet, but it is distinguised from the new species by having hairs on mesopleuron and a clearly different hypopygium, especially a larger left lobe of hypandrium (
|
||
<bibRefCitation id="EFC399C6FFD03D08FEB3FBA5E952648C" author="Disney" box="[301,579,1131,1156]" pageId="2" pageNumber="246" refString="Disney, R. H. L. & Withers, P. (2011) Scuttle flies (Diptera, Phoridae) reared from tree rotholes in France, including three new species of Megaselia Rondani. Fragmenta Faunistica, 54 (1), 29 - 41. http: // dx. doi. org / 10.3161 / 00159301 ff 2011.54.1.029" type="journal article" year="2011">Disney & Withers, 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348B7BCFFD03D08FF09FB41EF7864F8" pageId="2" pageNumber="246" type="materials_examined">
|
||
<paragraph id="8BEDE437FFD03D08FF09FB41EF8064C4" blockId="2.[151,1436,843,1264]" pageId="2" pageNumber="246">
|
||
<emphasis id="B9263825FFD03D08FF09FB41E91864A0" bold="true" box="[151,521,1167,1192]" pageId="2" pageNumber="246">
|
||
Material examined.
|
||
<typeStatus id="54E95A95FFD03D08FE02FB41E91864A0" box="[412,521,1167,1192]" pageId="2" pageNumber="246" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
♂
|
||
<collectingCountry id="F345A4A7FFD03D08FDDCFB41E99464A0" box="[578,645,1167,1192]" name="Spain" pageId="2" pageNumber="246">Spain</collectingCountry>
|
||
, Girona: Arbucias, Montseny Natural Park, beech forest, UTM 31TDG540279, 1250 m a.s.l., 19/
|
||
<date id="FFECC2F7FFD03D08FD91FB7DE9A964C4" box="[527,696,1203,1228]" pageId="2" pageNumber="246" value="1990-06-05" valueMax="1990-06-02" valueMin="1990-06-05">05-02/06/1990</date>
|
||
. Window trap. J.A. Barrientos leg (
|
||
<collectionCode id="ED437CF2FFD03D08FBDAFB7AEF9464C4" box="[1092,1157,1204,1228]" httpUri="http://grbio.org/cool/0md6-ynwr" name="Universidad Autonoma de Barcelona" pageId="2" pageNumber="246">UAB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEDE437FFD03D08FF59FB19EF7864F8" blockId="2.[151,1436,843,1264]" box="[199,1129,1239,1264]" pageId="2" pageNumber="246">
|
||
<emphasis id="B9263825FFD03D08FF59FB19EA2564F8" bold="true" box="[199,308,1239,1264]" pageId="2" pageNumber="246">
|
||
<typeStatus id="54E95A95FFD03D08FF59FB19EA2564F8" box="[199,308,1239,1264]" pageId="2" pageNumber="246" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
♂, same date, locality, habitat and capture method as
|
||
<typeStatus id="54E95A95FFD03D08FC0FFB19E8E264F8" box="[913,1011,1239,1264]" pageId="2" pageNumber="246" type="holotype">holotype</typeStatus>
|
||
(CCG-R).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |