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<document id="DE730B567D696C49F674F4920F4D0101" ID-CLB-Dataset="57602" ID-DOI="10.5852/ejt.2023.867.2109" ID-GBIF-Dataset="1d2570da-e150-4d75-94d8-bfc5813062bc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7891021" ID-ZooBank="8DEF295C-A8B1-4A6B-B873-B30949F64E07" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1683117081100" checkinUser="felipe" docAuthor="Iniesta, Luiz Felipe Moretti, Bouzan, Rodrigo Salvador &amp; Brescovit, Antonio Domingos" docDate="2023" docId="03F887BA1324B1794EADFDD1FBC951E8" docLanguage="en" docName="ejt-867_1-312.pdf.imf" docOrigin="European Journal of Taxonomy 867 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.867.2109" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudonannolene anapophysis Fontanetti 1996" docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFC1FFC21333B1614F56FFECFF9A553E" masterDocTitle="A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)" masterLastPageNumber="312" masterPageNumber="1" pageNumber="24" updateTime="1699515670513" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="62024205DEDF9D325C18D419A77CBB22">A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)</mods:title>
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<mods:namePart id="278ADC7CC7EF6A4D29CB1864A3EC0E8D">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:affiliation id="9819A6BC4A419647EE97D2F655E924C8">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="60BC34C9B05A405A857F0164DEA3980B">Bouzan, Rodrigo Salvador</mods:namePart>
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<mods:affiliation id="98E787B2EE39FA7A51E9711D9E787E03">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="27D78A08AA277F23B491BA048429E554">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="44BC9A1BB6F9095D8974FBEFA27E4B8C">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<taxonomicName id="4C514D2F1324B1764EADFDD1FBA35769" ID-CoL="6cc71d98-d5ce-47c6-9bf5-798a6b51bd32" authority="Fontanetti, 1996" authorityName="Fontanetti" authorityYear="1996" box="[507,1081,573,600]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anapophysis">
<emphasis id="B925EABE1324B1764EADFDD1FCEB5766" bold="true" box="[507,881,573,600]" italics="true" pageId="23" pageNumber="24">Pseudonannolene anapophysis</emphasis>
<bibRefCitation id="EFC04B5D1324B1764C2EFDD1FBA35769" author="Fontanetti C. S." box="[888,1081,573,599]" pageId="23" pageNumber="24" pagination="427 - 433" refId="ref101873" refString="Fontanetti C. S. 1996. Description of three cave diplopods of Pseudonannolene Silvestri (Diplopoda, Pseudonannolenida, Pseudonannolenidae). Revista Brasileira de Zoologia 13 (2): 427 - 433. https: // doi. org / 10.1590 / S 0101 - 81751996000200013" type="journal article" year="1996">Fontanetti, 1996</bibRefCitation>
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<subSubSection id="C34B65271324B1764E3EFD8DFB575745" box="[360,1229,609,636]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BEE36AC1324B1764E3EFD8DFB575745" blockId="23.[360,1229,573,636]" box="[360,1229,609,636]" pageId="23" pageNumber="24">
<figureCitation id="136A2A291324B1764E3EFD8DFE745745" box="[360,494,609,635]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="162.[189,232,764,790]" captionStartId-1="163.[189,232,1675,1701]" captionTargetBox-0="[189,1389,265,717]" captionTargetBox-1="[189,1398,270,1627]" captionTargetId-0="figure-13@162.[189,1398,265,717]" captionTargetId-1="figure-11@163.[189,1398,270,1627]" captionTargetPageId-0="162" captionTargetPageId-1="163" captionText-0="Fig. 48. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209), in lateral view. A. Anterior region. B. Posterior region. Scale bars = 1 mm." captionText-1="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7891143" figureDoi-1="http://doi.org/10.5281/zenodo.7891145" httpUri-0="https://zenodo.org/record/7891143/files/figure.png" httpUri-1="https://zenodo.org/record/7891145/files/figure.png" pageId="23" pageNumber="24">Figs 48–49</figureCitation>
,
<figureCitation id="136A2A291324B1764EACFD8DFDA65745" box="[506,572,609,635]" captionStart="Fig" captionStartId="266.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1743]" captionTargetId="figure-14@266.[250,1337,265,1743]" captionTargetPageId="266" captionText="Fig. 163. Stereoscopic images of antennae of species ofPseudonannolene Silvestri, 1895.A. P. albiventris Schubart, 1952. B. P. anapophysis Fontanetti, 1996. C. P. alata sp. nov. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. F. P. erikae Iniesta &amp; Ferreira, 2014. G. P. fontanettiae Iniesta &amp; Ferreira, 2014. H. P. halophila Schubart, 1949. I. P. inops Brölemann, 1929. J. P. leopoldoi Iniesta &amp; Ferreira, 2014. K. P. longicornis (Porat, 1888). L. P. magna Udulutsch &amp; Pietrobon, 2003. M. P. maritima Schubart, 1949. N. P. mesai Fontanetti, 2000. O. P. microzoporus Mauriès, 1987. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891405" httpUri="https://zenodo.org/record/7891405/files/figure.png" pageId="23" pageNumber="24">163B</figureCitation>
