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<document id="E5A360BD23058DF1BB72F56B127C906F" ID-DOI="10.5281/zenodo.181433" ID-GBIF-Dataset="c77d84f8-50e8-4bf3-9dbe-f74e48362846" ID-ISSN="1175-5326" ID-Zenodo-Dep="181433" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459833660273" checkinUser="plazi" docAuthor="Last, Peter R. &amp; White, William T." docDate="2008" docId="03F887AA9C1CFFD7FF3EFB44595BF9FC" docLanguage="en" docName="zt01734p026.pdf" docOrigin="Zootaxa 1734" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Squatina legnota Last &amp; White, 2008, new species" docType="treatment" docVersion="10" lastPageNumber="16" masterDocId="FFC1FFD29C17FFD8FFA9FFE3582EFFC0" masterDocTitle="Three new angel sharks (Chondrichthyes: Squatinidae) from the Indo-Australian region" masterLastPageNumber="26" masterPageNumber="1" pageNumber="12" updateTime="1698223152723" updateUser="plazi">
<mods:mods id="C560E03F652AD0D4B39B0B1377C1748B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DDDC84281FCF73F6F9E6F551FA7E4797">Three new angel sharks (Chondrichthyes: Squatinidae) from the Indo-Australian region</mods:title>
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<mods:namePart id="0D28B05ECC50DEEFAE79642BA0B48E44">Last, Peter R.</mods:namePart>
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<mods:namePart id="C2DE1284EF4907CBDBC10D97E1A9DD81">White, William T.</mods:namePart>
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<mods:date id="B00A338AC7E37070BC48B0F4EA1DA3E6">2008</mods:date>
<mods:detail id="4CD07E468E3791C78184AA8E5D58F2F0" type="volume">
<mods:number id="DF3ECDE1C6EEEED6DFC6C60347D33B33">1734</mods:number>
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<mods:classification id="AF6220032B97FEFB27410F238F86DB08">journal article</mods:classification>
<mods:identifier id="C80FC47DC9B7C748B1CF60F47B2B57C3" type="DOI">10.5281/zenodo.181433</mods:identifier>
<mods:identifier id="A61BA2D056A3952B306B6206AFD1D33A" type="GBIF-Dataset">c77d84f8-50e8-4bf3-9dbe-f74e48362846</mods:identifier>
<mods:identifier id="417FB31C656E1A6A15C8A34ABB3A9503" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6117500D4C068DB1205B7B7B1D884B32" type="Zenodo-Dep">181433</mods:identifier>
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<treatment id="03F887AA9C1CFFD7FF3EFB44595BF9FC" ID-DOI="http://doi.org/10.5281/zenodo.5689527" ID-GBIF-Taxon="119358309" ID-Zenodo-Dep="5689527" LSID="urn:lsid:plazi:treatment:03F887AA9C1CFFD7FF3EFB44595BF9FC" httpUri="http://treatment.plazi.org/id/03F887AA9C1CFFD7FF3EFB44595BF9FC" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="C34B65379C1CFFD3FF3EFB44599CFB24" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BEE36BC9C1CFFD3FF3EFB4459D5FB01" blockId="11.[151,507,1191,1292]" box="[151,507,1191,1217]" pageId="11" pageNumber="12">
<heading id="D0A681D09C1CFFD3FF3EFB4459D5FB01" bold="true" box="[151,507,1191,1217]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B925EAAE9C1CFFD3FF3EFB4459D5FB01" bold="true" box="[151,507,1191,1217]" pageId="11" pageNumber="12">
<taxonomicName id="4C514D3F9C1CFFD3FF3EFB44594AFB01" ID-CoL="6ZFHR" box="[151,356,1191,1217]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="legnota" status="sp. nov.">
<emphasis id="B925EAAE9C1CFFD3FF3EFB44594AFB01" bold="true" box="[151,356,1191,1217]" italics="true" pageId="11" pageNumber="12">Squatina legnota</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21657D59C1CFFD3FEC2FB4459D5FB01" box="[363,507,1191,1217]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36BC9C1CFFD3FF3EFB29599CFB24" blockId="11.[151,507,1191,1292]" box="[151,434,1226,1252]" pageId="11" pageNumber="12">Indonesian Angel Shark</paragraph>
</subSubSection>
<subSubSection id="C34B65379C1CFFD3FF3EFB115CCFFA99" pageId="11" pageNumber="12" type="description">
<paragraph id="8BEE36BC9C1CFFD3FF3EFB1159BBFACC" blockId="11.[151,507,1191,1292]" box="[151,405,1266,1292]" pageId="11" pageNumber="12">
<figureCitation id="136A2A399C1CFFD3FF3EFB1158CCFACC" box="[151,226,1266,1292]" captionStart="FIGURE 4" captionStartId="14.[151,255,1951,1975]" captionTargetBox="[264,1317,207,1901]" captionTargetId="figure@14.[252,1311,731,1356]" captionTargetPageId="14" captionText="FIGURE 4. Squatina legnota sp. nov.: A, dorsal view of holotype (MZB 15500, female 471 mm TL); B, dorsal view of non-type specimen (adult male, 1341 mm TL); and C, ventral view of holotype." httpUri="https://zenodo.org/record/181437/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
,
<figureCitation id="136A2A399C1CFFD3FF47FB1158D0FACC" box="[238,254,1266,1292]" captionStart="FIGURE 5" captionStartId="15.[151,255,581,605]" captionTargetBox="[539,1034,222,549]" captionTargetId="figure@15.[527,1068,194,565]" captionTargetPageId="15" captionText="FIGURE 5. Illustration of the anterior nasal flap of paratype of Squatina legnota sp. nov. (CSIRO H 6565 – 01). Illustration by Stephanie Barnes." httpUri="https://zenodo.org/record/181438/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
,
<figureCitation id="136A2A399C1CFFD3FEA2FB115937FACC" box="[267,281,1266,1292]" captionStart="FIGURE 8" captionStartId="22.[151,255,1939,1963]" captionTargetBox="[341,1188,213,1898]" captionTargetId="figure@22.[349,1200,769,1143]" captionTargetPageId="22" captionText="FIGURE 8. Lateral view of the tail and ventral view of the caudal keel of: A, Squatina albipunctata sp. nov. holotype (CSIRO H 6561 – 01, adult male 985 mm TL); B, Squatina legnota sp. nov. paratype (CSIRO H 6565 – 01, adult male 1252 mm TL); and C, Squatina pseudocellata sp. nov. (CSIRO H 2567 – 01, adult male 751 mm TL)." httpUri="https://zenodo.org/record/181441/files/figure.png" pageId="11" pageNumber="12">8</figureCitation>
b,
<tableCitation id="C6D303079C1CFFD3FE9EFB1159BEFACC" box="[311,400,1266,1292]" captionStart="TABLE 3" captionStartId="16.[151,239,151,175]" captionTargetBox="[159,1432,410,1969]" captionTargetPageId="16" captionText="TABLE 3 Proportional dimensions as percentages of total length for the holotype (MZB 15500), paratype (CSIRO H 6565 – 01) and one non-type of Squatina legnota n. sp. and for the holotype (CSIRO H 2567 – 01) and ranges for the 5 paratypes of Squatina pseudocellata n. sp." httpUri="http://table.plazi.org/id/DF2E66349C07FFC8FF3EFF745A74FF37" pageId="11" pageNumber="12" tableUuid="DF2E66349C07FFC8FF3EFF745A74FF37">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BEE36BC9C1CFFD3FF3EFAA15CCFFA99" blockId="11.[151,1249,1346,1369]" box="[151,1249,1346,1369]" pageId="11" pageNumber="12">
<taxonomicName id="4C514D3F9C1CFFD3FF3EFAA158D8FA99" box="[151,246,1346,1369]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B925EAAE9C1CFFD3FF3EFAA158D8FA99" box="[151,246,1346,1369]" italics="true" pageId="11" pageNumber="12">Squatina</emphasis>
</taxonomicName>
sp. 1:
<bibRefCitation id="EFC04B4D9C1CFFD3FE94FAA159D1FA99" author="White" box="[317,511,1346,1369]" pageId="11" pageNumber="26" refString="White, W. T., Last, P. R., Stevens, J. D., Yearsley, G. K., Fahmi &amp; Dharmadi (2006) Economically Important Sharks and Rays of Indonesia. ACIAR Monograph Series, No 124, ACIAR Publishing, Canberra, 329 pp." type="book" year="2006">
White
<emphasis id="B925EAAE9C1CFFD3FE2DFAA15994FA99" box="[388,442,1346,1369]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
, 2006
</bibRefCitation>
,
<emphasis id="B925EAAE9C1CFFD3FDA3FAA15C10FA99" box="[522,1086,1346,1369]" italics="true" pageId="11" pageNumber="12">
Economically Important Sharks &amp; Rays of
<collectingCountry id="F346762C9C1CFFD3FC7CFAA15C10FA99" box="[981,1086,1346,1369]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
</emphasis>
, pp 76, 77, fig.
</paragraph>
</subSubSection>
<subSubSection id="C34B65379C1CFFD3FF3EFA685DA4F984" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BEE36BC9C1CFFD3FF3EFA685975FA0C" blockId="11.[151,1436,1418,1964]" pageId="11" pageNumber="12">
<emphasis id="B925EAAE9C1CFFD3FF3EFA685924FA64" box="[151,266,1419,1444]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="54EA881E9C1CFFD3FF3EFA685928FA64" box="[151,262,1419,1444]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED40AE799C1CFFD3FEBCFA69597BFA64" box="[277,341,1418,1444]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="11" pageNumber="12">MZB</collectionCode>
15500, female
<quantity id="4CA99B599C1CFFD3FDB0FA695A51FA64" box="[537,639,1418,1444]" metricMagnitude="-1" metricUnit="m" metricValue="4.71" pageId="11" pageNumber="12" unit="mm" value="471.0">471 mm</quantity>
<collectionCode id="ED40AE799C1CFFD3FD23FA695A9FFA64" box="[650,689,1418,1444]" pageId="11" pageNumber="12">TL</collectionCode>
, Cilacap fish landing site, Central Java,
<collectingCountry id="F346762C9C1CFFD3FB01FA695D33FA64" box="[1192,1309,1418,1444]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
,
<geoCoordinate id="EE65507B9C1CFFD3FA84FA695DB8FA64" box="[1325,1430,1418,1444]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-7.7333336">07°44' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C1CFFD3FF3EFA515922FA0C" box="[151,268,1458,1484]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="109.0">109°00' E</geoCoordinate>
, 2004.
</paragraph>
<paragraph id="8BEE36BC9C1CFFD3FF6FFA385A62F9DC" blockId="11.[151,1436,1418,1964]" pageId="11" pageNumber="12">
<emphasis id="B925EAAE9C1CFFD3FF6FFA385917FA34" box="[198,313,1499,1524]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="54EA881E9C1CFFD3FF6FFA38591BFA34" box="[198,309,1499,1524]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectionCode id="ED40AE799C1CFFD3FEE8FA3959B9FA34" box="[321,407,1498,1524]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="11" pageNumber="12">CSIRO</collectionCode>
H 6565–01, adult male
<quantity id="4CA99B599C1CFFD3FD1DFA395B0AFA34" box="[692,804,1498,1524]" metricMagnitude="0" metricUnit="m" metricValue="1.252" pageId="11" pageNumber="12" unit="mm" value="1252.0">1252 mm</quantity>
<collectionCode id="ED40AE799C1CFFD3FC82FA395B7CFA34" box="[811,850,1498,1524]" pageId="11" pageNumber="12">TL</collectionCode>
, Tanjung Luar fish landing site, Lombok,
<collectingCountry id="F346762C9C1CFFD3FAE8FA395894F9DC" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
,
<geoCoordinate id="EE65507B9C1CFFD3FF6FF9E15905F9DC" box="[198,299,1538,1564]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="-8.75">08°45' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C1CFFD3FE91F9E15982F9DC" box="[312,428,1538,1564]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="116.583336">116°35' E</geoCoordinate>
,
<date id="FFEF107C9C1CFFD3FE10F9E15A66F9DC" box="[441,584,1538,1564]" pageId="11" pageNumber="12" value="2004-09-09">9 Sep. 2004</date>
.
</paragraph>
<paragraph id="8BEE36BC9C1CFFD3FF3EF9C85DA4F984" blockId="11.[151,1436,1418,1964]" box="[151,1418,1578,1604]" pageId="11" pageNumber="12">
<emphasis id="B925EAAE9C1CFFD3FF3EF9C85960F984" box="[151,334,1579,1604]" italics="true" pageId="11" pageNumber="12">Other material.</emphasis>
Adult male
<quantity id="4CA99B599C1CFFD3FE76F9C95A61F984" box="[479,591,1578,1604]" metricMagnitude="0" metricUnit="m" metricValue="1.341" pageId="11" pageNumber="12" unit="mm" value="1341.0">1341 mm</quantity>
<collectionCode id="ED40AE799C1CFFD3FDFFF9C95A53F984" box="[598,637,1578,1604]" pageId="11" pageNumber="12">TL</collectionCode>
, not retained, collected at same locality as
<typeStatus id="54EA881E9C1CFFD3FBDBF9C95CF4F984" box="[1138,1242,1578,1604]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
,
<date id="FFEF107C9C1CFFD3FB4CF9C95DA8F984" box="[1253,1414,1578,1604]" pageId="11" pageNumber="12" value="2005-03-10">10 Mar. 2005</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B65379C1CFFD3FF6FF9B15960F86C" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BEE36BC9C1CFFD3FF6FF9B15960F86C" blockId="11.[151,1436,1418,1964]" pageId="11" pageNumber="12">
<emphasis id="B925EAAE9C1CFFD3FF6FF9B15968F9AC" bold="true" box="[198,326,1618,1644]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
A moderately large
<taxonomicName id="4C514D3F9C1CFFD3FDE0F9B05A9FF9AC" box="[585,689,1619,1644]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B925EAAE9C1CFFD3FDE0F9B05A9FF9AC" box="[585,689,1619,1644]" italics="true" pageId="11" pageNumber="12">Squatina</emphasis>
</taxonomicName>
with the following characters: dorsal coloration simple, uniformly greyish brown; two dark subdorsal saddles; anterior ventral surface of pectoral fin blackish; no median row of scute-like denticles on trunk; anterior nasal flap with unfringed barbels; upper lip arch half-oval in shape (lip arch width 3.4–4.1% TL, height 0.9–1.2% TL); interorbital region weakly concave; pectoral–pelvic space 9.4–10.9% TL; pectoral-fin apex angular, not more than 120°; pelvic-fin free rear tips reaching first dorsal-fin origin; tail relatively long, pelvic-fin origin to ventral caudal lobe 3.8–4.0 times pectoral-fin base; pectoral fins moderately long, length 33.5–33.7% TL; dorsal fins not lobe-like; first dorsal-fin base longer than second dorsal-fin base; caudal fin of adult with angular apices; monospondylous centra 50 or 51; total vertebral centra 144.
</paragraph>
</subSubSection>
<subSubSection id="C34B65379C1BFFD7FF6CFF745BC4FD34" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="8BEE36BC9C1BFFD4FF6CFF745B78FE62" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="13">
<emphasis id="B925EAAE9C1BFFD4FF6CFF74597BFF71" bold="true" box="[197,341,151,177]" pageId="12" pageNumber="13">Description</emphasis>
. Body strongly depressed anteriorly, firm; trunk strongly depressed, almost ray-like, deepest at pectoral-fin base; not tapering abruptly at pelvic-fin insertion, tail strongly depressed, even at origin of caudal fin. Abdomen moderately elongate, pectoral–pelvic space 9.4 (10.9)% TL, 0.63 (0.74) of pre-branchial length; pelvic–caudal space 1.38 (1.28) times pelvic-fin length; tail moderately long, height at pelvic insertions 2.06 (2.24) in its width; caudal peduncle strongly depressed, short; lateral keels moderately developed, not greatly expanded anteriorly, originating below insertion of second dorsal fin, slightly below lateral midline, extending about two eye lengths past caudal-fin origin.
</paragraph>
<paragraph id="8BEE36BC9C1BFFD4FF6FFE5359B2FC3A" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="13">
Head broad, extended laterally, strongly depressed; oval in cross-section; subcircular, with weakly convex anterior margin when viewed from above (somewhat truncate in large adult male
<typeStatus id="54EA881E9C1BFFD4FBC1FE3B5CFCFE32" box="[1128,1234,472,498]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
); well-developed dermal folds present along ventrolateral margin of head, no lobes present near mouth corners; head width 19.9 (19.3)% TL, 3.17 (2.28) times head height. Snout very short, bluntly rounded in lateral view, horizontal preorbital length 3.5 (2.9)% TL; elevated slightly between nostril and eye; weak preorbital pit present, barely connected to snout apex by a shallow groove at midline; apex not indented anteriorly. Eye dorsolateral on head, small, suboval, length 2.4 (1.8)% TL, 6.11 (8.13) in pre-branchial length, interorbit weakly concave, no subocular pouches; widely separated, interorbital space 3.23 (4.45) times eye length, 2.26 (2.74) times horizontal preorbital length, 0.85 (0.93) times direct prespiracular length, 3.37 (3.06) times spiracle length. Spiracle of moderate size, cresentic, almost transverse to slightly oblique, well separated from eye, much larger than eye; anterior margin strongly concave, barely elevated above flattened posterior margin; posterior margin almost flat to forming a slight depression leading into spiracle; greatest diameter 3.31 (2.89) in interspiracular space. Gill slits large, pleated, lateral on head, subequal in length, located well forward of midbase of pectoral fin, intergill distance 7.4 (8.6)% TL, 2.68 (2.24) in head width; posterior lateral head fold partly overlapping anterior tip of pectoral fin.
</paragraph>
<paragraph id="8BEE36BC9C1BFFD4FF6FFBEB5A30F9C2" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="13">
Mouth large, terminal (or very slightly inferior), horizontally expanded, broadly arched, terminating just forward of mid-eye (slightly more posterior in large male
<typeStatus id="54EA881E9C1BFFD4FCFBFBD35B95FB8A" box="[850,955,1072,1098]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
), width 12.8 (12.1)% TL, 1.55 (1.59) in head width, 1.62 (1.50) times interorbital space; upper lip forming a broad, moderately convex border with snout; upper labial furrows very large, forming deep preoral groove from symphysis to posterior of jaw angle; lower labial furrows elongate, deep, separated from each other by less than their length; shorter postoral groove very deep, extending parallel to lower labial furrow from jaw angle, length slightly greater than half lower labial furrow length. Teeth widely spaced, unicuspid, small and dagger-like, conical without cusplets on a broad base; upper jaw with 3 small rows on each side of symphysis, midlateral teeth larger (about twice size of symphysial teeth) and more widely spaced than those adjacent; lower jaw edentate at symphysis, lateral teeth evenly spaced, slightly larger than those of upper jaw; no smaller, unpaired teeth at symphysis of upper jaw in
<typeStatus id="54EA881E9C1BFFD4FF4FFA7B5962FA72" box="[230,332,1432,1458]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(large male
<typeStatus id="54EA881E9C1BFFD4FE77FA7B5A6DFA72" box="[478,579,1432,1458]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
with well-developed symphysial teeth); anterior two lateral pairs at symphysis protruding anteriorly when mouth closed (5 rows in
<typeStatus id="54EA881E9C1BFFD4FCF7FA235BE6FA1A" box="[862,968,1472,1498]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
); teeth by row in
<typeStatus id="54EA881E9C1BFFD4FB35FA235D28FA1A" box="[1180,1286,1472,1498]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
,
<quantity id="4CA99B599C1BFFD4FABAFA235D7EFA1A" box="[1299,1360,1472,1498]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="12" pageNumber="13" unit="in" value="18.0">18 in</quantity>
upper jaw,
<quantity id="4CA99B599C1BFFD4FF67FA0B5924F9C2" box="[206,266,1512,1538]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="12" pageNumber="13" unit="in" value="18.0">18 in</quantity>
lower jaw, in
<typeStatus id="54EA881E9C1BFFD4FE19FA0B5A35F9C2" box="[432,539,1512,1538]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BEE36BC9C1BFFD4FF6FF9F35D01F8DA" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="13">Nostrils large, terminal on head, widely separated, internarial space 5.7 (6.2)% TL, 3.51 (3.12) in head width; adjacent upper lip of mouth. Anterior nasal flap large, overlapping mouth, with two well-developed, unfringed barbels; very large, basally flattened, posterolaterally directed barbel with an irregular membranous tip and large, membranous basal mesial lobe (partly concealed on inner basal edge of barbel); anteroventral margin of flap with a smaller, much more slender, semitubular barbel (capable of being unraveled) with a slightly expanded distal tip, directed mesially; posteroventral margin of nostril with a variably developed, membranous flap, margin sometimes irregular; no nasoral or circumnarial grooves; no dermal lobes.</paragraph>
<paragraph id="8BEE36BC9C1BFFD5FF6FF8CB5973FD52" blockId="12.[151,1437,151,2018]" lastBlockId="13.[151,1437,152,1892]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Dermal denticles covering entire dorsal surface of body, slightly less dense on distal edges of all fins (anterior apex of pectoral fins usually naked). Trunk denticles small, pungent, subconical, strongly raked posteriorly; anterior bases stellate, with clearly defined anterior ridges; similar in size along midline; well-separated, not imbricate; much larger on midline of trunk and tail than on pectoral and pelvic bases; no enlarged, thorn-like denticles on midline; pre- and postorbit with slightly enlarged denticles, much larger and more upright and strongly conical than those on dorsal midline of trunk and tail, more widely spaced than those above midorbit; similarly enlarged patches on postnarial snout. Adult male
<typeStatus id="54EA881E9C1AFFD5FB8BFF235CA9FF1A" box="[1058,1159,192,218]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
with partly embedded, malar thorn-like denticles along posteroanterior margin of pectoral fin; denticles slender, spear-shaped, directed posterolaterally or posteriorly; edge of pectoral and pelvic fin anterior margins with dense coverage of minute aspinous denticles (similar to those of ventral surface). Ventral surface largely naked; narrow longitudinal band extending along anterior margin of pectoral fin, sharply demarcated from rest of fin, greatest width slightly larger than orbit diameter; similar but less extensive band around apex of pelvic fin; post-pelvic tail mostly covered with denticles, commencing just anterior of free rear tip of pelvic fins and extending to caudal fin (in adult male
<typeStatus id="54EA881E9C1AFFD5FE15FE3B5A0AFE32" box="[444,548,472,498]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
, band better developed along pectoral and pelvic fins and tail, width of pectoral band about two eye lengths); denticles very small, dense, partially imbricate, aspinous, crowns circular with entire margins. Claspers of
<typeStatus id="54EA881E9C1AFFD5FD8AFDCB5AA6FD82" box="[547,648,552,578]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
largely naked, large ovoid patches on mid-ventral surface beside free rear tip of pelvic fin. Dorsal and caudal fins mostly with dense coverage of denticles (less so in
<typeStatus id="54EA881E9C1AFFD5FA9CFDB35DB5FDAA" box="[1333,1435,592,618]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
than adult male).
</paragraph>
<paragraph id="8BEE36BC9C1AFFD5FF6FFD435DB8FAF4" blockId="13.[151,1437,152,1892]" pageId="13" pageNumber="14">
Pectoral fins very large, expanded, base very fleshy, length 33.7 (33.6)% TL, 1.87 (1.77) times width; anterior margin almost straight to weakly convex, 1.39 (1.44) times inner margin; apex angular, angle slightly less than 120°; posterior margin slightly concave (almost straight in
<typeStatus id="54EA881E9C1AFFD5FC71FD095C6DFCC4" box="[984,1091,746,772]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
); inner margin very strongly convex, free rear tip broadly rounded. Pelvic fins large, elongate, length 23.2 (24.4)% TL, 1.45 (1.38) in pectoral-fin length; anterior margin convex, apex broadly rounded; posterior margin weakly convex; inner margin almost straight; free rear tip narrowly pointed (more bluntly pointed in
<typeStatus id="54EA881E9C1AFFD5FC74FCB95C69FCB4" box="[989,1095,858,884]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
), reaching origin of first dorsal fin; pelvic-fin origin anterior to pectoral-fin free rear tip. Claspers of adult male robust, elongate, expanded slightly subdistally, slightly depressed distally with narrowly rounded posterior margin, cartilage at glans opening not sharp-edged, outer margin 5.9% TL, 2.17 times base width. Dorsal fins similar in size and shape (first slightly larger in
<typeStatus id="54EA881E9C1AFFD5FE36FC135A24FBCA" box="[415,522,1008,1034]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
), strongly raked, close together, height of first 1.06 (1.17) times height of second; anterior margins straight to slightly convex; apices narrowly rounded to somewhat angular, well posterior to fin insertion; posterior margins convex (straight in
<typeStatus id="54EA881E9C1AFFD5FCCCFBD95BE1FB94" box="[869,975,1082,1108]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
), directed anteroventrally from top to bottom; inner margin almost parallel to dorsal surface, free rear tip obtuse, broadly rounded; posteriorly located, interdorsal space 1.63 (1.32) times first dorsal-fin base, 0.84 (0.79) times dorsal–caudal space. Caudal fin hypocercal, short, dorsal caudal margin 12.9 (15.2)% TL, 2.48 (2.06) in pelvic–caudal space, height 14.3 (17.0)% TL; lower lobe larger than upper lobe, preventral caudal margin 1.23 (1.06) times dorsal caudal margin; apices somewhat rounded (more angular in adult
<typeStatus id="54EA881E9C1AFFD5FCA4FB165B59FACF" box="[781,887,1269,1295]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
); dorsal caudal margin almost straight; preventral caudal margin slightly convex; postventral margin strongly notched, deepest slightly above its midlength.
</paragraph>
<paragraph id="8BEE36BC9C1AFFD5FF6FFAA3594EFABF" blockId="13.[151,1437,152,1892]" pageId="13" pageNumber="14">Vertebral counts: monospondylous centra 51 (50); precaudal centra 112 (113); caudal centra 32 (31); total centra 144 (144).</paragraph>
<paragraph id="8BEE36BC9C1AFFD5FF6CFA695CE2F934" blockId="13.[151,1437,152,1892]" pageId="13" pageNumber="14">
<emphasis id="B925EAAE9C1AFFD5FF6CFA69590CFA64" bold="true" box="[197,290,1418,1444]" pageId="13" pageNumber="14">Colour.</emphasis>
In preservative: Dorsal surface almost uniformly dark greyish brown; slightly darker over enlarged denticle patches, around eye and on snout; evidence of weak paler blotches on central trunk and tail. Ventral surface almost uniformly white, slightly darker on denticle band along anterior margin of pectoral fin; upper lip dusky; mouth dusky; spiracles pale. Pectoral and pelvic fins similar to body, outer margin pale, almost white; dorsal fins mostly greyish brown, posterior margins translucent; caudal fin similar to body, posterior margin pale to translucent. Adult male
<typeStatus id="54EA881E9C1AFFD5FD1AF9A65B36F99F" box="[691,792,1605,1631]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
(CSIRO H 6565–01) similar to
<typeStatus id="54EA881E9C1AFFD5FB35F9A65D2BF99F" box="[1180,1285,1605,1631]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, evidence of blackish subdorsal saddles; margins of pectoral and pelvic fins not white; denticle bands along pectoral and pelvic fins dark, somewhat blackish; darker areas on chin and behind gills; part of tail covered with denticles slightly darker than naked portion. Clasper dorsal surface bicoloured; anterior basal half white; posterior basal half and glans brownish; almost uniformly pale ventrally (denticulated area slightly darker).
</paragraph>
<paragraph id="8BEE36BC9C1AFFD5FF6FF8E35BA0F8A4" blockId="13.[151,1437,152,1892]" pageId="13" pageNumber="14">
Fresh coloration of adult male
<typeStatus id="54EA881E9C1AFFD5FD9BF8E35AB9F8DA" box="[562,663,1792,1818]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
similar, dark brownish subdorsal saddles much more evident. An additional male specimen (not retained) had similar general coloration but with large, dark brownish blotches and ocelli irregularly scattered over the dorsal surfaces (
<figureCitation id="136A2A399C1AFFD5FC80F8A95B5FF8A4" box="[809,881,1866,1892]" captionStart="FIGURE 4" captionStartId="14.[151,255,1951,1975]" captionTargetBox="[264,1317,207,1901]" captionTargetId="figure@14.[252,1311,731,1356]" captionTargetPageId="14" captionText="FIGURE 4. Squatina legnota sp. nov.: A, dorsal view of holotype (MZB 15500, female 471 mm TL); B, dorsal view of non-type specimen (adult male, 1341 mm TL); and C, ventral view of holotype." httpUri="https://zenodo.org/record/181437/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b).
</paragraph>
<caption id="DF2E66349C19FFD6FF3EF87C5BC4F81A" httpUri="https://zenodo.org/record/181437/files/figure.png" pageId="14" pageNumber="15" targetBox="[264,1317,207,1901]" targetPageId="14">
<paragraph id="8BEE36BC9C19FFD6FF3EF87C5BC4F81A" blockId="14.[151,1436,1951,2010]" pageId="14" pageNumber="15">
<emphasis id="B925EAAE9C19FFD6FF3EF87C5937F877" bold="true" box="[151,281,1951,1975]" pageId="14" pageNumber="15">FIGURE 4.</emphasis>
<taxonomicName id="4C514D3F9C19FFD6FE89F84359F8F877" box="[288,470,1952,1975]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="legnota" status="sp. nov.">
<emphasis id="B925EAAE9C19FFD6FE89F84359F8F877" box="[288,470,1952,1975]" italics="true" pageId="14" pageNumber="15">Squatina legnota</emphasis>
</taxonomicName>
<emphasis id="B925EAAE9C19FFD6FE75F87C5A1CF877" bold="true" box="[476,562,1951,1975]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A21657D59C19FFD6FE75F87C5A1CF877" box="[476,562,1951,1975]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, dorsal view of holotype (MZB 15500, female 471 mm TL); B, dorsal view of non-type specimen (adult male, 1341 mm TL); and C, ventral view of holotype.
</paragraph>
</caption>
<caption id="DF2E66349C18FFD7FF3EFDA65987FDBF" httpUri="https://zenodo.org/record/181438/files/figure.png" pageId="15" pageNumber="16" targetBox="[539,1034,222,549]" targetPageId="15">
<paragraph id="8BEE36BC9C18FFD7FF3EFDA65987FDBF" blockId="15.[151,1436,581,639]" pageId="15" pageNumber="16">
<emphasis id="B925EAAE9C18FFD7FF3EFDA6593CFD9D" bold="true" box="[151,274,581,605]" pageId="15" pageNumber="16">FIGURE 5</emphasis>
. Illustration of the anterior nasal flap of paratype of
<taxonomicName id="4C514D3F9C18FFD7FCEDFDA65BD7FD9C" box="[836,1017,581,604]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota" status="sp. nov.">
<emphasis id="B925EAAE9C18FFD7FCEDFDA65BD7FD9C" box="[836,1017,581,604]" italics="true" pageId="15" pageNumber="16">Squatina legnota</emphasis>
</taxonomicName>
<emphasis id="B925EAAE9C18FFD7FBA9FDA65C78FD9D" bold="true" box="[1024,1110,581,605]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A21657D59C18FFD7FBA9FDA65C78FD9D" box="[1024,1110,581,605]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CSIRO H 6565–01). Illustration by Stephanie Barnes.
</paragraph>
</caption>
<paragraph id="8BEE36BC9C18FFD7FF6CFD515BC4FD34" blockId="15.[151,1437,690,1596]" pageId="15" pageNumber="16">
<emphasis id="B925EAAE9C18FFD7FF6CFD5158D1FD0C" bold="true" box="[197,255,690,716]" pageId="15" pageNumber="16">Size.</emphasis>
Known from only four specimens, a female of
<quantity id="4CA99B599C18FFD7FCE2FD515B9EFD0C" box="[843,944,690,716]" metricMagnitude="-1" metricUnit="m" metricValue="4.71" pageId="15" pageNumber="16" unit="mm" value="471.0">471 mm</quantity>
TL (
<typeStatus id="54EA881E9C18FFD7FC58FD515C70FD0C" box="[1009,1118,690,716]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
), and three adult males of
<quantity id="4CA99B599C18FFD7FF3EFD39597EFD34" box="[151,336,730,756]" metricMagnitude="0" metricUnit="m" metricValue="1.2965" metricValueMax="1.341" metricValueMin="1.252" pageId="15" pageNumber="16" unit="mm" value="1296.5" valueMax="1341.0" valueMin="1252.0">
<date id="FFEF107C9C18FFD7FF3EFD395935FD34" box="[151,283,730,756]" pageId="15" pageNumber="16" value="1252" valueMax="1341">1252–1341</date>
mm
</quantity>
TL (the
<typeStatus id="54EA881E9C18FFD7FE1EFD395A32FD34" box="[439,540,730,756]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
and two other specimens not retained).
