treatments-xml/data/03/F8/87/03F887CAFF8DFFFFFC1BFD2CB536F82F.xml

<document id="8E7744E0B25E1CED57CDE698502024A9" ID-CLB-Dataset="7312" ID-DOI="10.1051/acarologia/20162189" ID-GBIF-Dataset="cdf059b4-657c-4580-98af-a3ae299d13f0" ID-ISSN="2107-7207" ID-Zenodo-Dep="4640182" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616776462960" checkinUser="carolina" docAuthor="Moraza, M. L., Kontschan, J., Sahoo, G. &amp; Ansari, Z. A." docDate="2016" docId="03F887CAFF8DFFFFFC1BFD2CB536F82F" docLanguage="en" docName="Acarologia.56.1.73-89.pdf" docOrigin="Acarologia 56 (1)" docSource="http://dx.doi.org/10.1051/acarologia/20162189" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Eutrachytes flagellatus Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari 2016, n. sp." docType="treatment" docVersion="8" lastPageNumber="85" masterDocId="FFC1FFB2FF8EFFF2FFA1FFE7B023FFB3" masterDocTitle="A new species of Eutrachytes (Acari: Uropodina: Eutrachytidae) associated with the Indian mangrove (Avicennia officinalis)" masterLastPageNumber="89" masterPageNumber="73" pageNumber="75" updateTime="1698935171382" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="3C3698198C7072FCB2E27FAC36D1C351">A new species of Eutrachytes (Acari: Uropodina: Eutrachytidae) associated with the Indian mangrove (Avicennia officinalis)</mods:title>
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<emphasis id="B925EACEFF8DFFF1FC1BFD2CB536FD50" bold="true" box="[954,1301,715,739]" pageId="3" pageNumber="75">
<taxonomicName id="4C514D5FFF8DFFF1FC1BFD2CB4EFFD50" authority="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari, 2016" authorityName="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari" authorityYear="2016" box="[954,1228,715,739]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="75" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B925EACEFF8DFFF1FC1BFD2CB4EFFD50" bold="true" box="[954,1228,715,739]" italics="true" pageId="3" pageNumber="75">Eutrachytes flagellatus</emphasis>
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Diagnosis — Adults with idiosoma discoidal in shape, as long as wide, with prominent anterior region and scabellum present; dorsal, marginal and peritrematal shields fused at the anteriormost region; posterior region of dorsal shield with a pair of marginal concave depressions present; submarginal shield with setae to the anterior level of coxa I in adult and nymphal instars; pygidial shield nude in all instars, only bearing one pair of poroids. Nymphal instars with a pairs of lateral flagellate tubular structures emerging from the peritreme. Hypostomal setae 
<emphasis id="B925EACEFF8DFFF1FB16FB6CB4F1FB10" box="[1207,1234,1163,1187]" italics="true" pageId="3" pageNumber="75">h2</emphasis>
longer than 
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and 
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divided in two pilose dissimilar branches. Larval podonotal shield with nine pairs of setae, three pairs of mesonothal scutellae; protonymph with five pairs of setae on podonotal shield, nude mesonotal and nude pygidial; deutonymphs with dorsal and marginal shield fused at anterior region and peritrematal shield free as in the protonymphal instar. Legs I-IV with pretarsus, a pair of well developed claws and three acuminate lobes; femora I-IV with ventral keel; femora II in the male with ventral setae modified as a conspicuous thick spine.
</paragraph>
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<emphasis id="B925EACEFF8DFFF1FCC1F9DBB394F9E7" bold="true" box="[864,951,1596,1620]" pageId="3" pageNumber="75">Female</emphasis>
</paragraph>
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Dorsal idiosoma (
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) — Idiosoma discoid in shape, almost as long (
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) as wide (
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), with smooth margin and humeral peritremal protrusions. Dorsal and marginal shields fused anteriorly at level of humeral protrusions, and fused with peritrematal shield at level of 
<emphasis id="B925EACEFF8DFFF1FB0BF8FCB4E1F880" box="[1194,1218,1819,1843]" italics="true" pageId="3" pageNumber="75">z2</emphasis>
. Anterior dorsal shield rounded acuminate, slightly bent to ventral region. Pygidial shield ca. 18 times wider than long, with posterior margin slightly lobulated, with one pair of poroids (lyrifissures) and at least three pairs of punctiform poroids. Submarginal shield rearchs the posterior margin of peritremes where bends ventrally covering part of distal peritremal loop (
<figureCitation id="136A2A59FF8BFFF7FF43FE24B14BFE68" box="[226,360,451,475]" captionStart="FIGURE" captionStartId="4.[159,171,1830,1849]" captionTargetBox="[232,1361,369,1746]" captionTargetId="figure-4@4.[227,1361,364,1798]" captionTargetPageId="4" captionText="FIGURE 1: Eutrachytes flagellatus n. sp., adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region." figureDoi="http://doi.org/10.5281/zenodo.4640184" httpUri="https://zenodo.org/record/4640184/files/figure.png" pageId="5" pageNumber="77">Figs 1B, 1D</figureCitation>
).
</paragraph>
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FIGURE 1: 
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<emphasis id="B925EACEFF8AFFF6FEA5F8C1B1EBF88A" box="[260,456,1830,1849]" italics="true" pageId="4" pageNumber="76">Eutrachytes flagellatus</emphasis>
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<taxonomicNameLabel id="A21657B5FF8AFFF6FE6CF8C1B222F88A" box="[461,513,1830,1849]" pageId="4" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
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, adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region.
</paragraph>
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<paragraph id="8BEE36DCFF8BFFF7FF64FE15B17EFC1C" blockId="5.[159,762,311,1948]" pageId="5" pageNumber="77">
Dorsal shield with lightly granulate cuticle, c.a 40 pairs of short, homogeneous in length, pilose setae, neotrichy present (podonotal region with several unpaired central setae 
<emphasis id="B925EACEFF8BFFF7FE48FDBBB1DDFDC0" box="[489,510,604,627]" italics="true" pageId="5" pageNumber="77">jx</emphasis>
, and duplicate setae 
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, 
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, 
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, 
<emphasis id="B925EACEFF8BFFF7FE89FD99B1AEFD25" box="[296,397,638,662]" italics="true" pageId="5" pageNumber="77">z4, z5, s5</emphasis>
; marginal shield with 11 pairs of setae (
<emphasis id="B925EACEFF8BFFF7FEAAFD46B160FD0A" box="[267,323,673,697]" italics="true" pageId="5" pageNumber="77">r2-r6</emphasis>
, 
<emphasis id="B925EACEFF8BFFF7FEF2FD46B1BAFD0A" box="[339,409,673,697]" italics="true" pageId="5" pageNumber="77">R1-R6</emphasis>
) similar in length and shape, and ca. 20 pair of setae with sparse and longer pilosity on the ventral submarginal region (setae 
<emphasis id="B925EACEFF8BFFF7FF3EFCECB0E6FC91" box="[159,197,779,802]" italics="true" pageId="5" pageNumber="77">UR</emphasis>
). Most dorsal setae associated to a punctiform pore like structure. Shield with a pair of concave marginal depressions at the posterior region which remain under the shield and open laterally over the marginal shield.
</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FF64FC21B183F952" blockId="5.[159,762,311,1948]" pageId="5" pageNumber="77">
Ventral idiosoma (
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) — Tritosternum with a strongly dentate base 
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long and three pilose laciniae free for most of length; medial lacinia longer (
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) than laterals (
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), devoid of basal pilosity and with bifid end (
<figureCitation id="136A2A59FF8BFFF7FE4BFBB5B266FBD9" box="[490,581,1106,1130]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2D</figureCitation>
). Sternal shield with anterior margin well delineated and a medial convexity. Sternal setae minute, smooth, poorly discernible; setae 
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anterior to epigynal shield, accompanied by a pair of elongated lyrifissures 
<emphasis id="B925EACEFF8BFFF7FD79FB39B2D9FB45" box="[728,762,1246,1270]" italics="true" pageId="5" pageNumber="77">iv1</emphasis>
and a pair of punctiform poroids (
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); 
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at both sides of epigynal shield, close to 
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, distance st1- st2 near one third st2-st3; 
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together with poroids 
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at margin of genital opening, and pores 
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; 
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behind the shield and the third pair of pores 
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at the posterior corners of the shield; genital setae 
<emphasis id="B925EACEFF8BFFF7FF3EFA33B09CFA58" box="[159,191,1492,1515]" italics="true" pageId="5" pageNumber="77">st5</emphasis>
behind coxae IV similar to sternal setae. Epigynal shield densely granulated as other idiosomal shields, subtriangular in shape, 
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long 
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wide, with straight posterior margin and acuminate anterior margin; one pair of liryfissures at the posterior region; shield located between coxae II and anterior margin of coxae IV. Endogynum with a pair or cup-like structures.
</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FF64F91FB3CEFD39" blockId="5.[159,762,311,1948]" lastBlockId="5.[826,1429,311,1948]" pageId="5" pageNumber="77">
Opisthogaster with three pairs of minute ventral setae (
<emphasis id="B925EACEFF8BFFF7FF57F8FCB13CF880" box="[246,287,1819,1843]" italics="true" pageId="5" pageNumber="77">JV1</emphasis>
and two more pairs &quot; 
<emphasis id="B925EACEFF8BFFF7FD8DF8FBB26FF880" box="[556,588,1820,1843]" italics="true" pageId="5" pageNumber="77">Vx</emphasis>
&quot; associated to glandular pores at the parapodal region of coxae IV), similar to sternal setae, and seven pairs at the posterior region laterals to anal opening which are similar in shape and length to the three circumanal setae such as in figure 1B; at least two pairs of liryfissures 
<emphasis id="B925EACEFF8BFFF7FC38FE99B39AFE26" box="[921,953,382,405]" italics="true" pageId="5" pageNumber="77">ivo</emphasis>
and eight pairs of pores (at least four are glandular pores). Anal opening framed, small and rounded (diameter 
<quantity id="4CA99B39FF8BFFF7FB82FE24B44FFE68" box="[1059,1132,451,476]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="77" unit="mm" value="16.0">16 µm</quantity>
), with nude anal valves; paranal and postanal setae ciliate, similar to other ventral lateral setae, and posterior to anal opening. Between setae 
<emphasis id="B925EACEFF8BFFF7FC4AFDCBB437FDF7" box="[1003,1044,556,580]" italics="true" pageId="5" pageNumber="77">JV1</emphasis>
and 
<emphasis id="B925EACEFF8BFFF7FBF3FDCAB450FDF7" box="[1106,1139,557,580]" italics="true" pageId="5" pageNumber="77">st5</emphasis>
a circular subcutaneous structure is observed with a pair of sigillar area at both sides of it.
</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FCC1FD79B3BAFC41" blockId="5.[826,1429,311,1948]" pageId="5" pageNumber="77">Endopodal elements between legs I-II, II-III and III-IV fused to sternal shield; endopodal I with a strong anterior apophysis with lateral corner acuminate and internal corner rounded; exopodal I-II as a hemispherical apophysis; exopodoal elements between coxae II-III and III-IV free from endopodal; parapodal IV semilunar in shape, well developed, at the botton of pedofosae IV. Pedofosae II, III and IV well delimited, separated by longitudinal bridges.</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FCC1FBE1B478FAA7" blockId="5.[826,1429,311,1948]" pageId="5" pageNumber="77">
Peritrematal shield with one pair of dorsal setae at both side of peritreme distal loop (
<figureCitation id="136A2A59FF8BFFF7FA86FBCEB5A4FBF2" box="[1319,1415,1065,1089]" captionStart="FIGURE" captionStartId="4.[159,171,1830,1849]" captionTargetBox="[232,1361,369,1746]" captionTargetId="figure-4@4.[227,1361,364,1798]" captionTargetPageId="4" captionText="FIGURE 1: Eutrachytes flagellatus n. sp., adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region." figureDoi="http://doi.org/10.5281/zenodo.4640184" httpUri="https://zenodo.org/record/4640184/files/figure.png" pageId="5" pageNumber="77">Fig. 1A</figureCitation>
). Respiratory stigmata situated in pedofosae III; from the stigmata the peritreme ascends to lateral conical protuberance at level of pedofossae II and descends for a shorter and narrow stretch; peritreme curved dorsally at level of the humeral conical protuberance (
<figureCitation id="136A2A59FF8BFFF7FC1CFB1CB46DFAA7" box="[957,1102,1275,1300]" captionStart="FIGURE" captionStartId="4.[159,171,1830,1849]" captionTargetBox="[232,1361,369,1746]" captionTargetId="figure-4@4.[227,1361,364,1798]" captionTargetPageId="4" captionText="FIGURE 1: Eutrachytes flagellatus n. sp., adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region." figureDoi="http://doi.org/10.5281/zenodo.4640184" httpUri="https://zenodo.org/record/4640184/files/figure.png" pageId="5" pageNumber="77">Figs. 1C, 1D</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FCC1FAC0B5ACF9EB" blockId="5.[826,1429,311,1948]" pageId="5" pageNumber="77">
Gnathosoma — Gnathotectum (
<figureCitation id="136A2A59FF8BFFF7FB70FAC0B56DFAF3" box="[1233,1358,1319,1344]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Figs. 2A,B</figureCitation>
). with a broad dentate base and medial projection with ventral surface provided of minute spines (
<figureCitation id="136A2A59FF8BFFF7FA8EFA89B5ABFA35" box="[1327,1416,1390,1414]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2A</figureCitation>
), lateral margins with 5-6 strong long spines (
<figureCitation id="136A2A59FF8BFFF7FA92FA76B5ABFA1A" box="[1331,1416,1425,1449]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2B</figureCitation>
), and a series of marginal strongly ciliate long setae; at the base of the medial projection, bent to ventral side, a strong dorsal bifid tectum (
<figureCitation id="136A2A59FF8BFFF7FB68FA1DB53DF9A1" box="[1225,1310,1530,1554]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2B</figureCitation>
); dorsally, at the basal region of central projection, three pairs of marginal long barbed straight setae (
<figureCitation id="136A2A59FF8BFFF7FAA6F9A7B5A0F9EB" box="[1287,1411,1600,1624]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Figs. 2A,B</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF8BFFF7FCC1F98BB5B6F82F" blockId="5.[826,1429,311,1948]" pageId="5" pageNumber="77">
Corniculi hornlike, short (
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), stout (
<quantity id="4CA99B39FF8BFFF7FA9DF98BB5A6F937" box="[1340,1413,1644,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="77" unit="mm" value="18.0">18 µm</quantity>
), inserted midway between insertions of subcapitular setae 
<emphasis id="B925EACEFF8BFFF7FC08F955B3E0F979" box="[937,963,1714,1738]" italics="true" pageId="5" pageNumber="77">h1</emphasis>
and 
<emphasis id="B925EACEFF8BFFF7FBA4F955B43CF979" box="[1029,1055,1714,1738]" italics="true" pageId="5" pageNumber="77">h2</emphasis>
, (
<figureCitation id="136A2A59FF8BFFF7FB98F955B4B8F979" box="[1081,1179,1714,1738]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2C</figureCitation>
); 
<emphasis id="B925EACEFF8BFFF7FB12F955B4EEF979" box="[1203,1229,1714,1738]" italics="true" pageId="5" pageNumber="77">h1</emphasis>
smooth, shorter (
<quantity id="4CA99B39FF8BFFF7FCE3F932B3ADF95E" box="[834,910,1749,1774]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="77" unit="mm" value="32.0">32 µm</quantity>
) than slightly pilose 
<emphasis id="B925EACEFF8BFFF7FB35F932B48CF95E" box="[1172,1199,1749,1773]" italics="true" pageId="5" pageNumber="77">h2</emphasis>
(
<quantity id="4CA99B39FF8BFFF7FB60F932B52DF95E" box="[1217,1294,1749,1774]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="77" unit="mm" value="40.0">40 µm</quantity>
); pilose 
<emphasis id="B925EACEFF8BFFF7FADBF932B5B6F95E" box="[1402,1429,1749,1773]" italics="true" pageId="5" pageNumber="77">h3</emphasis>
the longest setae (
<quantity id="4CA99B39FF8BFFF7FBABF91FB470F8A3" box="[1034,1107,1784,1809]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="77" unit="mm" value="64.0">64 µm</quantity>
), twice the length of 
<emphasis id="B925EACEFF8BFFF7FAEAF91FB545F8A3" box="[1355,1382,1784,1808]" italics="true" pageId="5" pageNumber="77">h1</emphasis>
, capitular setae (
<emphasis id="B925EACEFF8BFFF7FC7FF8FCB3DAF880" box="[990,1017,1819,1843]" italics="true" pageId="5" pageNumber="77">h4</emphasis>
) the shortest (
<quantity id="4CA99B39FF8BFFF7FB05F8FCB4CDF880" box="[1188,1262,1819,1844]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="77" unit="mm" value="20.0">20 µm</quantity>
) with a distal pilosity and a widened pilose process at the medial region (
<figureCitation id="136A2A59FF8BFFF7FC75F886B41CF8C9" box="[980,1087,1889,1914]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="5" pageNumber="77">Fig. 2C</figureCitation>
). Internal malae short with lateral margins pilose, extending to medial length
</paragraph>
<paragraph id="8BEE36DCFF88FFF4FF3EFF1DB16BFEBE" blockId="6.[159,328,250,269]" box="[159,328,250,269]" pageId="6" pageNumber="78">
Moraza M.L. 
<emphasis id="B925EACEFF88FFF4FEBCFF1CB16BFEBE" box="[285,328,250,269]" italics="true" pageId="6" pageNumber="78">et al.</emphasis>
</paragraph>
<caption id="DF2E6654FF88FFF4FF3EFBA2B4B4FBC0" ID-DOI="http://doi.org/10.5281/zenodo.4640186" ID-Zenodo-Dep="4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78" startId="6.[159,171,1093,1112]" targetBox="[227,1361,302,1061]" targetPageId="6">
<paragraph id="8BEE36DCFF88FFF4FF3EFBA2B4B4FBC0" blockId="6.[159,1429,1093,1140]" pageId="6" pageNumber="78">
FIGURE 2: 
<taxonomicName id="4C514D5FFF88FFF4FEA4FBA2B1EAFBEB" authority="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari, 2016" authorityName="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari" authorityYear="2016" box="[261,457,1093,1112]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="78" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B925EACEFF88FFF4FEA4FBA2B1EAFBEB" box="[261,457,1093,1112]" italics="true" pageId="6" pageNumber="78">Eutrachytes flagellatus</emphasis>
</taxonomicName>
<emphasis id="B925EACEFF88FFF4FE6EFBA2B220FBEB" bold="true" box="[463,515,1093,1112]" pageId="6" pageNumber="78">
<taxonomicNameLabel id="A21657B5FF88FFF4FE6EFBA2B220FBEB" box="[463,515,1093,1112]" pageId="6" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit.
</paragraph>
</caption>
<paragraph id="8BEE36DCFF88FFF4FF3EFB52B1C0F892" blockId="6.[159,762,1205,1825]" pageId="6" pageNumber="78">
of palpfemora (
<figureCitation id="136A2A59FF88FFF4FEFDFB52B19CFB7E" box="[348,447,1205,1229]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78">Fig. 2C</figureCitation>
). Deutosternum with two smooth and wide groves between 
<emphasis id="B925EACEFF88FFF4FD90FB3FB26FFB43" box="[561,588,1240,1264]" italics="true" pageId="6" pageNumber="78">h3</emphasis>
and 
<emphasis id="B925EACEFF88FFF4FD27FB3FB282FB43" box="[646,673,1240,1264]" italics="true" pageId="6" pageNumber="78">h4</emphasis>
. Palpal setation and form of setae as described by 
<bibRefCitation id="EFC04B2DFF88FFF4FD08FB1CB0C5FA85" author="Krantz G. W." pageId="6" pageNumber="78" pagination="62 - 70" refId="ref10281" refString="Krantz G. W. 1969 - The Mites of Quintana Roo. I. A new species of Eutrachytes from the Yucatan Peninsula, with observations on the classification of the genus - Ann. Entomol. Soc. Am., 62 (1): 62 - 70." type="journal article" year="1969">Krantz (1969)</bibRefCitation>
(2-5-5-14); palp- trochanter with inner setae (
<quantity id="4CA99B39FF88FFF4FF07FAA6B0D3FAEA" box="[166,240,1345,1370]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="78" unit="mm" value="47.0">47 µm</quantity>
) twice as long as external seta (
<figureCitation id="136A2A59FF88FFF4FDC7FAA6B2E3FAEA" box="[614,704,1345,1369]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78">Fig. 2C</figureCitation>
), the last spinose; palp-claw two-tined. Chelicerae (
<figureCitation id="136A2A59FF88FFF4FD6CFA83B09EFA2C" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78">Fig. 2E</figureCitation>
) with fixed digit edentate, (
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long) overlapping movable digit and nodulus present; digit with a socket subterminally which receives the hooked tip of the movable digit when the two are appressed (
<figureCitation id="136A2A59FF88FFF4FF06F9F4B0DCF99F" box="[167,255,1555,1580]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78">Fig. 2E</figureCitation>
); a densely pilose surface (brush) at masticatory region; terminally with a spine or lateral mucro. Movable digit of female 
<quantity id="4CA99B39FF88FFF4FE59F9BDB247F9C1" box="[504,612,1626,1650]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" metricValueMax="6.3" metricValueMin="6.1" pageId="6" pageNumber="78" unit="mm" value="62.0" valueMax="63.0" valueMin="61.0">61-63 µm</quantity>
long, with a dentate comb at distal region (c.a 17 teeth) followed by a prominent alveolar remnant (or two opposed teeth), and basal region smooth; dorsal cheliceral setae no discernible if present; without conspicuous arthrodial process (
<figureCitation id="136A2A59FF88FFF4FE20F8EEB1F4F892" box="[385,471,1801,1825]" captionStart="FIGURE" captionStartId="6.[159,171,1093,1112]" captionTargetBox="[227,1361,302,1061]" captionTargetId="figure-4@6.[227,1361,302,1061]" captionTargetPageId="6" captionText="FIGURE 2: Eutrachytes flagellatus n. sp., adult female: A – Detail of gnathotectum, ventral view; B – Detail of gnathotectum, dorsal view; C – Detail of subcapitulum; D – Sternapophisis; E – Chelicera, lateral view and detail of tip of fixed digit." figureDoi="http://doi.org/10.5281/zenodo.4640186" httpUri="https://zenodo.org/record/4640186/files/figure.png" pageId="6" pageNumber="78">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF88FFF5FF64F8D8B1EAFE4D" blockId="6.[159,762,1854,1948]" lastBlockId="7.[159,762,311,1949]" lastPageId="7" lastPageNumber="79" pageId="6" pageNumber="78">
Legs — Legs relatively short, clearly shorter than dorsal shield. Coxae I with two groups of coxal glands, dorsal concavity and laterodorsal surface with rounded prominences (
<figureCitation id="136A2A59FF88FFF4FB37FB52B4DBFB7E" box="[1174,1272,1205,1229]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="6" pageNumber="78">Fig. 3E</figureCitation>
); coxae II-IV with numerous ventro basal glandular openings; except tarsus I-IV, all other segments with a dorso distal striate tectum overlapping the articulation with the following segment; femora I-IV with a conspicuous ventral tectum (
<figureCitation id="136A2A59FF88FFF4FBCAFA83B4EFFACF" box="[1131,1228,1380,1404]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="6" pageNumber="78">Fig. 3A</figureCitation>
). Tarsus I with a pair of claws and a series of long terminal setae. Pretarsus of legs I to IV with slender claws at the end of a relatively long stalk; legs II-IV with three slender acuminate lobes (
<figureCitation id="136A2A59FF88FFF4FBCDFA17B4D0F9BA" box="[1132,1267,1520,1545]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="6" pageNumber="78">Figs. 3D,G</figureCitation>
). Tarsus II-IV about twice as long as tibia, telotarsus twice longer than basitarsus (
<quantity id="4CA99B39FF88FFF4FC5CF9D0B46DF9FC" box="[1021,1102,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="78" unit="mm" value="2.2">2.2 µm</quantity>
); apical ventral process absent; with 17 setae, apical setal processes 
<emphasis id="B925EACEFF88FFF4FA94F9BDB578F9C1" box="[1333,1371,1626,1650]" italics="true" pageId="6" pageNumber="78">ad1</emphasis>
, 
<emphasis id="B925EACEFF88FFF4FACCF9BDB5B6F9C1" box="[1389,1429,1626,1650]" italics="true" pageId="6" pageNumber="78">pd1</emphasis>
minute (&quot;d1&quot;), spinelike (
<figureCitation id="136A2A59FF88FFF4FBD1F99AB4F5F926" box="[1136,1238,1661,1685]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="6" pageNumber="78">Fig. 3B</figureCitation>
), and telotarsal seta 
<emphasis id="B925EACEFF88FFF4FCD3F947B3B7F90B" box="[882,916,1696,1720]" italics="true" pageId="6" pageNumber="78">md</emphasis>
twice as long as lateral setae 
<emphasis id="B925EACEFF88FFF4FAA9F947B50AF90B" box="[1288,1321,1696,1720]" italics="true" pageId="6" pageNumber="78">al1</emphasis>
, 
<emphasis id="B925EACEFF88FFF4FA9AF947B57FF90B" box="[1339,1372,1696,1720]" italics="true" pageId="6" pageNumber="78">pl1</emphasis>
; basitarsal setae feather-like, dorsal setae short (
<quantity id="4CA99B39FF88FFF4FAEEF924B343F94D" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="6" pageNumber="78" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9 – 10 µm</quantity>
), 
<emphasis id="B925EACEFF88FFF4FCD9F901B3BBF94D" box="[888,920,1766,1790]" italics="true" pageId="6" pageNumber="78">al3</emphasis>
reaching 
<emphasis id="B925EACEFF88FFF4FBACF901B40DF94D" box="[1037,1070,1766,1790]" italics="true" pageId="6" pageNumber="78">al2</emphasis>
setae (
<quantity id="4CA99B39FF88FFF4FBDEF901B522F94D" box="[1151,1281,1766,1790]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="6" pageNumber="78" unit="mm" value="30.0" valueMax="31.0" valueMin="29.0">29 – 31 µm</quantity>
) and 
<emphasis id="B925EACEFF88FFF4FAE7F901B54BF94D" box="[1350,1384,1766,1790]" italics="true" pageId="6" pageNumber="78">pl3</emphasis>
the longest (
<quantity id="4CA99B39FF88FFF4FC3DF8EEB436F892" box="[924,1045,1801,1825]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="5.3" metricValueMin="5.1" pageId="6" pageNumber="78" unit="mm" value="52.0" valueMax="53.0" valueMin="51.0">51 – 53 µm</quantity>
) reaching pair of lateral setae 
<emphasis id="B925EACEFF88FFF4FACFF8EEB5ADF892" box="[1390,1422,1801,1825]" italics="true" pageId="6" pageNumber="78">al1</emphasis>
, 
<emphasis id="B925EACEFF88FFF4FC9BF8CBB378F8F7" box="[826,859,1836,1860]" italics="true" pageId="6" pageNumber="78">pl1</emphasis>
(
<figureCitation id="136A2A59FF88FFF4FCCCF8CBB3D3F8F7" box="[877,1008,1836,1860]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="6" pageNumber="78">Figs. 3B,H</figureCitation>
). Other leg setae simple or slightly barbed. Full complement of setae: coxae, 2-2-2-1; trochanters, 5-5-5-5; femora, 9 (1 2/1 2/2 1) – 9 (1 2/1 2/2 1) – 7 (1 3/2 1) – 7 (1 3/2 1); genua, 8 (1 2/1 2/1 1) – 8 (1 2/1 2/1 1) – 8 (1 2/1 2/1 1) – 5 (1 2/0 2/0 0); tibiae, 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1). Leg lengths, excluding pretarsi: I 
<quantity id="4CA99B39FF89FFF5FEBDFE24B19BFE68" box="[284,440,451,476]" metricMagnitude="-1" metricUnit="m" metricValue="3.35" metricValueMax="3.4" metricValueMin="3.3" pageId="7" pageNumber="79" unit="mm" value="335.0" valueMax="340.0" valueMin="330.0">330 – 340 µm</quantity>
, II 
<quantity id="4CA99B39FF89FFF5FE40FE24B2A9FE68" box="[481,650,451,476]" metricMagnitude="0" metricUnit="m" metricValue="1.3225" metricValueMax="2.325" metricValueMin="0.32" pageId="7" pageNumber="79" unit="mm" value="1322.5" valueMax="2325.0" valueMin="320.0">320 – 2325 µm</quantity>
, III 
<quantity id="4CA99B39FF89FFF5FD1DFE24B0D7FE4D" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="7" pageNumber="79" unit="mm" value="305.0" valueMax="310.0" valueMin="300.0">300 – 310 µm</quantity>
, IV 
<quantity id="4CA99B39FF89FFF5FE8BFE01B1E1FE4D" box="[298,450,486,511]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="7" pageNumber="79" unit="mm" value="305.0" valueMax="310.0" valueMin="300.0">300 – 310 µm</quantity>
.
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FDF5B120FD99" blockId="7.[159,762,311,1949]" box="[197,259,530,554]" pageId="7" pageNumber="79">
<emphasis id="B925EACEFF89FFF5FF64FDF5B120FD99" bold="true" box="[197,259,530,554]" pageId="7" pageNumber="79">Male</emphasis>
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FDD9B2D2FDCA" blockId="7.[159,762,311,1949]" pageId="7" pageNumber="79">Dorsal idiosoma — Similar to female in shape; dorsal shielding and dorsal setae as for the female.</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FD6AB0FCFC0E" blockId="7.[159,762,311,1949]" pageId="7" pageNumber="79">
Ventral idiosoma — Anterior region of sternal region as in the female (
<figureCitation id="136A2A59FF89FFF5FE1DFD57B230FD7B" box="[444,531,688,712]" captionStart="FIGURE" captionStartId="4.[159,171,1830,1849]" captionTargetBox="[232,1361,369,1746]" captionTargetId="figure-4@4.[227,1361,364,1798]" captionTargetPageId="4" captionText="FIGURE 1: Eutrachytes flagellatus n. sp., adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region." figureDoi="http://doi.org/10.5281/zenodo.4640184" httpUri="https://zenodo.org/record/4640184/files/figure.png" pageId="7" pageNumber="79">Fig. 1E</figureCitation>
), with four pairs of minute sternal setae, two pairs of poroids (
<emphasis id="B925EACEFF89FFF5FD39FD34B299FD58" box="[664,698,723,747]" italics="true" pageId="7" pageNumber="79">iv1</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FD69FD34B2C9FD58" box="[712,746,723,747]" italics="true" pageId="7" pageNumber="79">iv3</emphasis>
), two or three pairs of glandular pores (
<emphasis id="B925EACEFF89FFF5FDDDFD11B28AFCBD" box="[636,681,758,782]" italics="true" pageId="7" pageNumber="79">gst1</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FD1DFD11B2C9FCBD" box="[700,746,758,782]" italics="true" pageId="7" pageNumber="79">gst3</emphasis>
), and one pair of punctiform pores; setae 
<emphasis id="B925EACEFF89FFF5FDDEFCFDB2BCFC82" box="[639,671,794,817]" italics="true" pageId="7" pageNumber="79">st5</emphasis>
behind genital opening betwee coxae IV, similar to other sternal setae. Genital opening (
<quantity id="4CA99B39FF89FFF5FDADFCB8B2A9FCC4" box="[524,650,863,888]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.4" metricValueMin="2.3" pageId="7" pageNumber="79" unit="mm" value="23.5" valueMax="24.0" valueMin="23.0">23 – 24 µm</quantity>
long) between legs III. Other ventral characteristics as in female.
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FC36B23EFC5A" blockId="7.[159,762,311,1949]" box="[197,541,977,1002]" pageId="7" pageNumber="79">Gnathosoma — As in female.</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FC19B2BFFAB9" blockId="7.[159,762,311,1949]" pageId="7" pageNumber="79">
Legs — Legs segments not strongly differentiated on either sex, except for a ventral thick claw like setae in femur II (
<figureCitation id="136A2A59FF89FFF5FE04FBA4B223FBEF" box="[421,512,1091,1116]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="7" pageNumber="79">Fig. 3C</figureCitation>
), spine like setae 
<emphasis id="B925EACEFF89FFF5FD73FBA3B2D9FBEF" box="[722,762,1092,1116]" italics="true" pageId="7" pageNumber="79">
pv 
<quantity id="4CA99B39FF89FFF5FD4EFBA3B2D9FBEF" box="[751,762,1092,1116]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="79" unit="in" value="1.0">1</quantity>
</emphasis>
in genua and tibia I-IV, and seta 
<emphasis id="B925EACEFF89FFF5FDBCFB80B266FBCC" box="[541,581,1127,1151]" italics="true" pageId="7" pageNumber="79">pv2</emphasis>
with a bulbose base in tasus II-IV (
<figureCitation id="136A2A59FF89FFF5FEDEFB6DB1FBFB11" box="[383,472,1162,1186]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="7" pageNumber="79">Fig. 3D</figureCitation>
). Anterior lateral ventral margin of the segments, except tarsi, strongly serrate (
<figureCitation id="136A2A59FF89FFF5FF7AFB37B176FB5B" box="[219,341,1232,1256]" captionStart="FIGURE" captionStartId="8.[159,171,1708,1727]" captionTargetBox="[231,1359,445,1641]" captionTargetId="figure-4@8.[227,1361,432,1676]" captionTargetPageId="8" captionText="FIGURE 3: Eutrachytes flagellatus n. sp.: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color." figureDoi="http://doi.org/10.5281/zenodo.4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="7" pageNumber="79">Figs. 3C,F</figureCitation>
). Legs length: I 
<quantity id="4CA99B39FF89FFF5FDB3FB37B288FB5B" box="[530,683,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="3.125" metricValueMax="3.15" metricValueMin="3.1" pageId="7" pageNumber="79" unit="mm" value="312.5" valueMax="315.0" valueMin="310.0">310 – 315 µm</quantity>
, II 
<quantity id="4CA99B39FF89FFF5FD73FB37B12BFAB8" metricMagnitude="-1" metricUnit="m" metricValue="3.125" metricValueMax="3.15" metricValueMin="3.1" pageId="7" pageNumber="79" unit="mm" value="312.5" valueMax="315.0" valueMin="310.0">310 – 315 µm</quantity>
, III 
<quantity id="4CA99B39FF89FFF5FE97FB14B1EDFAB8" box="[310,462,1267,1291]" metricMagnitude="-1" metricUnit="m" metricValue="2.925" metricValueMax="2.95" metricValueMin="2.9" pageId="7" pageNumber="79" unit="mm" value="292.5" valueMax="295.0" valueMin="290.0">290 – 295 µm</quantity>
, IV 
<quantity id="4CA99B39FF89FFF5FE5FFB14B2B6FAB8" box="[510,661,1267,1291]" metricMagnitude="-1" metricUnit="m" metricValue="3.025" metricValueMax="3.05" metricValueMin="3.0" pageId="7" pageNumber="79" unit="mm" value="302.5" valueMax="305.0" valueMin="300.0">300 – 305 µm</quantity>
.
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FAF8B20DFAE9" blockId="7.[159,762,311,1949]" pageId="7" pageNumber="79">
<emphasis id="B925EACEFF89FFF5FF64FAF8B144FA84" bold="true" box="[197,359,1311,1335]" pageId="7" pageNumber="79">Deutonymph</emphasis>
— Idiosoma 
<quantity id="4CA99B39FF89FFF5FDA4FAF8B279FA84" box="[517,602,1311,1335]" metricMagnitude="-1" metricUnit="m" metricValue="4.97" pageId="7" pageNumber="79" unit="mm" value="497.0">497 µm</quantity>
long, 
<quantity id="4CA99B39FF89FFF5FD05FAF8B2DAFA84" box="[676,761,1311,1335]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="79" unit="mm" value="420.0">420 µm</quantity>
wide at level of coxae III-IV (n=1).
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64FA89B236F9EB" blockId="7.[159,762,311,1949]" pageId="7" pageNumber="79">
Dorsal idiosoma — Dorsum (
<figureCitation id="136A2A59FF89FFF5FDBEFA89B25AFA35" box="[543,633,1390,1414]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="7" pageNumber="79">Fig. 4A</figureCitation>
) discoidal, with acuminate anterior region and humeral angles absent. Marginal shields completely differentiated, separated form dorsal shield by a fine suture which became indistinct anteriorly; pygidial shield undifferentiated except for the presence of a pair of poroids set on conical tubercles.
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FF64F98BB388FC33" blockId="7.[159,762,311,1949]" lastBlockId="7.[826,1429,311,896]" pageId="7" pageNumber="79">
Dorsal shield fused with marginal shields at level of setae 
<emphasis id="B925EACEFF89FFF5FEE0F968B175F914" box="[321,342,1679,1703]" italics="true" pageId="7" pageNumber="79">j3</emphasis>
, with granulate cuticle ornate with rounded shallow pits; shield holotrichous with 23 pairs of similarly short ciliate setae (
<figureCitation id="136A2A59FF89FFF5FDEFF932B28BF95E" box="[590,680,1749,1773]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="7" pageNumber="79">Fig. 4C</figureCitation>
) (
<emphasis id="B925EACEFF89FFF5FD60F932B2D1F95E" box="[705,754,1749,1773]" italics="true" pageId="7" pageNumber="79">j3-j6</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FF3EF91FB0F9F8A3" box="[159,218,1784,1808]" italics="true" pageId="7" pageNumber="79">z3-z6</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FF4DF91FB107F8A3" box="[236,292,1784,1808]" italics="true" pageId="7" pageNumber="79">s3-s6</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FE97F91FB149F8A3" box="[310,362,1784,1808]" italics="true" pageId="7" pageNumber="79">J1-J5</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FEDCF91FB1E7F8A3" box="[381,452,1784,1808]" italics="true" pageId="7" pageNumber="79">Z1-Z5</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FE74F91FB235F8A3" box="[469,534,1784,1808]" italics="true" pageId="7" pageNumber="79">S1-S5</emphasis>
), except posteriormost 
<emphasis id="B925EACEFF89FFF5FF41F8FBB0DCF880" box="[224,255,1820,1843]" italics="true" pageId="7" pageNumber="79">Z5</emphasis>
elongate and slightly curved (
<quantity id="4CA99B39FF89FFF5FDC4F8FCB28DF880" box="[613,686,1819,1844]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="79" unit="mm" value="87.0">87 µm</quantity>
long); sometimes two setae &quot;x&quot; between 
<emphasis id="B925EACEFF89FFF5FD88F8D8B21DF8E4" box="[553,574,1855,1879]" italics="true" pageId="7" pageNumber="79">j3</emphasis>
and 
<emphasis id="B925EACEFF89FFF5FDD9F8D8B2AFF8E4" box="[632,652,1855,1879]" italics="true" pageId="7" pageNumber="79">j4</emphasis>
present). Marginal shield with setae 
<emphasis id="B925EACEFF89FFF5FE41F885B1D6F8C9" box="[480,501,1890,1914]" italics="true" pageId="7" pageNumber="79">j2</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FDA2F885B23FF8C9" box="[515,540,1890,1914]" italics="true" pageId="7" pageNumber="79">z1</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FD8BF885B260F8C9" box="[554,579,1890,1914]" italics="true" pageId="7" pageNumber="79">z2</emphasis>
on the anterior acuminate region of shield, similar in shape to 
<emphasis id="B925EACEFF89FFF5FD44F862B2D9F82E" box="[741,762,1925,1949]" italics="true" pageId="7" pageNumber="79">j1</emphasis>
(
<quantity id="4CA99B39FF89FFF5FCE3FED0B3E9FEFC" box="[834,970,311,335]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="7" pageNumber="79" unit="mm" value="32.0" valueMax="34.0" valueMin="30.0">30 – 34 µm</quantity>
long); other marginal setae (
<emphasis id="B925EACEFF89FFF5FA8CFED0B567FEFC" box="[1325,1348,311,335]" italics="true" pageId="7" pageNumber="79">s2</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FAF7FED0B5ACFEFC" box="[1366,1423,311,335]" italics="true" pageId="7" pageNumber="79">r2-r6</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FC9BFEBDB3A2FEC1" box="[826,897,346,370]" italics="true" pageId="7" pageNumber="79">R1-R6</emphasis>
) apparently shorter, similar to setae on dorsal shield. Interior region of marginal shield lacks setae, only with three pairs of gland pores (including pygidial pores). Complement of pores as in figure 4A; dorsal shield with four pairs of lyrifissures, six pairs of glandular pores, and five pairs of punctiform poroids (the three pairs of glandular pores at the marginal region of dorsal shield may be related to concave formations in the adult); most setae on marginal shields associate to poroids set on conical tubercles. Submarginal region poorly sclerotized with 
<emphasis id="B925EACEFF89FFF5FC70FD3BB3C5FD47" box="[977,998,732,756]" italics="true" pageId="7" pageNumber="79">j1</emphasis>
and ca. 25 pair of curved and ciliate setae (setae 
<emphasis id="B925EACEFF89FFF5FC68FD18B3CCFCA5" box="[969,1007,767,790]" italics="true" pageId="7" pageNumber="79">UR</emphasis>
) similar in length and shape to anteriormost podonotal setae (
<figureCitation id="136A2A59FF89FFF5FB25FCC5B4FFFC89" box="[1156,1244,802,826]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="7" pageNumber="79">Fig. 4C</figureCitation>
); no discernible poroids in this region. All dorsal setae set on conical tubercles.