,
<figureCitation id="136A2A291324B1764D1EFD8DFD105742" box="[584,650,609,636]" captionStart="Fig" captionStartId="268.[189,232,1741,1767]" captionTargetBox="[348,1239,265,1699]" captionTargetId="figure-13@268.[348,1239,265,1699]" captionTargetPageId="268" captionText="Fig. 165. Stereoscopic images of midbody legs of species of Pseudonannolene Silvestri, 1895. A. P. ambuatinga Iniesta &amp; Ferreira, 2013. B. P. anapophysis Fontanetti, 1996. C. P. alata sp. nov. D. P. bovei Silvestri, 1895. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. curtipes Schubart, 1960. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M, P. leopoldoi Iniesta &amp; Ferreira, 2014. N. P. leucomelas Schubart, 1947. O. P. longicornis (Porat, 1888). P. P. lundi Iniesta &amp; Ferreira, 2015. Q. P. magna Udulutsch &amp; Pietrobon, 2003. R. P. maritima Schubart, 1949. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891411" httpUri="https://zenodo.org/record/7891411/files/figure.png" pageId="23" pageNumber="24">165B</figureCitation>
,
<figureCitation id="136A2A291324B1764DC0FD8DFD435745" box="[662,729,609,635]" captionStart="Fig" captionStartId="270.[189,232,1892,1918]" captionTargetBox="[213,1374,265,1851]" captionTargetId="figure-14@270.[213,1374,265,1851]" captionTargetPageId="270" captionText="Fig. 167. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. albiventris Schubart, 1952. B. P. ambuatinga Iniesta &amp; Ferreira, 2013. C. P. anapophysis Fontanetti, 1996. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891417" httpUri="https://zenodo.org/record/7891417/files/figure.png" pageId="23" pageNumber="24">167C</figureCitation>
,
<figureCitation id="136A2A291324B1764DB2FD8DFCB35745" box="[740,809,609,635]" captionStart="Fig" captionStartId="280.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1739]" captionTargetId="figure-14@280.[250,1337,265,1739]" captionTargetPageId="280" captionText="Fig. 177. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. albiventris Schubart, 1952. B. P. alegrensis Silvestri, 1897. C. P. ambuatinga Iniesta &amp; Ferreira, 2013. D. P. anapophysis Fontanetti, 1996. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. erikae Iniesta &amp; Ferreira, 2014. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M. P. leucocephalus Schubart, 1944. N. P. longicornis (Porat, 1888). O. P. lundi Iniesta &amp; Ferreira, 2015. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891439" httpUri="https://zenodo.org/record/7891439/files/figure.png" pageId="23" pageNumber="24">177D</figureCitation>
,
<figureCitation id="136A2A291324B1764C62FD8DFCF95745" box="[820,867,609,635]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="23" pageNumber="24">180</figureCitation>
; Supp. file 4:
<figureCitation id="136A2A291324B1764B50FD8DFB1B5745" box="[1030,1153,609,635]" captionStart="Fig" captionStartId="309.[242,285,707,733]" captionText="Fig. 204. Characters 29–31 and their character states. Coxae of first leg-pair of males. A. Pseudonannolene caatinga Iniesta &amp; Ferreira, 2014 (IBSP 1389). B. P. leucocephalus Schubart, 1944 (MZSP 1060). C. Epinannolene sp. (ICN). D. P. halophila Schubart, 1949 (IBSP 3297). E. P. anapophysis Fontanetti, 1996 (IBSP 5209). Scale bars: A, D–E = 0.5 mm; B–C = 0.2 mm." pageId="23" pageNumber="24">Figs 204E</figureCitation>
,
<figureCitation id="136A2A291324B1764BDAFD8DFB575745" box="[1164,1229,609,635]" captionStart="Fig" captionStartId="309.[242,285,1014,1040]" captionText="Fig. 206. Characters 35–37 and their character states. Prefemur of first leg-pair of males. A. Pseudonannolene anapophysis Fontanetti, 1996 (IBSP 5209). B. Holopodostreptus braueri Carl, 1918 (MNRJ). C. Choctella hubrichti Hoffman, 1965 (USNM). D. P. magna Udulutsch &amp; Pietrobon, 2003 (MZSP 941). Scale bars: A = 0.5 mm; B = 0.1 mm; D = 0.2 mm. Image C not to scale." pageId="23" pageNumber="24">206A</figureCitation>
</paragraph>
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<taxonomicName id="4C514D2F1324B1764FEBFD4DFCB45785" ID-CoL="6cc71d98-d5ce-47c6-9bf5-798a6b51bd32" authority="Fontanetti, 1996: 428" authorityName="Fontanetti" authorityPageNumber="428" authorityYear="1996" box="[189,814,672,699]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anapophysis">
<emphasis id="B925EABE1324B1764FEBFD4DFDB25785" box="[189,552,673,699]" italics="true" pageId="23" pageNumber="24">Pseudonannolene anapophysis</emphasis>
<bibRefCitation id="EFC04B5D1324B1764D79FD4DFCB45785" author="Fontanetti C. S." box="[559,814,672,699]" pageId="23" pageNumber="24" pagination="427 - 433" refId="ref101873" refString="Fontanetti C. S. 1996. Description of three cave diplopods of Pseudonannolene Silvestri (Diplopoda, Pseudonannolenida, Pseudonannolenidae). Revista Brasileira de Zoologia 13 (2): 427 - 433. https: // doi. org / 10.1590 / S 0101 - 81751996000200013" type="journal article" year="1996">Fontanetti, 1996: 428</bibRefCitation>
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, figs 1–4.
