297 lines
59 KiB
XML
297 lines
59 KiB
XML
<document id="078ACD841EF5EF9AEB798E6E7CDF5311" ID-CLB-Dataset="26653" ID-DOI="10.11646/zootaxa.4612.2.4" ID-GBIF-Dataset="099a649b-9b80-4d9d-bc71-441b4f6d603c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3234300" ID-ZooBank="7525A835-B189-4F33-B17B-59D722E50951" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559107549387" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03EC517274472453BBEFFA2DB711FBC1" docLanguage="en" docName="zootaxa.4612.2.4.pdf" docOrigin="Zootaxa 4612 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bicarinulodes Jałoszyński 2019, gen. n." docType="treatment" docVersion="10" lastPageNumber="217" masterDocId="FFD5290A744E245FBB78FF84B611FFD2" masterDocTitle="Taxonomy of the ' Afroeudesis group' of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="220" masterPageNumber="205" pageNumber="214" updateTime="1698732260275" updateUser="ExternalLinkService">
|
||
<mods:mods id="44D19D04A8A0259BC686E4813DFD53CA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0E0A1A2A8ED7585AF48E34ACF6702D45">
|
||
<mods:title id="95BC4E1532F7058320C9F947FCEF3FAC">Taxonomy of the ' Afroeudesis group' of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="98F6D19C661A0D48CA3095C5BA761335" type="personal">
|
||
<mods:role id="E8E857F5828B19619054FB12C7E08B85">
|
||
<mods:roleTerm id="440A41DB189BA597DF836F19CEE445F6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CC0CD01879136099B1F95A7EA243529B">Jałoszyński, Paweł</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="20EBC699B4C200154B279DC4E23EE00E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0CE0936D5DDB4F2CA965156B4BBBA256" type="host">
|
||
<mods:titleInfo id="B80F5749752032D1E988A2B881A7B582">
|
||
<mods:title id="8E1E7F8172149BB6FF324542B9737CC4">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="5B1FFB8E42ABE1B8F3E58F6C34CA871C">
|
||
<mods:date id="A87C7A857436D0FC561F065DDBD0D2EC">2019</mods:date>
|
||
<mods:detail id="0DC7871F7485C53FA108CC399C91E201" type="pubDate">
|
||
<mods:number id="FC9BCB3BFF748BEE2C8549906D6FEFE1">2019-05-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="8A003DF85A616A12A4AF6958F1B04481" type="volume">
|
||
<mods:number id="DAE981C3F4488F371EFE60DEC8800D97">4612</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="24CD42065E7472BDA2B1EB8580096136" type="issue">
|
||
<mods:number id="A673F2515710154E613D5848FF700C7B">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9864EB24CBFDE464EACEC543FBDF9957" unit="page">
|
||
<mods:start id="D7AF82BF51BE0A10621E6BC9B5E786D6">205</mods:start>
|
||
<mods:end id="F8D96A898760477B1239EDC6CDE00764">220</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="843E4C0D45A314D1CFE72B70771BB8F0">journal article</mods:classification>
|
||
<mods:identifier id="582F5A9144DAA57F3FD64685B300B2FF" type="CLB-Dataset">26653</mods:identifier>
|
||
<mods:identifier id="909650BF1A419C0271C11EA18676F6F2" type="DOI">10.11646/zootaxa.4612.2.4</mods:identifier>
|
||
<mods:identifier id="DEE71BF30ABEACB54029C0664D2AC71F" type="GBIF-Dataset">099a649b-9b80-4d9d-bc71-441b4f6d603c</mods:identifier>
|
||
<mods:identifier id="F0AE491D36412B4D8B60F7796792A992" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="5DFE21500C5282F3B5F8719A81C6E375" type="Zenodo-Dep">3234300</mods:identifier>
|
||
<mods:identifier id="9C92465FCF6EE53CBFDFA7549A890A85" type="ZooBank">7525A835-B189-4F33-B17B-59D722E50951</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EC517274472453BBEFFA2DB711FBC1" ID-GBIF-Taxon="157293831" LSID="urn:lsid:plazi:treatment:03EC517274472453BBEFFA2DB711FBC1" httpUri="http://treatment.plazi.org/id/03EC517274472453BBEFFA2DB711FBC1" lastPageId="12" lastPageNumber="217" pageId="9" pageNumber="214">
|
||
<subSubSection id="C35FB3EF74472456BBEFFA2DB78AFA11" box="[151,411,1449,1476]" pageId="9" pageNumber="214" type="nomenclature">
|
||
<paragraph id="8BFAE06474472456BBEFFA2DB78AFA11" blockId="9.[151,411,1449,1476]" box="[151,411,1449,1476]" pageId="9" pageNumber="214">
|
||
<heading id="D0B2570874472456BBEFFA2DB78AFA11" bold="true" box="[151,411,1449,1476]" fontSize="11" level="1" pageId="9" pageNumber="214" reason="1">
