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<document id="F8A8369EC6DB7658B7526A60EFE313E4" ID-DOI="10.5852/ejt.2024.921.2409" ID-GBIF-Dataset="b7115915-f7fd-4a41-8495-9e38bd514ad9" ID-ISSN="2118-9773" ID-Zenodo-Dep="10604105" ID-ZooBank="45B2A9CB-27F0-46B2-85F0-264F963B5D7E" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1706777316154" checkinUser="plazi" docAuthor="Pereira, Maria Paula, Gallão, Jonas E., Bichuette, Maria E. &amp; Pérez-González, Abel" docDate="2024" docId="03FB24562520112DFDC056818525FC27" docLanguage="en" docName="EJT.2024.921.36-63.pdf" docOrigin="European Journal of Taxonomy 921" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2409/10653" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Spaeleoleptes spaeleus H. Soares 1966" docType="treatment" docVersion="5" lastPageNumber="49" masterDocId="FFC25C2E25251120FFF0505A877CFFF8" masterDocTitle="Hidden in the caves: a new troglobitic species of Spaeleoleptes and the type species redescription (Opiliones, Laniatores)" masterLastPageNumber="63" masterPageNumber="36" pageNumber="41" updateTime="1707125972181" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D803B24F6F1E6726660544817C770AB8">Hidden in the caves: a new troglobitic species of Spaeleoleptes and the type species redescription (Opiliones, Laniatores)</mods:title>
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<mods:namePart id="8DC2262EE24B86B57D7864961C3794CB">Pereira, Maria Paula</mods:namePart>
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<mods:namePart id="BF256F76774382A3A587E3ACCE7604B0">Gallão, Jonas E.</mods:namePart>
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<mods:affiliation id="366324678496A6AA82F5283B49EB0874">Laboratório de Estudos Subterrâneos, Departamento de Ecologia e Biologia Evolutiva, Universidade Federal de São Carlos, Rodovia Washington Luís km 235, São Carlos, Brazil.</mods:affiliation>
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<mods:namePart id="84D06ED1A008DBE74F79B55D8D538D79">Bichuette, Maria E.</mods:namePart>
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<mods:affiliation id="4A9845FB75EE45473E226FA21A7C2D33">Laboratório de Estudos Subterrâneos, Departamento de Ecologia e Biologia Evolutiva, Universidade Federal de São Carlos, Rodovia Washington Luís km 235, São Carlos, Brazil.</mods:affiliation>
<mods:nameIdentifier id="AAB4F208BEA989D00CDCB022856F1D57" type="email">lina.cave@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2783FF9DA1C39D7C9BF5DBE7FC310D1B">Pérez-González, Abel</mods:namePart>
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<mods:date id="21A3F3D55617214A9DA578C175F30F26">2024</mods:date>
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<paragraph id="8BED954025201125FDC056818378F90D" blockId="5.[347,1241,1755,1852]" box="[560,1028,1755,1782]" pageId="5" pageNumber="41">
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<taxonomicName id="4C52EEC325201125FDC056818378F90D" ID-CoL="7WW8S" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[560,1028,1755,1782]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225201125FDC05681843DF90E" bold="true" box="[560,833,1755,1782]" italics="true" pageId="5" pageNumber="41">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B125201125FC9C56868378F90D" author="Soares" box="[876,1028,1755,1782]" firstAuthor="Soares" pageId="5" pageNumber="41" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
<uuid id="FFF4AF9525201125FD8B56A483A5F8E1" box="[635,1241,1790,1817]" pageId="5" pageNumber="41">AEA8884E-50DF-43AA-B4A3-C1AB8D4C0AB7</uuid>
</uri>
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<figureCitation id="136989C525201125FD1757788430F8C4" box="[743,844,1826,1852]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[189,233,1892,1918]" captionStartId-1="7.[189,231,1962,1988]" captionStartId-2="9.[189,232,1892,1918]" captionStartId-3="10.[189,231,1892,1918]" captionTargetBox-0="[189,1399,473,1834]" captionTargetBox-1="[189,1399,704,1914]" captionTargetBox-2="[189,1399,650,1852]" captionTargetBox-3="[313,1298,573,1821]" captionTargetId-0="figure-85@6.[189,1399,473,1834]" captionTargetId-1="figure-172@7.[189,1399,704,1914]" captionTargetId-2="figure-142@9.[189,1399,647,1852]" captionTargetId-3="figure-174@10.[263,1321,567,1835]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Fig. 3. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5. D. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium. E. Male pedipalp, mesal view." captionText-1="Fig. 4. Spaeleoleptes spaeleus H. Soares, 1966, female paratype (MZUSP 28462). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, left lateral view. D. Carapace, lateral view. E. Carapace, dorsal view." captionText-2="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." captionText-3="Fig. 6. Line drawings of male genitalia of Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Full genitalia, dorsal view. B. Detail of pars distalis, dorsal view. C. Detail of capsula interna, dorsal view. D. Detail of pars distalis, lateral view. E. Same, ventral view. Red lines: capsula interna. Blue lines: conductors. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10604113" figureDoi-1="http://doi.org/10.5281/zenodo.10604117" figureDoi-2="http://doi.org/10.5281/zenodo.10604119" figureDoi-3="http://doi.org/10.5281/zenodo.10604123" httpUri-0="https://zenodo.org/record/10604113/files/figure.png" httpUri-1="https://zenodo.org/record/10604117/files/figure.png" httpUri-2="https://zenodo.org/record/10604119/files/figure.png" httpUri-3="https://zenodo.org/record/10604123/files/figure.png" pageId="5" pageNumber="41">Figs 36</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BED954025201125FF4D5739840AF885" blockId="5.[189,886,1891,1918]" box="[189,886,1891,1918]" pageId="5" pageNumber="41">
<treatmentCitationGroup id="AB42B26E25201125FF4D5739840AF885" box="[189,886,1891,1918]" pageId="5" pageNumber="41">
<taxonomicName id="4C52EEC325201125FF4D573986B7F885" ID-CoL="7WW8S" authorityName="H. Soares" authorityYear="1966" box="[189,459,1891,1917]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225201125FF4D573986B7F885" box="[189,459,1891,1917]" italics="true" pageId="5" pageNumber="41">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
[sic] H.
<treatmentCitation id="0AF3B35125201125FDC157398478F885" author="Soares H. E. M." box="[561,772,1891,1917]" page="111" pageId="5" pageNumber="41" year="1966">
<bibRefCitation id="EFC3E8B125201125FDC157398478F885" author="Soares" box="[561,772,1891,1917]" firstAuthor="Soares" page="111" pageId="5" pageNumber="41" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966: 111</bibRefCitation>
</treatmentCitation>
, figs 79.
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</paragraph>
<paragraph id="8BED954025201125FF4D57FF86B5F81A" blockId="5.[189,1399,1957,2018]" pageId="5" pageNumber="41">
<treatmentCitationGroup id="AB42B26E25201125FF4D57FF86B5F81A" pageId="5" pageNumber="41">
<taxonomicName id="4C52EEC325201125FF4D57FF86ACF847" ID-CoL="7WW8S" authorityName="H. Soares" authorityYear="1966" box="[189,464,1957,1983]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225201125FF4D57FF86ACF847" box="[189,464,1957,1983]" italics="true" pageId="5" pageNumber="41">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
Gnaspini &amp; Trajano 1994: 582. —
<treatmentCitation id="0AF3B35125201125FC5F57FF839FF847" author="Pinto-da-Rocha R." box="[943,1251,1957,1984]" page="80" pageId="5" pageNumber="41" year="1995">
<bibRefCitation id="EFC3E8B125201125FC5F57FF839FF847" author="Pinto-da-Rocha" box="[943,1251,1957,1984]" firstAuthor="Pinto-da-Rocha" page="80" pageId="5" pageNumber="41" pagination="61 - 173" refId="ref10297" refString="Pinto-da-Rocha R. 1995. Sinopse da fauna cavernicola do Brasil. Papeis avulsos de Zoologia 39: 61 - 173." type="journal article" year="1995">Pinto-da-Rocha 1995: 80</bibRefCitation>
</treatmentCitation>
; 1996: 189. —
<treatmentCitation id="0AF3B35125201125FEE6579286B9F81A" author="Kury A. B." box="[278,453,1992,2018]" page="28" pageId="5" pageNumber="41" year="2003">
<bibRefCitation id="EFC3E8B125201125FEE6579286B9F81A" author="Kury" box="[278,453,1992,2018]" firstAuthor="Kury" page="28" pageId="5" pageNumber="41" refId="ref9413" refString="Kury A. B. 2003. Annotated Catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista iberica de Aracnologia, Zaragoza." type="book" year="2003">Kury 2003: 28</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="8BED954025231126FF4D515286D6FEDA" blockId="6.[189,426,264,290]" box="[189,426,264,290]" pageId="6" pageNumber="42">
<heading id="D0A5222C25231126FF4D515286D6FEDA" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="6" pageNumber="42" reason="3">
<emphasis id="B926495225231126FF4D515286D6FEDA" bold="true" box="[189,426,264,290]" pageId="6" pageNumber="42">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BED954025231126FF1A516D8620FEA9" blockId="6.[189,1399,311,408]" box="[234,348,311,337]" pageId="6" pageNumber="42">
<emphasis id="B926495225231126FF1A516D8620FEA9" bold="true" box="[234,348,311,337]" pageId="6" pageNumber="42">
<typeStatus id="54E92BE225231126FF1A516D8620FEA9" box="[234,348,311,337]" pageId="6" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BED954025231126FF4D510085F3FE60" blockId="6.[189,1399,311,408]" pageId="6" pageNumber="42">
<materialsCitation id="3B3A9F1D25231126FF4D510085F3FE60" ID-GBIF-Occurrence="4515792305" collectionCode="MZUSP" country="Brazil" latitude="-19.125278" location="Lapa Nova de Maquine cave" longLatPrecision="21" longitude="-44.351944" municipality="Cordisburgo" pageId="6" pageNumber="42" specimenCode="MZUSP 28463" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="F345D5D025231126FF4D5100865AFE8C" box="[189,294,346,372]" name="Brazil" pageId="6" pageNumber="42">BRAZIL</collectingCountry>
<specimenCount id="9D545EC925231126FEB551018622FE8D" box="[325,350,347,373]" count="1" pageId="6" pageNumber="42" type="male"></specimenCount>
;
<collectingRegion id="49965BA225231126FE9D5100856DFE8C" box="[365,529,346,372]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="42">Minas Gerais</collectingRegion>
,
<collectingMunicipality id="6B890F3A25231126FDD0510085C9FE8C" box="[544,693,346,372]" pageId="6" pageNumber="42">Cordisburgo</collectingMunicipality>
,
<location id="8E8DC39B25231126FD3351018358FE8D" LSID="urn:lsid:plazi:treatment:03FB24562520112DFDC056818525FC27:8E8DC39B25231126FD3351018358FE8D" box="[707,1060,346,373]" country="Brazil" latitude="-19.125278" longLatPrecision="21" longitude="-44.351944" municipality="Cordisburgo" name="Lapa Nova de Maquine cave" pageId="6" pageNumber="42" stateProvince="Minas Gerais">Lapa Nova de Maquiné cave</location>
;
<geoCoordinate id="EE66F38725231126FBC4510383B9FE8C" box="[1076,1221,345,373]" degrees="19" direction="south" minutes="07" orientation="latitude" pageId="6" pageNumber="42" precision="15" seconds="31" value="-19.125278">19°0731ʺ S</geoCoordinate>
,
<geoCoordinate id="EE66F38725231126FB255103820EFE8D" box="[1237,1394,345,373]" degrees="44" direction="west" minutes="21" orientation="longitude" pageId="6" pageNumber="42" precision="15" seconds="07" value="-44.351944">44°2107ʺ W</geoCoordinate>
;
<specimenCode id="DBF43D3B25231126FF4D51248612FE6F" box="[189,366,381,408]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="42">MZUSP 28463</specimenCode>
(ex:
<specimenCode id="DBF43D3B25231126FE5B512485FFFE60" box="[427,643,382,408]" collectionCode="MZUSP-OS" pageId="6" pageNumber="42">MZUSP-OS 3187</specimenCode>
).
</materialsCitation>
</paragraph>
<caption id="DF2DC5C825231126FF4D573E83F0F810" ID-DOI="http://doi.org/10.5281/zenodo.10604113" ID-Zenodo-Dep="10604113" httpUri="https://zenodo.org/record/10604113/files/figure.png" pageId="6" pageNumber="42" startId="6.[189,233,1892,1918]" targetBox="[189,1399,473,1834]" targetPageId="6">
<paragraph id="8BED954025231126FF4D573E83F0F810" blockId="6.[189,1398,1892,2025]" pageId="6" pageNumber="42">
<emphasis id="B926495225231126FF4D573E8668F887" bold="true" box="[189,276,1892,1919]" pageId="6" pageNumber="42">Fig. 3.</emphasis>
<taxonomicName id="4C52EEC325231126FED5573F8458F887" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[293,804,1893,1919]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225231126FED5573F853DF887" box="[293,577,1893,1919]" italics="true" pageId="6" pageNumber="42">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B125231126FD8E573F8458F887" author="Soares" box="[638,804,1893,1919]" firstAuthor="Soares" pageId="6" pageNumber="42" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
, male holotype (MZUSP 28463).
<emphasis id="B926495225231126FB15573E8386F886" bold="true" box="[1253,1274,1892,1918]" pageId="6" pageNumber="42">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B926495225231126FE9157D28609F85A" bold="true" box="[353,373,1928,1954]" pageId="6" pageNumber="42">B</emphasis>
. Habitus, ventral view.
<emphasis id="B926495225231126FD5957D285C2F85A" bold="true" box="[681,702,1928,1954]" pageId="6" pageNumber="42">C</emphasis>
. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5.