</paragraph>
<paragraph id="8BEE36DCFF89FFF5FCC1FC7AB48DFACE" blockId="7.[826,1429,925,1405]" pageId="7" pageNumber="79">
Ventral idiosoma (
<figureCitation id="136A2A59FF89FFF5FB9FFC7AB4BEFC06" box="[1086,1181,925,949]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="7" pageNumber="79">Fig. 4B</figureCitation>
) — Anterior acuminate prominence with four dentate marginal lobes. Sternal shield well sclerotized and ornamented with polygonal cells from setae 
<emphasis id="B925EACEFF89FFF5FBCEFBE1B4ACFBAD" box="[1135,1167,1030,1054]" italics="true" pageId="7" pageNumber="79">st2</emphasis>
to posterior margin of the shield; shield free from endopodal extensions between coxae II and III, with five pairs of setae 
<emphasis id="B925EACEFF89FFF5FACAFBABB342FB34" italics="true" pageId="7" pageNumber="79">st1- st5,</emphasis>
three pair of glandular pores (
<emphasis id="B925EACEFF89FFF5FB6BFB88B52DFB34" box="[1226,1294,1135,1159]" italics="true" pageId="7" pageNumber="79">gst1-3</emphasis>
), and three pairs of no glandular poroids (
<emphasis id="B925EACEFF89FFF5FB0CFB75B4ECFB19" box="[1197,1231,1170,1194]" italics="true" pageId="7" pageNumber="79">iv1</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FB41FB75B521FB19" box="[1248,1282,1170,1194]" italics="true" pageId="7" pageNumber="79">iv3</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FAB3FB74B517FB19" box="[1298,1332,1171,1194]" italics="true" pageId="7" pageNumber="79">iv5</emphasis>
); shield 
<quantity id="4CA99B39FF89FFF5FC9BFB52B3ACFB7E" box="[826,911,1205,1230]" metricMagnitude="-1" metricUnit="m" metricValue="2.46" pageId="7" pageNumber="79" unit="mm" value="246.0">246 µm</quantity>
long, 
<quantity id="4CA99B39FF89FFF5FC79FB52B403FB7E" box="[984,1056,1205,1230]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="79" unit="mm" value="76.0">76 µm</quantity>
wide at level of 
<emphasis id="B925EACEFF89FFF5FB42FB52B520FB7E" box="[1251,1283,1205,1229]" italics="true" pageId="7" pageNumber="79">st2</emphasis>
, 27 µm long at level of 
<emphasis id="B925EACEFF89FFF5FC16FB3EB3F4FB43" box="[951,983,1241,1264]" italics="true" pageId="7" pageNumber="79">st5</emphasis>
; eroded at level of 
<emphasis id="B925EACEFF89FFF5FB1CFB3EB4C6FB42" box="[1213,1253,1241,1265]" italics="true" pageId="7" pageNumber="79">gv1</emphasis>
and narrowed behind 
<emphasis id="B925EACEFF89FFF5FC33FB1BB391FAA7" box="[914,946,1276,1300]" italics="true" pageId="7" pageNumber="79">st4</emphasis>
: 
<emphasis id="B925EACEFF89FFF5FC60FB1BB3C2FAA0" box="[961,993,1276,1299]" italics="true" pageId="7" pageNumber="79">st5</emphasis>
and 
<emphasis id="B925EACEFF89FFF5FBBBFB1BB41FFAA0" box="[1050,1084,1276,1299]" italics="true" pageId="7" pageNumber="79">iv5</emphasis>
at the posterior margin of the shield at level of coxae IV (
<figureCitation id="136A2A59FF89FFF5FBD3FAF9B4E4FA84" box="[1138,1223,1310,1335]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="7" pageNumber="79">Fig. 4B</figureCitation>
). Setae 
<emphasis id="B925EACEFF89FFF5FABCFAF8B51DFA84" box="[1309,1342,1311,1335]" italics="true" pageId="7" pageNumber="79">st1</emphasis>
, 
<emphasis id="B925EACEFF89FFF5FAEAFAF8B5B6FA84" box="[1355,1429,1311,1335]" italics="true" pageId="7" pageNumber="79">st3-st5</emphasis>
minute and smooth (
<quantity id="4CA99B39FF89FFF5FB9BFAA5B4ACFAE9" box="[1082,1167,1346,1370]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="79" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 µm</quantity>
), 
<emphasis id="B925EACEFF89FFF5FB0BFAA5B4E9FAE9" box="[1194,1226,1346,1370]" italics="true" pageId="7" pageNumber="79">st2</emphasis>
rod-like, thicker and almost twice longer (
<quantity id="4CA99B39FF89FFF5FBC5FA83B4BDFACE" box="[1124,1182,1380,1405]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="79" unit="mm" value="5.0">5 µm</quantity>
).
</paragraph>
<paragraph id="8BEE36DCFF89FFF9FCC1FA7DB1CFFD39" blockId="7.[826,1429,1434,1949]" lastBlockId="11.[159,762,311,1949]" lastPageId="11" lastPageNumber="83" pageId="7" pageNumber="79">
Ventrianal shield well sclerotized and ornamented, almost three times wider than long (
<quantity id="4CA99B39FF89FFF5FAE4FA5AB343FA4B" metricMagnitude="-2" metricUnit="m" metricValue="6.8" metricValueMax="6.9" metricValueMin="6.7" pageId="7" pageNumber="79" unit="mm" value="68.0" valueMax="69.0" valueMin="67.0">67 – 69 µm</quantity>
long, 
<quantity id="4CA99B39FF89FFF5FC0DFA07B421FA4B" box="[940,1026,1504,1528]" metricMagnitude="-1" metricUnit="m" metricValue="2.62" pageId="7" pageNumber="79" unit="mm" value="262.0">262 µm</quantity>
wide), with pilose circumanal setae similar in length (ca. 
<quantity id="4CA99B39FF89FFF5FBFEF9E4B484F9A8" box="[1119,1191,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="79" unit="mm" value="27.0">27 µm</quantity>
), three pairs of ventral setae at the anterior margin of the shield, 
<emphasis id="B925EACEFF89FFF5FAE2F9C1B54EF98D" box="[1347,1389,1574,1598]" italics="true" pageId="7" pageNumber="79">JV3</emphasis>
(
<quantity id="4CA99B39FF89FFF5FAD8F9C1B3B5F9D2" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="7" pageNumber="79" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16 – 18 µm</quantity>
) poorly ciliate, 
<emphasis id="B925EACEFF89FFF5FBECF9AEB455F9D2" box="[1101,1142,1609,1633]" italics="true" pageId="7" pageNumber="79">JV4</emphasis>
and 
<emphasis id="B925EACEFF89FFF5FB10F9AEB4F9F9D3" box="[1201,1242,1609,1632]" italics="true" pageId="7" pageNumber="79">JV5</emphasis>
similar to other posterior most opisthogastric setae, two pairs of glandular pores, each associated with ventral setae 
<emphasis id="B925EACEFF89FFF5FC9BF955B347F979" box="[826,868,1714,1738]" italics="true" pageId="7" pageNumber="79">JV3</emphasis>
and 
<emphasis id="B925EACEFF89FFF5FC3BF954B3E7F979" box="[922,964,1715,1738]" italics="true" pageId="7" pageNumber="79">JV5</emphasis>
, and 
<emphasis id="B925EACEFF89FFF5FBA6F955B40DF979" box="[1031,1070,1714,1738]" italics="true" pageId="7" pageNumber="79">gv3</emphasis>
at the posterior margin at level of postanal setae; one pair of poroids associated to 
<emphasis id="B925EACEFF89FFF5FCFBF91FB3A0F8A3" box="[858,899,1784,1808]" italics="true" pageId="7" pageNumber="79">JV4</emphasis>
; cribrum narrow as in adult. Opisthogaster with five pairs of ventral setae on soft tegument, 
<emphasis id="B925EACEFF89FFF5FACAF8FCB5B6F880" box="[1387,1429,1819,1843]" italics="true" pageId="7" pageNumber="79">JV1</emphasis>
shortest (
<quantity id="4CA99B39FF89FFF5FC07F8D8B3C2F8E5" box="[934,993,1854,1879]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="79" unit="mm" value="8.0">8 µm</quantity>
) and smooth and four pairs of ciliate setae set on tubercles at the lateral sides (
<emphasis id="B925EACEFF89FFF5FABAF885B34FF82F" italics="true" pageId="7" pageNumber="79">ZV1, ZV3- ZV5</emphasis>
); two pairs of glandular pores and one pair of discernible poroids on soft opisthogastric cuticle (
<figureCitation id="136A2A59FF85FFF9FF06FEBDB121FEC1" box="[167,258,346,370]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="11" pageNumber="83">Fig. 4B</figureCitation>
). Endopodal strips between legs II and III present and free; endopodal between coxae III and IV fused with parapodal elements, and pedofossae IV well developed bearing one anterior liryfissure and one posterior glandular poroid: exopodal elements II and III fused and contiguous with exopodal I-II. One pair of poroids distinct on soft cuticle between legs III and IV. Rim of exopodal plate behind coxa IV inconspicuous.
</paragraph>
<caption id="DF2E6654FF86FFFAFF3EF94BB2AEF8A2" ID-DOI="http://doi.org/10.5281/zenodo.4640188" ID-Zenodo-Dep="4640188" httpUri="https://zenodo.org/record/4640188/files/figure.png" pageId="8" pageNumber="80" startId="8.[159,171,1708,1727]" targetBox="[231,1359,445,1641]" targetPageId="8">
<paragraph id="8BEE36DCFF86FFFAFF3EF94BB2AEF8A2" blockId="8.[159,1429,1707,1810]" pageId="8" pageNumber="80">
FIGURE 3: 
<taxonomicName id="4C514D5FFF86FFFAFEA4F94CB1E9F90D" authority="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari, 2016" authorityName="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari" authorityYear="2016" box="[261,458,1707,1726]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="80" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B925EACEFF86FFFAFEA4F94CB1E9F90D" box="[261,458,1707,1726]" italics="true" pageId="8" pageNumber="80">Eutrachytes flagellatus</emphasis>
</taxonomicName>
<emphasis id="B925EACEFF86FFFAFE71F94CB227F90D" bold="true" box="[464,516,1707,1726]" pageId="8" pageNumber="80">
<taxonomicNameLabel id="A21657B5FF86FFFAFE71F94CB227F90D" box="[464,516,1707,1726]" pageId="8" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – Female, leg IV, femur, genu and tibia, lateral view; B – Female, tarsus III, anterolateral view; C – Male, leg II, femur, genu and tibia, lateral view; D – Male, tarsus II, posterolateral view; E – Female, coxae I with glandular fields; F – Male, coxa and trochanter I, detail of serrate distal margins; G – Tarsus II, detail of pretarsus; H – Detail of denoted basitarsal feather-like setae. Dorsal setae in soli black color.
</paragraph>
</caption>
<caption id="DF2E6654FF87FFFBFF3EF8C1B325F8E6" ID-DOI="http://doi.org/10.5281/zenodo.4640192" ID-Zenodo-Dep="4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="9" pageNumber="81" startId="9.[159,171,1830,1849]" targetBox="[227,1359,364,1775]" targetPageId="9">
<paragraph id="8BEE36DCFF87FFFBFF3EF8C1B325F8E6" blockId="9.[159,1429,1830,1877]" pageId="9" pageNumber="81">
FIGURE 4: 
<taxonomicName id="4C514D5FFF87FFFBFEA5F8C1B1E4F88A" authority="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari, 2016" authorityName="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari" authorityYear="2016" box="[260,455,1830,1849]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="81" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B925EACEFF87FFFBFEA5F8C1B1E4F88A" box="[260,455,1830,1849]" italics="true" pageId="9" pageNumber="81">Eutrachytes flagellatus</emphasis>
</taxonomicName>
<emphasis id="B925EACEFF87FFFBFE6DF8C1B223F88A" bold="true" box="[460,512,1830,1849]" pageId="9" pageNumber="81">
<taxonomicNameLabel id="A21657B5FF87FFFBFE6DF8C1B223F88A" box="[460,512,1830,1849]" pageId="9" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure.
</paragraph>
</caption>
<caption id="DF2E6654FF84FFF8FF3EF8C1B2FDF8E6" ID-DOI="http://doi.org/10.5281/zenodo.4640194" ID-Zenodo-Dep="4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="10" pageNumber="82" startId="10.[159,171,1830,1849]" targetBox="[227,1354,365,1790]" targetPageId="10">
<paragraph id="8BEE36DCFF84FFF8FF3EF8C1B2FDF8E6" blockId="10.[159,1429,1830,1877]" pageId="10" pageNumber="82">
FIGURE 5: 
<taxonomicName id="4C514D5FFF84FFF8FEA9F8C1B1EEF88A" authority="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari, 2016" authorityName="Moraza &amp; Kontschan &amp; Sahoo &amp; Ansari" authorityYear="2016" box="[264,461,1830,1849]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="82" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B925EACEFF84FFF8FEA9F8C1B1EEF88A" box="[264,461,1830,1849]" italics="true" pageId="10" pageNumber="82">Eutrachytes flagellatus</emphasis>
</taxonomicName>
<emphasis id="B925EACEFF84FFF8FE75F8C1B22FF88A" bold="true" box="[468,524,1830,1849]" pageId="10" pageNumber="82">
<taxonomicNameLabel id="A21657B5FF84FFF8FE75F8C1B22FF88A" box="[468,524,1830,1849]" pageId="10" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view.