|
||
<emphasis id="B9313C7674472456BBEFFA2DB78AFA11" bold="true" box="[151,411,1449,1476]" pageId="9" pageNumber="214">
|
||
<taxonomicName id="4C459BE774472456BBEFFA2DB750FA16" ID-CoL="8XK2L" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,321,1449,1476]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="214" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9313C7674472456BBEFFA2DB750FA16" bold="true" box="[151,321,1449,1476]" italics="true" pageId="9" pageNumber="214">Bicarinulodes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202810D74472456BA30FA2DB78AFA11" box="[328,411,1449,1475]" pageId="9" pageNumber="214" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FB3EF74472456BBEFFA6AB578F9D7" box="[151,873,1516,1542]" pageId="9" pageNumber="214" type="materials_examined">
|
||
<paragraph id="8BFAE06474472456BBEFFA6AB578F9D7" blockId="9.[151,873,1516,1542]" box="[151,873,1516,1542]" pageId="9" pageNumber="214">
|
||
<typeStatus id="54FE5EC674472456BBEFFA6AB6D8F9D4" box="[151,201,1518,1542]" pageId="9" pageNumber="214">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C459BE774472456BA5EFA69B4BCF9D7" authority="Franz, 1988" authorityName="Franz" authorityYear="1988" box="[294,685,1517,1541]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="214" phylum="Arthropoda" rank="species" species="meridensis">
|
||
<emphasis id="B9313C7674472456BA5EFA69B43CF9D7" box="[294,557,1517,1541]" italics="true" pageId="9" pageNumber="214">Microscydmus meridensis</emphasis>
|
||
<bibRefCitation id="EFD49D9574472456B94CFA6AB4BCF9D7" author="Franz, H." box="[564,685,1517,1541]" pageId="9" pageNumber="214" pagination="9 - 84" refId="ref7458" refString="Franz, H. (1988) Nachtrag zur monographie der Scydmaeniden Venezuelas (Coleoptera, Scydmaenidae). Elytron, 1, 9 - 84." type="journal article" year="1988">Franz, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
(here designated).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FB3EF74472455BBEFF9B7B515FE59" lastPageId="10" lastPageNumber="215" pageId="9" pageNumber="214" type="diagnosis">
|
||
<paragraph id="8BFAE06474472455BBEFF9B7B515FE59" blockId="9.[151,1436,1587,2010]" lastBlockId="10.[151,1437,153,395]" lastPageId="10" lastPageNumber="215" pageId="9" pageNumber="214">
|
||
<emphasis id="B9313C7674472456BBEFF9B7B703F99C" bold="true" box="[151,274,1587,1614]" pageId="9" pageNumber="214">Diagnosis.</emphasis>
|
||
Body (Fig. 4) moderately stout, distinctly constricted between head and prothorax and between prothorax and elytra. Head (
|
||
<figureCitation id="137EFCE174472456BAF7F9D3B7F3F9A0" box="[399,482,1623,1650]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="9" pageNumber="214">Fig. 31</figureCitation>
|
||
) about as long as broad with short but distinct tempora; tempora, vertex, genae and postgenae lacking thick bristles; occipital constriction about as broad as vertex; submentum lacking lateral sutures; hypostomal ridges present and complete, obliquely running toward posterior tentorial pits where they are not connected; antennae with trimerous club and modified in males (with antennomere VII enlarged and projecting mesad); frons with shallow frontal impression; vertex lacking posteromedian impression; head lacking longitudinal groove and median subtriangular 'platform'; setae on frons and vertex largely symmetrical; maxillary palpomere IV slightly constricted before apex, so that its apical portion is broadened and truncate; pronotum (
|
||
<figureCitation id="137EFCE174472456BF14F8ABB2AFF898" box="[1132,1214,1839,1866]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="9" pageNumber="214">Fig. 31</figureCitation>
|
||
) broadest distinctly in front of middle, with unusual set of antebasal structures: two broad submedian longitudinal carinulae and two pairs of small lateral pits; lateral and sublateral pronotal carinae absent; thick bristles on sides of pronotum absent; prosternum (
|
||
<figureCitation id="137EFCE174472456BA5DF81FB764F864" box="[293,373,1947,1974]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="9" pageNumber="214">Fig. 33</figureCitation>