<emphasis id="B926495225231126FD0257F1847BF83D" bold="true" box="[754,775,1963,1989]" pageId="6" pageNumber="42">D</emphasis>
. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium.
<emphasis id="B926495225231126FCD057948448F810" bold="true" box="[800,820,1998,2024]" pageId="6" pageNumber="42">E</emphasis>
. Male pedipalp, mesal view.
</paragraph>
</caption>
<paragraph id="8BED954025221127FF1A51538620FEDB" blockId="7.[189,1261,265,326]" box="[234,348,265,291]" pageId="7" pageNumber="43">
<emphasis id="B926495225221127FF1A51538620FEDB" bold="true" box="[234,348,265,291]" pageId="7" pageNumber="43">
<typeStatus id="54E92BE225221127FF1A51538620FEDB" box="[234,348,265,291]" pageId="7" pageNumber="43" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BED954025221127FF4D51768391FEBE" blockId="7.[189,1261,265,326]" box="[189,1261,300,326]" pageId="7" pageNumber="43">
<materialsCitation id="3B3A9F1D25221127FF4D51768395FEBE" ID-GBIF-Occurrence="4515792301" box="[189,1257,300,326]" collectionCode="MZUSP" country="Brazil" latitude="-19.125278" location="Lapa Nova de Maquine cave" longLatPrecision="21" longitude="-44.351944" municipality="Cordisburgo" pageId="7" pageNumber="43" specimenCode="MZUSP 28462" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectingCountry id="F345D5D025221127FF4D5176865AFEBE" box="[189,294,300,326]" name="Brazil" pageId="7" pageNumber="43">BRAZIL</collectingCountry>
<specimenCount id="9D545EC925221127FECF5176862AFEBE" box="[319,342,300,326]" count="1" pageId="7" pageNumber="43" type="female"></specimenCount>
; same collection data as for holotYpe;
<specimenCode id="DBF43D3B25221127FCEB517684B0FEBE" box="[795,972,300,326]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="43">MZUSP 28462</specimenCode>
(ex:
<specimenCode id="DBF43D3B25221127FBF85176839DFEBE" box="[1032,1249,300,326]" collectionCode="MZUSP-OS" pageId="7" pageNumber="43">MZUSP-OS 3187</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C348C6CB25221128FF4D513785CAFE91" lastPageId="8" lastPageNumber="44" pageId="7" pageNumber="43" type="discussion">
<paragraph id="8BED954025221127FF4D51378553FE7F" blockId="7.[189,559,365,391]" box="[189,559,365,391]" pageId="7" pageNumber="43">
<heading id="D0A5222C25221127FF4D51378553FE7F" bold="true" box="[189,559,365,391]" fontSize="11" level="3" pageId="7" pageNumber="43" reason="3">
<emphasis id="B926495225221127FF4D51378553FE7F" bold="true" box="[189,559,365,391]" pageId="7" pageNumber="43">
Notes about the
<typeStatus id="54E92BE225221127FE78513786C2FE7F" box="[392,446,365,391]" pageId="7" pageNumber="43">type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED954025221128FF4D51C685CAFE91" blockId="7.[189,1399,412,649]" lastBlockId="8.[189,1399,265,362]" lastPageId="8" lastPageNumber="44" pageId="7" pageNumber="43">
<bibRefCitation id="EFC3E8B125221127FF4D51C6861DFE4E" author="Soares" box="[189,353,412,438]" firstAuthor="Soares" pageId="7" pageNumber="43" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares (1966)</bibRefCitation>
deposited the type material of
<taxonomicName id="4C52EEC325221127FD2351C6849EFE4E" authorityName="H. Soares" authorityYear="1966" box="[723,994,412,438]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225221127FD2351C6849EFE4E" box="[723,994,412,438]" italics="true" pageId="7" pageNumber="43">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
in the “Otto Schubart Collection” with the number 3187. In addition to a
<typeStatus id="54E92BE225221127FD8D51E58598FE21" box="[637,740,447,473]" pageId="7" pageNumber="43" type="holotype">holotype</typeStatus>
male,
<specimenCount id="9D545EC925221127FCC051E584CCFE21" box="[816,944,447,473]" count="1" pageId="7" pageNumber="43" type="female">one female</specimenCount>
<typeStatus id="54E92BE225221127FC4651E58367FE21" box="[950,1051,447,473]" pageId="7" pageNumber="43" type="paratype">paratype</typeStatus>
and other male
<typeStatus id="54E92BE225221127FB2351E58244FE21" box="[1235,1336,447,473]" pageId="7" pageNumber="43" type="paratype">paratype</typeStatus>
were deposited with the same number (3187) in this same collection. A third male and
<specimenCount id="9D545EC925221127FB8B51B98382FE04" box="[1147,1278,482,509]" count="2" pageId="7" pageNumber="43" type="female">two female</specimenCount>
<typeStatus id="54E92BE225221127FAF651B9820AFE05" box="[1286,1398,483,509]" pageId="7" pageNumber="43" type="paratype">paratypes</typeStatus>
were deposited in the collection “Coleção H. Soares” with the number 54 and this material was destroyed in the fire at the National Museum of
<collectingRegion id="49965BA225221127FD8E52728456FDBA" box="[638,810,552,578]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="43">Rio de Janeiro</collectingRegion>
(
<collectionCode id="ED430D8525221127FCCA527384F0FDBB" box="[826,908,553,579]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="43">MNRJ</collectionCode>
) in 2018, where the “Coleção H. Soares” was deposited. In 2003, Kury referred to the material type with the collection numbers
<collectionCode id="ED430D8525221127FB155216820AFD9E" box="[1253,1398,588,614]" pageId="7" pageNumber="43">MZUSP-OS</collectionCode>
3187. Today, the male
<typeStatus id="54E92BE225221127FE205235854BFD71" box="[464,567,623,649]" pageId="7" pageNumber="43" type="holotype">holotype</typeStatus>
and female
<typeStatus id="54E92BE225221127FD3E5235844FFD71" box="[718,819,623,649]" pageId="7" pageNumber="43" type="paratype">paratype</typeStatus>
from the “Otto Schubart Collection” (3187) are incorporated into the collection of the University of
<collectingRegion id="49965BA2252D1128FCD3515384E6FEDB" box="[803,922,265,291]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="44">São Paulo</collectingRegion>
Museum of Zoology (
<collectionCode id="ED430D85252D1128FB525153827BFEDB" box="[1186,1287,265,291]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="44">MZUSP</collectionCode>
) with the collection number:
<collectionCode id="ED430D85252D1128FE565176857BFEBE" box="[422,519,300,326]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="44">MZUSP</collectionCode>
28463 (male
<typeStatus id="54E92BE2252D1128FD5F51768464FEBE" box="[687,792,300,326]" pageId="8" pageNumber="44" type="holotype">holotype</typeStatus>
) and
<collectionCode id="ED430D85252D1128FCAD517684C2FEBE" box="[861,958,300,326]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="44">MZUSP</collectionCode>
28462 (
<specimenCount id="9D545EC9252D1128FBD2517683D4FEBE" box="[1058,1192,300,326]" count="1" pageId="8" pageNumber="44" type="female">one female</specimenCount>
<typeStatus id="54E92BE2252D1128FB4151768260FEBE" box="[1201,1308,300,326]" pageId="8" pageNumber="44" type="paratype">paratype</typeStatus>
). These
<specimenCount id="9D545EC9252D1128FF4D510A8616FE91" box="[189,362,335,362]" count="2" pageId="8" pageNumber="44" type="generic">two specimens</specimenCount>
were analyzed in this work.
</paragraph>
</subSubSection>
<caption id="DF2DC5C825221127FF4D57F0820BF810" ID-DOI="http://doi.org/10.5281/zenodo.10604117" ID-Zenodo-Dep="10604117" httpUri="https://zenodo.org/record/10604117/files/figure.png" pageId="7" pageNumber="43" startId="7.[189,231,1962,1988]" targetBox="[189,1399,704,1914]" targetPageId="7">
<paragraph id="8BED954025221127FF4D57F0820BF810" blockId="7.[189,1399,1962,2024]" pageId="7" pageNumber="43">
<emphasis id="B926495225221127FF4D57F08674F83D" bold="true" box="[189,264,1962,1989]" pageId="7" pageNumber="43">Fig. 4.</emphasis>
<taxonomicName id="4C52EEC325221127FEFD57F185ADF83D" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[269,721,1963,1989]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B926495225221127FEFD57F18568F83D" box="[269,532,1963,1989]" italics="true" pageId="7" pageNumber="43">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B125221127FDCB57F185ADF83D" author="Soares" box="[571,721,1963,1989]" firstAuthor="Soares" pageId="7" pageNumber="43" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
, female paratype (MZUSP 28462).
<emphasis id="B926495225221127FB9557F08306F83C" bold="true" box="[1125,1146,1962,1988]" pageId="7" pageNumber="43">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B926495225221127FF4D579487ACF810" bold="true" box="[189,208,1998,2024]" pageId="7" pageNumber="43">B</emphasis>
. Habitus, ventral view.
<emphasis id="B926495225221127FE2F57948688F810" bold="true" box="[479,500,1998,2024]" pageId="7" pageNumber="43">C</emphasis>
. Habitus, left lateral view.
<emphasis id="B926495225221127FCD657948447F810" bold="true" box="[806,827,1998,2024]" pageId="7" pageNumber="43">D</emphasis>
. Carapace, lateral view.
<emphasis id="B926495225221127FBA35794831AF810" bold="true" box="[1107,1126,1998,2024]" pageId="7" pageNumber="43">E</emphasis>
. Carapace, dorsal view.
</paragraph>
</caption>
<subSubSection id="C348C6CB252D1128FF4D51CB86B2FDBD" pageId="8" pageNumber="44" type="etymology">
<paragraph id="8BED9540252D1128FF4D51CB8638FE53" blockId="8.[189,324,401,427]" box="[189,324,401,427]" pageId="8" pageNumber="44">
<heading id="D0A5222C252D1128FF4D51CB8638FE53" bold="true" box="[189,324,401,427]" fontSize="11" level="3" pageId="8" pageNumber="44" reason="3">
<emphasis id="B9264952252D1128FF4D51CB8638FE53" bold="true" box="[189,324,401,427]" pageId="8" pageNumber="44">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BED9540252D1128FF4D51E586B2FDBD" blockId="8.[189,1399,447,581]" pageId="8" pageNumber="44">
The original species name is incorrect; it should be spelled spelaeus from Greek σπήλαιον (spelaion=cave, cavern). Although incorrect, the original name,
<taxonomicName id="4C52EEC3252D1128FC4751B88360FE04" authorityName="H. Soares" authorityYear="1966" box="[951,1052,482,508]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252D1128FC4751B88360FE04" box="[951,1052,482,508]" italics="true" pageId="8" pageNumber="44">spaeleus</emphasis>
</taxonomicName>
, cant be emended according to article 32.5.1 of the currently valid ICZN (1999), because it is considered as an incorrect latinization of the Greek σπήλαιον.