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<taxonomicName id="4C514D2F1324B1764FEBFD0CFDB657C4" ID-CoL="6cc71d98-d5ce-47c6-9bf5-798a6b51bd32" authorityName="Fontanetti" authorityYear="1996" box="[189,556,736,762]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anapophysis">
<emphasis id="B925EABE1324B1764FEBFD0CFDB657C4" box="[189,556,736,762]" italics="true" pageId="23" pageNumber="24">Pseudonannolene anapophysis</emphasis>
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–
<treatmentCitation id="0AF010BD1324B1764D07FD0CFC2857C5" author="Iniesta L. F. &amp; Ferreira R. L." box="[593,946,736,763]" page="92" pageId="23" pageNumber="24" year="2013">
<bibRefCitation id="EFC04B5D1324B1764D07FD0CFC2857C5" author="Iniesta L. F. &amp; Ferreira R. L." box="[593,946,736,763]" pageId="23" pageNumber="24" pagination="85 - 95" refId="ref103686" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 a. The first troglobitic Pseudonannolene from Brazilian iron ore caves (Spirostreptida: Pseudonannolenidae). Zootaxa 3669 (1): 85 - 95. https: // doi. org / 10.11646 / zootaxa. 3669.1.9" type="journal article" year="2013">Iniesta &amp; Ferreira 2013a: 92</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF010BD1324B1764C94FD0CFBD557C4" author="Iniesta L. F. &amp; Ferreira R. L." box="[962,1103,736,762]" page="366" pageId="23" pageNumber="24" year="2013">
<bibRefCitation id="EFC04B5D1324B1764C94FD0CFBD557C4" author="Iniesta L. F. &amp; Ferreira R. L." box="[962,1103,736,762]" pageId="23" pageNumber="24" pagination="357 - 369" refId="ref103738" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 b. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species. Zootaxa 3702 (4): 357 - 369. https: // doi. org / 10.11646 / zootaxa. 3702.4.3" type="journal article" year="2013">2013b: 366</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF010BD1324B1764B09FD0DFB4157C5" author="Iniesta L. F. &amp; Ferreira R. L." box="[1119,1243,737,763]" page="79" pageId="23" pageNumber="24" year="2013">
<bibRefCitation id="EFC04B5D1324B1764B09FD0DFB4157C5" author="Iniesta L. F. &amp; Ferreira R. L." box="[1119,1243,737,763]" pageId="23" pageNumber="24" pagination="75 - 80" refId="ref103802" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 c. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian iron ore caves (Spirostreptida: Pseudonannolenidae). Zootaxa 3716 (1): 75 - 80. https: // doi. org / 10.11646 / zootaxa. 3716.1.6" type="journal article" year="2013">2013c: 79</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF010BD1324B1764A45FD0CFE575622" author="Gallo J. S. &amp; Bichuette M. E." page="47" pageId="23" pageNumber="24" year="2019">
<bibRefCitation id="EFC04B5D1324B1764A45FD0CFE575622" author="Gallo J. S. &amp; Bichuette M. E." pageId="23" pageNumber="24" pagination="41 - 55" refId="ref102230" refString="Gallo J. S. &amp; Bichuette M. E. 2019. O que mudou na distribuicao dos diplopodes Pseudonannolene Silvestri, 1895 nas cavernas do Brasil 18 anos apos a sinopse de Trajano e colaboradores (2000)?. Espeleo-Tema 29 (1): 41 - 55. https: // doi. org / 10.3897 / aca. 1. e 30225" type="journal article" year="2019">Gallo &amp; Bichuette 2019: 47</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBFCC8FB8E5600" blockId="23.[189,1399,736,830]" box="[189,1044,803,830]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB4111821324B1764FEBFCC8FB8E5600" box="[189,1044,803,830]" pageId="23" pageNumber="24">
<taxonomicName id="4C514D2F1324B1764FEBFCC8FE2C5600" box="[189,438,804,830]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B925EABE1324B1764FEBFCC8FE145600" box="[189,398,804,830]" italics="true" pageId="23" pageNumber="24">Pseudonannolene</emphasis>
sp.
</taxonomicName>
“Igatu”–
<treatmentCitation id="0AF010BD1324B1764D7CFCC8FCF95600" author="Gallo J. S. &amp; Bichuette M. E." box="[554,867,804,830]" page="6" pageId="23" pageNumber="24" year="2017">
<bibRefCitation id="EFC04B5D1324B1764D7CFCC8FCF95600" author="Gallo J. S. &amp; Bichuette M. E." box="[554,867,804,830]" pageId="23" pageNumber="24" pagination="273 - 289" refId="ref102173" refString="Gallo J. S. &amp; Bichuette M. E. 2017. Is there correlation between photophobia and troglomorphism in Neotropical cave millipedes (Spirostreptida, Pseudonannolenidae)? Zoomorphology 137 (2): 273 - 289. https: // doi. org / 10.1007 / s 00435 - 017 - 0389 - 0" type="journal article" year="2017">Gallo &amp; Bichuette 2017: 6</bibRefCitation>
</treatmentCitation>
, figs 4f, 5f, 9h.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B65271324B1764FEBFC8FFAED56D0" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BEE36AC1324B1764FEBFC8FFEAC5643" blockId="23.[189,310,867,893]" box="[189,310,867,893]" pageId="23" pageNumber="24">
<heading id="D0A681C01324B1764FEBFC8FFEAC5643" bold="true" box="[189,310,867,893]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B925EABE1324B1764FEBFC8FFEAC5643" bold="true" box="[189,310,867,893]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBFC7CFAED56D0" blockId="23.[189,1399,912,1006]" pageId="23" pageNumber="24">
Males of
<taxonomicName id="4C514D2F1324B1764E65FC7DFE7D5694" authorityName="Fontanetti" authorityYear="1996" box="[307,487,912,938]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anapophysis">
<emphasis id="B925EABE1324B1764E65FC7DFE7D5694" box="[307,487,912,938]" italics="true" pageId="23" pageNumber="24">P. anapophysis</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C514D2F1324B1764D85FC7DFCAE5694" authorityName="* Silvestri" authorityYear="1895" box="[723,820,912,938]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bovei">
<emphasis id="B925EABE1324B1764D85FC7DFCAE5694" box="[723,820,912,938]" italics="true" pageId="23" pageNumber="24">P. bovei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D2F1324B1764C10FC7DFC545694" authorityName="Brolemann" authorityYear="1929" box="[838,974,912,938]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="caulleryi">