|
||
) with basisternal part vestigial; prosternal intercoxal process developed as a sharply marked but weakly elevated carina; procoxal cavities closed; notosternal sutures and hypomeral ridges complete; mesoscutel- lum (
|
||
<figureCitation id="137EFCE174442455BBACFF1DB736FF61" box="[212,295,153,179]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
) largely exposed between elytral bases; mesocoxal rests with diffuse marginal carinae; mesoventral intercoxal process (
|
||
<figureCitation id="137EFCE174442455BA0CFF39B7D6FF05" box="[372,455,188,215]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="10" pageNumber="215">Fig. 35</figureCitation>
|
||
) carinate and strongly elevated, narrowly separating mesocoxae, with broadened, flattened and smooth posterior portion, well-defined posterior tip and a narrow posteromedian notch; metaventral carinae present; anterior metaventral process present, well-defined, large, not divided into subrectangular anterior and triangular posterior portions, broadly subtriangular; metaventral intercoxal process (
|
||
<figureCitation id="137EFCE174442455BF17FEADB2D1FE91" box="[1135,1216,296,323]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="10" pageNumber="215">Fig. 35</figureCitation>
|
||
) with a pair of long spines separated by a long and narrow notch; each elytron (
|
||
<figureCitation id="137EFCE174442455B852FEC9B56BFEB5" box="[810,890,333,359]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
) with two vestigial asetose foveae; and aedeagus (
|
||
<figureCitation id="137EFCE174442455BBE7FEF5B70EFE59" box="[159,287,369,395]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="10" pageNumber="215">Figs 37–40</figureCitation>
|
||
) symmetrical with free, slender parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3AB0EC74442455BBEFFA84B7B9FAAA" httpUri="https://zenodo.org/record/3234308/files/figure.png" pageId="10" pageNumber="215" startId="10.[151,264,1280,1305]" targetBox="[151,1424,405,1268]" targetPageId="10">
|
||
<paragraph id="8BFAE06474442455BBEFFA84B7B9FAAA" blockId="10.[151,1437,1280,1401]" pageId="10" pageNumber="215">
|
||
<emphasis id="B9313C7674442455BBEFFA84B747FACB" bold="true" box="[151,342,1280,1305]" pageId="10" pageNumber="215">FIGURES 26–30.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C459BE774442455B9FDFA85B58CFACA" authority="Franz" authorityName="Franz" authorityYear="1980" box="[645,925,1281,1305]" class="Insecta" family="Staphylinidae" genus="Pseudoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="215" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B9313C7674442455B9FDFA85B548FACB" box="[645,857,1281,1305]" italics="true" pageId="10" pageNumber="215">Pseudoraphes minor</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
, male paratype. Pterothorax in ventral view (26); aedeagus in ventral (27, 29) and lateral (28, 30) views. Abbreviations: amvp, anterior metaventral process; mfa, median flattened area; mscr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35FB3EF74442453BBBFFA27B5A4FD21" lastPageId="12" lastPageNumber="217" pageId="10" pageNumber="215" type="description">
|
||
<paragraph id="8BFAE06474442455BBBFFA27B55FFA33" blockId="10.[151,1437,1443,2010]" pageId="10" pageNumber="215">
|
||
<emphasis id="B9313C7674442455BBBFFA27B748FA6C" bold="true" box="[199,345,1443,1470]" pageId="10" pageNumber="215">Description.</emphasis>
|
||
Body (Fig. 4) elongate but not very slender, distinctly constricted between head and prothorax and less so between prothorax and elytra, strongly convex, brown.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474442455BBBFFA6FB205F90F" blockId="10.[151,1437,1443,2010]" pageId="10" pageNumber="215">
|
||
Head (
|
||
<figureCitation id="137EFCE174442455BA6BFA6FB780F9D4" box="[275,401,1515,1542]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31, 33</figureCitation>
|
||
) with moderately large eyes; tempora short but distinct, rounded; frons impressed between antennal insertions, lacking longitudinal groove; vertex anteriorly confluent with frons, lacking posteromedian impression; numerous setae on frons and vertex largely symmetrical; gular plate (