</paragraph>
</subSubSection>
<subSubSection id="C348C6CB252D1128FF4D52308392FC7E" pageId="8" pageNumber="44" type="diagnosis">
<paragraph id="8BED9540252D1128FF4D5230864AFD7C" blockId="8.[189,310,618,644]" box="[189,310,618,644]" pageId="8" pageNumber="44">
<heading id="D0A5222C252D1128FF4D5230864AFD7C" bold="true" box="[189,310,618,644]" fontSize="11" level="3" pageId="8" pageNumber="44" reason="3">
<emphasis id="B9264952252D1128FF4D5230864AFD7C" bold="true" box="[189,310,618,644]" pageId="8" pageNumber="44">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BED9540252D1128FF4D52C38392FC7E" blockId="8.[189,1399,665,903]" pageId="8" pageNumber="44">
<taxonomicName id="4C52EEC3252D1128FF4D52C386B1FD4B" authorityName="H. Soares" authorityYear="1966" box="[189,461,665,691]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252D1128FF4D52C386B1FD4B" box="[189,461,665,691]" italics="true" pageId="8" pageNumber="44">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C52EEC3252D1128FD9F52C385B7FD4B" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[623,715,665,691]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B9264952252D1128FD9F52C385B7FD4B" box="[623,715,665,691]" italics="true" pageId="8" pageNumber="44">S. gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F429252D1128FD2552C3844EFD4B" box="[725,818,665,691]" pageId="8" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
mainly by the following combination of genital characters: apical lamina with a pointed medial projection; the presence of three lateral pairs of reduced setae ventrallY on the apical lamina; lateral projections of the capsula interna distallY pointed; and bY the ventral keel of the pars distalis with a round top in ventral view (
<figureCitation id="136989C5252D1128FC495358837DFCE4" box="[953,1025,770,796]" captionStart="Fig" captionStartId="10.[189,231,1892,1918]" captionTargetBox="[313,1298,573,1821]" captionTargetId="figure-174@10.[263,1321,567,1835]" captionTargetPageId="10" captionText="Fig. 6. Line drawings of male genitalia of Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Full genitalia, dorsal view. B. Detail of pars distalis, dorsal view. C. Detail of capsula interna, dorsal view. D. Detail of pars distalis, lateral view. E. Same, ventral view. Red lines: capsula interna. Blue lines: conductors. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10604123" httpUri="https://zenodo.org/record/10604123/files/figure.png" pageId="8" pageNumber="44">Fig. 6</figureCitation>
). Both species of
<taxonomicName id="4C52EEC3252D1128FB255358820AFCE4" authorityName="H. Soares" authorityYear="1966" box="[1237,1398,770,796]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9264952252D1128FB255358820AFCE4" box="[1237,1398,770,796]" italics="true" pageId="8" pageNumber="44">Spaeleoleptes</emphasis>
</taxonomicName>
have a very similar external morphology, but
<taxonomicName id="4C52EEC3252D1128FD1B537C840DFCB8" authorityName="H. Soares" authorityYear="1966" box="[747,881,806,832]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252D1128FD1B537C840DFCB8" box="[747,881,806,832]" italics="true" pageId="8" pageNumber="44">S. spaeleus</emphasis>
</taxonomicName>
can be distinguished by having the tibia I modified, horse saddle-shaped, in retrolateral view (
<figureCitation id="136989C5252D1128FCD3531384D6FC9C" box="[803,938,841,868]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1399,650,1852]" captionTargetId="figure-142@9.[189,1399,647,1852]" captionTargetPageId="9" captionText="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="8" pageNumber="44">Figs 5AB</figureCitation>
); the two large pointed tubercles in the post ocularium region; and bY the bodY being covered bY large setiferous tubercles (
<figureCitation id="136989C5252D1128FB67533683A2FC7E" box="[1175,1246,876,902]" captionStart="Fig" captionStartId="6.[189,233,1892,1918]" captionTargetBox="[189,1399,473,1834]" captionTargetId="figure-85@6.[189,1399,473,1834]" captionTargetPageId="6" captionText="Fig. 3. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5. D. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium. E. Male pedipalp, mesal view." figureDoi="http://doi.org/10.5281/zenodo.10604113" httpUri="https://zenodo.org/record/10604113/files/figure.png" pageId="8" pageNumber="44">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348C6CB252D112DFF4D53F78554FC88" lastPageId="13" lastPageNumber="49" pageId="8" pageNumber="44" type="description">
<paragraph id="8BED9540252D1128FF4D53F78617FC3F" blockId="8.[189,363,941,967]" box="[189,363,941,967]" pageId="8" pageNumber="44">
<heading id="D0A5222C252D1128FF4D53F78617FC3F" bold="true" box="[189,363,941,967]" fontSize="11" level="3" pageId="8" pageNumber="44" reason="3">
<emphasis id="B9264952252D1128FF4D53F78617FC3F" bold="true" box="[189,363,941,967]" pageId="8" pageNumber="44">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BED9540252D1128FF1A53868590FC0E" blockId="8.[189,1399,988,1085]" box="[234,748,988,1014]" pageId="8" pageNumber="44">
<emphasis id="B9264952252D1128FF1A53868647FC0E" bold="true" box="[234,315,988,1014]" pageId="8" pageNumber="44">Male (</emphasis>
<typeStatus id="54E92BE2252D1128FECB538686D9FC0E" box="[315,421,988,1014]" pageId="8" pageNumber="44" type="holotype">holotype</typeStatus>
, MZUSP 28463) (
<figureCitation id="136989C5252D1128FD8B53868598FC0E" box="[635,740,988,1014]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[189,233,1892,1918]" captionStartId-1="7.[189,231,1962,1988]" captionStartId-2="9.[189,232,1892,1918]" captionStartId-3="10.[189,231,1892,1918]" captionTargetBox-0="[189,1399,473,1834]" captionTargetBox-1="[189,1399,704,1914]" captionTargetBox-2="[189,1399,650,1852]" captionTargetBox-3="[313,1298,573,1821]" captionTargetId-0="figure-85@6.[189,1399,473,1834]" captionTargetId-1="figure-172@7.[189,1399,704,1914]" captionTargetId-2="figure-142@9.[189,1399,647,1852]" captionTargetId-3="figure-174@10.[263,1321,567,1835]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Fig. 3. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5. D. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium. E. Male pedipalp, mesal view." captionText-1="Fig. 4. Spaeleoleptes spaeleus H. Soares, 1966, female paratype (MZUSP 28462). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, left lateral view. D. Carapace, lateral view. E. Carapace, dorsal view." captionText-2="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." captionText-3="Fig. 6. Line drawings of male genitalia of Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Full genitalia, dorsal view. B. Detail of pars distalis, dorsal view. C. Detail of capsula interna, dorsal view. D. Detail of pars distalis, lateral view. E. Same, ventral view. Red lines: capsula interna. Blue lines: conductors. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10604113" figureDoi-1="http://doi.org/10.5281/zenodo.10604117" figureDoi-2="http://doi.org/10.5281/zenodo.10604119" figureDoi-3="http://doi.org/10.5281/zenodo.10604123" httpUri-0="https://zenodo.org/record/10604113/files/figure.png" httpUri-1="https://zenodo.org/record/10604117/files/figure.png" httpUri-2="https://zenodo.org/record/10604119/files/figure.png" httpUri-3="https://zenodo.org/record/10604123/files/figure.png" pageId="8" pageNumber="44">Figs 36</figureCitation>
)
</paragraph>
<paragraph id="8BED9540252D1128FF4D53A583D0FBC5" blockId="8.[189,1399,988,1085]" pageId="8" pageNumber="44">MEASUREMENTS (in millimeters). Total bodY length = 2.2; carapace length =0.78; scutum magnum length =1.9; carapace maximum width=1.0; mesotergal scute maximum width=1.50.</paragraph>
<paragraph id="8BED9540252D1128FF4D543E86FBFB59" blockId="8.[189,1399,1124,1185]" pageId="8" pageNumber="44">
BODY. Entirely covered by setiferous tubercles with the exception of the carapace, chelicerae, and pedipalp (
<figureCitation id="136989C5252D1128FEC154DD860BFB59" box="[305,375,1159,1185]" captionStart="Fig" captionStartId="6.[189,233,1892,1918]" captionTargetBox="[189,1399,473,1834]" captionTargetId="figure-85@6.[189,1399,473,1834]" captionTargetPageId="6" captionText="Fig. 3. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5. D. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium. E. Male pedipalp, mesal view." figureDoi="http://doi.org/10.5281/zenodo.10604113" httpUri="https://zenodo.org/record/10604113/files/figure.png" pageId="8" pageNumber="44">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BED9540252D1128FF4D549385D3F937" blockId="8.[189,1399,1224,1744]" pageId="8" pageNumber="44">
DORSUM. Scutum magnum hourglass-shaped with the carapace slightly narrower than the mesotergal shield. Left and right anterior margin of the carapace is armed with two large pointed tubercles close to each other. Mesotergal shield trapezoidal, with areas increasing in width in an anteroposterior direction, with area I being the smallest and area IV the largest (along transversal axis). Areas IV and V with the same width (
<figureCitation id="136989C5252D1128FE75550F86B2FA97" box="[389,462,1365,1391]" captionStart="Fig" captionStartId="6.[189,233,1892,1918]" captionTargetBox="[189,1399,473,1834]" captionTargetId="figure-85@6.[189,1399,473,1834]" captionTargetPageId="6" captionText="Fig. 3. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, right lateral view. Square indicatesglandular/ sensorial? region of leg I, more detail in Fig. 5. D. Carapace, lateral view. Arrows indicate the pointed tubercles on the posterior region of the ocularium. E. Male pedipalp, mesal view." figureDoi="http://doi.org/10.5281/zenodo.10604113" httpUri="https://zenodo.org/record/10604113/files/figure.png" pageId="8" pageNumber="44">Fig. 3</figureCitation>
). Anterior margin of carapace straight, with no pronounced cheliceral sockets. Posterior margin of scutum straight. Each antero-lateral margin of the carapace is armed with two large, pointed tubercles close to each other. Massive ocularium and frontal hump well marked near the anterior margin of the carapace. Ocularium rounded, with several small granules on the surface. Two eyes on the lateral surface of the ocularium with reduced cornea, no visible retina. Carapace in lateral view with a posterior ocularium region slightly convex, armed with two rounded-tipped tubercles. Deep sulcus I. Mesotergal scutum with five distinguishable areas of approximatelY the same size (along anteroposterior axis). Sulci IIIV shallow (not well marked), especiallY the sulci III and IV. Sulci IIIV complete, extending from center to edge of mesotergum. Areas IV and free tergites each with a transversal row of pointed setiferous tubercles. Coxa IV visible in dorsal view, armed with several prominent setiferous tubercles on the retrolateral-dorsal surface.
</paragraph>
<paragraph id="8BED9540252D1128FF4D56AD8250F8A0" blockId="8.[189,1399,1783,1880]" pageId="8" pageNumber="44">VENTER. Free sternites each with a transverse row of pointed setiferous tubercles. Anal operculum covered bY manY low, robust setiferous tubercles of the same size as those of the free tergites. Coxa IV longer than wide, with low setiferous tubercles on the ventral surface. Spiracles free, not concealed bY coxa IV.</paragraph>
<paragraph id="8BED9540252D1128FF4D572584ADF844" blockId="8.[189,1398,1919,1980]" pageId="8" pageNumber="44">CHELICERA. Basichelicerite unarmed, with a well-marked rounded bulla. Cheliceral hand unarmed, normal, neither swollen nor hypertelic, covered with some sensilla.</paragraph>
<paragraph id="8BED9540252C1129FF4D5153843FFE0F" blockId="9.[189,1398,265,503]" pageId="9" pageNumber="45">
PEDIPALP. Raptorial morphotYpe (sensu
<bibRefCitation id="EFC3E8B1252C1129FD6251538423FEDB" author="Wolff" box="[658,863,265,291]" etAl="et al." firstAuthor="Wolff" pageId="9" pageNumber="45" pagination="558 - 601" refId="ref10703" refString="Wolff J. O., Schonhofer A. L., Martens J., Wijnhoven H., TaYlor C. K. &amp; Gorb S. N. 2016. The evolution of pedipalps and glandular hairs as predatory devices in harvestmen (Arachnida, Opiliones). Zoological Journal of the Linnean Society 177: 558 - 601. https: // doi. org / 10.1111 / zoj. 12375" type="journal article" year="2016">
Wolff
<emphasis id="B9264952252C1129FD2C5150846BFEDB" box="[732,791,265,291]" italics="true" pageId="9" pageNumber="45">et al.</emphasis>
2016
</bibRefCitation>
). Coxa short, armed with one large setiferous tubercle ventrally. Trochanter globular, with one setiferous tubercle and two pronounced granules ventrally. Femur armed ventrally with two proximal and two smaller medial major spines (i.e., stiff pointed bristles in highly elevated sockets, sensu
<bibRefCitation id="EFC3E8B1252C1129FD04512884C6FE74" author="Wolff" box="[756,954,370,396]" etAl="et al." firstAuthor="Wolff" pageId="9" pageNumber="45" pagination="558 - 601" refId="ref10703" refString="Wolff J. O., Schonhofer A. L., Martens J., Wijnhoven H., TaYlor C. K. &amp; Gorb S. N. 2016. The evolution of pedipalps and glandular hairs as predatory devices in harvestmen (Arachnida, Opiliones). Zoological Journal of the Linnean Society 177: 558 - 601. https: // doi. org / 10.1111 / zoj. 12375" type="journal article" year="2016">
Wolff
<emphasis id="B9264952252C1129FCCB51298408FE74" box="[827,884,370,396]" italics="true" pageId="9" pageNumber="45">et al.</emphasis>
2016
</bibRefCitation>
); dorsallY with a row of small granules and the subdistal-mesal surface with one major spine. Patella armed with one major spine on the mesal surface. Tibia armed ventrally with three ectal and three mesal major spines. Tarsus armed ventrally with three ectal and three mesal major spines. Claw present.
</paragraph>
<paragraph id="8BED9540252C112AFF4D5244832DFE0F" blockId="9.[189,1399,542,603]" lastBlockId="10.[189,1399,265,503]" lastPageId="10" lastPageNumber="46" pageId="9" pageNumber="45">
LEGS. Leg measurements in
<tableCitation id="C6D0A0FB252C1129FDF95244851DFDC0" box="[521,609,542,568]" captionStart="Table 1" captionStartId="11.[188,257,629,658]" captionTargetBox="[204,1902,759,1654]" captionText="Table 1 (continued on next page). Body and leg measurements (in mm) of analyzed specimens of Spaeleoleptes spaeleus H. Soares, 1966 and Spaeleoleptes gimli sp. nov. Abbreviations: tr=trochanter; fe=femur; pa=patella; ti=tibia; me=metatarsus; ta=tarsus." httpUri="http://table.plazi.org/id/DF2DC5C8252E112BFF4C522F814DFD4C" pageId="9" pageNumber="45" tableUuid="DF2DC5C8252E112BFF4C522F814DFD4C">Table 1</tableCitation>
. Legs covered with large setiferous tubercles from the coxae to the middle of the metatarsus. Legs I and II sexuallY dimorphic, males with thickened tibia I and modified distal region horse saddle-shaped in retrolateral view. This modified glandular/sensorial? region has several “droplet-like” pegs present mainly on the retrolateral surface, but they may also be present dorsally and extend from the subproximal region to the subdistal region of the tibia I, occupying the entire medial area (
<figureCitation id="136989C5252F112AFF365128865EFE75" box="[198,290,370,397]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1399,650,1852]" captionTargetId="figure-142@9.[189,1399,647,1852]" captionTargetPageId="9" captionText="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="10" pageNumber="46">Fig. 5B</figureCitation>
). Tibia I also expanded retrolaterally in its medial region, in dorsal view (
<figureCitation id="136989C5252F112AFB755128839FFE75" box="[1157,1251,370,397]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1399,650,1852]" captionTargetId="figure-142@9.[189,1399,647,1852]" captionTargetPageId="9" captionText="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="10" pageNumber="46">Fig. 5A</figureCitation>
). Males also have legs II with a glandular/sensorial? region with “droplet-like” pegs on the ventral surface, medially on the patella II and proximally on the tibia II (
<figureCitation id="136989C5252F112AFD4151E38471FE2C" box="[689,781,441,468]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1399,650,1852]" captionTargetId="figure-142@9.[189,1399,647,1852]" captionTargetPageId="9" captionText="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="10" pageNumber="46">Fig. 5D</figureCitation>
). Patella and tibia II slightly expanded ventrally in the glandular/sensorial? region, in lateral view (
<figureCitation id="136989C5252F112AFD4551868473FE0F" box="[693,783,476,503]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1399,650,1852]" captionTargetId="figure-142@9.[189,1399,647,1852]" captionTargetPageId="9" captionText="Fig. 5. Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463), glandular/sensorial? region. A. Modified region of right tibia I, dorsal view. B. Detail of glandular/sensorial? region on right tibia I, retrolateral view. C. Modified region of patella and tibia II, lateral view. D. Detail of glandular/ sensorial? region on patella and tibia II, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="10" pageNumber="46">Fig. 5C</figureCitation>
). Tarsal formula: 4(2):?:5:5.