</paragraph>
</caption>
<paragraph id="8BEE36DCFF85FFF9FF64FD7CB12EFC7F" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="83">
Peritrematal shield from coxae I to anterior margin of coxae III ornate as exopodal elements; respiratory stigmata at level of coxae II, peritremes extending to a point between coxae I and II and continues outside the body given way to a long, flagellate and tubular structure (
<figureCitation id="136A2A59FF85FFF9FE7BFCACB213FCD0" box="[474,560,843,867]" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="11" pageNumber="83">Fig. 4B</figureCitation>
). This hollow cuticular structure has heavily ringed surface and its diameter decreases progressively at the distal end (
<figureCitation id="136A2A59FF85FFF9FF06FC53B123FC7F" box="[167,256,948,972]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="11" pageNumber="83">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF85FFF9FF64FC3AB171FB32" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="83">Gnathosoma — Gnathotectum, chelicerae and other mouthpart structures, corniculi and adjacent structures as in adult female; palpi similar to those in adult female, including similar form of setae on palptrochanter.</paragraph>
<paragraph id="8BEE36DCFF85FFF9FF64FB74B2D9FAEA" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="83">
Legs — Pretarsal structures, chaetotaxy and form and shape of leg setae similar to those on adult female, except that femora I-IV lack ventral tectum. Legs length: I 
<quantity id="4CA99B39FF85FFF9FEE9FB1BB1C2FAA0" box="[328,481,1275,1300]" metricMagnitude="-1" metricUnit="m" metricValue="3.35" metricValueMax="3.4" metricValueMin="3.3" pageId="11" pageNumber="83" unit="mm" value="335.0" valueMax="340.0" valueMin="330.0">330 – 340 µm</quantity>
, II 
<quantity id="4CA99B39FF85FFF9FDA9FB1BB282FAA0" box="[520,673,1275,1300]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="11" pageNumber="83" unit="mm" value="295.0" valueMax="300.0" valueMin="290.0">290 – 300 µm</quantity>
, III 
<quantity id="4CA99B39FF85FFF9FD73FB1BB129FA84" metricMagnitude="-1" metricUnit="m" metricValue="3.025" metricValueMax="3.05" metricValueMin="3.0" pageId="11" pageNumber="83" unit="mm" value="302.5" valueMax="305.0" valueMin="300.0">300 – 305 µm</quantity>
, IV 
<quantity id="4CA99B39FF85FFF9FE9DFAF9B1F4FA84" box="[316,471,1310,1335]" metricMagnitude="-1" metricUnit="m" metricValue="3.275" metricValueMax="3.3" metricValueMin="3.25" pageId="11" pageNumber="83" unit="mm" value="327.5" valueMax="330.0" valueMin="325.0">325 – 330 µm</quantity>
. Chaetotaxy of legs I-IV as in adult, including fanlike setae on basitarsi II-IV.
</paragraph>
<paragraph id="8BEE36DCFF85FFF9FF64FA8CB22AFA15" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="83">
<emphasis id="B925EACEFF85FFF9FF64FA8CB17EFA30" bold="true" box="[197,349,1387,1411]" pageId="11" pageNumber="83">Protonymph</emphasis>
— Idiosoma 
<quantity id="4CA99B39FF85FFF9FE5EFA8CB275FA30" box="[511,598,1387,1411]" metricMagnitude="-1" metricUnit="m" metricValue="3.02" pageId="11" pageNumber="83" unit="mm" value="302.0">302 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FD02FA8CB2DAFA30" box="[675,761,1387,1411]" metricMagnitude="-1" metricUnit="m" metricValue="2.42" pageId="11" pageNumber="83" unit="mm" value="242.0">242 µm</quantity>
wide at level of coxae III (n=1).
</paragraph>
<paragraph id="8BEE36DCFF85FFF9FF64FA50B1D5F89E" blockId="11.[159,762,311,1949]" pageId="11" pageNumber="83">
Dorsal idiosoma — Dorsum (
<figureCitation id="136A2A59FF85FFF9FD9CFA50B28EFA7C" box="[573,685,1463,1487]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="11" pageNumber="83">Fig. 5A</figureCitation>
) with well sclerotized rhomboidal podonotal shield with rounded corners and eroded lateral margins; shield ca. 
<quantity id="4CA99B39FF85FFF9FF1CF9C7B134F98B" box="[189,279,1568,1592]" metricMagnitude="-1" metricUnit="m" metricValue="1.71" pageId="11" pageNumber="83" unit="mm" value="171.0">171 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FEC7F9C7B19DF98B" box="[358,446,1568,1592]" metricMagnitude="-1" metricUnit="m" metricValue="1.38" pageId="11" pageNumber="83" unit="mm" value="138.0">138 µm</quantity>
wide, with central region punctate and lateral regions ornate with round shallow pits; one pair of nude, slightly ornate mesonotal shield at both sides of posterior region of podonotal shield, 
<quantity id="4CA99B39FF85FFF9FEBFF94BB180F977" box="[286,419,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" metricValueMax="5.3" metricValueMin="4.9" pageId="11" pageNumber="83" unit="mm" value="51.0" valueMax="53.0" valueMin="49.0">49 – 53 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FE53F94BB254F977" box="[498,631,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="11" pageNumber="83" unit="mm" value="25.5" valueMax="26.0" valueMin="25.0">25 – 26 µm</quantity>
wide, and nude pygidial shield narrow (
<quantity id="4CA99B39FF85FFF9FDA8F928B2B3F954" box="[521,656,1743,1768]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="11" pageNumber="83" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13 – 14 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FD7EF928B0E6F8B9" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="83" unit="mm" value="84.0">84 µm</quantity>
wide), as a concave strip, and with one pair of poroids at the lateral corners.
</paragraph>
<paragraph id="8BEE36DCFF85FFF9FF64F8D9B455FC75" blockId="11.[159,762,311,1949]" lastBlockId="11.[826,1429,311,966]" pageId="11" pageNumber="83">
Dorsal setation holotrichous, with addition of setae 
<emphasis id="B925EACEFF85FFF9FF43F885B0D4F8C9" box="[226,247,1890,1914]" italics="true" pageId="11" pageNumber="83">j2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FEA6F885B13EF8C9" box="[263,285,1890,1914]" italics="true" pageId="11" pageNumber="83">J1</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FE8CF885B16FF8C9" box="[301,332,1890,1914]" italics="true" pageId="11" pageNumber="83">Z1</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FEFDF885B158F8C9" box="[348,379,1890,1914]" italics="true" pageId="11" pageNumber="83">Z2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FE2AF885B180F8CA" box="[395,419,1890,1913]" italics="true" pageId="11" pageNumber="83">s5</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FE12F885B1EDF8C9" box="[435,462,1890,1914]" italics="true" pageId="11" pageNumber="83">S2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FE7EF885B1D5F8C9" box="[479,502,1890,1914]" italics="true" pageId="11" pageNumber="83">r2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FDA7F885B23DF8C9" box="[518,542,1890,1914]" italics="true" pageId="11" pageNumber="83">r3</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FD8FF885B266F8CA" box="[558,581,1890,1913]" italics="true" pageId="11" pageNumber="83">r5</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FDF4F885B256F8C9" box="[597,629,1890,1914]" italics="true" pageId="11" pageNumber="83">R1</emphasis>
; paravertical poroids associate to 
<emphasis id="B925EACEFF85FFF9FE1EF862B1F7F82E" box="[447,468,1925,1949]" italics="true" pageId="11" pageNumber="83">j1</emphasis>
distinct and tuberculate (
<figureCitation id="136A2A59FF85FFF9FCE3FED0B383FEFC" box="[834,928,311,335]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="11" pageNumber="83">Fig. 5A</figureCitation>
). Podonotal shield with five pair of ciliate setae (
<emphasis id="B925EACEFF85FFF9FC27FEBDB39BFEC1" box="[902,952,346,370]" italics="true" pageId="11" pageNumber="83">j3-j6</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC68FEBDB3C1FEC2" box="[969,994,346,369]" italics="true" pageId="11" pageNumber="83">z5</emphasis>
), similar in length and shape to larval setae and four pairs of poroids (three pairs appear to be glandular pores). Other podonotal and opisthonotal setae on soft cuticle, including setae 
<emphasis id="B925EACEFF85FFF9FC9BFE01B373FE4D" box="[826,848,486,510]" italics="true" pageId="11" pageNumber="83">J1</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FCFCFE01B350FE4D" box="[861,883,486,510]" italics="true" pageId="11" pageNumber="83">J2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC20FE01B3BCFE4D" box="[897,927,486,510]" italics="true" pageId="11" pageNumber="83">Z1</emphasis>
similar to setae on podonotal shield (short and curved), with sparse and long ciliae and set in conical tubercles; other dorsal setae longer, slightly curved with few ciliae; setae 
<emphasis id="B925EACEFF85FFF9FB35FDA8B48EFDD4" box="[1172,1197,591,615]" italics="true" pageId="11" pageNumber="83">z4</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FB1DFDA8B4F0FDD4" box="[1212,1235,591,615]" italics="true" pageId="11" pageNumber="83">r3</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FB43FDA8B521FDD4" box="[1250,1282,591,615]" italics="true" pageId="11" pageNumber="83">R1</emphasis>
(
<quantity id="4CA99B39FF85FFF9FAB0FDA8B5A6FDD4" box="[1297,1413,591,616]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="83" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9 – 11 µm</quantity>
); other podonotal setae, 
<emphasis id="B925EACEFF85FFF9FBEBFD94B449FD39" box="[1098,1130,627,650]" italics="true" pageId="11" pageNumber="83">Z5</emphasis>
and 
<emphasis id="B925EACEFF85FFF9FB07FD94B4E1FD38" box="[1190,1218,627,651]" italics="true" pageId="11" pageNumber="83">S2</emphasis>
(ca. 