|
||
<figureCitation id="137EFCE174442455B887F9B7B243F99F" box="[1023,1106,1587,1613]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 33</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BF24F9B0B269F99F" box="[1116,1144,1588,1613]" italics="true" pageId="10" pageNumber="215">gp</emphasis>
|
||
) subtrapezoidal; posterior tentorial pits (
|
||
<figureCitation id="137EFCE174442455BA49F9D3B792F9A3" box="[305,387,1623,1649]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 33</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BAF6F9DCB7A3F9A3" box="[398,434,1624,1649]" italics="true" pageId="10" pageNumber="215">ptp</emphasis>
|
||
) in front of transverse impression demarcating 'neck' region ventrally, minute; hypostomal ridges (
|
||
<figureCitation id="137EFCE174442455BB95F9FFB750F947" box="[237,321,1659,1685]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 33</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BA37F9FFB779F947" box="[335,360,1659,1685]" italics="true" pageId="10" pageNumber="215">hr</emphasis>
|
||
) complete, reaching level of posterior tentorial pits, not connected; submentum (
|
||
<figureCitation id="137EFCE174442455BE7BF9FFB349F947" box="[1283,1368,1659,1685]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 33</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BE1EF9F8B382F947" box="[1382,1427,1660,1685]" italics="true" pageId="10" pageNumber="215">smn</emphasis>
|
||
) lacking lateral sutures. Antennae (Figs 4, 31–32) slender, with distinct trimerous club, in males modified, with antennomere VII (
|
||
<figureCitation id="137EFCE174442455BA32F947B78CF90F" box="[330,413,1731,1757]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 32</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BAD1F940B7D4F90C" box="[425,453,1732,1758]" italics="true" pageId="10" pageNumber="215">a7</emphasis>
|
||
) enlarged and with a mesal subtriangular projection.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474442455BBBFF96CB34FF80B" blockId="10.[151,1437,1443,2010]" pageId="10" pageNumber="215">
|
||
Pronotum (
|
||
<figureCitation id="137EFCE174442455BA3EF963B789F8D0" box="[326,408,1767,1794]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
) bell-shaped, broadest anterior to middle; lacking lateral and sublateral carinae, with a pair of longitudinal, broad and flattened carinulae (
|
||
<figureCitation id="137EFCE174442455B9DDF88FB4E8F8F7" box="[677,761,1803,1829]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455B87DF888B53BF8F7" box="[773,810,1804,1829]" italics="true" pageId="10" pageNumber="215">car</emphasis>
|
||
) shortly in front of basal margin, and with two pairs of small antebasal lateral pits: inner (
|
||
<figureCitation id="137EFCE174442455B966F8ABB462F89B" box="[542,627,1839,1865]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455B9F8F8B4B487F89B" box="[640,662,1840,1865]" italics="true" pageId="10" pageNumber="215">ip</emphasis>
|
||
) and outer (
|
||
<figureCitation id="137EFCE174442455B85BF8ABB566F89B" box="[803,887,1839,1865]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 31</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455B8FCF8B4B5B1F89B" box="[900,928,1840,1865]" italics="true" pageId="10" pageNumber="215">op</emphasis>
|
||
); transverse antebasal groove distinct, broad and diffuse, divided by carinulae into three impressions. Prosternum with vestigial basisternal region (
|
||
<figureCitation id="137EFCE174442455BE6DF8D7B386F8BF" box="[1301,1431,1875,1901]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Figs 33–34</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BBEFF8F3B6A9F843" box="[151,184,1911,1937]" italics="true" pageId="10" pageNumber="215">bst</emphasis>
|
||
), prosternal process carinate but weakly elevated (
|
||
<figureCitation id="137EFCE174442455B996F8F3B550F843" box="[750,833,1911,1937]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 34</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455B835F8FCB565F843" box="[845,884,1912,1937]" italics="true" pageId="10" pageNumber="215">psp</emphasis>
|
||
); notosternal sutures (