<emphasis id="B925EABE1324B1764C10FC7DFC545694" box="[838,974,912,938]" italics="true" pageId="23" pageNumber="24">P. caulleryi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D2F1324B1764CB7FC7DFBD85694" authorityName="Brolemann" authorityYear="1929" baseAuthorityName="Brolemann" baseAuthorityYear="1929" box="[993,1090,913,938]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="B925EABE1324B1764CB7FC7DFBD85694" box="[993,1090,913,938]" italics="true" pageId="23" pageNumber="24">P. inops</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C514D2F1324B1764BDFFC7DFEFE56F2" authority="Iniesta &amp; Ferreira, 2014" authorityName="Iniesta &amp; Ferreira" authorityYear="2014" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="xavieri">
<emphasis id="B925EABE1324B1764BDFFC7DFB615694" box="[1161,1275,913,938]" italics="true" pageId="23" pageNumber="24">P. xavieri</emphasis>
<bibRefCitation id="EFC04B5D1324B1764A50FC7CFEFE56F2" author="Iniesta L. F. &amp; Ferreira R. L." pageId="23" pageNumber="24" pagination="361 - 397" refId="ref103858" refString="Iniesta L. F. M &amp; Ferreira R. L. 2014. New species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves with comments on the potential distribution of the genus in South America (Spirostreptida: Pseudonannolenidae). Zootaxa 3846 (3): 361 - 397. https: // doi. org / 10.11646 / zootaxa. 3846.3.3" type="journal article" year="2014">Iniesta &amp; Ferreira, 2014</bibRefCitation>
</taxonomicName>
by having solenomere with subtriangular ectal process directed horizontally (
<figureCitation id="136A2A291324B1764BADFC5EFAF356F2" box="[1275,1385,946,972]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="23" pageNumber="24">Fig. 49D</figureCitation>
), but can be easily distinguished by the absence of a prefemoral process on the first leg-pair (
<figureCitation id="136A2A291324B1764B8DFC38FAF056D0" box="[1243,1386,980,1006]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="23" pageNumber="24">Fig. 49A–B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271324B1764FEBFBF8FBE15143" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BEE36AC1324B1764FEBFBF8FEDE5110" blockId="23.[189,324,1044,1070]" box="[189,324,1044,1070]" pageId="23" pageNumber="24">
<heading id="D0A681C01324B1764FEBFBF8FEDE5110" bold="true" box="[189,324,1044,1070]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B925EABE1324B1764FEBFBF8FEDE5110" bold="true" box="[189,324,1044,1070]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBFBACFBE15143" blockId="23.[189,1398,1088,1149]" pageId="23" pageNumber="24">
Named after the Greek prefix ‘
<emphasis id="B925EABE1324B1764D74FBADFDA55164" box="[546,575,1089,1114]" italics="true" pageId="23" pageNumber="24">an</emphasis>
-’ = ‘without’, and ‘
<emphasis id="B925EABE1324B1764C72FBACFC035164" box="[804,921,1088,1114]" italics="true" pageId="23" pageNumber="24">apophysis</emphasis>
’. Unspecified in the original description, but likely to be related to the absence of a prefemoral process on the first leg-pair.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271324B1764FEBFB4EFD33536A" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BEE36AC1324B1764FEBFB4EFE305182" blockId="23.[189,426,1186,1212]" box="[189,426,1186,1212]" pageId="23" pageNumber="24">
<heading id="D0A681C01324B1764FEBFB4EFE305182" bold="true" box="[189,426,1186,1212]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B925EABE1324B1764FEBFB4EFE305182" bold="true" box="[189,426,1186,1212]" pageId="23" pageNumber="24">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1324B1764FBCFB23FEDE5012" blockId="23.[189,1398,1231,1324]" pageId="23" pageNumber="24">
<materialsCitation id="3B393CF11324B1764FBCFB23FEDE5012" ID-GBIF-Occurrence="4101309688" collectingDate="1987-01" collectionCode="MZSP" collectorName="F. Chaimowicz" country="Brazil" latitude="-12.540361" location="cave Lapao" longLatPrecision="1" longitude="-41.40271" municipality="Lencois" pageId="23" pageNumber="24" specimenCode="MZSP 940" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="B925EABE1324B1764FBCFB23FEC651D7" bold="true" box="[234,348,1231,1257]" pageId="23" pageNumber="24">
<typeStatus id="54EA880E1324B1764FBCFB23FEC651D7" box="[234,348,1231,1257]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F346763C1324B1764FEBFB1DFEBC5035" box="[189,294,1265,1291]" name="Brazil" pageId="23" pageNumber="24">BRAZIL</collectingCountry>
•
<specimenCount id="9D57FD251324B1764E14FB1DFEC15035" box="[322,347,1265,1291]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
;
<collectingRegion id="4995F84E1324B1764E3FFB1CFE2A5034" box="[361,432,1264,1290]" country="Brazil" name="Bahia" pageId="23" pageNumber="24">Bahia</collectingRegion>
,
<collectingMunicipality id="6B8AACD61324B1764EEBFB1CFD875034" box="[445,541,1264,1290]" pageId="23" pageNumber="24">Lençóis</collectingMunicipality>
,
<location id="8E8E60771324B1764D7CFB1DFD2C5035" LSID="urn:lsid:plazi:treatment:03F887BA1324B1794EADFDD1FBC951E8:8E8E60771324B1764D7CFB1DFD2C5035" box="[554,694,1265,1291]" country="Brazil" latitude="-12.540361" longLatPrecision="1" longitude="-41.40271" municipality="Lencois" name="cave Lapao" pageId="23" pageNumber="24" stateProvince="Bahia">cave Lapão</location>
; [
<geoCoordinate id="EE65506B1324B1764D9CFB1CFCCC5035" box="[714,854,1264,1291]" degrees="12.540361" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-12.540361">-12.540361</geoCoordinate>
,
<geoCoordinate id="EE65506B1324B1764C32FB1CFC6B5035" box="[868,1009,1264,1291]" degrees="41.402709" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-41.40271">-41.402709</geoCoordinate>
];
<date id="FFEF106C1324B1764B53FB1DFBE05034" box="[1029,1146,1264,1291]" pageId="23" pageNumber="24" value="1987-01">
<collectingDate id="EFABE9841324B1764B53FB1DFBE05034" box="[1029,1146,1264,1291]" pageId="23" pageNumber="24" value="1987-01">Jan. 1987</collectingDate>
</date>
;
<collectorName id="26A4537A1324B1764BDEFB1DFAA05034" box="[1160,1338,1264,1291]" pageId="23" pageNumber="24">F. Chaimowicz</collectorName>
leg.;