<quantity id="4CA99B39FF85FFF9FAA1FD94B56AFD39" box="[1280,1353,626,651]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="83" unit="mm" value="22.0">22 µm</quantity>
long), 
<emphasis id="B925EACEFF85FFF9FC9BFD71B371FD1E" box="[826,850,662,685]" italics="true" pageId="11" pageNumber="83">s5</emphasis>
and other opisthonotal setae longer (
<quantity id="4CA99B39FF85FFF9FAA6FD71B5A6FD1E" box="[1287,1413,661,686]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="4.0" metricValueMin="2.9" pageId="11" pageNumber="83" unit="mm" value="34.5" valueMax="40.0" valueMin="29.0">29 – 40 µm</quantity>
). Soft tegument with 18 pairs of poroids as in figure 23 (eight pododontal and 10 opisthonotal); pores associate at the base of dorsal setae 
<emphasis id="B925EACEFF85FFF9FB77FD18B4C8FCA4" box="[1238,1259,767,791]" italics="true" pageId="11" pageNumber="83">j1</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FB5DFD18B530FCA4" box="[1276,1299,767,791]" italics="true" pageId="11" pageNumber="83">r2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FA85FD18B51EFCA4" box="[1316,1341,767,791]" italics="true" pageId="11" pageNumber="83">z4</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FAEFFD18B545FCA4" box="[1358,1382,767,791]" italics="true" pageId="11" pageNumber="83">s4</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FAD7FD18B5ADFCA5" box="[1398,1422,767,790]" italics="true" pageId="11" pageNumber="83">s5</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC9BFCC5B371FC89" box="[826,850,802,826]" italics="true" pageId="11" pageNumber="83">s6</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FCC2FCC5B35BFC89" box="[867,888,802,826]" italics="true" pageId="11" pageNumber="83">J2</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC2BFCC5B383FC8A" box="[906,928,802,825]" italics="true" pageId="11" pageNumber="83">J5</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC10FCC5B3EEFC89" box="[945,973,802,826]" italics="true" pageId="11" pageNumber="83">S4</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC7FFCC5B3D9FC8A" box="[990,1018,802,825]" italics="true" pageId="11" pageNumber="83">S5</emphasis>
. Peritremal shields narrow, laterally with a conical peritremal extension at level of setae 
<emphasis id="B925EACEFF85FFF9FCDCFC8FB3B6FC33" box="[893,917,872,896]" italics="true" pageId="11" pageNumber="83">s4</emphasis>
and 
<emphasis id="B925EACEFF85FFF9FC72FC8FB3C9FCCC" box="[979,1002,872,895]" italics="true" pageId="11" pageNumber="83">r5</emphasis>
, of which emerges a tubular flagellate tubular structure similar to the subsequent deutonymphal instar (
<figureCitation id="136A2A59FF85FFF9FBB5FC49B44AFC75" box="[1044,1129,942,966]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="11" pageNumber="83">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF85FFF9FCC1FC04B3B3F892" blockId="11.[826,1429,995,1825]" pageId="11" pageNumber="83">
Ventral idiosoma (
<figureCitation id="136A2A59FF85FFF9FBE4FC04B48CFC48" box="[1093,1199,995,1019]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="11" pageNumber="83">Fig. 5B</figureCitation>
) — Sternal shield as in deutonymph, well sclerotized, pear like in shape, 
<quantity id="4CA99B39FF85FFF9FC2FFBCEB3C7FBF2" box="[910,996,1065,1090]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="83" unit="mm" value="115.0">115 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FB91FBCEB459FBF2" box="[1072,1146,1065,1090]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="83" unit="mm" value="82.0">82 µm</quantity>
wide and level of setae 
<emphasis id="B925EACEFF85FFF9FC9BFBABB379FBD7" box="[826,858,1100,1124]" italics="true" pageId="11" pageNumber="83">st2</emphasis>
; shield entire, without endopodal extensions between coxae II-III and III-IV, and with sternal setae 
<emphasis id="B925EACEFF85FFF9FC9BFB75B3A0FB19" box="[826,899,1170,1194]" italics="true" pageId="11" pageNumber="83">st1-st3</emphasis>
on rounded rounded tubercles, and poroids 
<emphasis id="B925EACEFF85FFF9FC9BFB52B37FFB7E" box="[826,860,1205,1229]" italics="true" pageId="11" pageNumber="83">iv1</emphasis>
; setae 
<emphasis id="B925EACEFF85FFF9FC17FB52B3F4FB7E" box="[950,983,1205,1229]" italics="true" pageId="11" pageNumber="83">st1</emphasis>
(
<quantity id="4CA99B39FF85FFF9FC4BFB52B478FB7E" box="[1002,1115,1205,1230]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="11" pageNumber="83" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4 – 5 µm</quantity>
long) and 
<emphasis id="B925EACEFF85FFF9FB46FB52B52BFB7E" box="[1255,1288,1205,1229]" italics="true" pageId="11" pageNumber="83">st3</emphasis>
(
<quantity id="4CA99B39FF85FFF9FABAFB52B5AFFB7E" box="[1307,1420,1205,1230]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="11" pageNumber="83" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6 – 7 µm</quantity>
) shorter than 
<emphasis id="B925EACEFF85FFF9FC75FB3EB3D6FB42" box="[980,1013,1241,1265]" italics="true" pageId="11" pageNumber="83">st2</emphasis>
(
<quantity id="4CA99B39FF85FFF9FBA7FB3EB4A8FB43" box="[1030,1163,1240,1265]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="11" pageNumber="83" unit="mm" value="28.5" valueMax="29.0" valueMin="28.0">28 – 29 µm</quantity>
); thin and smooth 
<emphasis id="B925EACEFF85FFF9FAD5FB3EB5B6FB43" box="[1396,1429,1241,1264]" italics="true" pageId="11" pageNumber="83">st5</emphasis>
behind coxae IV (
<quantity id="4CA99B39FF85FFF9FBA9FB1BB44FFAA0" box="[1032,1132,1275,1300]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="11" pageNumber="83" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4 – 5 µm</quantity>
long). Intercoxal soft cuticle with a subcutaneous clear spot between coxae IV. Opisthogaster with well sclerotized subpentagonal anal shield of moderate size (
<quantity id="4CA99B39FF85FFF9FB7FFA83B509FACE" box="[1246,1322,1380,1405]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="83" unit="mm" value="65.0">65 µm</quantity>
long, 
<quantity id="4CA99B39FF85FFF9FADBFA82B343FA13" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="11" pageNumber="83" unit="mm" value="89.0">89 µm</quantity>
wide), relatively longer than in subsequent deutonymphal instar, and with three circumanal pilose setae similar in shape and length to 
<emphasis id="B925EACEFF85FFF9FB74FA29B4DCFA56" box="[1237,1279,1486,1509]" italics="true" pageId="11" pageNumber="83">JV5</emphasis>
(
<quantity id="4CA99B39FF85FFF9FAADFA29B5A6FA55" box="[1292,1413,1486,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="11" pageNumber="83" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15 – 16 µm</quantity>
), glands 
<emphasis id="B925EACEFF85FFF9FC30FA16B39AF9BA" box="[913,953,1521,1545]" italics="true" pageId="11" pageNumber="83">gv3</emphasis>
on the margin posterior to postanal setae, and cribum. Opistogaster with one pair of porelike structures and two pairs of pilose setae (
<emphasis id="B925EACEFF85FFF9FAC4F9D0B5ADF9FC" box="[1381,1422,1591,1615]" italics="true" pageId="11" pageNumber="83">JV1</emphasis>
, 
<emphasis id="B925EACEFF85FFF9FC9BF9BCB340F9C1" box="[826,867,1627,1650]" italics="true" pageId="11" pageNumber="83">JV5</emphasis>
) on tubercles. Rim of exopodal plate not discernible behind coxae IV, but inguinal gland pores 
<emphasis id="B925EACEFF85FFF9FC9BF947B341F90B" box="[826,866,1696,1720]" italics="true" pageId="11" pageNumber="83">gv2</emphasis>
and poroids present there. A pair of poroids lateral to coxae III-IV. Reduce peritrematal shields with short and narrow peritremes between coxae II and III.
</paragraph>
<paragraph id="8BEE36DCFF85FFFEFCC1F8D9B23FFE0B" blockId="11.[826,1429,1854,1948]" lastBlockId="12.[159,762,311,1948]" lastPageId="12" lastPageNumber="84" pageId="11" pageNumber="83">Gnathosoma — Gnathotectum and tritosternal base as in subsequent instars. Form of corniculus and internal malae much as in subsequent instars; deutosternum similar to those in deutonymphs. Palps with normal complement of setae (1-2-5-12); chelicera not clearly discernible in a single available specimen; palp-trochanter nude.</paragraph>
<paragraph id="8BEE36DCFF82FFFEFF64FE28B21DFCF5" blockId="12.[159,762,311,1948]" pageId="12" pageNumber="84">Legs — Legs I to IV with pretarsi, welldeveloped claws and pulvillus. Legs complement of setae as follows: coxae, 2-2-2-1; trochanters 5-5-5- 5; femora 9 – 9 – 6 – 6; genua 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1) – 5 (0 2/0 2/0 1); tibiae 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1). Basitarsus II-IV as long as telotarsus excluding the pretarsus. Coxa, trochanter, femora, genua and tibia I-IV with ventrolateral distal margins strongly serrate. Leg setae generally simple or pilose, not markedly differentiated.</paragraph>
<paragraph id="8BEE36DCFF82FFFEFF64FCBAB0C7FC2B" blockId="12.[159,762,311,1948]" pageId="12" pageNumber="84">
<emphasis id="B925EACEFF82FFFEFF64FCBAB12AFCC6" bold="true" box="[197,265,861,885]" pageId="12" pageNumber="84">Larva</emphasis>
— Idiosoma 
<quantity id="4CA99B39FF82FFFEFE10FCBAB229FCC6" box="[433,522,861,885]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="84" unit="mm" value="310.0">310 µm</quantity>
long, 
<quantity id="4CA99B39FF82FFFEFDFAFCBBB297FCC6" box="[603,692,860,885]" metricMagnitude="-1" metricUnit="m" metricValue="2.26" pageId="12" pageNumber="84" unit="mm" value="226.0">226 µm</quantity>
wide (n=1).
</paragraph>
<paragraph id="8BEE36DCFF82FFFEFF64FC48B563FE26" blockId="12.[159,762,311,1948]" lastBlockId="12.[826,1429,311,1948]" pageId="12" pageNumber="84">
Dorsal idiosoma — Dorsal shielding clearly delimited and surface discernibly ornamented as on nymphal instars (
<figureCitation id="136A2A59FF82FFFEFED5FC12B1FBFBBE" box="[372,472,1013,1037]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="12" pageNumber="84">Fig. 5C</figureCitation>
). Body dorsum with 20 pairs of setae set on cylindrical tubercles; nine pairs of setae (
<emphasis id="B925EACEFF82FFFEFEA5FBDCB13AFBE0" box="[260,281,1083,1107]" italics="true" pageId="12" pageNumber="84">j1</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFE84FBDCB174FBE0" box="[293,343,1083,1107]" italics="true" pageId="12" pageNumber="84">j3-j6</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFEC2FBDCB15FFBE0" box="[355,380,1083,1107]" italics="true" pageId="12" pageNumber="84">z2</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFE29FBDCB181FBE0" box="[392,418,1083,1107]" italics="true" pageId="12" pageNumber="84">z4</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFE0FFBDCB1E4FBE1" box="[430,455,1083,1106]" italics="true" pageId="12" pageNumber="84">z5</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFE72FBDCB1C8FBE0" box="[467,491,1083,1107]" italics="true" pageId="12" pageNumber="84">s4</emphasis>
) on rhomboid podonotal shield (
<quantity id="4CA99B39FF82FFFEFE83FBB9B15EFBC5" box="[290,381,1118,1142]" metricMagnitude="-1" metricUnit="m" metricValue="2.12" pageId="12" pageNumber="84" unit="mm" value="212.0">212 µm</quantity>
long, 
<quantity id="4CA99B39FF82FFFEFE70FBB9B208FBC5" box="[465,555,1118,1142]" metricMagnitude="-1" metricUnit="m" metricValue="1.5699999999999998" pageId="12" pageNumber="84" unit="mm" value="157.0">157 µm</quantity>
wide at level of seta 
<emphasis id="B925EACEFF82FFFEFF74FB66B0CEFB2A" box="[213,237,1153,1177]" italics="true" pageId="12" pageNumber="84">s4</emphasis>
); setae 
<emphasis id="B925EACEFF82FFFEFEE9FB66B142FB2B" box="[328,353,1153,1176]" italics="true" pageId="12" pageNumber="84">z5</emphasis>
on the parallel lines connecting 
<emphasis id="B925EACEFF82FFFEFD44FB66B2D9FB2A" box="[741,762,1153,1177]" italics="true" pageId="12" pageNumber="84">j1</emphasis>
and 
<emphasis id="B925EACEFF82FFFEFF75FB43B0CAFB0F" box="[212,233,1188,1212]" italics="true" pageId="12" pageNumber="84">j6</emphasis>
; 11 pairs of setae on soft unsclerotized cuticle, podonotal pair 
<emphasis id="B925EACEFF82FFFEFE21FB20B1BBFB6C" box="[384,408,1223,1247]" italics="true" pageId="12" pageNumber="84">s6</emphasis>
and opisthonotal 
<emphasis id="B925EACEFF82FFFEFDCBFB20B283FB6C" box="[618,672,1223,1247]" italics="true" pageId="12" pageNumber="84">J2-J5</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFD0DFB20B2D0FB6C" box="[684,755,1223,1247]" italics="true" pageId="12" pageNumber="84">Z3-Z5</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFF3EFB0DB0FCFAB1" box="[159,223,1258,1282]" italics="true" pageId="12" pageNumber="84">S3-S5</emphasis>
; at least eighth pairs of poroids (four podonotal and three opisthonotal) of which one pair of no glandular pores is associated with a conspicuous protuberance on lateral margin of podonotal shield between setae 
<emphasis id="B925EACEFF82FFFEFEE8FA90B141FA3C" box="[329,354,1399,1423]" italics="true" pageId="12" pageNumber="84">z4</emphasis>
and 
<emphasis id="B925EACEFF82FFFEFE3DFA90B197FA3C" box="[412,436,1399,1423]" italics="true" pageId="12" pageNumber="84">s4</emphasis>
; other conspicuous glandular poroid on soft opisthonotal cuticle behind 
<emphasis id="B925EACEFF82FFFEFD75FA7DB2D0FA01" box="[724,755,1434,1458]" italics="true" pageId="12" pageNumber="84">Z3</emphasis>
, and three pairs of poroids on soft cuticle at level of coxae II and III. Setae 
<emphasis id="B925EACEFF82FFFEFE7FFA07B1D0FA4B" box="[478,499,1504,1528]" italics="true" pageId="12" pageNumber="84">j1</emphasis>
smooth divided into two dissimilar branches, short branch (
<quantity id="4CA99B39FF82FFFEFDCAF9E4B296F9A8" box="[619,693,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="84" unit="mm" value="24.0">24 µm</quantity>
long) half the length of the longest (
<quantity id="4CA99B39FF82FFFEFE5EF9C1B26BF98D" box="[511,584,1574,1598]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="12" pageNumber="84" unit="mm" value="59.0">59 µm</quantity>
); setae 
<emphasis id="B925EACEFF82FFFEFD00F9C1B2D9F98E" box="[673,762,1574,1598]" italics="true" pageId="12" pageNumber="84">j3-j6, z5</emphasis>
shortened (
<quantity id="4CA99B39FF82FFFEFE80F9AEB1B9F9D2" box="[289,410,1609,1633]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="12" pageNumber="84" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10 – 12 µm</quantity>
), with long ciliate and curved; marginal setae on the shield (
<emphasis id="B925EACEFF82FFFEFDA6F98BB23CF937" box="[519,543,1644,1668]" italics="true" pageId="12" pageNumber="84">z2</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFD90F98BB269F937" box="[561,586,1644,1668]" italics="true" pageId="12" pageNumber="84">z4</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFDFAF98BB250F937" box="[603,627,1644,1668]" italics="true" pageId="12" pageNumber="84">s4</emphasis>
) elongated and ciliate, 
<emphasis id="B925EACEFF82FFFEFE89F968B163F914" box="[296,320,1679,1703]" italics="true" pageId="12" pageNumber="84">z2</emphasis>
(
<quantity id="4CA99B39FF82FFFEFEF1F968B1F2F914" box="[336,465,1679,1704]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="12" pageNumber="84" unit="mm" value="36.0" valueMax="37.0" valueMin="35.0">35 – 37 µm</quantity>
) shorter than 
<emphasis id="B925EACEFF82FFFEFDDBF968B2B0F914" box="[634,659,1679,1703]" italics="true" pageId="12" pageNumber="84">z4</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFD03F968B299F914" box="[674,698,1679,1703]" italics="true" pageId="12" pageNumber="84">s4</emphasis>
(
<quantity id="4CA99B39FF82FFFEFD68F968B0CAF979" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="12" pageNumber="84" unit="mm" value="50.0" valueMax="51.0" valueMin="49.0">49 – 51 µm</quantity>
); 11 pairs on soft tegument smooth or with few long cilia, 
<emphasis id="B925EACEFF82FFFEFEE9F932B17EF95E" box="[328,349,1749,1773]" italics="true" pageId="12" pageNumber="84">J2</emphasis>
the shortest (
<quantity id="4CA99B39FF82FFFEFE58F932B256F95E" box="[505,629,1749,1774]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="12" pageNumber="84" unit="mm" value="26.0" valueMax="27.0" valueMin="25.0">25 – 27 µm</quantity>
), 
<emphasis id="B925EACEFF82FFFEFD2AF932B281F95E" box="[651,674,1749,1773]" italics="true" pageId="12" pageNumber="84">s6</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFD0EF932B2EDF95E" box="[687,718,1749,1773]" italics="true" pageId="12" pageNumber="84">Z3</emphasis>
, 
<emphasis id="B925EACEFF82FFFEFD7AF932B2D9F95E" box="[731,762,1749,1773]" italics="true" pageId="12" pageNumber="84">Z4</emphasis>
(
<quantity id="4CA99B39FF82FFFEFF06F91FB108F8A3" box="[167,299,1784,1809]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="12" pageNumber="84" unit="mm" value="34.0" valueMax="35.0" valueMin="33.0">33 – 35 µm</quantity>
), other setae similar in length (
<quantity id="4CA99B39FF82FFFEFD05F91FB0E6F880" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="12" pageNumber="84" unit="mm" value="48.0" valueMax="51.0" valueMin="45.0">45 – 51 µm</quantity>
). Three pairs of mesonotal ovoid scutellae discernible, posteriomost the largest (
<quantity id="4CA99B39FF82FFFEFDE3F8D9B2ADF8E5" box="[578,654,1854,1879]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="84" unit="mm" value="12.0">12 µm</quantity>
long, 
<quantity id="4CA99B39FF82FFFEFD7EF8D9B0E6F8CA" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="84" unit="mm" value="24.0">24 µm</quantity>
wide). A second pair of tubercular structures are present at level of setae 
<emphasis id="B925EACEFF82FFFEFE52F863B229F82F" box="[499,522,1924,1948]" italics="true" pageId="12" pageNumber="84">s6</emphasis>
which resemble se- tal bases. Pygidial shield small (
<quantity id="4CA99B39FF82FFFEFB14FED0B517FEFC" box="[1205,1332,311,335]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="12" pageNumber="84" unit="mm" value="12.0" valueMax="14.0" valueMin="10.0">10 – 14 µm</quantity>
long by 
<quantity id="4CA99B39FF82FFFEFC9BFEBDB38BFEC1" box="[826,936,346,370]" metricMagnitude="-2" metricUnit="m" metricValue="7.85" metricValueMax="7.9" metricValueMin="7.8" pageId="12" pageNumber="84" unit="mm" value="78.5" valueMax="79.0" valueMin="78.0">78-79 µm</quantity>
wide) and nude, with two pairs of anal sigillae. Soft cuticle with granulate striation.