|
||
<figureCitation id="137EFCE174442455BF15F8F3B2FFF843" box="[1133,1262,1911,1937]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Figs 33–34</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BF82F8FCB30FF843" box="[1274,1310,1912,1937]" italics="true" pageId="10" pageNumber="215">nss</emphasis>
|
||
) complete; hypomeral ridges (
|
||
<figureCitation id="137EFCE174442455BA1FF81FB7F8F867" box="[359,489,1947,1973]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Figs 33–34</figureCitation>
|
||
:
|
||
<emphasis id="B9313C7674442455BA8AF81FB406F867" box="[498,535,1947,1973]" italics="true" pageId="10" pageNumber="215">hyr</emphasis>
|
||
) complete; inner (adcoxal) portion of hypomeron relatively narrow and glabrous, outer portion (confluent with side of pronotum) lacking thick bristles. Procoxal cavities (
|
||
<figureCitation id="137EFCE174442455BF01F83BB2DDF80B" box="[1145,1228,1983,2009]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="10" pageNumber="215">Fig. 34</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674442455BFA0F844B2EEF80B" box="[1240,1279,1984,2009]" italics="true" pageId="10" pageNumber="215">pcc</emphasis>
|
||
) closed.
|
||
</paragraph>
|
||
<caption id="DF3AB0EC74452454BBEFF8A8B43EF837" httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="11" pageNumber="216" startId="11.[151,264,1836,1861]" targetBox="[151,1436,206,1783]" targetPageId="11">
|
||
<paragraph id="8BFAE06474452454BBEFF8A8B43EF837" blockId="11.[151,1437,1836,2021]" pageId="11" pageNumber="216">
|
||
<emphasis id="B9313C7674452454BBEFF8A8B742F897" bold="true" box="[151,339,1836,1861]" pageId="11" pageNumber="216">FIGURES 31–34.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C459BE774452454B90EF8A9B5D9F897" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2019" baseAuthorityName="Jałoszyński" baseAuthorityYear="2019" box="[630,968,1837,1861]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="216" phylum="Arthropoda" rank="species" species="meridensis" status="comb. nov.">
|
||
<emphasis id="B9313C7674452454B90EF8A9B569F897" box="[630,888,1837,1861]" italics="true" pageId="11" pageNumber="216">Bicarinulodes meridensis</emphasis>
|
||
(Franz)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202810D74452454B8B5F8A9B239F897" box="[973,1064,1837,1861]" pageId="11" pageNumber="216" rank="species">
|
||
<emphasis id="B9313C7674452454B8B5F8A9B239F897" bold="true" box="[973,1064,1837,1861]" pageId="11" pageNumber="216">comb. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFAE06474422453BBBFFF1DB6D4FE91" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
Mesoventrite with a pair of transverse impressions behind its anterior ridge that function as procoxal rests (
|
||
<figureCitation id="137EFCE174422453BE12FF1DB6F1FF05" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Figs 35–36</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BB96FF3AB702FF05" box="[238,275,190,215]" italics="true" pageId="12" pageNumber="217">pcr</emphasis>
|
||
); mesoventral intercoxal process (
|
||
<figureCitation id="137EFCE174422453B9E6FF39B533FF05" box="[670,802,188,215]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Figs 35–36</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453B848FF3AB57BFF05" box="[816,874,190,215]" italics="true" pageId="12" pageNumber="217">msvp</emphasis>
|
||
) carinate, long, strongly elevated, with distinctly broadened posterior portion bearing a median flattened area (
|
||
<figureCitation id="137EFCE174422453B845FF65B59EFF29" box="[829,911,224,251]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Fig. 35</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453B8E2FF65B5D5FF29" box="[922,964,225,251]" italics="true" pageId="12" pageNumber="217">mfa</emphasis>
|
||
) that is smooth and devoid of setae, posterior tip of process with subtriangular emargination; mesoventrite at each side with deep dorsolateral fovea (
|
||
<figureCitation id="137EFCE174422453BE3CFE81B386FECD" box="[1348,1431,261,287]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Fig. 36</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BBEFFEADB6A4FE91" box="[151,181,297,323]" italics="true" pageId="12" pageNumber="217">dlf</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFEC9B6CEFE59" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