</paragraph>
<caption id="DF2DC5C8252C1129FF4D573E8451F810" ID-DOI="http://doi.org/10.5281/zenodo.10604119" ID-Zenodo-Dep="10604119" httpUri="https://zenodo.org/record/10604119/files/figure.png" pageId="9" pageNumber="45" startId="9.[189,232,1892,1918]" targetBox="[189,1399,650,1852]" targetPageId="9">
<paragraph id="8BED9540252C1129FF4D573E8451F810" blockId="9.[189,1399,1892,2025]" pageId="9" pageNumber="45">
<emphasis id="B9264952252C1129FF4D573E8671F887" bold="true" box="[189,269,1892,1919]" pageId="9" pageNumber="45">Fig. 5.</emphasis>
<taxonomicName id="4C52EEC3252C1129FEE7573F858EF887" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[279,754,1893,1919]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252C1129FEE7573F855AF887" box="[279,550,1893,1919]" italics="true" pageId="9" pageNumber="45">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B1252C1129FDA6573F858EF887" author="Soares" box="[598,754,1893,1919]" firstAuthor="Soares" pageId="9" pageNumber="45" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
, male holotype (MZUSP 28463), glandular/sensorial? region.
<emphasis id="B9264952252C1129FEE657D28657F85A" bold="true" box="[278,299,1928,1954]" pageId="9" pageNumber="45">A</emphasis>
. Modified region of right tibia I, dorsal view.
<emphasis id="B9264952252C1129FCB557D28425F85A" bold="true" box="[837,857,1928,1954]" pageId="9" pageNumber="45">B</emphasis>
. Detail of glandular/sensorial? region on right tibia I, retrolateral view.
<emphasis id="B9264952252C1129FE1157F1868AF83D" bold="true" box="[481,502,1963,1989]" pageId="9" pageNumber="45">C</emphasis>
. Modified region of patella and tibia II, lateral view.
<emphasis id="B9264952252C1129FB9B57F183FCF83D" bold="true" box="[1131,1152,1963,1989]" pageId="9" pageNumber="45">D</emphasis>
. Detail of glandular/ sensorial? region on patella and tibia II, ventral view.
</paragraph>
</caption>
<caption id="DF2DC5C8252F112AFF4D573E85E5F811" ID-DOI="http://doi.org/10.5281/zenodo.10604123" ID-Zenodo-Dep="10604123" httpUri="https://zenodo.org/record/10604123/files/figure.png" pageId="10" pageNumber="46" startId="10.[189,231,1892,1918]" targetBox="[313,1298,573,1821]" targetPageId="10">
<paragraph id="8BED9540252F112AFF4D573E85E5F811" blockId="10.[189,1399,1892,2025]" pageId="10" pageNumber="46">
<emphasis id="B9264952252F112AFF4D573E867BF887" bold="true" box="[189,263,1892,1919]" pageId="10" pageNumber="46">Fig. 6.</emphasis>
Line drawings of male genitalia of
<taxonomicName id="4C52EEC3252F112AFD68573F8320F887" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[664,1116,1893,1919]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="46" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252F112AFD68573F84E2F887" box="[664,926,1893,1919]" italics="true" pageId="10" pageNumber="46">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B1252F112AFC36573F8320F887" author="Soares" box="[966,1116,1893,1919]" firstAuthor="Soares" pageId="10" pageNumber="46" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
, male holotype (MZUSP 28463).
<emphasis id="B9264952252F112AFEED57D2864EF85A" bold="true" box="[285,306,1928,1954]" pageId="10" pageNumber="46">A</emphasis>
. Full genitalia, dorsal view.
<emphasis id="B9264952252F112AFD8857D285F7F85A" bold="true" box="[632,651,1928,1954]" pageId="10" pageNumber="46">B</emphasis>
. Detail of pars distalis, dorsal view.
<emphasis id="B9264952252F112AFBDE57D2833FF85A" bold="true" box="[1070,1091,1928,1954]" pageId="10" pageNumber="46">C</emphasis>
. Detail of capsula interna, dorsal view.
<emphasis id="B9264952252F112AFEA557F18616F83D" bold="true" box="[341,362,1963,1989]" pageId="10" pageNumber="46">D</emphasis>
. Detail of pars distalis, lateral view.
<emphasis id="B9264952252F112AFCD357F1844AF83D" bold="true" box="[803,822,1963,1989]" pageId="10" pageNumber="46">E</emphasis>
. Same, ventral view. Red lines: capsula interna. Blue lines: conductors. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF2DC5C8252E112BFF4C522F814DFD4C" ID-Table-UUID="DF2DC5C8252E112BFF4C522F814DFD4C" httpUri="http://table.plazi.org/id/DF2DC5C8252E112BFF4C522F814DFD4C" pageId="11" pageNumber="47" startId="11.[188,257,629,658]" targetBox="[204,1902,759,1654]" targetIsTable="true" targetPageId="11">
<paragraph id="8BED9540252E112BFF4C522F814DFD4C" blockId="11.[188,1941,629,692]" pageId="11" pageNumber="47">
<emphasis id="B9264952252E112BFF4C522F8667FD6A" bold="true" box="[188,283,629,658]" pageId="11" pageNumber="47">Table 1</emphasis>
(continued on next page). Body and leg measurements (in mm) of analyzed specimens of
<taxonomicName id="4C52EEC3252E112BFA8E522F801CFD69" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[1406,1888,629,657]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252E112BFA8E522F81ECFD69" box="[1406,1680,629,657]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B1252E112BF934522F801CFD69" author="Soares" box="[1732,1888,629,657]" firstAuthor="Soares" pageId="11" pageNumber="47" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C52EEC3252E112BFF4C52C286DCFD4C" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[188,416,664,692]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B9264952252E112BFF4C52C286DCFD4C" box="[188,416,664,692]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F429252E112BFE5852C2857FFD4C" box="[424,515,664,692]" pageId="11" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: tr=trochanter; fe=femur; pa=patella; ti =tibia; me= metatarsus; ta= tarsus.
</paragraph>
</caption>
<paragraph id="8BED9540252E112BFE1452AD81D1F98E" pageId="11" pageNumber="47">
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<tr id="35629702252EEEDFFF3C52AD8012FCA9" box="[204,1902,759,849]" gridrow="0" isHeader="true" pageId="11" pageNumber="47" rowspan-0="1">
<th id="76B3FE7E252EEEDFFE1352AD85A5FCA9" box="[483,729,759,849]" gridcol="1" gridrow="0" pageId="11" pageNumber="47">
<taxonomicName id="4C52EEC3252E112BFE1452AD85A5FCE9" authorityName="H. Soares" authorityYear="1966" box="[484,729,759,785]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252E112BFE1452AD85A5FCE9" box="[484,729,759,785]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
male holotype (MZUSP 28463)
</th>
<th id="76B3FE7E252EEEDFFCFA52AD837CFCA9" box="[778,1024,759,849]" gridcol="2" gridrow="0" pageId="11" pageNumber="47">
<taxonomicName id="4C52EEC3252E112BFCFA52AD837CFCE9" authorityName="H. Soares" authorityYear="1966" box="[778,1024,759,785]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B9264952252E112BFCFA52AD837CFCE9" box="[778,1024,759,785]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
female paratype (MZUSP 28462)
</th>
<th id="76B3FE7E252EEEDFFBC152AD825DFCA9" box="[1073,1313,759,849]" gridcol="3" gridrow="0" pageId="11" pageNumber="47">
<taxonomicName id="4C52EEC3252E112BFBC152AD827DFCE9" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1073,1281,759,785]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B9264952252E112BFBC152AD827DFCE9" box="[1073,1281,759,785]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F429252E112BFBC1534D83F8FCC9" box="[1073,1156,791,817]" pageId="11" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
male holotype (LES 00341)
</th>
<th id="76B3FE7E252EEEDFFAA752AD8120FCA9" box="[1367,1628,759,849]" gridcol="4" gridrow="0" pageId="11" pageNumber="47">
<taxonomicName id="4C52EEC3252E112BFAA852AD8154FCE9" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1368,1576,759,785]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B9264952252E112BFAA852AD8154FCE9" box="[1368,1576,759,785]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F429252E112BFAA8534D82D7FCC9" box="[1368,1451,791,817]" pageId="11" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
female paratype (LES 027941)
</th>
<th id="76B3FE7E252EEEDFF98E52AD8012FCA9" box="[1662,1902,759,849]" gridcol="5" gridrow="0" pageId="11" pageNumber="47">
<taxonomicName id="4C52EEC3252E112BF98F52AD8033FCE9" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1663,1871,759,785]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="47" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B9264952252E112BF98F52AD8033FCE9" box="[1663,1871,759,785]" italics="true" pageId="11" pageNumber="47">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F429252E112BF98F534D81ADFCC9" box="[1663,1745,791,817]" pageId="11" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
male paratype (LES 027941)
</th>
</tr>
<tr id="35629702252EEEDFFF3C532D8012FC69" box="[204,1902,887,913]" gridrow="1" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C532D86BBFC69" box="[204,455,887,913]" gridcol="0" gridrow="1" pageId="11" pageNumber="47">total body length</th>
<td id="76B3FE7E252EEEDFFE13532D85A5FC69" box="[483,729,887,913]" gridcol="1" gridrow="1" pageId="11" pageNumber="47">2.20</td>
<td id="76B3FE7E252EEEDFFCFA532D837CFC69" box="[778,1024,887,913]" gridcol="2" gridrow="1" pageId="11" pageNumber="47">2.30</td>
<td id="76B3FE7E252EEEDFFBC1532D825DFC69" box="[1073,1313,887,913]" gridcol="3" gridrow="1" pageId="11" pageNumber="47">2.18</td>
<td id="76B3FE7E252EEEDFFAA7532D8120FC69" box="[1367,1628,887,913]" gridcol="4" gridrow="1" pageId="11" pageNumber="47">2.30</td>
<td id="76B3FE7E252EEEDFF98E532D8012FC69" box="[1662,1902,887,913]" gridcol="5" gridrow="1" pageId="11" pageNumber="47">2.05</td>
</tr>
<tr id="35629702252EEEDFFF3C53C58012FC41" box="[204,1902,927,953]" gridrow="2" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C53C586BBFC41" box="[204,455,927,953]" gridcol="0" gridrow="2" pageId="11" pageNumber="47">scutum magnum length</th>
<td id="76B3FE7E252EEEDFFE1353C585A5FC41" box="[483,729,927,953]" gridcol="1" gridrow="2" pageId="11" pageNumber="47">1.90</td>
<td id="76B3FE7E252EEEDFFCFA53C5837CFC41" box="[778,1024,927,953]" gridcol="2" gridrow="2" pageId="11" pageNumber="47">2.00</td>
<td id="76B3FE7E252EEEDFFBC153C5825DFC41" box="[1073,1313,927,953]" gridcol="3" gridrow="2" pageId="11" pageNumber="47">1.84</td>
<td id="76B3FE7E252EEEDFFAA753C58120FC41" box="[1367,1628,927,953]" gridcol="4" gridrow="2" pageId="11" pageNumber="47">1.80</td>
<td id="76B3FE7E252EEEDFF98E53C58012FC41" box="[1662,1902,927,953]" gridcol="5" gridrow="2" pageId="11" pageNumber="47">1.77</td>
</tr>
<tr id="35629702252EEEDFFF3C539C8012FC18" box="[204,1902,966,992]" gridrow="3" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C539C86BBFC18" box="[204,455,966,992]" gridcol="0" gridrow="3" pageId="11" pageNumber="47">carapace length</th>
<td id="76B3FE7E252EEEDFFE13539C85A5FC18" box="[483,729,966,992]" gridcol="1" gridrow="3" pageId="11" pageNumber="47">0.78</td>
<td id="76B3FE7E252EEEDFFCFA539C837CFC18" box="[778,1024,966,992]" gridcol="2" gridrow="3" pageId="11" pageNumber="47">0.80</td>
<td id="76B3FE7E252EEEDFFBC1539C825DFC18" box="[1073,1313,966,992]" gridcol="3" gridrow="3" pageId="11" pageNumber="47">0.77</td>