</paragraph>
</subSubSection>
<subSubSection id="C34B65379C18FFD7FF6FFCE15D46FCAC" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BEE36BC9C18FFD7FF6FFCE15D46FCAC" blockId="15.[151,1437,690,1596]" pageId="15" pageNumber="16">
<emphasis id="B925EAAE9C18FFD7FF6FFCE15949FCDC" bold="true" box="[198,359,770,796]" pageId="15" pageNumber="16">Distribution.</emphasis>
Occurs off southern
<collectingCountry id="F346762C9C18FFD7FDF6FCE15AFBFCDC" box="[607,725,770,796]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
. Observed or collected from 4 fish landing sites, Palabuhanratu in West Java (
<geoCoordinate id="EE65507B9C18FFD7FEDEFCC959F1FC84" box="[375,479,810,836]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-6.9833336">06°59' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C18FFD7FE47FCC95A48FC84" box="[494,614,810,836]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="106.53333">106°32' E</geoCoordinate>
), Cilacap in Central Java (
<geoCoordinate id="EE65507B9C18FFD7FC0EFCC95C20FC84" box="[935,1038,810,836]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-7.7333336">07°44' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C18FFD7FBB4FCC95CB8FC84" box="[1053,1174,810,836]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="109.0">109°00' E</geoCoordinate>
), Kedonganan in Bali (
<geoCoordinate id="EE65507B9C18FFD7FF09FCB1592BFCAC" box="[160,261,850,876]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-8.75">08°45' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C18FFD7FEBBFCB159A6FCAC" box="[274,392,850,876]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="115.166664">115°10' E</geoCoordinate>
), and Tanjung Luar in Lombok (
<geoCoordinate id="EE65507B9C18FFD7FCAFFCB15B42FCAC" box="[774,876,850,876]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-8.8">08°48' S</geoCoordinate>
,
<geoCoordinate id="EE65507B9C18FFD7FCD1FCB15BC0FCAC" box="[888,1006,850,876]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="116.46667">116°28' E</geoCoordinate>
). Depth information not known.
</paragraph>
</subSubSection>
<subSubSection id="C34B65379C18FFD7FF6CFC99595BF9FC" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BEE36BC9C18FFD7FF6CFC995A53FC7C" blockId="15.[151,1437,690,1596]" pageId="15" pageNumber="16">
<emphasis id="B925EAAE9C18FFD7FF6CFC99597CFC54" bold="true" box="[197,338,890,916]" pageId="15" pageNumber="16">Etymology.</emphasis>
Derived from Greek
<emphasis id="B925EAAE9C18FFD7FDF8FC985A9AFC54" box="[593,692,891,916]" italics="true" pageId="15" pageNumber="16">legnotos</emphasis>
meaning ‘having a coloured border’ with reference to the dark anterior edges of pectoral and pelvic fins.
</paragraph>
<paragraph id="8BEE36BC9C18FFD7FF6CFC29595BF9FC" blockId="15.[151,1437,690,1596]" pageId="15" pageNumber="16">
<emphasis id="B925EAAE9C18FFD7FF6CFC29595CFC24" bold="true" box="[197,370,970,996]" pageId="15" pageNumber="16">Comparisons.</emphasis>
<taxonomicName id="4C514D3F9C18FFD7FED2FC285A6CFC24" box="[379,578,971,996]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota">
<emphasis id="B925EAAE9C18FFD7FED2FC285A6CFC24" box="[379,578,971,996]" italics="true" pageId="15" pageNumber="16">Squatina legnota</emphasis>
</taxonomicName>
differs from all Australian angel sharks in having a simple anterior nasal flap with unfringed (rather than strongly fringed) barbels. It also has a less complex color pattern being uniformly coloured apart from its two darker subdorsal saddles (rather than covered with spots, flecks, blotches or ocelli). The scaled anterior ventral margin of the pectoral fin is black (otherwise pale or whitish). Four species of
<taxonomicName id="4C514D3F9C18FFD7FF42FB88597DFB44" box="[235,339,1131,1156]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B925EAAE9C18FFD7FF42FB88597DFB44" box="[235,339,1131,1156]" italics="true" pageId="15" pageNumber="16">Squatina</emphasis>
</taxonomicName>
occur in the western North Pacific:
<taxonomicName id="4C514D3F9C18FFD7FD50FB885C38FB44" authority="Shen &amp; Ting" authorityName="Shen &amp; Ting" box="[761,1046,1130,1156]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="formosa">
<emphasis id="B925EAAE9C18FFD7FD50FB885B58FB44" box="[761,886,1131,1156]" italics="true" pageId="15" pageNumber="16">S. formosa</emphasis>
Shen &amp; Ting
</taxonomicName>
,
<taxonomicName id="4C514D3F9C18FFD7FB8BFB885D27FB44" authority="Bleeker" authorityName="Bleeker" box="[1058,1289,1130,1156]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="japonica">
<emphasis id="B925EAAE9C18FFD7FB8BFB885C8BFB44" box="[1058,1189,1131,1156]" italics="true" pageId="15" pageNumber="16">S. japonica</emphasis>
Bleeker
</taxonomicName>
,
<taxonomicName id="4C514D3F9C18FFD7FABCFB8858CCFB6C" authority="Regan" authorityName="Regan" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nebulosa">
<emphasis id="B925EAAE9C18FFD7FABCFB885DB5FB44" box="[1301,1435,1131,1156]" italics="true" pageId="15" pageNumber="16">S. nebulosa</emphasis>
Regan
</taxonomicName>
and
<taxonomicName id="4C514D3F9C18FFD7FE88FB705B33FB6C" authority="Walsh &amp; Ebert, 2007" authorityName="Walsh &amp; Ebert" authorityYear="2007" box="[289,797,1170,1196]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="tergocellatoides">
<emphasis id="B925EAAE9C18FFD7FE88FB7059D3FB6C" box="[289,509,1171,1196]" italics="true" pageId="15" pageNumber="16">S. tergocellatoides</emphasis>
(
<bibRefCitation id="EFC04B4D9C18FFD7FDA6FB715B3BFB6C" author="Walsh" box="[527,789,1170,1196]" pageId="15" pageNumber="26" refString="Walsh, J. H. &amp; Ebert, D. A. (2007) A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae). Zootaxa, 1551, 31 - 47." type="journal article" year="2007">Walsh &amp; Ebert, 2007</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C514D3F9C18FFD7FC85FB705BD9FB6C" box="[812,1015,1171,1196]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota">
<emphasis id="B925EAAE9C18FFD7FC85FB705BD9FB6C" box="[812,1015,1171,1196]" italics="true" pageId="15" pageNumber="16">Squatina legnota</emphasis>
</taxonomicName>
does not possess a row of median scutes or spines.
<bibRefCitation id="EFC04B4D9C18FFD7FEC8FB595A16FB14" author="Compagno" box="[353,568,1210,1236]" pageId="15" pageNumber="25" refString="Compagno, L. J. V. (1984) FAO Species Catalogue. Vol. 4, Sharks of the World. An annotated and illustrated catalogue of shark species known to date. FAO Fisheries Synopsis No. 125. vol. 4, pt. 1 (noncarcharhinoids), pp. viii, 1 - 250, pt. 2 (Carcharhiniformes), pp. x, 251 - 655." type="journal article" year="1984">Compagno (1984)</bibRefCitation>
claimed that all 4 western North Pacific species have a row of spines on the midline of the back and tail, but according to
<bibRefCitation id="EFC04B4D9C18FFD7FD75FB015BCAFB3C" author="Walsh" box="[732,996,1250,1276]" pageId="15" pageNumber="26" refString="Walsh, J. H. &amp; Ebert, D. A. (2007) A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae). Zootaxa, 1551, 31 - 47." type="journal article" year="2007">Walsh &amp; Ebert (2007)</bibRefCitation>
, this state is only present in
<emphasis id="B925EAAE9C18FFD7FA9BFB0058ECFAE4" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C514D3F9C18FFD7FA9BFB005890FAE4" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="japonica">S. japonica</taxonomicName>
.
</emphasis>
The three remaining western North Pacific species have different colour patterns to
<emphasis id="B925EAAE9C18FFD7FB07FAE858D9FA8C" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C514D3F9C18FFD7FB07FAE85D09FAE4" box="[1198,1319,1291,1316]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota">S. legnota</taxonomicName>
.
<taxonomicName id="4C514D3F9C18FFD7FA9DFAE858D9FA8C" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="formosa">Squatina formosa</taxonomicName>
</emphasis>
and
<taxonomicName id="4C514D3F9C18FFD7FE87FAD0599BFA8C" box="[302,437,1331,1356]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nebulosa">
<emphasis id="B925EAAE9C18FFD7FE87FAD0599BFA8C" box="[302,437,1331,1356]" italics="true" pageId="15" pageNumber="16">S. nebulosa</emphasis>
</taxonomicName>
are both covered with black and white spots and appear to have fewer vertebrae (total central 137–139 and 138–139 respectively vs.
<quantity id="4CA99B599C18FFD7FD69FAB95B22FAB4" box="[704,780,1370,1396]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="15" pageNumber="16" unit="in" value="144.0">144 in</quantity>
<taxonomicName id="4C514D3F9C18FFD7FCBDFAB85BA4FAB4" box="[788,906,1371,1396]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota">
<emphasis id="B925EAAE9C18FFD7FCBDFAB85BA4FAB4" box="[788,906,1371,1396]" italics="true" pageId="15" pageNumber="16">S. legnota</emphasis>
</taxonomicName>
).
<emphasis id="B925EAAE9C18FFD7FC0DFAB85CF9FAB4" box="[932,1239,1371,1396]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C514D3F9C18FFD7FC0DFAB85CFDFAB4" box="[932,1235,1371,1396]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="tergocellatoides">Squatina tergocellatoides</taxonomicName>
,
</emphasis>
which is known from nearby Borneo (
<bibRefCitation id="EFC04B4D9C18FFD7FE3FFA615A49FA5C" author="Yano" box="[406,615,1410,1436]" pageId="15" pageNumber="26" refString="Yano, K., Ahmad, A., Gambang, A. C., Idris, A. H., Solahuddin, A. R. &amp; Aznan, Z. (2005) Sharks and Rays of Malaysia and Brunei Darussalam. SEAFDEC-MFRDMD / SP / 12. Kuala Terengganu. 557 pp." type="book" year="2005">
Yano
<emphasis id="B925EAAE9C18FFD7FE75FA605A36FA5C" box="[476,536,1411,1436]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
, 2005
</bibRefCitation>
), has large blackish spots on the pectoral fin and origins of the dorsal fins, but lacks distinct subdorsal saddles. Based on comments by
<bibRefCitation id="EFC04B4D9C18FFD7FC3AFA495CB7FA04" author="Walsh" box="[915,1177,1450,1476]" pageId="15" pageNumber="26" refString="Walsh, J. H. &amp; Ebert, D. A. (2007) A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae). Zootaxa, 1551, 31 - 47." type="journal article" year="2007">Walsh &amp; Ebert (2007)</bibRefCitation>
, its pelvic-fin tips and dorsal fin appear to be more widely spaced and it seems to have more ornate nasal barbels.
<taxonomicName id="4C514D3F9C18FFD7FB7DFA305DB2FA2C" box="[1236,1436,1491,1516]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="legnota">
<emphasis id="B925EAAE9C18FFD7FB7DFA305DB2FA2C" box="[1236,1436,1491,1516]" italics="true" pageId="15" pageNumber="16">Squatina legnota</emphasis>
</taxonomicName>
also possesses more vertebrae than the four species of
<taxonomicName id="4C514D3F9C18FFD7FC8CFA185BA3F9D4" box="[805,909,1531,1556]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B925EAAE9C18FFD7FC8CFA185BA3F9D4" box="[805,909,1531,1556]" italics="true" pageId="15" pageNumber="16">Squatina</emphasis>
</taxonomicName>
from the western North Pacific,
<emphasis id="B925EAAE9C18FFD7FAB0FA185D13F9D4" box="[1305,1341,1531,1556]" italics="true" pageId="15" pageNumber="16">i.e.</emphasis>
144 vs. &lt;140, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>
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