</paragraph>
<paragraph id="8BEE36DCFF82FFFEFCC1FE4EB4FCFB76" blockId="12.[826,1429,311,1948]" pageId="12" pageNumber="84">
Ventral idiosoma (
<figureCitation id="136A2A59FF82FFFEFBE9FE4EB49AFE72" box="[1096,1209,425,449]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="12" pageNumber="84">Fig. 5D</figureCitation>
) — Tritosternum normally developed, with wide triangular base with slightly serrate margin and three long pilose laciniae. Presternal region with striate soft cuticle. Sternal shield distinguishable, trapezoidal in shape, ca. 
<quantity id="4CA99B39FF82FFFEFCC3FDBFB389FDC3" box="[866,938,600,625]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="84" unit="mm" value="94.0">94 µm</quantity>
long, 
<quantity id="4CA99B39FF82FFFEFC53FDBFB419FDC3" box="[1010,1082,600,625]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="84" unit="mm" value="75.0">75 µm</quantity>
wide at level of 
<emphasis id="B925EACEFF82FFFEFB5DFDBFB53EFDC3" box="[1276,1309,600,624]" italics="true" pageId="12" pageNumber="84">st1</emphasis>
, and 41 at level of setae 
<emphasis id="B925EACEFF82FFFEFC79FD9CB3DAFD20" box="[984,1017,635,659]" italics="true" pageId="12" pageNumber="84">st3</emphasis>
; anterior margin truncate, straight; setae 
<emphasis id="B925EACEFF82FFFEFCDDFD79B3BEFD05" box="[892,925,670,694]" italics="true" pageId="12" pageNumber="84">st1</emphasis>
(
<quantity id="4CA99B39FF82FFFEFC0DFD79B43DFD05" box="[940,1054,670,695]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="12" pageNumber="84" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9 – 10 µm</quantity>
long), 
<emphasis id="B925EACEFF82FFFEFBD0FD79B4B1FD05" box="[1137,1170,670,694]" italics="true" pageId="12" pageNumber="84">st2</emphasis>
(
<quantity id="4CA99B39FF82FFFEFB00FD79B502FD05" box="[1185,1313,670,695]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="12" pageNumber="84" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21 – 22 µm</quantity>
long), 
<emphasis id="B925EACEFF82FFFEFAD5FD79B5B6FD05" box="[1396,1429,670,694]" italics="true" pageId="12" pageNumber="84">st3</emphasis>
(
<quantity id="4CA99B39FF82FFFEFCE2FD25B388FD6A" box="[835,939,705,730]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="12" pageNumber="84" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3 – 4 µm</quantity>
long), and poroids 
<emphasis id="B925EACEFF82FFFEFB3BFD25B49FFD69" box="[1178,1212,706,730]" italics="true" pageId="12" pageNumber="84">iv1</emphasis>
. Anal shield well sclerotized, slightly ornate, widened, subtriangular in shape, wider (
<quantity id="4CA99B39FF82FFFEFBA0FCEFB46FFC93" box="[1025,1100,776,800]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="12" pageNumber="84" unit="mm" value="92.0">92 µm</quantity>
at level of glands 
<emphasis id="B925EACEFF82FFFEFA86FCEFB56CFC93" box="[1319,1359,776,800]" italics="true" pageId="12" pageNumber="84">gv3</emphasis>
) than long (
<quantity id="4CA99B39FF82FFFEFCDAFCCCB3D6FCF0" box="[891,1013,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="12" pageNumber="84" unit="mm" value="39.5" valueMax="40.0" valueMin="39.0">39 – 40 µm</quantity>
); adanal setae (
<quantity id="4CA99B39FF82FFFEFB06FCCCB4CDFCF0" box="[1191,1262,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="84" unit="mm" value="17.0">17 µm</quantity>
long) inserted at level to anal opening, and longer than postanal spine-like seta (
<quantity id="4CA99B39FF82FFFEFC54FC96B410FC3A" box="[1013,1075,881,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="84" unit="mm" value="4.0">4 µm</quantity>
); adanal gland pores (
<emphasis id="B925EACEFF82FFFEFAE4FC96B556FC3A" box="[1349,1397,881,905]" italics="true" pageId="12" pageNumber="84">gv3)</emphasis>
at level of postanal seta with well sclerotized cuticular ring. Opisthogaster with a pair of discernible subcutaneous horn-like structures behind posterior margins of coxae III; a pair of discernible pore-like structures, a pair of subcutaneous clear areas, two pairs of well developed opisthogastric setae anterolateral to shield, 
<emphasis id="B925EACEFF82FFFEFBA0FB81B408FBCD" box="[1025,1067,1126,1150]" italics="true" pageId="12" pageNumber="84">JV1</emphasis>
(
<quantity id="4CA99B39FF82FFFEFB9AFB81B4A6FBCD" box="[1083,1157,1126,1151]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="84" unit="mm" value="14.0">14 µm</quantity>
long) and 
<emphasis id="B925EACEFF82FFFEFAA9FB80B512FBCD" box="[1288,1329,1127,1150]" italics="true" pageId="12" pageNumber="84">JV5</emphasis>
(
<quantity id="4CA99B39FF82FFFEFAE0FB81B343FB12" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="12" pageNumber="84" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19 – 20 µm</quantity>
long), and one pair of poroids 
<emphasis id="B925EACEFF82FFFEFB7EFB6DB4DCFB12" box="[1247,1279,1162,1185]" italics="true" pageId="12" pageNumber="84">ivo</emphasis>
; one pair of poroid posterior to anal shield (
<emphasis id="B925EACEFF82FFFEFB0CFB4AB4ECFB77" box="[1197,1231,1197,1220]" italics="true" pageId="12" pageNumber="84">ivp</emphasis>
).
</paragraph>
<paragraph id="8BEE36DCFF82FFFEFCC1FB3FB35FFA2C" blockId="12.[826,1429,311,1948]" pageId="12" pageNumber="84">Gnathosoma — Gnathotectum as in subsequent instars. Form of corniculus, internal malae as in subsequent instars. Deutosternum similar to those in nymphs. Palpus with normal larval complement of setae; palp-trochanter nude, plap-genua with five setae.</paragraph>
<paragraph id="8BEE36DCFF82FFFEFCC1FA53B571F8C3" blockId="12.[826,1429,311,1948]" pageId="12" pageNumber="84">
Legs — Legs I to III with pretarsi, welldeveloped claws and pulvillus. Legs length: I 
<quantity id="4CA99B39FF82FFFEFAC5FA30B3B3F9A1" metricMagnitude="-1" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.2" metricValueMin="1.19" pageId="12" pageNumber="84" unit="mm" value="119.5" valueMax="120.0" valueMin="119.0">119- 120 µm</quantity>
, II 
<quantity id="4CA99B39FF82FFFEFC1BFA1DB432F9A1" box="[954,1041,1530,1554]" metricMagnitude="-1" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="84" unit="mm" value="116.0">116 µm</quantity>
, III 
<quantity id="4CA99B39FF82FFFEFBE2FA1DB4B9F9A1" box="[1091,1178,1530,1554]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="12" pageNumber="84" unit="mm" value="118.0">118 µm</quantity>
. Larval complement of setae as follows: coxae, 2-2-2; trochanters 3-3-3; femora 8 (1 2/1 2/1 1) – 7 (1 2/1 2/0 1) – 6 (1 2/0 2/0 1); genua 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1); tibiae 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1). Coxa I-III with ventral rim serrate; coxae I with glandular field conspicuous; genua and tibia I, II, III with distal rim serrated lateroventrally. Leg setae generally simple, not markedly differentiated except for fan-like setae on basitarsus II-III. Other legs characteristics as in protonymphal instar.
</paragraph>
<paragraph id="8BEE36DCFF82FFFFFCC1F863B146FE26" blockId="12.[826,1429,311,1948]" lastBlockId="13.[159,762,311,1949]" lastPageId="13" lastPageNumber="85" pageId="12" pageNumber="84">Etymology — The specific name &quot;flagellatus&quot; is adjectival and refers to the conspicuous flagellate tubular structure associate to the peritreme in nymphal instars.</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64FE4BB241FD0A" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="85">
<materialsCitation id="3B393C81FF83FFFFFF64FE4BB14CFDC0" ID-GBIF-Occurrence="3077733301" collectingDate="2014-01-25" collectionCode="PSU" collectorName="Gobardhan Sahoo" country="India" county="Long" latitude="15.505167" location="Studied" longLatPrecision="1" longitude="73.87875" municipality="Lat" pageId="13" pageNumber="85" specimenCount="2" typeStatus="holotype">
<location id="8E8E6007FF83FFFFFF64FE4BB102FE77" LSID="urn:lsid:plazi:treatment:03F887CAFF8DFFFFFC1BFD2CB536F82F:8E8E6007FF83FFFFFF64FE4BB102FE77" box="[197,289,428,452]" country="India" county="Long" latitude="15.505167" longLatPrecision="1" longitude="73.87875" municipality="Lat" name="Studied" pageId="13" pageNumber="85">Studied</location>
material — 
<typeStatus id="54EA887EFF83FFFFFE0CFE4BB23CFE77" box="[429,543,428,452]" pageId="13" pageNumber="85" type="holotype">Holotype</typeStatus>
: adult female, 
<location id="8E8E6007FF83FFFFFD68FE4BB2DAFE77" LSID="urn:lsid:plazi:treatment:03F887CAFF8DFFFFFC1BFD2CB536F82F:8E8E6007FF83FFFFFD68FE4BB2DAFE77" box="[713,761,428,452]" country="India" county="Long" latitude="15.505167" longLatPrecision="1" longitude="73.87875" municipality="Lat" name="Goa" pageId="13" pageNumber="85">Goa</location>
(
<collectingCountry id="F346764CFF83FFFFFF06FE28B0CBFE54" box="[167,232,463,487]" name="India" pageId="13" pageNumber="85">India</collectingCountry>
), 
<location id="8E8E6007FF83FFFFFF5EFE28B1B4FE54" LSID="urn:lsid:plazi:treatment:03F887CAFF8DFFFFFC1BFD2CB536F82F:8E8E6007FF83FFFFFF5EFE28B1B4FE54" box="[255,407,463,487]" country="India" county="Long" latitude="15.505167" longLatPrecision="1" longitude="73.87875" municipality="Lat" name="Divar Island" pageId="13" pageNumber="85">Divar Island</location>
, 
<location id="8E8E6007FF83FFFFFE07FE28B232FE54" LSID="urn:lsid:plazi:treatment:03F887CAFF8DFFFFFC1BFD2CB536F82F:8E8E6007FF83FFFFFE07FE28B232FE54" box="[422,529,463,487]" country="India" county="Long" latitude="15.505167" longLatPrecision="1" longitude="73.87875" municipality="Lat" name="Mandovi" pageId="13" pageNumber="85">Mandovi</location>
estuary, 
<collectingMunicipality id="6B8AACA6FF83FFFFFD21FE28B288FE54" box="[640,683,463,487]" pageId="13" pageNumber="85">Lat</collectingMunicipality>
/ 
<collectingCounty id="628F4E50FF83FFFFFD15FE28B2D5FE54" box="[692,758,463,487]" pageId="13" pageNumber="85">Long</collectingCounty>
: 
<geoCoordinate id="EE65501BFF83FFFFFF3EFE15B116FDB9" box="[159,309,498,522]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="13" pageNumber="85" precision="1" seconds="18.6" value="15.505167">15°30’18.6&quot;N</geoCoordinate>
<geoCoordinate id="EE65501BFF83FFFFFE9FFE15B1F2FDB9" box="[318,465,498,522]" degrees="73" direction="east" minutes="52" orientation="longitude" pageId="13" pageNumber="85" precision="1" seconds="43.5" value="73.87875">73°52’43.5&quot;E</geoCoordinate>
, collected on 
<date id="FFEF101CFF83FFFFFDD8FE15B0F4FD9E" pageId="13" pageNumber="85" value="2014-01-25">
<collectingDate id="EFABE9F4FF83FFFFFDD8FE15B0F4FD9E" pageId="13" pageNumber="85" value="2014-01-25">25 January 2014</collectingDate>
</date>
, leg. 
<collectorName id="26A4530AFF83FFFFFE86FDF2B221FD9E" box="[295,514,533,557]" pageId="13" pageNumber="85">Gobardhan Sahoo</collectorName>
; water temperature, pH, and salinity of the site was 29 °C, 7.81, 17 
<collectionCode id="ED40AE19FF83FFFFFF3EFDBCB0F1FDC0" box="[159,210,603,627]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="13" pageNumber="85" type="Museum">PSU</collectionCode>
respectively.
</materialsCitation>
<materialsCitation id="3B393C81FF83FFFFFE24FDBBB27EFD0A" ID-GBIF-Occurrence="3077733302" collectingDate="2014-01-25" collectionCode="PSU" collectorName="Gobardhan Sahoo" country="India" county="Long" latitude="15.505167" location="Studied" longLatPrecision="1" longitude="73.87875" municipality="Lat" pageId="13" pageNumber="85" specimenCount="2" typeStatus="paratype">
<typeStatus id="54EA887EFF83FFFFFE24FDBBB1DEFDC7" box="[389,509,604,628]" pageId="13" pageNumber="85" type="paratype">Paratypes</typeStatus>
: three females, four males, three dutonymphs, two protonymphs and one larva, with same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64FD37B23EFCC7" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="85">
Remarks — Currently 13 species belong to genus 
<taxonomicName id="4C514D5FFF83FFFFFF3EFD14B103FCB8" authorityName="Berlese" authorityYear="1914" box="[159,288,755,779]" class="Arachnida" family="Uropodidae" genus="Eutrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B925EACEFF83FFFFFF3EFD14B103FCB8" box="[159,288,755,779]" italics="true" pageId="13" pageNumber="85">Eutrachytes</emphasis>
</taxonomicName>
. The new species found in Goa, 
<collectingCountry id="F346764CFF83FFFFFD79FD13B0E5FC9D" name="India" pageId="13" pageNumber="85">India</collectingCountry>
, has a pair of concave depressions at the posterior dorsal region, which was not observed earlier within the family 
<taxonomicName id="4C514D5FFF83FFFFFED0FCBBB239FCC7" box="[369,538,860,884]" family="Eutracyhtidae" pageId="13" pageNumber="85" rank="family">Eutracyhtidae</taxonomicName>
.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64FC6CB245FB9C" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="85">
The presence of the complete ontogenetic instars of this species allows us to track back to the larval instar the structural development of idiosomal chaetotaxy and shielding and to speculate on other elements such as the 
<emphasis id="B925EACEFF83FFFFFE37FBF0B27CFB9C" box="[406,607,1047,1071]" italics="true" pageId="13" pageNumber="85">flagellate structure</emphasis>
.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64FBA0B108F9FA" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="85">
In the larva and nymphal instars the complement of dorsal and ventral setae is readily recognized and denotable. However, rather than irregularly or asymmetrically multiple, adults display a dorsal and ventral hypertrichy which appears to be a duplicative neotrichy derived secondarily. This hypertrichy of paired setae seems to be in the form of setal duplications, such that symmetrically duplicated pairs can be recognized (in figure 1, outlined duplication fields are speculative). Similar dorsal neotrichy occurs in other Gamasina mites (
<bibRefCitation id="EFC04B2DFF83FFFFFF06FA2FB22EFA53" author="Moraza M. L. &amp; Lindquist E. E." box="[167,525,1480,1504]" pageId="13" pageNumber="85" pagination="301 - 351" refId="ref10646" refString="Moraza M. L., Lindquist E. E. 2015 - Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoidea) - Zootaxa 3931 (3): 301 - 351 doi: 10.11646 / zootaxa. 3931.3.1" type="journal article" year="2015">Moraza and Lindquist, 2015</bibRefCitation>
). Other unpaired, asymmetrical setae (some of them present in the deutonymphal instar) complete the hypertrichy in this species.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64F987B12AF8F9" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="85">
Opisthogastric setae 
<emphasis id="B925EACEFF83FFFFFE1CF987B1C5F9CB" box="[445,486,1632,1656]" italics="true" pageId="13" pageNumber="85">JV2</emphasis>
and 
<emphasis id="B925EACEFF83FFFFFD84F987B27BF9CB" box="[549,600,1632,1656]" italics="true" pageId="13" pageNumber="85">ZV2</emphasis>
are absent in the larval and subsequent instars of this and other species of the family (see 
<bibRefCitation id="EFC04B2DFF83FFFFFE6AF940B240F90D" author="Krantz G. W." box="[459,611,1702,1726]" pageId="13" pageNumber="85" pagination="62 - 70" refId="ref10281" refString="Krantz G. W. 1969 - The Mites of Quintana Roo. I. A new species of Eutrachytes from the Yucatan Peninsula, with observations on the classification of the genus - Ann. Entomol. Soc. Am., 62 (1): 62 - 70." type="journal article" year="1969">Krantz, 1969</bibRefCitation>
). The pair of larval opisthogastric pore-like structures (
<figureCitation id="136A2A59FF83FFFFFD32F92EB2D2F952" box="[659,753,1737,1761]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="13" pageNumber="85">Fig. 5D</figureCitation>
) remains in the protonymph lateral to 
<emphasis id="B925EACEFF83FFFFFDEEF90AB25AF8B6" box="[591,633,1773,1797]" italics="true" pageId="13" pageNumber="85">JV1</emphasis>
, and a pair of clear areas is distinguishable between coxae IV (
<figureCitation id="136A2A59FF83FFFFFF06F8D4B0DFF8F9" box="[167,252,1842,1867]" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="13" pageNumber="85">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFF64F885B51BFC89" blockId="13.[159,762,311,1949]" lastBlockId="13.[826,1429,311,826]" pageId="13" pageNumber="85">
In the larva, at the lateral margin of podonotal shield between setae 
<emphasis id="B925EACEFF83FFFFFE06F862B1E3F82E" box="[423,448,1925,1949]" italics="true" pageId="13" pageNumber="85">z4</emphasis>
and 
<emphasis id="B925EACEFF83FFFFFDA4F862B23EF82E" box="[517,541,1925,1949]" italics="true" pageId="13" pageNumber="85">s4</emphasis>
, a pair of conical tubercles with a pore-like structure at the base are present; a second pair of tubercular structures are present on soft tegument at level of setae 
<emphasis id="B925EACEFF83FFFFFA9FFE9AB575FE26" box="[1342,1366,381,405]" italics="true" pageId="13" pageNumber="85">s6</emphasis>
(
<figureCitation id="136A2A59FF83FFFFFAC9FE9AB37EFE0B" captionStart="FIGURE" captionStartId="10.[159,171,1830,1849]" captionTargetBox="[227,1354,365,1790]" captionTargetId="figure-4@10.[227,1361,364,1798]" captionTargetPageId="10" captionText="FIGURE 5: Eutrachytes flagellatus n. sp.: A – Protonymph, idiosoma, dorsal view; B – Protonymph, idiosoma, ventral view; C – Larva, idiosoma, dorsal view; D – Larva, idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4640194" httpUri="https://zenodo.org/record/4640194/files/figure.png" pageId="13" pageNumber="85">Fig. 5C</figureCitation>
). In the protonymph, dorsal shield is reduced leaving dorsal setae 
<emphasis id="B925EACEFF83FFFFFB84FE24B419FE68" box="[1061,1082,451,475]" italics="true" pageId="13" pageNumber="85">j1</emphasis>
, 
<emphasis id="B925EACEFF83FFFFFBE7FE24B47CFE68" box="[1094,1119,451,475]" italics="true" pageId="13" pageNumber="85">z2</emphasis>
, 
<emphasis id="B925EACEFF83FFFFFBCAFE24B4A7FE68" box="[1131,1156,451,475]" italics="true" pageId="13" pageNumber="85">z4</emphasis>
and 
<emphasis id="B925EACEFF83FFFFFB1DFE24B4F7FE68" box="[1212,1236,451,475]" italics="true" pageId="13" pageNumber="85">s4</emphasis>
on the soft tegument and a humeral dorsolateral protuberance between setae 
<emphasis id="B925EACEFF83FFFFFC6FFDEEB3C5FD92" box="[974,998,521,545]" italics="true" pageId="13" pageNumber="85">s4</emphasis>
and 
<emphasis id="B925EACEFF83FFFFFB85FDEDB41FFD92" box="[1060,1084,522,545]" italics="true" pageId="13" pageNumber="85">r5</emphasis>
related to the peritreme; the flagellate structure is well developed. In the deutonymph, the dorsolateral protuberances have been reduced to small nodes in the soft unsclerotized submarginal region of idiosoma, nodes that remain related to the peritreme tips in the adult. Out of the two pairs of protuberances described, whether either pair of protuberances could be the possible origin of the peritremal flagellum is uncertain.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFCC1FC6CB4ECFB6C" blockId="13.[826,1429,907,1247]" pageId="13" pageNumber="85">
Dorsal shield marginal concavities are surrounded by opisthonotal setae 
<emphasis id="B925EACEFF83FFFFFB05FC49B4E3FC75" box="[1188,1216,942,966]" italics="true" pageId="13" pageNumber="85">S3</emphasis>
, 
<emphasis id="B925EACEFF83FFFFFB6CFC49B536FC75" box="[1229,1301,942,966]" italics="true" pageId="13" pageNumber="85">Z3, S4</emphasis>
and a conspicuous non glandular poroid at their bases (
<figureCitation id="136A2A59FF83FFFFFAC9FC36B37EFBBE" captionStart="FIGURE" captionStartId="4.[159,171,1830,1849]" captionTargetBox="[232,1361,369,1746]" captionTargetId="figure-4@4.[227,1361,364,1798]" captionTargetPageId="4" captionText="FIGURE 1: Eutrachytes flagellatus n. sp., adult female: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Peritrematal shield with detail of distal tip of the peritreme; D – Detail of peritreme; 5. Male, detail of sternitigenital region." figureDoi="http://doi.org/10.5281/zenodo.4640184" httpUri="https://zenodo.org/record/4640184/files/figure.png" pageId="13" pageNumber="85">Fig. 1A</figureCitation>
). In the deutonymphal instar, three pairs of glandular pores and a pair of poroids are located at the same level than such concavities in the adult (
<figureCitation id="136A2A59FF83FFFFFAC9FBDCB343FBC5" captionStart="FIGURE" captionStartId="9.[159,171,1830,1849]" captionTargetBox="[227,1359,364,1775]" captionTargetId="figure-6@9.[227,1361,364,1798]" captionTargetPageId="9" captionText="FIGURE 4: Eutrachytes flagellatus n. sp., Deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of notated dorsal setae; D – Detail of peritreme and flagellate tracheal structure." figureDoi="http://doi.org/10.5281/zenodo.4640192" httpUri="https://zenodo.org/record/4640192/files/figure.png" pageId="13" pageNumber="85">Fig. 4A</figureCitation>
). The transition of these glandular poroids associated with the concavities may be complete in the adult instar, and if this is the case, the function of such structures would be clarified.
</paragraph>
<paragraph id="8BEE36DCFF83FFFFFCC1FAD7B536F82F" blockId="13.[826,1429,1328,1948]" pageId="13" pageNumber="85">
Regarding the flagellated structure being present only in nymphal instars, its structural relationship with the peritreme suggests its possible role in the breathing process, such as providing an air duct to the stigmata when the mite is immersed. However, if that were true, its absence in adults would be difficult to explain as they share the habitat. It is possible that immature instars need this &quot;snorkel&quot; because they live immersed in water while adults do not immerse. Other Gamasina mites which live immersed in fluid have similar structures: 
<taxonomicName id="4C514D5FFF83FFFFFC60F954B429F979" authorityName="Oudemans" authorityYear="1904" box="[961,1034,1715,1738]" class="Arachnida" family="Varroidae" genus="Varroa" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B925EACEFF83FFFFFC60F954B429F979" box="[961,1034,1715,1738]" italics="true" pageId="13" pageNumber="85">Varroa</emphasis>
</taxonomicName>
(immersed inside a cell in a beehive) has an emergent peritreme (Bruce 
<emphasis id="B925EACEFF83FFFFFAAEF931B560F95E" box="[1295,1347,1749,1773]" italics="true" pageId="13" pageNumber="85">et al.</emphasis>
, 1997), and the semi-aquatic 
<taxonomicName id="4C514D5FFF83FFFFFBE2F91FB52FF8A3" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1091,1292,1784,1808]" class="Arachnida" family="Ascidae" genus="Platyseius" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="85" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B925EACEFF83FFFFFBE2F91FB52FF8A3" box="[1091,1292,1784,1808]" italics="true" pageId="13" pageNumber="85">Platyseius italicus</emphasis>
</taxonomicName>
has an expanded peritreme with a plastron (Hilton, 1971). In future, more exhaustive anatomical and physiological study is required to gain some deep insights into the nature and function of this structure.
</paragraph>
</subSubSection>
</treatment>
</document>