<specimenCode id="DBF79ED71324B1764FEBFAFEFEDA5012" box="[189,320,1298,1324]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24" type="Museum">MZSP 940</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE36AC1324B1764FBCFABEFB1D50B0" blockId="23.[189,1159,1362,1422]" pageId="23" pageNumber="24">
<materialsCitation id="3B393CF11324B1764FBCFABEFB1950B0" ID-GBIF-Occurrence="4101310047" collectingDate="1987-01" collectionCode="MZSP" collectorName="F. Chaimowicz" country="Brazil" latitude="-12.540361" location="BRAZIL" longLatPrecision="1" longitude="-41.40271" municipality="Lencois" pageId="23" pageNumber="24" specimenCode="MZSP 940" specimenCount="4" specimenCount-female="1" specimenCount-immature="2" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="B925EABE1324B1764FBCFABEFEFD5052" bold="true" box="[234,359,1362,1388]" pageId="23" pageNumber="24">
<typeStatus id="54EA880E1324B1764FBCFABEFEFD5052" box="[234,359,1362,1388]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
(total:
<specimenCount id="9D57FD251324B1764EEBFABEFE715052" box="[445,491,1362,1388]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD251324B1764EA1FABEFDB95052" box="[503,547,1362,1388]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D57FD251324B1764D64FABEFD205053" box="[562,698,1362,1389]" pageId="23" pageNumber="24" type="immature">1 immature</specimenCount>
)
<collectingCountry id="F346763C1324B1764FEBFA98FEBC50B0" box="[189,294,1396,1422]" name="Brazil" pageId="23" pageNumber="24">BRAZIL</collectingCountry>
•
<specimenCount id="9D57FD251324B1764E69FA98FDA450B3" box="[319,574,1395,1422]" pageId="23" pageNumber="24" type="immature">1 ♂, 1 ♀, 1 immature</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBF79ED71324B1764B56FA98FB1950B0" box="[1024,1155,1396,1422]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24" type="Museum">MZSP 940</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36AC1324B1764FBCFA5FFD4650F0" blockId="23.[189,1399,1459,1620]" box="[234,732,1459,1486]" pageId="23" pageNumber="24">
<emphasis id="B925EABE1324B1764FBCFA5FFE3C50F3" bold="true" box="[234,422,1459,1485]" pageId="23" pageNumber="24">Other material</emphasis>
(total:
<specimenCount id="9D57FD251324B1764EADFA5FFDA550F0" box="[507,575,1459,1486]" pageId="23" pageNumber="24" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D57FD251324B1764D1DFA5FFD4950F0" box="[587,723,1459,1486]" pageId="23" pageNumber="24" type="immature">1 immature</specimenCount>
)
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBFA39FD33536A" blockId="23.[189,1399,1459,1620]" pageId="23" pageNumber="24">
<materialsCitation id="3B393CF11324B1764FEBFA39FCE9532E" ID-GBIF-Occurrence="4101309540" collectingDate="2010-01-03" collectionCode="R" collectorName="R. L. Ferreira" country="Brazil" elevation="397" latitude="-12.561843" location="cave Lapao de Lendres" longLatPrecision="1" longitude="-41.38981" municipality="Lencois" pageId="23" pageNumber="24" specimenCode="ISLA 20617" specimenCount="2" specimenCount-immature="1" specimenCount-male="1" stateProvince="Bahia">
<collectingCountry id="F346763C1324B1764FEBFA39FEBC50D1" box="[189,294,1493,1519]" name="Brazil" pageId="23" pageNumber="24">BRAZIL</collectingCountry>
–
<emphasis id="B925EABE1324B1764E1AFA39FE0F50D1" bold="true" box="[332,405,1493,1519]" pageId="23" pageNumber="24">
<collectingRegion id="4995F84E1324B1764E1AFA39FE0F50D1" box="[332,405,1493,1519]" country="Brazil" name="Bahia" pageId="23" pageNumber="24">Bahia</collectingRegion>
</emphasis>
•
<specimenCount id="9D57FD251324B1764EE1FA39FE7050D1" box="[439,490,1493,1519]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD251324B1764EADFA39FD1050D1" box="[507,650,1493,1519]" pageId="23" pageNumber="24" type="immature">1 immature</specimenCount>
;
<collectingMunicipality id="6B8AACD61324B1764DCCFA39FD6050D1" box="[666,762,1493,1519]" pageId="23" pageNumber="24">Lençóis</collectingMunicipality>
,
<location id="8E8E60771324B1764C5CFA39FBB050D1" LSID="urn:lsid:plazi:treatment:03F887BA1324B1794EADFDD1FBC951E8:8E8E60771324B1764C5CFA39FBB050D1" box="[778,1066,1493,1519]" country="Brazil" latitude="-12.561843" longLatPrecision="1" longitude="-41.38981" municipality="Lencois" name="cave Lapao de Lendres" pageId="23" pageNumber="24" stateProvince="Bahia">cave Lapão de Lendres</location>
; [
<geoCoordinate id="EE65506B1324B1764B14FA39FB5450CE" box="[1090,1230,1493,1520]" degrees="12.561843" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-12.561843">-12.561843</geoCoordinate>
,
<geoCoordinate id="EE65506B1324B1764B88FA38FAF150D1" box="[1246,1387,1492,1519]" degrees="41.389809" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-41.38981">-41.389809</geoCoordinate>
];
<elevation id="007CD19F1324B1764FEBFA1BFED7532E" box="[189,333,1526,1553]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="23" pageNumber="24" unit="m" value="397.0">
<quantity id="4CA99B491324B1764FEBFA1BFE90532F" box="[189,266,1527,1553]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="23" pageNumber="24" unit="m" value="397.0">397 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1324B1764E08FA1BFE74532F" box="[350,494,1527,1553]" pageId="23" pageNumber="24" value="2010-01-03">
<collectingDate id="EFABE9841324B1764E08FA1BFE74532F" box="[350,494,1527,1553]" pageId="23" pageNumber="24" value="2010-01-03">3 Jan. 2010</collectingDate>
</date>
;
<collectorName id="26A4537A1324B1764EABFA1BFD02532E" box="[509,664,1526,1553]" pageId="23" pageNumber="24">R.L. Ferreira</collectorName>
leg.;
<specimenCode id="DBF79ED71324B1764DB6FA1BFCE9532E" box="[736,883,1526,1553]" pageId="23" pageNumber="24">ISLA 20617</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11324B1764CC5FA1BFC1F530D" ID-GBIF-Occurrence="4101309965" collectingDate="2012-01-16" collectionCode="R" collectorName="I. L. F. Magalhaes" country="Brazil" elevation="397" latitude="-12.561843" location="cave Lapao" longLatPrecision="1" longitude="-41.38981" municipality="Lencois" pageId="23" pageNumber="24" specimenCode="IBSP 5209" specimenCount="1" specimenCount-immature="1" specimenCount-male="1" stateProvince="Bahia">