Mesoscutellum (
|
||
<figureCitation id="137EFCE174422453BAFBFEC9B7C9FEB5" box="[387,472,333,359]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="12" pageNumber="217">Fig. 31</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BA9DFECAB400FEB5" box="[485,529,334,359]" italics="true" pageId="12" pageNumber="217">msc</emphasis>
|
||
) exposed between elytral bases in intact beetles, subtriangular, about as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFE11B2B9FDC9" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
Metaventrite with massive subtrapezoidal anterior metaventral process (
|
||
<figureCitation id="137EFCE174422453B89EFE11B268FE7D" box="[998,1145,404,431]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Figs 35–36B</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BFFCFE12B2ADFE7D" box="[1156,1212,406,431]" italics="true" pageId="12" pageNumber="217">amtp</emphasis>
|
||
); each side of metaventrite with metaventral carina (
|
||
<figureCitation id="137EFCE174422453B968FE3DB485FE01" box="[528,660,440,467]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Figs 35–36</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453B9D9FE3EB4C7FE01" box="[673,726,442,467]" italics="true" pageId="12" pageNumber="217">mtvc</emphasis>
|
||
), carinae are parallel to each other; mesocoxal rests very deep and filled with setae, their posterior marginal carina diffuse but complete, postmarginal carina absent; metaventral intercoxal process (
|
||
<figureCitation id="137EFCE174422453BA09FD85B7D4FDC9" box="[369,453,512,539]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Fig. 35</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BAA9FD86B416FDC9" box="[465,519,514,539]" italics="true" pageId="12" pageNumber="217">mtvp</emphasis>
|
||
) with long lateral spines, indistinctly separating metacoxae.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFDA1B212FDB1" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
Elytra (Figs 4, 31) oval, each elytron with one pair of vestigial, asetose basal elytral foveae (
|
||
<figureCitation id="137EFCE174422453BFB4FDA1B30EFDED" box="[1228,1311,549,575]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1836,1861]" captionTargetBox="[151,1436,206,1783]" captionTargetId="figure@11.[151,1436,206,1783]" captionTargetPageId="11" captionText="FIGURES 31–34. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Head, prothorax and elytral base in dorsal view (31); antennomeres VI–VIII of right antenna in ventral view (32); head and prothorax in ventral view (33); prothorax in ventral view (34). Abbreviations: a6–8, antennomere 6–8; bef, basal elytral fovea; bst, basisternal portion of prosternum; car, carinulae; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ip, inner pit; md, mandible; mn, mentum; msc, mesoscutellum; nss, notosternal suture; op, outer pit; pcc, procoxal cavity; pcr, procoxal rest; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/3234310/files/figure.png" pageId="12" pageNumber="217">Fig. 31</figureCitation>
|
||
;
|
||
<emphasis id="B9313C7674422453BE53FDA1B35CFDED" box="[1323,1357,549,575]" italics="true" pageId="12" pageNumber="217">bef</emphasis>
|
||
), lacking basal impression, and with distinct humeral callus. Hind wings developed.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFDE9B207FD55" blockId="12.[151,1437,153,1043]" box="[199,1046,621,647]" pageId="12" pageNumber="217">Legs do not differ from other genera of Glandulariini, unmodified in males.</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFD15B5A5FD79" blockId="12.[151,1437,153,1043]" box="[199,948,657,683]" pageId="12" pageNumber="217">Abdomen unmodified, with sternites III and IV subequal in length.</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFD31B5A4FD21" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
Aedeagus (
|
||