<td id="76B3FE7E252EEEDFFAA7539C8120FC18" box="[1367,1628,966,992]" gridcol="4" gridrow="3" pageId="11" pageNumber="47">0.80</td>
<td id="76B3FE7E252EEEDFF98E539C8012FC18" box="[1662,1902,966,992]" gridcol="5" gridrow="3" pageId="11" pageNumber="47">0.73</td>
</tr>
<tr id="35629702252EEEDFFF3C53B48012FBD0" box="[204,1902,1006,1064]" gridrow="4" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C53B486BBFBD0" box="[204,455,1006,1064]" gridcol="0" gridrow="4" pageId="11" pageNumber="47">carapace maximum width</th>
<td id="76B3FE7E252EEEDFFE1353B485A5FBD0" box="[483,729,1006,1064]" gridcol="1" gridrow="4" pageId="11" pageNumber="47">1.00</td>
<td id="76B3FE7E252EEEDFFCFA53B4837CFBD0" box="[778,1024,1006,1064]" gridcol="2" gridrow="4" pageId="11" pageNumber="47">1.00</td>
<td id="76B3FE7E252EEEDFFBC153B4825DFBD0" box="[1073,1313,1006,1064]" gridcol="3" gridrow="4" pageId="11" pageNumber="47">1.07</td>
<td id="76B3FE7E252EEEDFFAA753B48120FBD0" box="[1367,1628,1006,1064]" gridcol="4" gridrow="4" pageId="11" pageNumber="47">1.10</td>
<td id="76B3FE7E252EEEDFF98E53B48012FBD0" box="[1662,1902,1006,1064]" gridcol="5" gridrow="4" pageId="11" pageNumber="47">1.10</td>
</tr>
<tr id="35629702252EEEDFFF3C546C8012FB88" box="[204,1902,1078,1136]" gridrow="5" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C546C86BBFB88" box="[204,455,1078,1136]" gridcol="0" gridrow="5" pageId="11" pageNumber="47">mesotergal scute maximum width</th>
<td id="76B3FE7E252EEEDFFE13546C85A5FB88" box="[483,729,1078,1136]" gridcol="1" gridrow="5" pageId="11" pageNumber="47">1.50</td>
<td id="76B3FE7E252EEEDFFCFA546C837CFB88" box="[778,1024,1078,1136]" gridcol="2" gridrow="5" pageId="11" pageNumber="47">1.60</td>
<td id="76B3FE7E252EEEDFFBC1546C825DFB88" box="[1073,1313,1078,1136]" gridcol="3" gridrow="5" pageId="11" pageNumber="47">1.50</td>
<td id="76B3FE7E252EEEDFFAA7546C8120FB88" box="[1367,1628,1078,1136]" gridcol="4" gridrow="5" pageId="11" pageNumber="47">1.60</td>
<td id="76B3FE7E252EEEDFF98E546C8012FB88" box="[1662,1902,1078,1136]" gridcol="5" gridrow="5" pageId="11" pageNumber="47">1.51</td>
</tr>
<tr id="35629702252EEEDFFF3C54248012FB60" box="[204,1902,1150,1176]" gridrow="6" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C542486BBFB60" box="[204,455,1150,1176]" gridcol="0" gridrow="6" pageId="11" pageNumber="47">Pedipalp tr</th>
<td id="76B3FE7E252EEEDFFE13542485A5FB60" box="[483,729,1150,1176]" gridcol="1" gridrow="6" pageId="11" pageNumber="47">0.25</td>
<td id="76B3FE7E252EEEDFFCFA5424837CFB60" box="[778,1024,1150,1176]" gridcol="2" gridrow="6" pageId="11" pageNumber="47">0.30</td>
<td id="76B3FE7E252EEEDFFBC15424825DFB60" box="[1073,1313,1150,1176]" gridcol="3" gridrow="6" pageId="11" pageNumber="47">0.24</td>
<td id="76B3FE7E252EEEDFFAA754248120FB60" box="[1367,1628,1150,1176]" gridcol="4" gridrow="6" pageId="11" pageNumber="47">0.20</td>
<td id="76B3FE7E252EEEDFF98E54248012FB60" box="[1662,1902,1150,1176]" gridcol="5" gridrow="6" pageId="11" pageNumber="47">0.25</td>
</tr>
<tr id="35629702252EEEDFFF3C54FC8012FB38" box="[204,1902,1190,1216]" gridrow="7" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C54FC86BBFB38" box="[204,455,1190,1216]" gridcol="0" gridrow="7" pageId="11" pageNumber="47">fe</th>
<td id="76B3FE7E252EEEDFFE1354FC85A5FB38" box="[483,729,1190,1216]" gridcol="1" gridrow="7" pageId="11" pageNumber="47">0.96</td>
<td id="76B3FE7E252EEEDFFCFA54FC837CFB38" box="[778,1024,1190,1216]" gridcol="2" gridrow="7" pageId="11" pageNumber="47">1.01</td>
<td id="76B3FE7E252EEEDFFBC154FC825DFB38" box="[1073,1313,1190,1216]" gridcol="3" gridrow="7" pageId="11" pageNumber="47">0.84</td>
<td id="76B3FE7E252EEEDFFAA754FC8120FB38" box="[1367,1628,1190,1216]" gridcol="4" gridrow="7" pageId="11" pageNumber="47">0.80</td>
<td id="76B3FE7E252EEEDFF98E54FC8012FB38" box="[1662,1902,1190,1216]" gridcol="5" gridrow="7" pageId="11" pageNumber="47">0.83</td>
</tr>
<tr id="35629702252EEEDFFF3C54948012FB10" box="[204,1902,1230,1256]" gridrow="8" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C549486BBFB10" box="[204,455,1230,1256]" gridcol="0" gridrow="8" pageId="11" pageNumber="47">pa</th>
<td id="76B3FE7E252EEEDFFE13549485A5FB10" box="[483,729,1230,1256]" gridcol="1" gridrow="8" pageId="11" pageNumber="47">0.43</td>
<td id="76B3FE7E252EEEDFFCFA5494837CFB10" box="[778,1024,1230,1256]" gridcol="2" gridrow="8" pageId="11" pageNumber="47">0.50</td>
<td id="76B3FE7E252EEEDFFBC15494825DFB10" box="[1073,1313,1230,1256]" gridcol="3" gridrow="8" pageId="11" pageNumber="47">0.37</td>
<td id="76B3FE7E252EEEDFFAA754948120FB10" box="[1367,1628,1230,1256]" gridcol="4" gridrow="8" pageId="11" pageNumber="47">0.30</td>
<td id="76B3FE7E252EEEDFF98E54948012FB10" box="[1662,1902,1230,1256]" gridcol="5" gridrow="8" pageId="11" pageNumber="47">0.40</td>
</tr>
<tr id="35629702252EEEDFFF3C54AF8012FAF7" box="[204,1902,1269,1295]" gridrow="9" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C54AF86BBFAF7" box="[204,455,1269,1295]" gridcol="0" gridrow="9" pageId="11" pageNumber="47">ti</th>
<td id="76B3FE7E252EEEDFFE1354AF85A5FAF7" box="[483,729,1269,1295]" gridcol="1" gridrow="9" pageId="11" pageNumber="47">0.71</td>
<td id="76B3FE7E252EEEDFFCFA54AF837CFAF7" box="[778,1024,1269,1295]" gridcol="2" gridrow="9" pageId="11" pageNumber="47">0.70</td>
<td id="76B3FE7E252EEEDFFBC154AF825DFAF7" box="[1073,1313,1269,1295]" gridcol="3" gridrow="9" pageId="11" pageNumber="47">0.62</td>
<td id="76B3FE7E252EEEDFFAA754AF8120FAF7" box="[1367,1628,1269,1295]" gridcol="4" gridrow="9" pageId="11" pageNumber="47">0.60</td>
<td id="76B3FE7E252EEEDFF98E54AF8012FAF7" box="[1662,1902,1269,1295]" gridcol="5" gridrow="9" pageId="11" pageNumber="47">0.69</td>
</tr>
<tr id="35629702252EEEDFFF3C55478012FACF" box="[204,1902,1309,1335]" gridrow="10" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C554786BBFACF" box="[204,455,1309,1335]" gridcol="0" gridrow="10" pageId="11" pageNumber="47">ta</th>
<td id="76B3FE7E252EEEDFFE13554785A5FACF" box="[483,729,1309,1335]" gridcol="1" gridrow="10" pageId="11" pageNumber="47">0.51</td>
<td id="76B3FE7E252EEEDFFCFA5547837CFACF" box="[778,1024,1309,1335]" gridcol="2" gridrow="10" pageId="11" pageNumber="47">0.61</td>
<td id="76B3FE7E252EEEDFFBC15547825DFACF" box="[1073,1313,1309,1335]" gridcol="3" gridrow="10" pageId="11" pageNumber="47">0.55</td>
<td id="76B3FE7E252EEEDFFAA755478120FACF" box="[1367,1628,1309,1335]" gridcol="4" gridrow="10" pageId="11" pageNumber="47">0.50</td>
<td id="76B3FE7E252EEEDFF98E55478012FACF" box="[1662,1902,1309,1335]" gridcol="5" gridrow="10" pageId="11" pageNumber="47">0.51</td>
</tr>
<tr id="35629702252EEEDFFF3C551F8012FAA7" box="[204,1902,1349,1375]" gridrow="11" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C551F86BBFAA7" box="[204,455,1349,1375]" gridcol="0" gridrow="11" pageId="11" pageNumber="47">Total</th>
<td id="76B3FE7E252EEEDFFE13551F85A5FAA7" box="[483,729,1349,1375]" gridcol="1" gridrow="11" pageId="11" pageNumber="47">2.86</td>
<td id="76B3FE7E252EEEDFFCFA551F837CFAA7" box="[778,1024,1349,1375]" gridcol="2" gridrow="11" pageId="11" pageNumber="47">3.12</td>
<td id="76B3FE7E252EEEDFFBC1551F825DFAA7" box="[1073,1313,1349,1375]" gridcol="3" gridrow="11" pageId="11" pageNumber="47">2.62</td>
<td id="76B3FE7E252EEEDFFAA7551F8120FAA7" box="[1367,1628,1349,1375]" gridcol="4" gridrow="11" pageId="11" pageNumber="47">2.40</td>
<td id="76B3FE7E252EEEDFF98E551F8012FAA7" box="[1662,1902,1349,1375]" gridcol="5" gridrow="11" pageId="11" pageNumber="47">2.68</td>
</tr>
<tr id="35629702252EEEDFFF3C55378012FA7F" box="[204,1902,1389,1415]" gridrow="12" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C553786BBFA7F" box="[204,455,1389,1415]" gridcol="0" gridrow="12" pageId="11" pageNumber="47">Leg I tr</th>
<td id="76B3FE7E252EEEDFFE13553785A5FA7F" box="[483,729,1389,1415]" gridcol="1" gridrow="12" pageId="11" pageNumber="47">0.30</td>
<td id="76B3FE7E252EEEDFFCFA5537837CFA7F" box="[778,1024,1389,1415]" gridcol="2" gridrow="12" pageId="11" pageNumber="47">0.33</td>
<td id="76B3FE7E252EEEDFFBC15537825DFA7F" box="[1073,1313,1389,1415]" gridcol="3" gridrow="12" pageId="11" pageNumber="47">0.21</td>
<td id="76B3FE7E252EEEDFFAA755378120FA7F" box="[1367,1628,1389,1415]" gridcol="4" gridrow="12" pageId="11" pageNumber="47">0.20</td>
<td id="76B3FE7E252EEEDFF98E55378012FA7F" box="[1662,1902,1389,1415]" gridcol="5" gridrow="12" pageId="11" pageNumber="47">0.24</td>
</tr>
<tr id="35629702252EEEDFFF3C55CF8012FA57" box="[204,1902,1429,1455]" gridrow="13" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C55CF86BBFA57" box="[204,455,1429,1455]" gridcol="0" gridrow="13" pageId="11" pageNumber="47">fe</th>
<td id="76B3FE7E252EEEDFFE1355CF85A5FA57" box="[483,729,1429,1455]" gridcol="1" gridrow="13" pageId="11" pageNumber="47">1.70</td>
<td id="76B3FE7E252EEEDFFCFA55CF837CFA57" box="[778,1024,1429,1455]" gridcol="2" gridrow="13" pageId="11" pageNumber="47">1.64</td>
<td id="76B3FE7E252EEEDFFBC155CF825DFA57" box="[1073,1313,1429,1455]" gridcol="3" gridrow="13" pageId="11" pageNumber="47">1.29</td>
<td id="76B3FE7E252EEEDFFAA755CF8120FA57" box="[1367,1628,1429,1455]" gridcol="4" gridrow="13" pageId="11" pageNumber="47">0.90</td>
<td id="76B3FE7E252EEEDFF98E55CF8012FA57" box="[1662,1902,1429,1455]" gridcol="5" gridrow="13" pageId="11" pageNumber="47">1.12</td>
</tr>
<tr id="35629702252EEEDFFF3C55E68012FA2E" box="[204,1902,1468,1494]" gridrow="14" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C55E686BBFA2E" box="[204,455,1468,1494]" gridcol="0" gridrow="14" pageId="11" pageNumber="47">pa</th>
<td id="76B3FE7E252EEEDFFE1355E685A5FA2E" box="[483,729,1468,1494]" gridcol="1" gridrow="14" pageId="11" pageNumber="47">0.58</td>
<td id="76B3FE7E252EEEDFFCFA55E6837CFA2E" box="[778,1024,1468,1494]" gridcol="2" gridrow="14" pageId="11" pageNumber="47">0.60</td>
<td id="76B3FE7E252EEEDFFBC155E6825DFA2E" box="[1073,1313,1468,1494]" gridcol="3" gridrow="14" pageId="11" pageNumber="47">0.35</td>
<td id="76B3FE7E252EEEDFFAA755E68120FA2E" box="[1367,1628,1468,1494]" gridcol="4" gridrow="14" pageId="11" pageNumber="47">0.50</td>
<td id="76B3FE7E252EEEDFF98E55E68012FA2E" box="[1662,1902,1468,1494]" gridcol="5" gridrow="14" pageId="11" pageNumber="47">0.38</td>
</tr>
<tr id="35629702252EEEDFFF3C55BE8012FA06" box="[204,1902,1508,1534]" gridrow="15" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C55BE86BBFA06" box="[204,455,1508,1534]" gridcol="0" gridrow="15" pageId="11" pageNumber="47">ti</th>
<td id="76B3FE7E252EEEDFFE1355BE85A5FA06" box="[483,729,1508,1534]" gridcol="1" gridrow="15" pageId="11" pageNumber="47">1.26</td>
<td id="76B3FE7E252EEEDFFCFA55BE837CFA06" box="[778,1024,1508,1534]" gridcol="2" gridrow="15" pageId="11" pageNumber="47">1.23</td>
<td id="76B3FE7E252EEEDFFBC155BE825DFA06" box="[1073,1313,1508,1534]" gridcol="3" gridrow="15" pageId="11" pageNumber="47">0.88</td>
<td id="76B3FE7E252EEEDFFAA755BE8120FA06" box="[1367,1628,1508,1534]" gridcol="4" gridrow="15" pageId="11" pageNumber="47">0.80</td>
<td id="76B3FE7E252EEEDFF98E55BE8012FA06" box="[1662,1902,1508,1534]" gridcol="5" gridrow="15" pageId="11" pageNumber="47">0.83</td>
</tr>
<tr id="35629702252EEEDFFF3C56568012F9DE" box="[204,1902,1548,1574]" gridrow="16" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C565686BBF9DE" box="[204,455,1548,1574]" gridcol="0" gridrow="16" pageId="11" pageNumber="47">me</th>
<td id="76B3FE7E252EEEDFFE13565685A5F9DE" box="[483,729,1548,1574]" gridcol="1" gridrow="16" pageId="11" pageNumber="47">1.78</td>
<td id="76B3FE7E252EEEDFFCFA5656837CF9DE" box="[778,1024,1548,1574]" gridcol="2" gridrow="16" pageId="11" pageNumber="47">1.63</td>
<td id="76B3FE7E252EEEDFFBC15656825DF9DE" box="[1073,1313,1548,1574]" gridcol="3" gridrow="16" pageId="11" pageNumber="47">1.17</td>
<td id="76B3FE7E252EEEDFFAA756568120F9DE" box="[1367,1628,1548,1574]" gridcol="4" gridrow="16" pageId="11" pageNumber="47">1.10</td>
<td id="76B3FE7E252EEEDFF98E56568012F9DE" box="[1662,1902,1548,1574]" gridcol="5" gridrow="16" pageId="11" pageNumber="47">1.05</td>
</tr>
<tr id="35629702252EEEDFFF3C566E8012F9B6" box="[204,1902,1588,1614]" gridrow="17" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C566E86BBF9B6" box="[204,455,1588,1614]" gridcol="0" gridrow="17" pageId="11" pageNumber="47">ta</th>
<td id="76B3FE7E252EEEDFFE13566E85A5F9B6" box="[483,729,1588,1614]" gridcol="1" gridrow="17" pageId="11" pageNumber="47">1.16</td>
<td id="76B3FE7E252EEEDFFCFA566E837CF9B6" box="[778,1024,1588,1614]" gridcol="2" gridrow="17" pageId="11" pageNumber="47">1.20</td>
<td id="76B3FE7E252EEEDFFBC1566E825DF9B6" box="[1073,1313,1588,1614]" gridcol="3" gridrow="17" pageId="11" pageNumber="47">0.77</td>
<td id="76B3FE7E252EEEDFFAA7566E8120F9B6" box="[1367,1628,1588,1614]" gridcol="4" gridrow="17" pageId="11" pageNumber="47">0.70</td>
<td id="76B3FE7E252EEEDFF98E566E8012F9B6" box="[1662,1902,1588,1614]" gridcol="5" gridrow="17" pageId="11" pageNumber="47">0.75</td>
</tr>
<tr id="35629702252EEEDFFF3C56068012F98E" box="[204,1902,1628,1654]" gridrow="18" pageId="11" pageNumber="47">
<th id="76B3FE7E252EEEDFFF3C560686BBF98E" box="[204,455,1628,1654]" gridcol="0" gridrow="18" pageId="11" pageNumber="47">Total</th>
<td id="76B3FE7E252EEEDFFE13560685A5F98E" box="[483,729,1628,1654]" gridcol="1" gridrow="18" pageId="11" pageNumber="47">6.78</td>
<td id="76B3FE7E252EEEDFFCFA5606837CF98E" box="[778,1024,1628,1654]" gridcol="2" gridrow="18" pageId="11" pageNumber="47">6.63</td>
<td id="76B3FE7E252EEEDFFBC15606825DF98E" box="[1073,1313,1628,1654]" gridcol="3" gridrow="18" pageId="11" pageNumber="47">4.67</td>
<td id="76B3FE7E252EEEDFFAA756068120F98E" box="[1367,1628,1628,1654]" gridcol="4" gridrow="18" pageId="11" pageNumber="47">4.20</td>
<td id="76B3FE7E252EEEDFF98E56068012F98E" box="[1662,1902,1628,1654]" gridcol="5" gridrow="18" pageId="11" pageNumber="47">4.37</td>
</tr>
</table>
</paragraph>
<caption id="DF2DC5C82529112CFF4C521E8239FD7B" isContinuationCaption="true" pageId="12" pageNumber="48" startId="12.[188,257,580,609]" targetBox="[198,1896,754,1703]" targetIsTable="true" targetPageId="12">
<paragraph id="8BED95402529112CFF4C521E8239FD7B" blockId="12.[188,1941,580,643]" pageId="12" pageNumber="48">
<emphasis id="B92649522529112CFF4C521E866BFD99" bold="true" box="[188,279,580,609]" pageId="12" pageNumber="48">Table 1</emphasis>
(continued). Body and leg measurements (in mm) of analyzed specimens of
<taxonomicName id="4C52EEC32529112CFB54521E810AFD98" authority="H. Soares, 1966" authorityName="H. Soares" authorityYear="1966" box="[1188,1654,580,608]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B92649522529112CFB54521E82CEFD98" box="[1188,1458,580,608]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes spaeleus</emphasis>
H.
<bibRefCitation id="EFC3E8B12529112CFA2E521E810AFD98" author="Soares" box="[1502,1654,580,608]" firstAuthor="Soares" pageId="12" pageNumber="48" pagination="103 - 115" refId="ref10502" refString="Soares H. E. M. 1966. Novos opilioes da colecao &quot; Otto Schubart &quot; (Opiliones: Cosmetidae, Gonyleptidae, Phalangodidae). Papeis avulsos do Departamento de Zoologia da Secretaria de Agricultura 18 (11): 103 - 115." type="journal article" year="1966">Soares, 1966</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C52EEC32529112CF940521E80E9FD98" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1712,1941,580,608]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B92649522529112CF940521E80E9FD98" box="[1712,1941,580,608]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F4292529112CFF4C523D866AFD7B" box="[188,278,615,643]" pageId="12" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: tr =trochanter; fe =femur; pa=patella; ti =tibia; me=metatarsus; ta =tarsus.
</paragraph>
</caption>
<paragraph id="8BED95402529112CFE2E52A881D4F95F" pageId="12" pageNumber="48">
<table id="F95267E02529EEDFFF3652A88014F95F" box="[198,1896,754,1703]" colsContinueFrom="11.[204,1902,759,1654]" gridcols="6" gridrows="22" pageId="12" pageNumber="48">
<tr id="356297022529EEDFFF3652A88014FCB4" box="[198,1896,754,844]" gridrow="0" isHeader="true" pageId="12" pageNumber="48" rowspan-0="1">
<th id="76B3FE7E2529EEDFFE2D52A885AFFCB4" box="[477,723,754,844]" gridcol="1" gridrow="0" pageId="12" pageNumber="48">
<taxonomicName id="4C52EEC32529112CFE2E52A885AFFCF4" authorityName="H. Soares" authorityYear="1966" box="[478,723,754,780]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B92649522529112CFE2E52A885AFFCF4" box="[478,723,754,780]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
male holotype (MZUSP 28463)
</th>
<th id="76B3FE7E2529EEDFFCF452A88486FCB4" box="[772,1018,754,844]" gridcol="2" gridrow="0" pageId="12" pageNumber="48">
<taxonomicName id="4C52EEC32529112CFCF552A88486FCF4" authorityName="H. Soares" authorityYear="1966" box="[773,1018,754,780]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="spaeleus">
<emphasis id="B92649522529112CFCF552A88486FCF4" box="[773,1018,754,780]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes spaeleus</emphasis>
</taxonomicName>
female paratype (MZUSP 28462)
</th>
<th id="76B3FE7E2529EEDFFBDB52A88260FCB4" box="[1067,1308,754,844]" gridcol="3" gridrow="0" pageId="12" pageNumber="48">
<taxonomicName id="4C52EEC32529112CFBDB52A88380FCF4" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1067,1276,754,780]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B92649522529112CFBDB52A88380FCF4" box="[1067,1276,754,780]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F4292529112CFBDB53488302FCD4" box="[1067,1150,786,812]" pageId="12" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
male holotype (LES 00341)
</th>
<th id="76B3FE7E2529EEDFFAA252A8812AFCB4" box="[1362,1622,754,844]" gridcol="4" gridrow="0" pageId="12" pageNumber="48">
<taxonomicName id="4C52EEC32529112CFAA252A8815FFCF4" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1362,1571,754,780]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B92649522529112CFAA252A8815FFCF4" box="[1362,1571,754,780]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F4292529112CFAA2534882D9FCD4" box="[1362,1445,786,812]" pageId="12" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
female paratype (LES 027941)
</th>
<th id="76B3FE7E2529EEDFF98952A88014FCB4" box="[1657,1896,754,844]" gridcol="5" gridrow="0" pageId="12" pageNumber="48">
<taxonomicName id="4C52EEC32529112CF98952A88035FCF4" authority="Pereira &amp; Bichuette &amp; Pérez-González, 2024" authorityName="Pereira &amp; Bichuette &amp; Pérez-González" authorityYear="2024" box="[1657,1865,754,780]" class="Arachnida" family="Escadabiidae" genus="Spaeleoleptes" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="48" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="B92649522529112CF98952A88035FCF4" box="[1657,1865,754,780]" italics="true" pageId="12" pageNumber="48">Spaeleoleptes gimli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215F4292529112CF989534881B0FCD4" box="[1657,1740,786,812]" pageId="12" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
male paratype (LES 027941)
</th>
</tr>
<tr id="356297022529EEDFFF36532B8014FC73" box="[198,1896,881,907]" gridrow="1" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36532B86FFFC73" box="[198,387,881,907]" gridcol="0" gridrow="1" pageId="12" pageNumber="48">Leg II tr</th>
<td id="76B3FE7E2529EEDFFE2D532B85AFFC73" box="[477,723,881,907]" gridcol="1" gridrow="1" pageId="12" pageNumber="48">0.38</td>
<td id="76B3FE7E2529EEDFFCF4532B8486FC73" box="[772,1018,881,907]" gridcol="2" gridrow="1" pageId="12" pageNumber="48">0.40</td>
<td id="76B3FE7E2529EEDFFBDB532B8260FC73" box="[1067,1308,881,907]" gridcol="3" gridrow="1" pageId="12" pageNumber="48">0.25</td>
<td id="76B3FE7E2529EEDFFAA2532B812AFC73" box="[1362,1622,881,907]" gridcol="4" gridrow="1" pageId="12" pageNumber="48">0.30</td>
<td id="76B3FE7E2529EEDFF989532B8014FC73" box="[1657,1896,881,907]" gridcol="5" gridrow="1" pageId="12" pageNumber="48">0.24</td>
</tr>
<tr id="356297022529EEDFFF3653C38014FC4B" box="[198,1896,921,947]" gridrow="2" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3653C386FFFC4B" box="[198,387,921,947]" gridcol="0" gridrow="2" pageId="12" pageNumber="48">fe</th>
<td id="76B3FE7E2529EEDFFE2D53C385AFFC4B" box="[477,723,921,947]" gridcol="1" gridrow="2" pageId="12" pageNumber="48">2.40</td>
<td id="76B3FE7E2529EEDFFCF453C38486FC4B" box="[772,1018,921,947]" gridcol="2" gridrow="2" pageId="12" pageNumber="48">2.43</td>
<td id="76B3FE7E2529EEDFFBDB53C38260FC4B" box="[1067,1308,921,947]" gridcol="3" gridrow="2" pageId="12" pageNumber="48">1.68</td>
<td id="76B3FE7E2529EEDFFAA253C3812AFC4B" box="[1362,1622,921,947]" gridcol="4" gridrow="2" pageId="12" pageNumber="48">1.50</td>
<td id="76B3FE7E2529EEDFF98953C38014FC4B" box="[1657,1896,921,947]" gridcol="5" gridrow="2" pageId="12" pageNumber="48">1.57</td>
</tr>
<tr id="356297022529EEDFFF36539B8014FC23" box="[198,1896,961,987]" gridrow="3" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36539B86FFFC23" box="[198,387,961,987]" gridcol="0" gridrow="3" pageId="12" pageNumber="48">pa</th>
<td id="76B3FE7E2529EEDFFE2D539B85AFFC23" box="[477,723,961,987]" gridcol="1" gridrow="3" pageId="12" pageNumber="48">0.76</td>
<td id="76B3FE7E2529EEDFFCF4539B8486FC23" box="[772,1018,961,987]" gridcol="2" gridrow="3" pageId="12" pageNumber="48">0.82</td>
<td id="76B3FE7E2529EEDFFBDB539B8260FC23" box="[1067,1308,961,987]" gridcol="3" gridrow="3" pageId="12" pageNumber="48">0.59</td>
<td id="76B3FE7E2529EEDFFAA2539B812AFC23" box="[1362,1622,961,987]" gridcol="4" gridrow="3" pageId="12" pageNumber="48">0.60</td>
<td id="76B3FE7E2529EEDFF989539B8014FC23" box="[1657,1896,961,987]" gridcol="5" gridrow="3" pageId="12" pageNumber="48">0.48</td>
</tr>
<tr id="356297022529EEDFFF3653B38014FBFB" box="[198,1896,1001,1027]" gridrow="4" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3653B386FFFBFB" box="[198,387,1001,1027]" gridcol="0" gridrow="4" pageId="12" pageNumber="48">ti</th>
<td id="76B3FE7E2529EEDFFE2D53B385AFFBFB" box="[477,723,1001,1027]" gridcol="1" gridrow="4" pageId="12" pageNumber="48">-</td>
<td id="76B3FE7E2529EEDFFCF453B38486FBFB" box="[772,1018,1001,1027]" gridcol="2" gridrow="4" pageId="12" pageNumber="48">2.04</td>
<td id="76B3FE7E2529EEDFFBDB53B38260FBFB" box="[1067,1308,1001,1027]" gridcol="3" gridrow="4" pageId="12" pageNumber="48">1.39</td>
<td id="76B3FE7E2529EEDFFAA253B3812AFBFB" box="[1362,1622,1001,1027]" gridcol="4" gridrow="4" pageId="12" pageNumber="48">1.30</td>
<td id="76B3FE7E2529EEDFF98953B38014FBFB" box="[1657,1896,1001,1027]" gridcol="5" gridrow="4" pageId="12" pageNumber="48">1.32</td>
</tr>
<tr id="356297022529EEDFFF36544A8014FBD2" box="[198,1896,1040,1066]" gridrow="5" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36544A86FFFBD2" box="[198,387,1040,1066]" gridcol="0" gridrow="5" pageId="12" pageNumber="48">me</th>
<td id="76B3FE7E2529EEDFFE2D544A85AFFBD2" box="[477,723,1040,1066]" gridcol="1" gridrow="5" pageId="12" pageNumber="48">-</td>
<td id="76B3FE7E2529EEDFFCF4544A8486FBD2" box="[772,1018,1040,1066]" gridcol="2" gridrow="5" pageId="12" pageNumber="48">2.11</td>
<td id="76B3FE7E2529EEDFFBDB544A8260FBD2" box="[1067,1308,1040,1066]" gridcol="3" gridrow="5" pageId="12" pageNumber="48">1.57</td>
<td id="76B3FE7E2529EEDFFAA2544A812AFBD2" box="[1362,1622,1040,1066]" gridcol="4" gridrow="5" pageId="12" pageNumber="48">1.50</td>
<td id="76B3FE7E2529EEDFF989544A8014FBD2" box="[1657,1896,1040,1066]" gridcol="5" gridrow="5" pageId="12" pageNumber="48">1.40</td>
</tr>
<tr id="356297022529EEDFFF3654628014FBAA" box="[198,1896,1080,1106]" gridrow="6" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36546286FFFBAA" box="[198,387,1080,1106]" gridcol="0" gridrow="6" pageId="12" pageNumber="48">ta</th>
<td id="76B3FE7E2529EEDFFE2D546285AFFBAA" box="[477,723,1080,1106]" gridcol="1" gridrow="6" pageId="12" pageNumber="48">-</td>
<td id="76B3FE7E2529EEDFFCF454628486FBAA" box="[772,1018,1080,1106]" gridcol="2" gridrow="6" pageId="12" pageNumber="48">2.98</td>
<td id="76B3FE7E2529EEDFFBDB54628260FBAA" box="[1067,1308,1080,1106]" gridcol="3" gridrow="6" pageId="12" pageNumber="48">1.57</td>
<td id="76B3FE7E2529EEDFFAA25462812AFBAA" box="[1362,1622,1080,1106]" gridcol="4" gridrow="6" pageId="12" pageNumber="48">1.30</td>
<td id="76B3FE7E2529EEDFF98954628014FBAA" box="[1657,1896,1080,1106]" gridcol="5" gridrow="6" pageId="12" pageNumber="48">1.48</td>
</tr>
<tr id="356297022529EEDFFF36543A8014FB82" box="[198,1896,1120,1146]" gridrow="7" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36543A86FFFB82" box="[198,387,1120,1146]" gridcol="0" gridrow="7" pageId="12" pageNumber="48">Total</th>
<td id="76B3FE7E2529EEDFFE2D543A85AFFB82" box="[477,723,1120,1146]" gridcol="1" gridrow="7" pageId="12" pageNumber="48">-</td>
<td id="76B3FE7E2529EEDFFCF4543A8486FB82" box="[772,1018,1120,1146]" gridcol="2" gridrow="7" pageId="12" pageNumber="48">10.78</td>
<td id="76B3FE7E2529EEDFFBDB543A8260FB82" box="[1067,1308,1120,1146]" gridcol="3" gridrow="7" pageId="12" pageNumber="48">7.05</td>
<td id="76B3FE7E2529EEDFFAA2543A812AFB82" box="[1362,1622,1120,1146]" gridcol="4" gridrow="7" pageId="12" pageNumber="48">6.50</td>
<td id="76B3FE7E2529EEDFF989543A8014FB82" box="[1657,1896,1120,1146]" gridcol="5" gridrow="7" pageId="12" pageNumber="48">6.49</td>
</tr>
<tr id="356297022529EEDFFF3654D28014FB5A" box="[198,1896,1160,1186]" gridrow="8" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3654D286FFFB5A" box="[198,387,1160,1186]" gridcol="0" gridrow="8" pageId="12" pageNumber="48">Leg III tr</th>
<td id="76B3FE7E2529EEDFFE2D54D285AFFB5A" box="[477,723,1160,1186]" gridcol="1" gridrow="8" pageId="12" pageNumber="48">0.37</td>
<td id="76B3FE7E2529EEDFFCF454D28486FB5A" box="[772,1018,1160,1186]" gridcol="2" gridrow="8" pageId="12" pageNumber="48">0.36</td>
<td id="76B3FE7E2529EEDFFBDB54D28260FB5A" box="[1067,1308,1160,1186]" gridcol="3" gridrow="8" pageId="12" pageNumber="48">0.28</td>
<td id="76B3FE7E2529EEDFFAA254D2812AFB5A" box="[1362,1622,1160,1186]" gridcol="4" gridrow="8" pageId="12" pageNumber="48">0.20</td>
<td id="76B3FE7E2529EEDFF98954D28014FB5A" box="[1657,1896,1160,1186]" gridcol="5" gridrow="8" pageId="12" pageNumber="48">0.27</td>
</tr>
<tr id="356297022529EEDFFF3654EA8014FB32" box="[198,1896,1200,1226]" gridrow="9" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3654EA86FFFB32" box="[198,387,1200,1226]" gridcol="0" gridrow="9" pageId="12" pageNumber="48">fe</th>
<td id="76B3FE7E2529EEDFFE2D54EA85AFFB32" box="[477,723,1200,1226]" gridcol="1" gridrow="9" pageId="12" pageNumber="48">1.76</td>
<td id="76B3FE7E2529EEDFFCF454EA8486FB32" box="[772,1018,1200,1226]" gridcol="2" gridrow="9" pageId="12" pageNumber="48">1.85</td>
<td id="76B3FE7E2529EEDFFBDB54EA8260FB32" box="[1067,1308,1200,1226]" gridcol="3" gridrow="9" pageId="12" pageNumber="48">1.24</td>
<td id="76B3FE7E2529EEDFFAA254EA812AFB32" box="[1362,1622,1200,1226]" gridcol="4" gridrow="9" pageId="12" pageNumber="48">0.80</td>
<td id="76B3FE7E2529EEDFF98954EA8014FB32" box="[1657,1896,1200,1226]" gridcol="5" gridrow="9" pageId="12" pageNumber="48">1.12</td>
</tr>
<tr id="356297022529EEDFFF36548D8014FB09" box="[198,1896,1239,1265]" gridrow="10" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36548D86FFFB09" box="[198,387,1239,1265]" gridcol="0" gridrow="10" pageId="12" pageNumber="48">pa</th>
<td id="76B3FE7E2529EEDFFE2D548D85AFFB09" box="[477,723,1239,1265]" gridcol="1" gridrow="10" pageId="12" pageNumber="48">0.62</td>
<td id="76B3FE7E2529EEDFFCF4548D8486FB09" box="[772,1018,1239,1265]" gridcol="2" gridrow="10" pageId="12" pageNumber="48">0.59</td>
<td id="76B3FE7E2529EEDFFBDB548D8260FB09" box="[1067,1308,1239,1265]" gridcol="3" gridrow="10" pageId="12" pageNumber="48">0.33</td>
<td id="76B3FE7E2529EEDFFAA2548D812AFB09" box="[1362,1622,1239,1265]" gridcol="4" gridrow="10" pageId="12" pageNumber="48">0.50</td>
<td id="76B3FE7E2529EEDFF989548D8014FB09" box="[1657,1896,1239,1265]" gridcol="5" gridrow="10" pageId="12" pageNumber="48">0.37</td>
</tr>
<tr id="356297022529EEDFFF3654A58014FAE1" box="[198,1896,1279,1305]" gridrow="11" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3654A586FFFAE1" box="[198,387,1279,1305]" gridcol="0" gridrow="11" pageId="12" pageNumber="48">ti</th>
<td id="76B3FE7E2529EEDFFE2D54A585AFFAE1" box="[477,723,1279,1305]" gridcol="1" gridrow="11" pageId="12" pageNumber="48">1.44</td>
<td id="76B3FE7E2529EEDFFCF454A58486FAE1" box="[772,1018,1279,1305]" gridcol="2" gridrow="11" pageId="12" pageNumber="48">1.41</td>
<td id="76B3FE7E2529EEDFFBDB54A58260FAE1" box="[1067,1308,1279,1305]" gridcol="3" gridrow="11" pageId="12" pageNumber="48">0.95</td>
<td id="76B3FE7E2529EEDFFAA254A5812AFAE1" box="[1362,1622,1279,1305]" gridcol="4" gridrow="11" pageId="12" pageNumber="48">0.90</td>
<td id="76B3FE7E2529EEDFF98954A58014FAE1" box="[1657,1896,1279,1305]" gridcol="5" gridrow="11" pageId="12" pageNumber="48">0.86</td>
</tr>
<tr id="356297022529EEDFFF36557D8014FAB9" box="[198,1896,1319,1345]" gridrow="12" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36557D86FFFAB9" box="[198,387,1319,1345]" gridcol="0" gridrow="12" pageId="12" pageNumber="48">me</th>
<td id="76B3FE7E2529EEDFFE2D557D85AFFAB9" box="[477,723,1319,1345]" gridcol="1" gridrow="12" pageId="12" pageNumber="48">1.83</td>
<td id="76B3FE7E2529EEDFFCF4557D8486FAB9" box="[772,1018,1319,1345]" gridcol="2" gridrow="12" pageId="12" pageNumber="48">1.77</td>
<td id="76B3FE7E2529EEDFFBDB557D8260FAB9" box="[1067,1308,1319,1345]" gridcol="3" gridrow="12" pageId="12" pageNumber="48">1.16</td>
<td id="76B3FE7E2529EEDFFAA2557D812AFAB9" box="[1362,1622,1319,1345]" gridcol="4" gridrow="12" pageId="12" pageNumber="48">1.10</td>
<td id="76B3FE7E2529EEDFF989557D8014FAB9" box="[1657,1896,1319,1345]" gridcol="5" gridrow="12" pageId="12" pageNumber="48">1.15</td>
</tr>
<tr id="356297022529EEDFFF3655158014FA91" box="[198,1896,1359,1385]" gridrow="13" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36551586FFFA91" box="[198,387,1359,1385]" gridcol="0" gridrow="13" pageId="12" pageNumber="48">ta</th>
<td id="76B3FE7E2529EEDFFE2D551585AFFA91" box="[477,723,1359,1385]" gridcol="1" gridrow="13" pageId="12" pageNumber="48">1.92</td>
<td id="76B3FE7E2529EEDFFCF455158486FA91" box="[772,1018,1359,1385]" gridcol="2" gridrow="13" pageId="12" pageNumber="48">1.89</td>
<td id="76B3FE7E2529EEDFFBDB55158260FA91" box="[1067,1308,1359,1385]" gridcol="3" gridrow="13" pageId="12" pageNumber="48">1.11</td>
<td id="76B3FE7E2529EEDFFAA25515812AFA91" box="[1362,1622,1359,1385]" gridcol="4" gridrow="13" pageId="12" pageNumber="48">1.10</td>
<td id="76B3FE7E2529EEDFF98955158014FA91" box="[1657,1896,1359,1385]" gridcol="5" gridrow="13" pageId="12" pageNumber="48">0.98</td>
</tr>
<tr id="356297022529EEDFFF36552D8014FA69" box="[198,1896,1399,1425]" gridrow="14" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36552D86FFFA69" box="[198,387,1399,1425]" gridcol="0" gridrow="14" pageId="12" pageNumber="48">Total</th>
<td id="76B3FE7E2529EEDFFE2D552D85AFFA69" box="[477,723,1399,1425]" gridcol="1" gridrow="14" pageId="12" pageNumber="48">7.94</td>
<td id="76B3FE7E2529EEDFFCF4552D8486FA69" box="[772,1018,1399,1425]" gridcol="2" gridrow="14" pageId="12" pageNumber="48">7.87</td>
<td id="76B3FE7E2529EEDFFBDB552D8260FA69" box="[1067,1308,1399,1425]" gridcol="3" gridrow="14" pageId="12" pageNumber="48">5.07</td>
<td id="76B3FE7E2529EEDFFAA2552D812AFA69" box="[1362,1622,1399,1425]" gridcol="4" gridrow="14" pageId="12" pageNumber="48">4.60</td>
<td id="76B3FE7E2529EEDFF989552D8014FA69" box="[1657,1896,1399,1425]" gridcol="5" gridrow="14" pageId="12" pageNumber="48">4.75</td>
</tr>
<tr id="356297022529EEDFFF3655C58014FA41" box="[198,1896,1439,1465]" gridrow="15" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3655C586FFFA41" box="[198,387,1439,1465]" gridcol="0" gridrow="15" pageId="12" pageNumber="48">Leg IV tr</th>
<td id="76B3FE7E2529EEDFFE2D55C585AFFA41" box="[477,723,1439,1465]" gridcol="1" gridrow="15" pageId="12" pageNumber="48">0.37</td>
<td id="76B3FE7E2529EEDFFCF455C58486FA41" box="[772,1018,1439,1465]" gridcol="2" gridrow="15" pageId="12" pageNumber="48">0.44</td>
<td id="76B3FE7E2529EEDFFBDB55C58260FA41" box="[1067,1308,1439,1465]" gridcol="3" gridrow="15" pageId="12" pageNumber="48">0.32</td>
<td id="76B3FE7E2529EEDFFAA255C5812AFA41" box="[1362,1622,1439,1465]" gridcol="4" gridrow="15" pageId="12" pageNumber="48">0.40</td>
<td id="76B3FE7E2529EEDFF98955C58014FA41" box="[1657,1896,1439,1465]" gridcol="5" gridrow="15" pageId="12" pageNumber="48">0.27</td>
</tr>
<tr id="356297022529EEDFFF36559C8014FA18" box="[198,1896,1478,1504]" gridrow="16" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36559C86FFFA18" box="[198,387,1478,1504]" gridcol="0" gridrow="16" pageId="12" pageNumber="48">fe</th>
<td id="76B3FE7E2529EEDFFE2D559C85AFFA18" box="[477,723,1478,1504]" gridcol="1" gridrow="16" pageId="12" pageNumber="48">2.36</td>
<td id="76B3FE7E2529EEDFFCF4559C8486FA18" box="[772,1018,1478,1504]" gridcol="2" gridrow="16" pageId="12" pageNumber="48">2.39</td>
<td id="76B3FE7E2529EEDFFBDB559C8260FA18" box="[1067,1308,1478,1504]" gridcol="3" gridrow="16" pageId="12" pageNumber="48">1.58</td>
<td id="76B3FE7E2529EEDFFAA2559C812AFA18" box="[1362,1622,1478,1504]" gridcol="4" gridrow="16" pageId="12" pageNumber="48">1.70</td>
<td id="76B3FE7E2529EEDFF989559C8014FA18" box="[1657,1896,1478,1504]" gridcol="5" gridrow="16" pageId="12" pageNumber="48">1.39</td>
</tr>
<tr id="356297022529EEDFFF3655B48014F9F0" box="[198,1896,1518,1544]" gridrow="17" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3655B486FFF9F0" box="[198,387,1518,1544]" gridcol="0" gridrow="17" pageId="12" pageNumber="48">pa</th>
<td id="76B3FE7E2529EEDFFE2D55B485AFF9F0" box="[477,723,1518,1544]" gridcol="1" gridrow="17" pageId="12" pageNumber="48">0.69</td>
<td id="76B3FE7E2529EEDFFCF455B48486F9F0" box="[772,1018,1518,1544]" gridcol="2" gridrow="17" pageId="12" pageNumber="48">0.73</td>
<td id="76B3FE7E2529EEDFFBDB55B48260F9F0" box="[1067,1308,1518,1544]" gridcol="3" gridrow="17" pageId="12" pageNumber="48">0.60</td>
<td id="76B3FE7E2529EEDFFAA255B4812AF9F0" box="[1362,1622,1518,1544]" gridcol="4" gridrow="17" pageId="12" pageNumber="48">0.40</td>
<td id="76B3FE7E2529EEDFF98955B48014F9F0" box="[1657,1896,1518,1544]" gridcol="5" gridrow="17" pageId="12" pageNumber="48">1.48</td>
</tr>
<tr id="356297022529EEDFFF36564C8014F9C8" box="[198,1896,1558,1584]" gridrow="18" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36564C86FFF9C8" box="[198,387,1558,1584]" gridcol="0" gridrow="18" pageId="12" pageNumber="48">ti</th>
<td id="76B3FE7E2529EEDFFE2D564C85AFF9C8" box="[477,723,1558,1584]" gridcol="1" gridrow="18" pageId="12" pageNumber="48">2.06</td>
<td id="76B3FE7E2529EEDFFCF4564C8486F9C8" box="[772,1018,1558,1584]" gridcol="2" gridrow="18" pageId="12" pageNumber="48">1.93</td>
<td id="76B3FE7E2529EEDFFBDB564C8260F9C8" box="[1067,1308,1558,1584]" gridcol="3" gridrow="18" pageId="12" pageNumber="48">1.30</td>
<td id="76B3FE7E2529EEDFFAA2564C812AF9C8" box="[1362,1622,1558,1584]" gridcol="4" gridrow="18" pageId="12" pageNumber="48">1.30</td>
<td id="76B3FE7E2529EEDFF989564C8014F9C8" box="[1657,1896,1558,1584]" gridcol="5" gridrow="18" pageId="12" pageNumber="48">1.24</td>
</tr>
<tr id="356297022529EEDFFF3656648014F9A0" box="[198,1896,1598,1624]" gridrow="19" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36566486FFF9A0" box="[198,387,1598,1624]" gridcol="0" gridrow="19" pageId="12" pageNumber="48">me</th>
<td id="76B3FE7E2529EEDFFE2D566485AFF9A0" box="[477,723,1598,1624]" gridcol="1" gridrow="19" pageId="12" pageNumber="48">2.34</td>
<td id="76B3FE7E2529EEDFFCF456648486F9A0" box="[772,1018,1598,1624]" gridcol="2" gridrow="19" pageId="12" pageNumber="48">2.32</td>
<td id="76B3FE7E2529EEDFFBDB56648260F9A0" box="[1067,1308,1598,1624]" gridcol="3" gridrow="19" pageId="12" pageNumber="48">1.59</td>
<td id="76B3FE7E2529EEDFFAA25664812AF9A0" box="[1362,1622,1598,1624]" gridcol="4" gridrow="19" pageId="12" pageNumber="48">1.60</td>
<td id="76B3FE7E2529EEDFF98956648014F9A0" box="[1657,1896,1598,1624]" gridcol="5" gridrow="19" pageId="12" pageNumber="48">1.59</td>
</tr>
<tr id="356297022529EEDFFF36563C8014F978" box="[198,1896,1638,1664]" gridrow="20" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF36563C86FFF978" box="[198,387,1638,1664]" gridcol="0" gridrow="20" pageId="12" pageNumber="48">ta</th>
<td id="76B3FE7E2529EEDFFE2D563C85AFF978" box="[477,723,1638,1664]" gridcol="1" gridrow="20" pageId="12" pageNumber="48">2.45</td>
<td id="76B3FE7E2529EEDFFCF4563C8486F978" box="[772,1018,1638,1664]" gridcol="2" gridrow="20" pageId="12" pageNumber="48">2.42</td>
<td id="76B3FE7E2529EEDFFBDB563C8260F978" box="[1067,1308,1638,1664]" gridcol="3" gridrow="20" pageId="12" pageNumber="48">1.28</td>
<td id="76B3FE7E2529EEDFFAA2563C812AF978" box="[1362,1622,1638,1664]" gridcol="4" gridrow="20" pageId="12" pageNumber="48">1.30</td>
<td id="76B3FE7E2529EEDFF989563C8014F978" box="[1657,1896,1638,1664]" gridcol="5" gridrow="20" pageId="12" pageNumber="48">1.16</td>
</tr>
<tr id="356297022529EEDFFF3656D78014F95F" box="[198,1896,1677,1703]" gridrow="21" pageId="12" pageNumber="48">
<th id="76B3FE7E2529EEDFFF3656D786FFF95F" box="[198,387,1677,1703]" gridcol="0" gridrow="21" pageId="12" pageNumber="48">Total</th>
<td id="76B3FE7E2529EEDFFE2D56D785AFF95F" box="[477,723,1677,1703]" gridcol="1" gridrow="21" pageId="12" pageNumber="48">10.27</td>
<td id="76B3FE7E2529EEDFFCF456D78486F95F" box="[772,1018,1677,1703]" gridcol="2" gridrow="21" pageId="12" pageNumber="48">10.23</td>
<td id="76B3FE7E2529EEDFFBDB56D78260F95F" box="[1067,1308,1677,1703]" gridcol="3" gridrow="21" pageId="12" pageNumber="48">6.67</td>
<td id="76B3FE7E2529EEDFFAA256D7812AF95F" box="[1362,1622,1677,1703]" gridcol="4" gridrow="21" pageId="12" pageNumber="48">6.70</td>
<td id="76B3FE7E2529EEDFF98956D78014F95F" box="[1657,1896,1677,1703]" gridcol="5" gridrow="21" pageId="12" pageNumber="48">7.13</td>
</tr>
</table>
</paragraph>
<paragraph id="8BED95402528112DFF4D5153858DFDE1" blockId="13.[189,1399,265,537]" pageId="13" pageNumber="49">
MALE GENITALIA (
<figureCitation id="136989C52528112DFE63515386A1FEDB" box="[403,477,265,291]" captionStart="Fig" captionStartId="10.[189,231,1892,1918]" captionTargetBox="[313,1298,573,1821]" captionTargetId="figure-174@10.[263,1321,567,1835]" captionTargetPageId="10" captionText="Fig. 6. Line drawings of male genitalia of Spaeleoleptes spaeleus H. Soares, 1966, male holotype (MZUSP 28463). A. Full genitalia, dorsal view. B. Detail of pars distalis, dorsal view. C. Detail of capsula interna, dorsal view. D. Detail of pars distalis, lateral view. E. Same, ventral view. Red lines: capsula interna. Blue lines: conductors. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10604123" httpUri="https://zenodo.org/record/10604123/files/figure.png" pageId="13" pageNumber="49">Fig. 6</figureCitation>
). Pars distalis swollen, separated from the pars basalis by a constriction. The apical region of pars basalis extends over the pars distalis dorsally, forming a structure similar to a small round-tipped plate in the region of constriction that separates pars basalis and pars distalis. Pars distalis with a ventral keel with round top in ventral view. Pars distalis with abroad apical lamina partially surrounding the glans ventrolaterally. Apical lamina with three lateral pairs of reduced setae ventrally. Apical lamina with a pointed medial projection. Four ventral setae reduced on the pars distalis. Glans with a pair of very robust, thickened, and folded conductors, partially covering the capsula interna. Capsula interna with lateral projections pointed.
</paragraph>
<paragraph id="8BED95402528112DFF1A521B8599FDA3" blockId="13.[189,1399,577,674]" box="[234,741,577,603]" pageId="13" pageNumber="49">
<emphasis id="B92649522528112DFF1A521B8639FDA3" bold="true" box="[234,325,577,603]" pageId="13" pageNumber="49">Female</emphasis>
(
<typeStatus id="54E92BE22528112DFEA5521B86C3FDA3" box="[341,447,577,603]" pageId="13" pageNumber="49" type="paratype">paratype</typeStatus>
, MZUSP 28462) (
<figureCitation id="136989C52528112DFD65521B85A0FDA3" box="[661,732,577,603]" captionStart="Fig" captionStartId="7.[189,231,1962,1988]" captionTargetBox="[189,1399,704,1914]" captionTargetId="figure-172@7.[189,1399,704,1914]" captionTargetPageId="7" captionText="Fig. 4. Spaeleoleptes spaeleus H. Soares, 1966, female paratype (MZUSP 28462). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, left lateral view. D. Carapace, lateral view. E. Carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10604117" httpUri="https://zenodo.org/record/10604117/files/figure.png" pageId="13" pageNumber="49">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BED95402528112DFF4D523E821EFD59" blockId="13.[189,1399,577,674]" pageId="13" pageNumber="49">
MEASUREMENTS (in millimeters).Total bodY length=2.3;carapace length=0.8;scutum magnum length=2.0; carapace maximum width=1.0; mesotergal scute maximum width=1.60. Leg measurements in
<tableCitation id="C6D0A0FB2528112DFAF652DD8220FD59" box="[1286,1372,647,673]" captionStart="Table 1" captionStartId="11.[188,257,629,658]" captionTargetBox="[204,1902,759,1654]" captionText="Table 1 (continued on next page). Body and leg measurements (in mm) of analyzed specimens of Spaeleoleptes spaeleus H. Soares, 1966 and Spaeleoleptes gimli sp. nov. Abbreviations: tr=trochanter; fe=femur; pa=patella; ti=tibia; me=metatarsus; ta=tarsus." httpUri="http://table.plazi.org/id/DF2DC5C8252E112BFF4C522F814DFD4C" pageId="13" pageNumber="49" tableUuid="DF2DC5C8252E112BFF4C522F814DFD4C">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BED95402528112DFF4D52938554FC88" blockId="13.[189,1398,713,880]" pageId="13" pageNumber="49">
BODY. Similar in appearance to the males but without the modified region on the tibia I, II and patella II. Ocularium with no small granules on the surface. Left antero-lateral margin of the carapace is armed with two large pointed tubercles close to each other, and the right antero-lateral margin of the carapace is armed with three large pointed tubercles close to each other (
<figureCitation id="136989C52528112DFAF753688215FCB4" box="[1287,1385,818,844]" captionStart="Fig" captionStartId="7.[189,231,1962,1988]" captionTargetBox="[189,1399,704,1914]" captionTargetId="figure-172@7.[189,1399,704,1914]" captionTargetPageId="7" captionText="Fig. 4. Spaeleoleptes spaeleus H. Soares, 1966, female paratype (MZUSP 28462). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, left lateral view. D. Carapace, lateral view. E. Carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10604117" httpUri="https://zenodo.org/record/10604117/files/figure.png" pageId="13" pageNumber="49">Fig. 4E</figureCitation>
). Tarsal formula: 4(2):5(3):5:5.
</paragraph>
</subSubSection>
<subSubSection id="C348C6CB2528112DFF4D53CD8525FC27" box="[189,601,919,992]" pageId="13" pageNumber="49" type="distribution">
<paragraph id="8BED95402528112DFF4D53CD862AFC49" blockId="13.[189,342,919,945]" box="[189,342,919,945]" pageId="13" pageNumber="49">
<heading id="D0A5222C2528112DFF4D53CD862AFC49" bold="true" box="[189,342,919,945]" fontSize="11" level="3" pageId="13" pageNumber="49" reason="3">
<emphasis id="B92649522528112DFF4D53CD862AFC49" bold="true" box="[189,342,919,945]" pageId="13" pageNumber="49">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BED95402528112DFF4D539C8525FC27" blockId="13.[189,601,965,992]" box="[189,601,965,992]" pageId="13" pageNumber="49">
Known only from the
<typeStatus id="54E92BE22528112DFE32539C8689FC18" box="[450,501,966,992]" pageId="13" pageNumber="49">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>