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<document ID-DOI="http://doi.org/10.5281/zenodo.5411924" ID-GBIF-Dataset="03a6dd3f-a52a-4d60-bc0d-944c00931276" ID-GBIF-Taxon="123195631" ID-Zenodo-Dep="5411924" approvalRequired="73" approvalRequired_for_bibRefs="41" approvalRequired_for_document="1" approvalRequired_for_matCits="6" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="3" checkinTime="1468511911299" checkinUser="plazi" docAuthor="Lien Thi Phuong Nguyen &amp; James M. Carpenter" docDate="2016" docId="03ECDE0DFFDE7741F2AC9C7B75E43F43" docLanguage="en" docName="entScience.12218.pdf" docOrigin="Entomological Science 2016" docStyle="DocumentStyle{}" docTitle="Zethus propodeus Nguyen &amp; Carpenter, sp. nov" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="8" masterDocId="FFD5A675FFD87746F23B995F74033D3E" masterDocTitle="Taxonomic review of the genus Zethus Fabricius (Hymenoptera: Vespidae: Eumeninae) from Vietnam with descriptions of four new species" masterLastPageNumber="9" masterPageNumber="1" pageId="6" pageNumber="7" updateTime="1643498771858" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic review of the genus Zethus Fabricius (Hymenoptera: Vespidae: Eumeninae) from Vietnam with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lien Thi Phuong Nguyen</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>James M. Carpenter</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Entomological Science</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>2016</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>9</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1111/ens.12218</mods:identifier>
<mods:identifier type="GBIF-Dataset">03a6dd3f-a52a-4d60-bc0d-944c00931276</mods:identifier>
<mods:identifier type="Zenodo-Dep">269807</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5411924" ID-GBIF-Taxon="123195631" ID-Zenodo-Dep="5411924" LSID="urn:lsid:plazi:treatment:03ECDE0DFFDE7741F2AC9C7B75E43F43" httpUri="http://treatment.plazi.org/id/03ECDE0DFFDE7741F2AC9C7B75E43F43" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<heading bold="true" fontSize="11" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" pageId="6" pageNumber="7">
<subSubSection box="[151,749,1316,1343]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph blockId="6.[151,758,1316,1479]" box="[151,749,1316,1343]" pageId="6" pageNumber="7">
<taxonomicName authority="Nguyen &amp; Carpenter" authorityName="Nguyen &amp; Carpenter" box="[151,646,1316,1343]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="propodeus" status="sp. nov">
<emphasis bold="true" box="[151,375,1316,1343]" italics="true" pageId="6" pageNumber="7">Zethus propodeus</emphasis>
Nguyen &amp; Carpenter
</taxonomicName>
,
<taxonomicNameLabel box="[659,749,1317,1343]" pageId="6" pageNumber="7" rank="species">sp. nov</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection box="[151,308,1348,1375]" pageId="6" pageNumber="7" type="description">
<paragraph blockId="6.[151,758,1316,1479]" box="[151,308,1348,1375]" pageId="6" pageNumber="7">
(
<figureCitation box="[159,299,1348,1375]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Figs
<number box="[218,299,1348,1375]" pageId="6" pageNumber="7" value="30.5" valueMax="32.0" valueMin="29.0">29–32</number>
</figureCitation>
)
</paragraph>
</subSubSection>
</emphasis>
</heading>
<subSubSection pageId="6" pageNumber="7" type="materials_examined">
<paragraph blockId="6.[151,758,1316,1479]" pageId="6" pageNumber="7">
<materialsCitation ID-GBIF-Occurrence="1275388306" collectingDate="2012-08-08" collectionCode="IEBR" collectorName="LTP Nguyen &amp; IEC-c" country="Vietnam" elevation="1000" latitude="22.541666" location="Pi Oac NR" longitude="105.8833" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus box="[151,251,1390,1414]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[151,251,1390,1414]" italics="true" pageId="6" pageNumber="7">Holotype</emphasis>
</typeStatus>
,
<specimenCount box="[270,289,1391,1414]" pageId="6" pageNumber="7" type="female">♀</specimenCount>
,
<emphasis bold="true" box="[309,323,1391,1414]" pageId="6" pageNumber="7">“</emphasis>
<collectingCountry box="[322,446,1390,1414]" name="Vietnam" pageId="6" pageNumber="7">VIETNAM</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03ECDE0DFFDE7741F2AC9C7B75E43F43:8E9A39C0FFDE7740F3F49C31765238B8" box="[463,593,1390,1414]" country="Vietnam" latitude="22.541666" name="Pi Oac NR" pageId="6" pageNumber="7">Pi Oac NR</location>
,
<location LSID="urn:lsid:plazi:treatment:03ECDE0DFFDE7741F2AC9C7B75E43F43:8E9A39C0FFDE7740F0599C3176F238B8" box="[610,753,1390,1414]" country="Vietnam" latitude="22.541666" name="Thanh Cong" pageId="6" pageNumber="7">Thanh Cong</location>
,
<location LSID="urn:lsid:plazi:treatment:03ECDE0DFFDE7741F2AC9C7B75E43F43:8E9A39C0FFDE7740F2AC9CD075253898" box="[151,294,1422,1447]" country="Vietnam" latitude="22.541666" name="Nguyen Binh" pageId="6" pageNumber="7">Nguyen Binh</location>
,
<location LSID="urn:lsid:plazi:treatment:03ECDE0DFFDE7741F2AC9C7B75E43F43:8E9A39C0FFDE7740F3099CD1759D3899" box="[306,414,1422,1447]" country="Vietnam" latitude="22.541666" name="Cao Bang" pageId="6" pageNumber="7">Cao Bang</location>
,
<geoCoordinate box="[425,537,1422,1446]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="92" value="22.541666">
<number box="[425,453,1422,1446]" pageId="6" pageNumber="7" value="22.0">22</number>
°
<number box="[461,510,1422,1446]" pageId="6" pageNumber="7" value="32.5">32.5</number>
<emphasis bold="true" box="[510,517,1423,1446]" pageId="6" pageNumber="7">’</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[544,622,1422,1446]" pageId="6" pageNumber="7">
<number box="[544,585,1422,1446]" pageId="6" pageNumber="7" value="105.0">105</number>
°
<number box="[594,622,1422,1446]" pageId="6" pageNumber="7" value="53.0">53</number>
</geoCoordinate>
<emphasis bold="true" box="[623,630,1423,1446]" pageId="6" pageNumber="7">’</emphasis>
,
<elevation box="[643,723,1422,1446]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="1000.0">
<quantity box="[643,723,1422,1446]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="1000.0">1000 m</quantity>
</elevation>
,
<collectingDate pageId="6" pageNumber="7" value="2012-08-08">
<date pageId="6" pageNumber="7" value="2012-08-08">
<number box="[737,751,1422,1446]" pageId="6" pageNumber="7" value="8.0">8</number>
. viii.
<number box="[192,249,1454,1478]" pageId="6" pageNumber="7" value="2012.0">2012</number>
</date>
</collectingDate>
,
<collectorName box="[259,393,1455,1479]" pageId="6" pageNumber="7">LTP Nguyen</collectorName>
&amp;
<collectorName box="[430,490,1454,1478]" pageId="6" pageNumber="7">IEC-c</collectorName>
<emphasis bold="true" box="[489,503,1455,1478]" pageId="6" pageNumber="7">”</emphasis>
[
<collectionCode box="[516,580,1455,1478]" pageId="6" pageNumber="7">IEBR</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="description">
<paragraph blockId="6.[151,758,1525,1910]" box="[151,288,1525,1551]" pageId="6" pageNumber="7">
<heading bold="true" box="[151,288,1525,1551]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[151,288,1525,1551]" italics="true" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[151,758,1525,1910]" lastBlockId="6.[810,1417,1150,1911]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[151,226,1567,1591]" italics="true" pageId="6" pageNumber="7">Female</emphasis>
.
<typeStatus box="[243,347,1566,1590]" pageId="6" pageNumber="7">Holotype</typeStatus>
. Body length
<quantity box="[499,574,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="mm" value="17.0">17 mm</quantity>
; forewing length
<quantity box="[151,248,1598,1622]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="6" pageNumber="7" unit="mm" value="14.5">14.5 mm</quantity>
. Structure as in
<taxonomicName box="[420,533,1599,1623]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis bold="true" box="[420,533,1599,1623]" italics="true" pageId="6" pageNumber="7">Z. dolosus</emphasis>
</taxonomicName>
and
<emphasis bold="true" pageId="6" pageNumber="7">
<taxonomicName box="[588,720,1599,1623]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="angulatus" status="sp. nov.">
<emphasis bold="true" box="[588,720,1599,1623]" italics="true" pageId="6" pageNumber="7">Z. angulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but differs as follows: body covered with coarser and more uniform punctures. Head in frontal view wider than high,
<number box="[262,296,1694,1718]" pageId="6" pageNumber="7" value="1.2">1.2</number>
× as wide as high (
<figureCitation box="[481,556,1694,1718]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number box="[526,556,1694,1718]" pageId="6" pageNumber="7" value="29.0">29</number>
</figureCitation>
). Inner eye margins in frontal view further apart from each other at vertex than at clypeus. Clypeus in frontal view
<number box="[593,627,1758,1782]" pageId="6" pageNumber="7" value="1.2">1.2</number>
× as wide as long, emarginated at basal margin and concave medially, apical margin nearly truncated (
<figureCitation box="[498,577,1822,1846]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number box="[547,577,1822,1846]" pageId="6" pageNumber="7" value="29.0">29</number>
</figureCitation>
). Mandible with four teeth, the third and fourth teeth truncated. FI
<number box="[703,716,1854,1878]" pageId="6" pageNumber="7" value="2.0">2</number>
× as long as its maximum width, FII
<emphasis bold="true" box="[518,530,1887,1910]" pageId="6" pageNumber="7">–</emphasis>
III as long as wide, FIV
<emphasis bold="true" box="[850,862,1151,1174]" pageId="6" pageNumber="7">–</emphasis>
IX wider than long, FX bullet-shaped, as long as its basal width. Propodeum (
<figureCitation box="[1094,1172,1182,1206]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number box="[1144,1172,1182,1206]" pageId="6" pageNumber="7" value="30.0">30</number>
</figureCitation>
) rugose, dull, without lateral carina, with
<number box="[1021,1061,1214,1238]" pageId="6" pageNumber="7" value="4.5" valueMax="5.0" valueMin="4.0">
4
<emphasis bold="true" box="[1035,1047,1215,1238]" pageId="6" pageNumber="7">–</emphasis>
5
</number>
oblique striae along each side of medial carina, lateral side with small and sparse punctures, without striae, submarginal carina produced into round and short lamella above propodeal valvulae. Metasomal segment I proportionally shorter than in
<taxonomicName class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis bold="true" italics="true" pageId="6" pageNumber="7">Z. dolosus</emphasis>
</taxonomicName>
,
<number box="[908,942,1374,1398]" pageId="6" pageNumber="7" value="2.6">2.6</number>
× as long as wide, in dorsal view gradually widening from one-fourth of the distance from base to basal half, then gradually narrowly toward apical margin (
<figureCitation box="[818,893,1470,1494]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number box="[864,893,1470,1494]" pageId="6" pageNumber="7" value="31.0">31</number>
</figureCitation>
), with maximum width
<number box="[1150,1184,1470,1494]" pageId="6" pageNumber="7" value="2.6">2.6</number>
× its basal width, with a medial carina which runs from basal margin to onefourth the length of the tergum, swollen part
<number box="[1289,1323,1534,1558]" pageId="6" pageNumber="7" value="2.1">2.1</number>
× as long as wide in dorsal view, tergum and sternum fused, area between two sides with undefined border, with small and clearly defined punctures, in lateral view dorsal and ventral margin not parallel. Metasomal segment II with rather long petiole at base (Figs 31,32), TII in lateral view slightly concave near base, then gradually convex to near apical margin, slightly depressed before apical margin, SII in lateral view gradually convex from base to apical margin (
<figureCitation box="[900,976,1822,1846]" captionStart="Figures 29 – 32" captionStartId="6.[810,878,996,1018]" captionTargetBox="[821,1405,231,983]" captionTargetId="figure@6.[819,1407,229,986]" captionTargetPageId="6" captionText="Figures 29 – 32 Zethus propodeus sp. nov., holotype ♀. 29 Head, frontal view. 30 Propodeum, dorsal view. 31 TI &amp; II, dorsal view. 32 Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269813/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number box="[946,976,1822,1846]" pageId="6" pageNumber="7" value="32.0">32</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[810,1417,1150,1911]" box="[836,1323,1854,1879]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[836,901,1855,1879]" italics="true" pageId="6" pageNumber="7">Color.</emphasis>
Black. Propodeal valvulae dark brown.
</paragraph>
</subSubSection>
<subSubSection box="[836,1194,1886,1911]" pageId="6" pageNumber="7" type="distribution">
<paragraph blockId="6.[810,1417,1150,1911]" box="[836,1194,1886,1911]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[836,966,1887,1911]" italics="true" pageId="6" pageNumber="7">Distribution</emphasis>
.
<collectingCountry box="[979,1072,1886,1910]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
: Cao Bang.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="discussion">
<paragraph blockId="7.[172,778,229,637]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[197,302,229,253]" italics="true" pageId="7" pageNumber="8">Remarks.</emphasis>
This species can be distinguished from the similar species
<taxonomicName authority="Gusenleitner, 2010" authorityName="Gusenleitner" authorityYear="2010" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis bold="true" box="[405,616,261,285]" italics="true" pageId="7" pageNumber="8">Zethus malayanus</emphasis>
<bibRefCitation author="Gusenleitner J" journalOrPublisher="Linzer Biologische Beitrage" pageId="7" pageNumber="8" pagination="695 - 709" part="42" refString="Gusenleitner J (2010) Bemerkenswerte Faltenwespen-Funde aus der orientalischen Region Teil 5 (Hymenoptera: Vespidae, Eumeninae). Linzer Biologische Beitrage 42, 695 - 709." title="Bemerkenswerte Faltenwespen-Funde aus der orientalischen Region Teil 5 (Hymenoptera: Vespidae, Eumeninae)" type="journal article" year="2010">
Gusenleitner,
<number box="[172,229,293,317]" pageId="7" pageNumber="8" value="2010.0">2010</number>
</bibRefCitation>
</taxonomicName>
and other known species of the genus by the combination of the following characters: body covered with rather uniform punctures and long hairs, metasomal segment I in lateral view with dorsal and ventral margin not parallel, propodeum rugose, without lateral carinae, submarginal carina produced into short and round lamella above propodeal valvulae, metasomal segment II with rather long petiole at base.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="etymology">
<paragraph blockId="7.[172,778,229,637]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[197,319,581,605]" italics="true" pageId="7" pageNumber="8">Etymology.</emphasis>
The specific name refers to the propodeum with its diagnostic characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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