<figureCitation id="137EFCE174422453BA3BFD31B7D3FD1D" box="[323,450,693,719]" captionStart="FIGURES 35–40" captionStartId="12.[151,264,1885,1910]" captionTargetBox="[151,1436,1068,1850]" captionTargetId="figure@12.[151,1436,1068,1850]" captionTargetPageId="12" captionText="FIGURES 35–40. Morphological structures of Bicarinulodes meridensis (Franz) comb. n., male holotype. Pterothorax in ventral view (35–36); aedeagus in ventral (37, 39) and lateral (38, 40) views. Abbreviations: amtp, anterior metaventral process; dlf, dorsolateral fovea; mfa, median flattened area; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/3234312/files/figure.png" pageId="12" pageNumber="217">Figs 37–40</figureCitation>
|
||
) slender, weakly sclerotized, with symmetrical median lobe and symmetrical, but weakly sclerotized endophallic structures; parameres free, bearing apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FB3EF74422453BBBFFD79B34EFCC5" box="[199,1375,765,791]" pageId="12" pageNumber="217" type="etymology">
|
||
<paragraph id="8BFAE06474422453BBBFFD79B34EFCC5" blockId="12.[151,1437,153,1043]" box="[199,1375,765,791]" pageId="12" pageNumber="217">
|
||
<emphasis id="B9313C7674422453BBBFFD79B75FFCC5" bold="true" box="[199,334,765,791]" pageId="12" pageNumber="217">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C459BE774422453BAB2FD79B476FCC5" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[458,615,765,791]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="214" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9313C7674422453BAB2FD79B476FCC5" box="[458,615,765,791]" italics="true" pageId="12" pageNumber="217">Bicarinulodes</emphasis>
|
||
</taxonomicName>
|
||
reflects the unique antebasal pronotal carinulae. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FB3EF74422453BBBFFCA5B711FBC1" pageId="12" pageNumber="217" type="discussion">
|
||
<paragraph id="8BFAE06474422453BBBFFCA5B22EFC19" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
<emphasis id="B9313C7674422453BBBFFCA5B72AFCE9" bold="true" box="[199,315,801,827]" pageId="12" pageNumber="217">Remarks.</emphasis>
|
||
<taxonomicName id="4C459BE774422453BA3BFCA5B7F1FCE9" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[323,480,801,827]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="214" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9313C7674422453BA3BFCA5B7F1FCE9" box="[323,480,801,827]" italics="true" pageId="12" pageNumber="217">Bicarinulodes</emphasis>
|
||
</taxonomicName>
|
||
is more similar to
|
||
<taxonomicName id="4C459BE774422453B9C1FCA5B52DFCE9" authorityName="Jałoszyński" authorityYear="2019" box="[697,828,801,827]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9313C7674422453B9C1FCA5B52DFCE9" box="[697,828,801,827]" italics="true" pageId="12" pageNumber="217">Afroeudesis</emphasis>
|
||
</taxonomicName>
|
||
than to the
|
||
<emphasis id="B9313C7674422453B8BDFCA5B6AFFC8D" italics="true" pageId="12" pageNumber="217">
|
||
<taxonomicName id="4C459BE774422453B8BDFCA5B25AFCE9" box="[965,1099,801,827]" class="Insecta" family="Staphylinidae" genus="Meridaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">Meridaphes</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C459BE774422453BF2CFCA5B2E2FCE9" authorityName="Franz" authorityYear="1980" box="[1108,1267,801,827]" class="Insecta" family="Staphylinidae" genus="Pseudoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">Pseudoraphes</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C459BE774422453BF84FCA5B6AFFC8D" authorityName="Franz" authorityYear="1985" class="Insecta" family="Staphylinidae" genus="Stenichnoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">Stenichnoconnus</taxonomicName>
|
||
</emphasis>
|
||
group of genera, but it clearly differs from all subgenera of
|
||
<taxonomicName id="4C459BE774422453B815FCC1B5E1FC8D" authorityName="Jałoszyński" authorityYear="2019" box="[877,1008,837,863]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9313C7674422453B815FCC1B5E1FC8D" box="[877,1008,837,863]" italics="true" pageId="12" pageNumber="217">Afroeudesis</emphasis>
|
||
</taxonomicName>
|
||
in the lack of the conspicuous subtriangular and scaly-sculptured 'platform' on the frons and vertex.