<specimenCount id="9D57FD251324B1764CC5FA1BFC5F532F" box="[915,965,1527,1553]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B8AACD61324B1764C82FA1AFBAE532E" box="[980,1076,1526,1552]" pageId="23" pageNumber="24">Lençóis</collectingMunicipality>
,
<location id="8E8E60771324B1764B15FA1BFB4A532F" LSID="urn:lsid:plazi:treatment:03F887BA1324B1794EADFDD1FBC951E8:8E8E60771324B1764B15FA1BFB4A532F" box="[1091,1232,1527,1553]" country="Brazil" latitude="-12.561843" longLatPrecision="1" longitude="-41.38981" municipality="Lencois" name="cave Lapao" pageId="23" pageNumber="24" stateProvince="Bahia">cave Lapão</location>
; [
<geoCoordinate id="EE65506B1324B1764BB0FA1AFAE8532F" box="[1254,1394,1526,1553]" degrees="12.540361" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-12.540361">-12.540361</geoCoordinate>
,
<geoCoordinate id="EE65506B1324B1764FEBF9FBFED0530C" box="[189,330,1559,1586]" degrees="41.402709" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-41.40271">-41.402709</geoCoordinate>
];
<date id="FFEF106C1324B1764E0AF9F4FE6E530C" box="[348,500,1560,1587]" pageId="23" pageNumber="24" value="2012-01-16">
<collectingDate id="EFABE9841324B1764E0AF9F4FE6E530C" box="[348,500,1560,1587]" pageId="23" pageNumber="24" value="2012-01-16">16 Jan. 2012</collectingDate>
</date>
;
<collectorName id="26A4537A1324B1764D56F9F5FD5F530C" box="[512,709,1560,1587]" pageId="23" pageNumber="24">I.L.F. Magalhães</collectorName>
leg.;
<specimenCode id="DBF79ED71324B1764C51F9F4FC1F530D" box="[775,901,1560,1587]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="23" pageNumber="24">IBSP 5209</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11324B1764CCBF9F4FD3F536A" ID-GBIF-Occurrence="4101309752" collectingDate="1991-09-03" collectionCode="MZSP" collectorName="E. Trajano" country="Iceland" latitude="-12.540361" location="cave Lapao" longLatPrecision="1" longitude="-41.40271" municipality="Lencois" pageId="23" pageNumber="24" specimenCode="MZSP 1006" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D57FD251324B1764CCBF9F4FC56530D" box="[925,972,1560,1587]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; same locality data as for preceding;
<date id="FFEF106C1324B1764FEBF9D6FED6536A" box="[189,332,1594,1620]" pageId="23" pageNumber="24" value="1991-09-03">
<collectingDate id="EFABE9841324B1764FEBF9D6FED6536A" box="[189,332,1594,1620]" pageId="23" pageNumber="24" value="1991-09-03">3 Sep. 1991</collectingDate>
</date>
;
<collectorName id="26A4537A1324B1764E0EF9D6FE48536A" box="[344,466,1594,1620]" pageId="23" pageNumber="24">E. Trajano</collectorName>
leg.;
<specimenCode id="DBF79ED71324B1764D42F9D6FD3F536A" box="[532,677,1594,1620]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24" type="Museum">MZSP 1006</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271324B1794FEBF995FAEC5156" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="8BEE36AC1324B1764FEBF995FE0953AA" blockId="23.[189,403,1657,1684]" box="[189,403,1657,1684]" pageId="23" pageNumber="24">
<heading id="D0A681C01324B1764FEBF995FE0953AA" bold="true" box="[189,403,1657,1684]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B925EABE1324B1764FEBF995FE0953AA" bold="true" box="[189,403,1657,1684]" pageId="23" pageNumber="24">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBF94BFEED53DD" blockId="23.[189,1398,1702,1763]" pageId="23" pageNumber="24">
<smallCapsWord id="8D08A0701324B1764FEBF94BFEF45381" baselines="1722,1722" box="[189,366,1703,1729]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="23" pageNumber="24">MEASUREMENTS</smallCapsWord>
. 60 body rings (1–2 apodous + telson). Males: body length
<quantity id="4CA99B491324B1764B78F94BFB1E53FF" box="[1070,1156,1703,1729]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="24" unit="mm" value="90.0">90 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B491324B1764E7DF924FEE853DD" box="[299,370,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBF8E4FEA9527D" blockId="23.[189,1398,1800,1859]" pageId="23" pageNumber="24">
<smallCapsWord id="8D08A0701324B1764FEBF8E4FE93521E" baselines="1819,1819" box="[189,265,1800,1826]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="23" pageNumber="24">COLOR</smallCapsWord>
. Body color greyish; collum darker; metazonites with a light posterior band; antennae and legs brownish.
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBF886FB7952F9" blockId="23.[189,1398,1897,1991]" pageId="23" pageNumber="24">
<smallCapsWord id="8D08A0701324B1764FEBF886FF6652BC" baselines="1917,1917" box="[189,252,1898,1924]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="23" pageNumber="24">HEAD</smallCapsWord>
. Antennae long (
<figureCitation id="136A2A291324B1764E8DF885FDC652BA" box="[475,604,1897,1924]" captionStart="Fig" captionStartId="266.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1743]" captionTargetId="figure-14@266.[250,1337,265,1743]" captionTargetPageId="266" captionText="Fig. 163. Stereoscopic images of antennae of species ofPseudonannolene Silvestri, 1895.A. P. albiventris Schubart, 1952. B. P. anapophysis Fontanetti, 1996. C. P. alata sp. nov. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. F. P. erikae Iniesta &amp; Ferreira, 2014. G. P. fontanettiae Iniesta &amp; Ferreira, 2014. H. P. halophila Schubart, 1949. I. P. inops Brölemann, 1929. J. P. leopoldoi Iniesta &amp; Ferreira, 2014. K. P. longicornis (Porat, 1888). L. P. magna Udulutsch &amp; Pietrobon, 2003. M. P. maritima Schubart, 1949. N. P. mesai Fontanetti, 2000. O. P. microzoporus Mauriès, 1987. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891405" httpUri="https://zenodo.org/record/7891405/files/figure.png" pageId="23" pageNumber="24">Fig. 163B</figureCitation>
), just reaching back to end of ring 6 when extended dorsally; antennomeres elongated; relative antennomere lengths 1&lt;2&lt;3≈4&lt;5≈6&gt;7. Mandibular cardo with ventral margin swollen. Ommatidial cluster well-developed, elliptical; ca 23 ommatidia in 4 rows.
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFEE5FED754B7" blockId="24.[187,1399,265,393]" pageId="24" pageNumber="25">
<smallCapsWord id="8D08A070132BB1794FEBFEE5FF67541F" baselines="284,284" box="[189,253,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="24" pageNumber="25">BODY</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794E55FEE2FEDB541F" baselines="284" box="[259,321,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="24" pageNumber="25">RINGS</smallCapsWord>
. Collum with lateral lobes broadly rounded, with ca 9 striae, slightly curved ectad (
<figureCitation id="136A2A29132BB1794BA8FEE5FAF3541D" box="[1278,1385,265,291]" captionStart="Fig" captionStartId="162.[189,232,764,790]" captionTargetBox="[189,1389,265,717]" captionTargetId="figure-13@162.[189,1398,265,717]" captionTargetPageId="162" captionText="Fig. 48. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209), in lateral view. A. Anterior region. B. Posterior region. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891143" httpUri="https://zenodo.org/record/7891143/files/figure.png" pageId="24" pageNumber="25">Fig. 48A</figureCitation>
). Very faintly constriction between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae. Anterior sterna in midbody rings subrectangular, without transverse striae (
<figureCitation id="136A2A29132BB1794F90FE82FEDA54B7" box="[198,320,366,393]" captionStart="Fig" captionStartId="270.[189,232,1892,1918]" captionTargetBox="[213,1374,265,1851]" captionTargetId="figure-14@270.[213,1374,265,1851]" captionTargetPageId="270" captionText="Fig. 167. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. albiventris Schubart, 1952. B. P. ambuatinga Iniesta &amp; Ferreira, 2013. C. P. anapophysis Fontanetti, 1996. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891417" httpUri="https://zenodo.org/record/7891417/files/figure.png" pageId="24" pageNumber="25">Fig. 167C</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFE5CFADF570E" blockId="24.[187,1399,431,560]" pageId="24" pageNumber="25">
<smallCapsWord id="8D08A070132BB1794FEBFE5CFF6054F6" baselines="451,451" box="[189,250,432,458]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="First" pageId="24" pageNumber="25">FIRST</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794E50FE59FEFF54F6" baselines="451,451,451" box="[262,357,432,458]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="24" pageNumber="25">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794E27FE59FE1154F6" baselines="451" box="[369,395,437,456]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="24" pageNumber="25">OF</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794EC0FE59FE4654F6" baselines="451" box="[406,476,437,456]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="24" pageNumber="25">MALES</smallCapsWord>
. Coxae (
<emphasis id="B925EABE132BB1794D19FE5CFDF154F4" bold="true" box="[591,619,432,458]" italics="true" pageId="24" pageNumber="25">cx</emphasis>
) short (less than half of remaining podomere lengths), with the base slightly arched, densely setose, and apically projected (
<figureCitation id="136A2A29132BB1794C29FE3DFC7754D5" box="[895,1005,465,491]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49A</figureCitation>
); prefemoral process (
<emphasis id="B925EABE132BB1794BA5FE3DFA8D54D5" bold="true" box="[1267,1303,465,491]" italics="true" pageId="24" pageNumber="25">prf</emphasis>
) almost vestigial, with mesal region of prefemur whitish, covered by long setae, and ectal region more sclerotized and slightly projected apically (
<figureCitation id="136A2A29132BB1794D78FDFAFD03570E" box="[558,665,534,560]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49B</figureCitation>
); remaining podomeres with setae along the mesal region.
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFDBAFE4C578A" blockId="24.[188,1398,598,693]" pageId="24" pageNumber="25">
<smallCapsWord id="8D08A070132BB1794FEBFDBAFE8E5750" baselines="617,617" box="[189,276,598,624]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Second" pageId="24" pageNumber="25">SECOND</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794E76FDB7FE1A5751" baselines="617,617,618" box="[288,384,598,624]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="24" pageNumber="25">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794EDAFDB7FE3C5750" baselines="617" box="[396,422,603,622]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="24" pageNumber="25">OF</smallCapsWord>
<smallCapsWord id="8D08A070132BB1794EE4FDB7FE625750" baselines="617" box="[434,504,603,622]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="24" pageNumber="25">MALES</smallCapsWord>
. Coxa (
<emphasis id="B925EABE132BB1794D08FDBAFDE0574E" bold="true" box="[606,634,598,624]" italics="true" pageId="24" pageNumber="25">cx</emphasis>
) rounded; penis (
<emphasis id="B925EABE132BB1794C00FDBAFCEF574E" bold="true" box="[854,885,598,624]" italics="true" pageId="24" pageNumber="25">pn</emphasis>
) located at proximal region, rounded, not extended basally (
<figureCitation id="136A2A29132BB1794ED9FD94FE6357AC" box="[399,505,632,658]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49C</figureCitation>
); prefemur slightly compressed dorsoventrally; remaining podomeres setose, with long setae mesally.
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFD37FEFA5138" blockId="24.[188,1399,731,1030]" pageId="24" pageNumber="25">
<smallCapsWord id="8D08A070132BB1794FEBFD37FEA957CD" baselines="750,750" box="[189,307,731,757]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="24" pageNumber="25">GONOPODS</smallCapsWord>
. Gonocoxa (
<emphasis id="B925EABE132BB1794E9DFD37FE6F57CB" bold="true" box="[459,501,731,757]" italics="true" pageId="24" pageNumber="25">gcx</emphasis>
) elongated, almost twice as long as telopodite, with the base slightly arched; flattened antero-posteriorly (
<figureCitation id="136A2A29132BB1794D59FD11FD045629" box="[527,670,765,791]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49D–F</figureCitation>
); with rows of papillae mesally. Seminal groove (
<emphasis id="B925EABE132BB1794BA4FD11FA965629" bold="true" box="[1266,1292,765,791]" italics="true" pageId="24" pageNumber="25">sg</emphasis>
) curved; arising medially on mesal cavity and terminating apically on the seminal apophysis (
<emphasis id="B925EABE132BB1794BF4FCF3FB265607" bold="true" box="[1186,1212,799,825]" italics="true" pageId="24" pageNumber="25">sa</emphasis>
); protruding on squamous region of solenomere. Shoulder (
<emphasis id="B925EABE132BB1794D9EFCACFD7E5665" bold="true" box="[712,740,832,859]" italics="true" pageId="24" pageNumber="25">sh</emphasis>
) long, subtriangular. Telopodite (
<emphasis id="B925EABE132BB1794B20FCADFB175665" bold="true" box="[1142,1165,833,859]" italics="true" pageId="24" pageNumber="25">tp</emphasis>
) as wide as half of
<emphasis id="B925EABE132BB1794FE9FC8FFF7D5643" bold="true" box="[191,231,867,893]" italics="true" pageId="24" pageNumber="25">gcx</emphasis>
, separated from
<emphasis id="B925EABE132BB1794EE7FC8FFE575640" bold="true" box="[433,461,867,894]" italics="true" pageId="24" pageNumber="25">sh</emphasis>
by deep depression (
<figureCitation id="136A2A29132BB1794D9AFC8FFCA15643" box="[716,827,867,893]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49D</figureCitation>
); solenomere (
<emphasis id="B925EABE132BB1794CBDFC8FFC655640" bold="true" box="[1003,1023,867,894]" italics="true" pageId="24" pageNumber="25">sl</emphasis>
) with subtriangular apicomesal process (
<emphasis id="B925EABE132BB1794E73FC6AFEC3569E" bold="true" box="[293,345,902,928]" italics="true" pageId="24" pageNumber="25">amp</emphasis>
); ectal process (
<emphasis id="B925EABE132BB1794D41FC6AFDA9569E" bold="true" box="[535,563,902,928]" italics="true" pageId="24" pageNumber="25">ep</emphasis>
) subtriangular, elongated and perpendicular to
<emphasis id="B925EABE132BB1794B01FC6AFB11569E" bold="true" box="[1111,1163,902,928]" italics="true" pageId="24" pageNumber="25">amp</emphasis>
;
<emphasis id="B925EABE132BB1794BCDFC6AFB2F569E" bold="true" box="[1179,1205,902,928]" italics="true" pageId="24" pageNumber="25">sa</emphasis>
located at mesal portion, visible apically. Internal branch (
<emphasis id="B925EABE132BB1794DF0FC4BFD2756FC" bold="true" box="[678,701,935,962]" italics="true" pageId="24" pageNumber="25">ib</emphasis>
) shovel-shaped, rounded and slightly curved apically, with horizontal plate rounded; setae restricted to the apical region of
<emphasis id="B925EABE132BB1794C83FC25FC7656DA" bold="true" box="[981,1004,969,996]" italics="true" pageId="24" pageNumber="25">ib</emphasis>
, exceeding seminal region of
<emphasis id="B925EABE132BB1794A35FC25FAED56DA" bold="true" box="[1379,1399,969,996]" italics="true" pageId="24" pageNumber="25">sl</emphasis>
(
<figureCitation id="136A2A29132BB1794F90FC00FEC95138" box="[198,339,1004,1030]" captionStart="Fig" captionStartId="163.[189,232,1675,1701]" captionTargetBox="[189,1398,270,1627]" captionTargetId="figure-11@163.[189,1398,270,1627]" captionTargetPageId="163" captionText="Fig. 49. Pseudonannolene anapophysis Fontanetti, 1996, ♂ (IBSP 5209). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, E–F = 0.5 mm; B, D = 0.2 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891145" httpUri="https://zenodo.org/record/7891145/files/figure.png" pageId="24" pageNumber="25">Fig. 49D–F</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFBC1FAEC5156" blockId="24.[187,1398,1068,1129]" pageId="24" pageNumber="25">
<smallCapsWord id="8D08A070132BB1794FEBFBC1FE89517B" baselines="1088,1088" box="[189,275,1069,1095]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="24" pageNumber="25">VULVAE</smallCapsWord>
. As typical for the genus. Bursa subtriangular, glabrous (
<figureCitation id="136A2A29132BB1794CF5FBC0FB875178" box="[931,1053,1068,1094]" captionStart="Fig" captionStartId="280.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1739]" captionTargetId="figure-14@280.[250,1337,265,1739]" captionTargetPageId="280" captionText="Fig. 177. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. albiventris Schubart, 1952. B. P. alegrensis Silvestri, 1897. C. P. ambuatinga Iniesta &amp; Ferreira, 2013. D. P. anapophysis Fontanetti, 1996. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. erikae Iniesta &amp; Ferreira, 2014. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M. P. leucocephalus Schubart, 1944. N. P. longicornis (Porat, 1888). O. P. lundi Iniesta &amp; Ferreira, 2015. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891439" httpUri="https://zenodo.org/record/7891439/files/figure.png" pageId="24" pageNumber="25">Fig. 177D</figureCitation>
); internal valve subtriangular, with mesal region rounded; operculum narrow, constricted medially; external valve wide, subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C34B6527132BB1794FEBFB62FBC951E8" box="[189,1107,1166,1239]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BEE36AC132BB1794FEBFB62FECC5196" blockId="24.[189,342,1166,1192]" box="[189,342,1166,1192]" pageId="24" pageNumber="25">
<heading id="D0A681C0132BB1794FEBFB62FECC5196" bold="true" box="[189,342,1166,1192]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B925EABE132BB1794FEBFB62FECC5196" bold="true" box="[189,342,1166,1192]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC132BB1794FEBFB51FBC951E8" blockId="24.[189,1107,1212,1239]" box="[189,1107,1212,1239]" pageId="24" pageNumber="25">
Known only from the central region of the Brazilian State of
<collectingRegion id="4995F84E132BB1794CDDFB50FC5551E8" box="[907,975,1212,1238]" country="Brazil" name="Bahia" pageId="24" pageNumber="25">Bahia</collectingRegion>
(
<figureCitation id="136A2A29132BB1794C89FB50FBDC51E8" box="[991,1094,1212,1238]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="24" pageNumber="25">Fig. 180</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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