|
||
<taxonomicName id="4C459BE774422453B81DFCEDB213FC51" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[869,1026,873,899]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="214" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9313C7674422453B81DFCEDB213FC51" box="[869,1026,873,899]" italics="true" pageId="12" pageNumber="217">Bicarinulodes</emphasis>
|
||
</taxonomicName>
|
||
has two autapomorphies, the pair of short, flat submedian carinulae on the pronotal base, and the modified antennomere VII in males. For these reasons,
|
||
<taxonomicName id="4C459BE774422453BBEFFC35B7A8FC19" authorityName="Franz" authorityYear="1988" box="[151,441,945,971]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="species" species="meridensis">
|
||
<emphasis id="B9313C7674422453BBEFFC35B7A8FC19" box="[151,441,945,971]" italics="true" pageId="12" pageNumber="217">Microscydmus meridensis</emphasis>
|
||
</taxonomicName>
|
||
is not placed in
|
||
<taxonomicName id="4C459BE774422453B90AFC35B4E4FC19" authorityName="Jałoszyński" authorityYear="2019" box="[626,757,945,971]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9313C7674422453B90AFC35B4E4FC19" box="[626,757,945,971]" italics="true" pageId="12" pageNumber="217">Afroeudesis</emphasis>
|
||
</taxonomicName>
|
||
, but in a separate new genus.
|
||
</paragraph>
|
||
<paragraph id="8BFAE06474422453BBBFFC50B711FBC1" blockId="12.[151,1437,153,1043]" pageId="12" pageNumber="217">
|
||
<emphasis id="B9313C7674422453BBBFFC50B425FC3D" bold="true" box="[199,564,980,1007]" pageId="12" pageNumber="217">Composition and distribution.</emphasis>
|
||
<taxonomicName id="4C459BE774422453B945FC51B4CBFC3D" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[573,730,981,1007]" class="Insecta" family="Staphylinidae" genus="Bicarinulodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="214" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9313C7674422453B945FC51B4CBFC3D" box="[573,730,981,1007]" italics="true" pageId="12" pageNumber="217">Bicarinulodes</emphasis>
|
||
</taxonomicName>
|
||
comprises only its
|
||
<typeStatus id="54FE5EC674422453B8C4FC51B5FDFC3D" box="[956,1004,981,1007]" pageId="12" pageNumber="217">type</typeStatus>
|
||
species known to occur in
|
||
<collectingCountry id="F352A0F474422453BE51FC51B38DFC3D" box="[1321,1436,981,1007]" name="Venezuela" pageId="12" pageNumber="217">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="137EFCE174422453BBD8FC7DB6E2FBC1" box="[160,243,1017,1043]" captionStart="FIGURE 41" captionStartId="13.[151,250,1995,2020]" captionTargetBox="[161,1417,1304,1961]" captionTargetId="figure@13.[151,1436,1269,1969]" captionTargetPageId="13" captionText="FIGURE 41. Distribution of 'Afroeudesis group'." httpUri="https://zenodo.org/record/3234314/files/figure.png" pageId="12" pageNumber="217">Fig. 41</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |