427 lines
48 KiB
XML
427 lines
48 KiB
XML
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<mods:title id="46BEE6944011C161146EC2CF32EE9E8C">Six new species and a key to 12 species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from India, Nepal and Pakistan</mods:title>
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<mods:namePart id="E97FCE08D5B9D07535197A1502AE160A">Huang, Tian</mods:namePart>
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<mods:affiliation id="527DE4B199B30952DE2D204B40248BE7">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:namePart id="D76ED947964DF01FB8C2CEA78920D851">Janák, Jiří</mods:namePart>
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<mods:affiliation id="09BE68FDAB939A72E24C6C17F9449B95">CZ- 41762 Rtyně nad Bílinou, č. 4, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="466E53E9931BE986FEA8797FF8BF0D2C" type="email">janak.jiri1@gmail.com</mods:nameIdentifier>
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<mods:namePart id="58DE6BE0F5F662291467C07CF0563790">Zhou, Hong-Zhang</mods:namePart>
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<mods:affiliation id="74BBEB70A8BA49CE57D99A39B32FDFB9">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:nameIdentifier id="996637F6F15455474F942C779B426560" type="email">zhouhz@ioz.ac.cn</mods:nameIdentifier>
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<mods:date id="A3E28A00043C5E9F10691ECB86B321DB">2024</mods:date>
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<treatment id="03E287E6DF1DFFA5FF1FFF4FE54BA289" ID-DOI="http://doi.org/10.5281/zenodo.14248199" ID-GBIF-Taxon="248070642" ID-Zenodo-Dep="14248199" LSID="urn:lsid:plazi:treatment:03E287E6DF1DFFA5FF1FFF4FE54BA289" httpUri="http://treatment.plazi.org/id/03E287E6DF1DFFA5FF1FFF4FE54BA289" lastPageId="15" lastPageNumber="516" pageId="12" pageNumber="513" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae" scope_tribe="Diochini">
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3.
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<taxonomicName id="4C4B4D73DF1DFFA6FF4FFF4FE3D4A6FD" authority="Huang, Janak & Zhou" authorityName="Huang, Janak & Zhou" authorityYear="2024" box="[199,674,150,178]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="setifer" status="sp. nov.">
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<emphasis id="B93FEAE2DF1DFFA6FF4FFF4FE009A6FD" bold="true" box="[199,383,151,178]" italics="true" pageId="12" pageNumber="513">Diochus setifer</emphasis>
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Huang, Janák & Zhou
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,
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<taxonomicNameLabel id="A20C5799DF1DFFA6FD27FF4FE27AA6FD" box="[687,780,151,177]" pageId="12" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="13702A75DF1DFFA6FF17FF63E18DA699" box="[159,251,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1556,1580]" captionTargetBox="[158,1435,490,1525]" captionTargetId="figure-176@4.[151,1436,488,1533]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Diochus species from Bhutan, India, Nepal, Myanmar, Pakistan and Thailand in dorsal view: A. D. guizhouensis Shuai, Nozaki & Tang, 2021; B. D. setatus sp. nov.; C. D. setifer sp. nov.; D. D. emarginatus sp. nov.; E. D. cameroni sp. nov.; F. D. ceylonicus sp. nov., 1929; G. D. conicollis (Motschulsky, 1858); H. D. punctipennis (Motschulsky, 1858); I. D. keralanus sp. nov.; J. D. antennatus (Motschulsky, 1858); K. D. bicoloripennis Coiffait, 1981; L. D. ochraceus Cameron, 1926 (Scale bar: 0.5 mm; photos of six new species based on the holotype respectively)." figureDoi="http://doi.org/10.5281/zenodo.14240053" httpUri="https://zenodo.org/record/14240053/files/figure.png" pageId="12" pageNumber="513">Figs 1C</figureCitation>
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; 4-1; 4-2)
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locality.
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<collectingCountry id="F35C7660DF1DFFA6FEB4FF25E0F6A75B" box="[316,384,253,279]" name="India" pageId="12" pageNumber="513">India</collectingCountry>
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:
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:
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.
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<emphasis id="B93FEAE2DF1DFFA6FF4FFEF9E001A776" bold="true" box="[199,375,288,315]" pageId="12" pageNumber="513">Type material.</emphasis>
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:
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:
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:
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,
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,
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,
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(
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)
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.
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:
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:
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:
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,
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, same data as holotype (
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)
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</materialsCitation>
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;
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,
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<collectingMunicipality id="6B90AC8ADF1DFFA6FAD3FE9CE1A6A7CE" country="India" name="Khasi Hills" pageId="12" pageNumber="513" stateProvince="Meghalaya">Khasi Hills</collectingMunicipality>
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,
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,
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,
|
||
<date id="FFF51030DF1DFFA6FE3BFEBFE368A7CE" box="[435,542,359,386]" pageId="12" pageNumber="513" value="1978-11-05">
|
||
<collectingDate id="EFB1E9D8DF1DFFA6FE3BFEBFE368A7CE" box="[435,542,359,386]" pageId="12" pageNumber="513" value="1978-11-05">5. XI. 78</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BE5326DF1DFFA6FDA5FEB0E3A4A7CE" box="[557,722,360,386]" pageId="12" pageNumber="513">Besuchet-Löbl</collectorName>
|
||
(
|
||
<collectionCode id="ED5AAE35DF1DFFA6FD6CFEB0E236A7CE" box="[740,832,360,386]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="513" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B233CADDF1DFFA6FCDEFEB0E0C7A7E9" ID-GBIF-Occurrence="4986903383" collectingDate="1978-11-01" collectionCode="MHNG" collectorName="Besuchet-Lobl" country="India" elevation="800" location="Garo Hills" pageId="12" pageNumber="513" specimenCount="1" specimenCount-female="1" stateProvince="Meghalaya" typeStatus="paratype">
|
||
<specimenCount id="9D4DFD79DF1DFFA6FCDEFEB0E2CBA7CE" box="[854,957,360,386]" count="1" pageId="12" pageNumber="513" type="female">1 female</specimenCount>
|
||
,
|
||
<location id="8E94602BDF1DFFA6FC43FEB0E530A7CE" LSID="urn:lsid:plazi:treatment:03E287E6DF1DFFA5FF1FFF4FE54BA289:8E94602BDF1DFFA6FC43FEB0E530A7CE" box="[971,1094,360,386]" country="India" name="Garo Hills" pageId="12" pageNumber="513" stateProvince="Meghalaya">Garo Hills</location>
|
||
,
|
||
<location id="8E94602BDF1DFFA6FBDCFEB0E5FDA7CE" LSID="urn:lsid:plazi:treatment:03E287E6DF1DFFA5FF1FFF4FE54BA289:8E94602BDF1DFFA6FBDCFEB0E5FDA7CE" box="[1108,1163,360,386]" country="India" name="Tura" pageId="12" pageNumber="513" stateProvince="Meghalaya">Tura</location>
|
||
,
|
||
<quantity id="4CB39B15DF1DFFA6FB11FEB0E46BA7CE" box="[1177,1309,360,386]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="12" pageNumber="513" unit="m" value="800.0" valueMax="900.0" valueMin="700.0">
|
||
<elevation id="0066D1C3DF1DFFA6FB11FEB0E46BA7CE" box="[1177,1309,360,386]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="12" pageNumber="513" unit="m" value="800.0" valueMax="900.0" valueMin="700.0">700–900 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFF51030DF1DFFA6FAA4FEB0E4E1A7CE" box="[1324,1431,360,386]" pageId="12" pageNumber="513" value="1978-11-01">
|
||
<collectingDate id="EFB1E9D8DF1DFFA6FAA4FEB0E4E1A7CE" box="[1324,1431,360,386]" pageId="12" pageNumber="513" value="1978-11-01">1. XI. 78</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BE5326DF1DFFA6FF1FFE53E04AA7E9" box="[151,316,395,421]" pageId="12" pageNumber="513">Besuchet-Löbl</collectorName>
|
||
(
|
||
<collectionCode id="ED5AAE35DF1DFFA6FEC5FE53E0DFA7E9" box="[333,425,395,421]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="513" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B233CADDF1DFFA6FE35FE53E4EEA7E9" ID-GBIF-Occurrence="4986903343" box="[445,1432,395,422]" collectingDate="1978-10-22" collectionCode="MHNG, JJRC" collectorName="Besuchet-Lobl" country="India" elevation="200" location="Manas" municipality="Manas" pageId="12" pageNumber="513" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Assam" typeStatus="paratype">
|
||
<collectingRegion id="498FF812DF1DFFA6FE35FE53E378A7E9" box="[445,526,395,421]" country="India" name="Assam" pageId="12" pageNumber="513">Assam</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6B90AC8ADF1DFFA6FD92FE54E311A7EA" box="[538,615,396,422]" pageId="12" pageNumber="513">Manas</collectingMunicipality>
|
||
,
|
||
<specimenCount id="9D4DFD79DF1DFFA6FDFDFE53E3A6A7E9" box="[629,720,395,421]" count="3" pageId="12" pageNumber="513" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4DFD79DF1DFFA6FD56FE53E239A7E9" box="[734,847,395,421]" count="8" pageId="12" pageNumber="513" type="female">8 females</specimenCount>
|
||
,
|
||
<quantity id="4CB39B15DF1DFFA6FCD4FE53E2DEA7EA" box="[860,936,395,422]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="513" unit="m" value="200.0">
|
||
<elevation id="0066D1C3DF1DFFA6FCD4FE53E2DEA7EA" box="[860,936,395,422]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="513" unit="m" value="200.0">200 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFF51030DF1DFFA6FC3EFE53E552A7E9" box="[950,1060,395,422]" pageId="12" pageNumber="513" value="1978-10-22">
|
||
<collectingDate id="EFB1E9D8DF1DFFA6FC3EFE53E552A7E9" box="[950,1060,395,422]" pageId="12" pageNumber="513" value="1978-10-22">22. X. 78</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BE5326DF1DFFA6FBBBFE53E5AEA7E9" box="[1075,1240,395,421]" pageId="12" pageNumber="513">Besuchet-Löbl</collectorName>
|
||
(
|
||
<collectionCode id="ED5AAE35DF1DFFA6FB60FE53E434A7E9" box="[1256,1346,395,421]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="513" type="Museum">MHNG</collectionCode>
|
||
,
|
||
<collectionCode id="ED5AAE35DF1DFFA6FAD8FE53E4E6A7E9" box="[1360,1424,395,421]" pageId="12" pageNumber="513">JJRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B233CADDF1DFFA6FF1FFE76E54FA785" ID-GBIF-Occurrence="4986903424" box="[151,1081,430,457]" collectingDate="1979-10-24" collectionCode="UP, MHNG" collectorName="I. Lobl" country="India" elevation="1500" location="Garhwal" pageId="12" pageNumber="513" specimenCount="1" specimenCount-male="1" stateProvince="Assam" typeStatus="paratype">
|
||
<location id="8E94602BDF1DFFA6FF1FFE76E181A784" LSID="urn:lsid:plazi:treatment:03E287E6DF1DFFA5FF1FFF4FE54BA289:8E94602BDF1DFFA6FF1FFE76E181A784" box="[151,247,430,456]" country="India" name="Garhwal" pageId="12" pageNumber="513" stateProvince="Assam">Garhwal</location>
|
||
(
|
||
<collectionCode id="ED5AAE35DF1DFFA6FE8EFE77E046A785" box="[262,304,431,457]" country="Papua New Guinea" name="University of Papua and New Guinea" pageId="12" pageNumber="513">UP</collectionCode>
|
||
):
|
||
<specimenCount id="9D4DFD79DF1DFFA6FEC9FE77E0FBA784" box="[321,397,430,457]" count="1" pageId="12" pageNumber="513" type="male">1 male</specimenCount>
|
||
,
|
||
<location id="8E94602BDF1DFFA6FE1FFE76E3D4A784" LSID="urn:lsid:plazi:treatment:03E287E6DF1DFFA5FF1FFF4FE54BA289:8E94602BDF1DFFA6FE1FFE76E3D4A784" box="[407,674,430,456]" country="India" name="au-dessous de Bhatwari" pageId="12" pageNumber="513" stateProvince="Assam">au-dessous de Bhatwari</location>
|
||
,
|
||
<quantity id="4CB39B15DF1DFFA6FD23FE76E277A785" box="[683,769,430,457]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="513" unit="m" value="1500.0">
|
||
<elevation id="0066D1C3DF1DFFA6FD23FE76E277A785" box="[683,769,430,457]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="513" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFF51030DF1DFFA6FC83FE77E204A785" box="[779,882,431,457]" pageId="12" pageNumber="513" value="1979-10-24">
|
||
<collectingDate id="EFB1E9D8DF1DFFA6FC83FE77E204A785" box="[779,882,431,457]" pageId="12" pageNumber="513" value="1979-10-24">24. X. 79</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BE5326DF1DFFA6FCF5FE77E2BEA784" box="[893,968,430,457]" pageId="12" pageNumber="513">I. Löbl</collectorName>
|
||
(
|
||
<collectionCode id="ED5AAE35DF1DFFA6FC5DFE77E547A785" box="[981,1073,431,457]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="513" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54F08852DF1DFFA6FBCAFE76E5DCA784" box="[1090,1194,430,456]" pageId="12" pageNumber="513" type="holotype">Holotype</typeStatus>
|
||
with an identification label: “
|
||
<typeStatus id="54F08852DF1DFFA6FF62FE0AE0E5A7A0" box="[234,403,466,492]" pageId="12" pageNumber="513" type="holotype">HOLOTYPUS</typeStatus>
|
||
<taxonomicName id="4C4B4D73DF1DFFA6FE16FE0AE33AA7A0" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[414,588,466,492]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="setifer" status="sp. nov.">Diochus setifer</taxonomicName>
|
||
<taxonomicNameLabel id="A20C5799DF1DFFA6FDDFFE0BE3C7A7A1" box="[599,689,467,493]" pageId="12" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Huang, Janák &
|
||
<determinerName id="7E28DA34DF1DFFA6FC0BFE0AE2C8A7A0" box="[899,958,466,492]" pageId="12" pageNumber="513">Zhou</determinerName>
|
||
det. 2024”.
|
||
<typeStatus id="54F08852DF1DFFA6FBDBFE0BE5B6A7A1" box="[1107,1216,467,493]" pageId="12" pageNumber="513" type="paratype">Paratypes</typeStatus>
|
||
with identification labels: “
|
||
<typeStatus id="54F08852DF1DFFA6FF79FE2EE0E4A45C" box="[241,402,502,528]" pageId="12" pageNumber="513" type="paratype">PARATYPUS</typeStatus>
|
||
<taxonomicName id="4C4B4D73DF1DFFA6FE10FE2DE335A443" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[408,579,501,527]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="setifer" status="sp. nov.">Diochus setifer</taxonomicName>
|
||
<taxonomicNameLabel id="A20C5799DF1DFFA6FDC2FE2EE3D6A45C" box="[586,672,502,528]" pageId="12" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Huang, Janák &
|
||
<determinerName id="7E28DA34DF1DFFA6FCEBFE2DE2E8A443" box="[867,926,501,527]" pageId="12" pageNumber="513">Zhou</determinerName>
|
||
det. 2024”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351657BDF1DFFA5FF4FFDC1E0DDA5A1" lastPageId="15" lastPageNumber="516" pageId="12" pageNumber="513" type="description">
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFDC1E53AA436" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFDC1E008A47F" bold="true" box="[199,382,537,563]" pageId="12" pageNumber="513">Measurements.</emphasis>
|
||
BL =
|
||
<quantity id="4CB39B15DF1DFFA6FE40FDC1E359A47F" box="[456,559,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="12" pageNumber="513" unit="mm" value="5.05">5.05 mm</quantity>
|
||
, FL =
|
||
<quantity id="4CB39B15DF1DFFA6FDF3FDC1E394A47F" box="[635,738,537,563]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" pageId="12" pageNumber="513" unit="mm" value="2.39">2.39 mm</quantity>
|
||
, HL =
|
||
<quantity id="4CB39B15DF1DFFA6FCBBFDC1E2ECA47F" box="[819,922,537,563]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="12" pageNumber="513" unit="mm" value="0.64">0.64 mm</quantity>
|
||
, HW =
|
||
<quantity id="4CB39B15DF1DFFA6FC7DFDC1E52AA47F" box="[1013,1116,537,564]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="513" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, EyL =
|
||
<quantity id="4CB39B15DF1DFFA6FB3FFDC1E468A47F" box="[1207,1310,537,563]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="513" unit="mm" value="0.16">0.16 mm</quantity>
|
||
, TL =
|
||
<quantity id="4CB39B15DF1DFFA6FAE3FDC1E1B3A41B" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="12" pageNumber="513" unit="mm" value="0.36">0.36 mm</quantity>
|
||
, ANL =
|
||
<quantity id="4CB39B15DF1DFFA6FEAEFDE5E0FBA41B" box="[294,397,573,599]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="12" pageNumber="513" unit="mm" value="1.32">1.32 mm</quantity>
|
||
, ANT1 =
|
||
<quantity id="4CB39B15DF1DFFA6FE75FDE5E315A41B" box="[509,611,573,599]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="513" unit="mm" value="0.21">0.21 mm</quantity>
|
||
, ANT2 =
|
||
<quantity id="4CB39B15DF1DFFA6FD5CFDE5E24CA41B" box="[724,826,573,599]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="513" unit="mm" value="0.13">0.13 mm</quantity>
|
||
, ANT3 =
|
||
<quantity id="4CB39B15DF1DFFA6FC22FDE5E567A41B" box="[938,1041,573,599]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="513" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, ANT4 =
|
||
<quantity id="4CB39B15DF1DFFA6FB09FDE5E591A41B" box="[1153,1255,573,599]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="513" unit="mm" value="0.1">0.10 mm</quantity>
|
||
, ANT 11 =
|
||
<quantity id="4CB39B15DF1DFFA6FAE3FDE5E1B3A436" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="513" unit="mm" value="0.13">0.13 mm</quantity>
|
||
, PL =
|
||
<quantity id="4CB39B15DF1DFFA6FE86FDB8E002A436" box="[270,372,608,634]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="12" pageNumber="513" unit="mm" value="0.91">0.91 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CB39B15DF1DFFA6FE4FFDB8E35BA436" box="[455,557,608,634]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="513" unit="mm" value="0.74">0.74 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CB39B15DF1DFFA6FDF0FDB8E3A8A436" box="[632,734,608,634]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="513" unit="mm" value="0.87">0.87 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CB39B15DF1DFFA6FCBAFDB8E2EEA436" box="[818,920,608,634]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="12" pageNumber="513" unit="mm" value="0.96">0.96 mm</quantity>
|
||
, SL =
|
||
<quantity id="4CB39B15DF1DFFA6FC69FDB8E53EA436" box="[993,1096,608,634]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="513" unit="mm" value="0.6">0.60 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFD5BE2E7A4A8" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFD5BE02FA4D1" bold="true" box="[199,345,643,669]" pageId="12" pageNumber="513">Description.</emphasis>
|
||
Body long, narrowed anteriorly and posteriorly, medium sized. Body dark brown, abdominal intersegmental membrane lighter. Legs brown, tarsi distinctly lighter. Antennae dark brown to yellowish brown, apex of each antennomere lighter. Maxillary and labial palpi brown.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFD36E02FA2C2" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFD36E071A544" bold="true" box="[199,263,750,776]" pageId="12" pageNumber="513">Head</emphasis>
|
||
(Fig 4-1A). Round square in shape, 1.28 times as long as wide. Tempora slightly widened backwards, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5–7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of tempus and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter shorter than half the length of tempora (eye: tempora = 0.44). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (
|
||
<quantity id="4CB39B15DF1DFFA6FE9BFBF5E00CA20B" box="[275,378,1069,1096]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="513" unit="mm" value="0.15">0.15 mm</quantity>
|
||
) distinctly longer than distance from antenna to eye (
|
||
<quantity id="4CB39B15DF1DFFA6FC5AFBF5E54AA20B" box="[978,1084,1069,1095]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="513" unit="mm" value="0.09">0.09 mm</quantity>
|
||
). Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent at basal 1/3, then extended to base.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFB4FE3BDA350" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFB4FE04CA2FD" bold="true" box="[199,314,1175,1201]" pageId="12" pageNumber="513">Antennae</emphasis>
|
||
(Fig 4-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate; antennomere 3 slightly longer than 2; length of antennomere 4 about thrice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFAFDE26CA3CA" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFAFDE020A373" bold="true" box="[199,342,1317,1343]" pageId="12" pageNumber="513">Mouthparts</emphasis>
|
||
(Fig 4-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin is about 1/3 the width at the widest point. Mandibles (Figs 4-1D) falciform, left one with a sharp tooth and two blunt teeth on inner margin, right one with two teeth.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFA48E3F3A3BC" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFA48E072A3E6" bold="true" box="[199,260,1424,1450]" pageId="12" pageNumber="513">Neck</emphasis>
|
||
(Fig 4-1A). Cylindrical, shiny, only bearing shallow microsculpture, width
|
||
<quantity id="4CB39B15DF1DFFA6FBF8FA48E5AFA3E6" box="[1136,1241,1424,1450]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="513" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FFA22E23DA0A5" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FFA22E037A058" bold="true" box="[199,321,1530,1556]" pageId="12" pageNumber="513">Prothorax</emphasis>
|
||
(Fig 4-1A). Pronotum dark brown, elongate (PL to PW ratio 1.24), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FF92AE591A11F" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FF92AE021A140" bold="true" box="[199,343,1778,1804]" pageId="12" pageNumber="513">Pterothorax</emphasis>
|
||
(Fig 4-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; the paired oblique furrows short but deep; the carina delimiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1DFFA6FF4FF885E5CFA1AD" blockId="12.[151,1437,253,2018]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93FEAE2DF1DFFA6FF4FF885E065A13B" bold="true" box="[199,275,1885,1911]" pageId="12" pageNumber="513">Elytra</emphasis>
|
||
(Fig 4-1A). Elytra dark brown, slightly transverse (EL to EW ratio 0.91), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.
|
||
</paragraph>
|
||
<caption id="DF346678DF1CFFA7FF1FF925E34EA111" ID-DOI="http://doi.org/10.5281/zenodo.14240069" ID-Zenodo-Dep="14240069" httpUri="https://zenodo.org/record/14240069/files/figure.png" pageId="13" pageNumber="514" startId="13.[151,250,1789,1813]" targetBox="[176,1411,191,1766]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BF436F0DF1CFFA7FF1FF925E34EA111" blockId="13.[151,1437,1789,1885]" pageId="13" pageNumber="514">
|
||
<emphasis id="B93FEAE2DF1CFFA7FF1FF925E05CA159" bold="true" box="[151,298,1789,1813]" pageId="13" pageNumber="514">FIGURE 4-1.</emphasis>
|
||
<taxonomicName id="4C4B4D73DF1CFFA7FEBCF925E0A6A159" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[308,464,1789,1813]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="setifer" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1CFFA7FEBCF925E0A6A159" box="[308,464,1789,1813]" italics="true" pageId="13" pageNumber="514">Diochus setifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1CFFA7FE51F925E35BA159" bold="true" box="[473,557,1789,1813]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="A20C5799DF1CFFA7FE51F925E35BA159" box="[473,557,1789,1813]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: holotype: A. forebody, dorsal view; B. antennae; C. proleg; D. mandibles; E. male sternite VIII; F. male tergite VIII; G. male sternite IX; H. male tergites IX–X; I. aedeagus, dorsal view; J. aedeagus, ventral view. (Scale bars: A = 0.5 mm, B–J = 0.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF346678DF1FFFA4FF1FF84FE1B8A1BB" ID-DOI="http://doi.org/10.5281/zenodo.14240071" ID-Zenodo-Dep="14240071" httpUri="https://zenodo.org/record/14240071/files/figure.png" pageId="14" pageNumber="515" startId="14.[151,250,1943,1967]" targetBox="[170,1417,186,1913]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BF436F0DF1FFFA4FF1FF84FE1B8A1BB" blockId="14.[151,1436,1943,2039]" pageId="14" pageNumber="515">
|
||
<emphasis id="B93FEAE2DF1FFFA4FF1FF84FE05FA1E3" bold="true" box="[151,297,1943,1967]" pageId="14" pageNumber="515">FIGURE 4-2.</emphasis>
|
||
<taxonomicName id="4C4B4D73DF1FFFA4FEB8F84FE0BDA1E3" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[304,459,1943,1967]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="setifer" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1FFFA4FEB8F84FE0BDA1E3" box="[304,459,1943,1967]" italics="true" pageId="14" pageNumber="515">Diochus setifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1FFFA4FE5AF84FE352A1E3" bold="true" box="[466,548,1943,1967]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="A20C5799DF1FFFA4FE5AF84FE352A1E3" box="[466,548,1943,1967]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A. aedeagus, dorsal view; B. male sternite VIII; C. male tergite VIII; D. male tergites IX–X; E. male sternite IX; F. female sternite VIII; G. female sternite X; H. female sternite IX; I. spermatheca. (Scale bars: 0.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFF4FE0EFA6B5" blockId="15.[151,1437,151,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFF4FE189A6FD" bold="true" box="[199,255,151,177]" pageId="15" pageNumber="516">Legs</emphasis>
|
||
(Fig 4-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly longer than 2−4 together, that of meso-, meta- nearly equal to the length of each 2−3.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFEDAE09EA79D" blockId="15.[151,1437,151,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFEDAE04AA750" bold="true" box="[199,316,258,284]" pageId="15" pageNumber="516">Abdomen</emphasis>
|
||
. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFE03E3C5A575" blockId="15.[151,1437,151,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFE03E072A7B9" bold="true" box="[199,260,475,501]" pageId="15" pageNumber="516">Male</emphasis>
|
||
(Figs 4-1E–J; 4-2A–E). Head with a small round elevation between eyes. Posterior margin of sternite VIII emarginate medially, also emarginate on both sides, with several setae on both sides oriented diagonally obliquely backwards (Figs 4-1E, L; 4-2B). Posterior margin of tergite VIII arcuate (Figs 4-1F; 4-2C). Tergite IX (Figs 4- 1H; 4-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 4-1G; 4-2E) asymmetrical, widest near basal 1/3 to 1/2; basal margin straight, apical margin normally emarginate; width of apical margin distinctly shorter than basal margin. Tergite X (Figs 4-1H; 4-2D) symmetrical, triangular. Aedeagus (Figs 4-1I−J; 4-2A) symmetrical, medium-sized, ca.
|
||
<quantity id="4CB39B15DF1EFFA5FDC6FD6BE3C4A481" box="[590,690,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="516" unit="mm" value="0.6">0.60 mm</quantity>
|
||
long, normally sclerotized. Parameres symmetrical, thin and long, reaching to apical tenth of median lobe. Internal structures sclerotized, paired strip-shaped sclerites and paired rectangular sclerites on both sides, in black color; a rectangular sclerite and a V-shaped sclerite in middle, brown color. Sperm pump coiled, thinned toward apex.
|
||
</paragraph>
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFC9BE0DDA5A1" blockId="15.[151,1437,151,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFC9BE068A511" bold="true" box="[199,286,835,861]" pageId="15" pageNumber="516">Female</emphasis>
|
||
(Figs 4-2F−I). Head without elevation between eyes. Sternite VIII (Fig 4-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 4-2H) symmetrical, with deep concaved margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 4-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized, thin and long, with several bends (Fig 4-2I).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351657BDF1EFFA5FF4FFC2EE0DFA25D" box="[199,425,1014,1041]" pageId="15" pageNumber="516" type="distribution">
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFC2EE0DFA25D" blockId="15.[151,1437,151,1221]" box="[199,425,1014,1041]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFC2EE017A25C" bold="true" box="[199,353,1014,1040]" pageId="15" pageNumber="516">Distribution.</emphasis>
|
||
<collectingCountry id="F35C7660DF1EFFA5FEEFFC2FE0D2A25D" box="[359,420,1015,1041]" name="India" pageId="15" pageNumber="516">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351657BDF1EFFA5FF4FFBC2E353A2ED" pageId="15" pageNumber="516" type="diagnosis">
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFBC2E353A2ED" blockId="15.[151,1437,151,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFBC2E034A278" bold="true" box="[199,322,1050,1076]" pageId="15" pageNumber="516">Diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="4C4B4D73DF1EFFA5FDF5FBC3E258A278" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[637,814,1051,1076]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="emarginatus" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1EFFA5FDF5FBC3E258A278" box="[637,814,1051,1076]" italics="true" pageId="15" pageNumber="516">D. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1EFFA5FCB1FBC3E2E1A279" bold="true" box="[825,919,1051,1077]" pageId="15" pageNumber="516">
|
||
<taxonomicNameLabel id="A20C5799DF1EFFA5FCB1FBC3E2E1A279" box="[825,919,1051,1077]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4B4D73DF1EFFA5FC5EFBC3E513A278" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[982,1125,1051,1076]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="cameroni" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1EFFA5FC5EFBC3E513A278" box="[982,1125,1051,1076]" italics="true" pageId="15" pageNumber="516">D. cameroni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1EFFA5FBF8FBC3E5B8A279" bold="true" box="[1136,1230,1051,1077]" pageId="15" pageNumber="516">
|
||
<taxonomicNameLabel id="A20C5799DF1EFFA5FBF8FBC3E5B8A279" box="[1136,1230,1051,1077]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in body size and coloration. The main difference is the setae of male sternite VIII: in this species, the posterior margin is with numerous normal setae on both sides, while in
|
||
<taxonomicName id="4C4B4D73DF1EFFA5FD27FBBBE22BA230" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[687,861,1123,1148]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="emarginatus" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1EFFA5FD27FBBBE22BA230" box="[687,861,1123,1148]" italics="true" pageId="15" pageNumber="516">D. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1EFFA5FCEEFBBBE2B6A231" bold="true" box="[870,960,1123,1149]" pageId="15" pageNumber="516">
|
||
<taxonomicNameLabel id="A20C5799DF1EFFA5FCEEFBBBE2B6A231" box="[870,960,1123,1149]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with fascicular setae on both sides and in
|
||
<taxonomicName id="4C4B4D73DF1EFFA5FF1FFB5FE054A2EC" authorityName="Huang & Janák & Zhou" authorityYear="2024" box="[151,290,1159,1184]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="cameroni" status="sp. nov.">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF1FFB5FE054A2EC" box="[151,290,1159,1184]" italics="true" pageId="15" pageNumber="516">D. cameroni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE2DF1EFFA5FEA1FB5FE0F5A2ED" bold="true" box="[297,387,1159,1185]" pageId="15" pageNumber="516">
|
||
<taxonomicNameLabel id="A20C5799DF1EFFA5FEA1FB5FE0F5A2ED" box="[297,387,1159,1185]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with no setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351657BDF1EFFA5FF4FFB73E54BA289" box="[199,1085,1195,1221]" pageId="15" pageNumber="516" type="etymology">
|
||
<paragraph id="8BF436F0DF1EFFA5FF4FFB73E54BA289" blockId="15.[151,1437,151,1221]" box="[199,1085,1195,1221]" pageId="15" pageNumber="516">
|
||
<emphasis id="B93FEAE2DF1EFFA5FF4FFB73E038A289" bold="true" box="[199,334,1195,1221]" pageId="15" pageNumber="516">Etymology.</emphasis>
|
||
The name refers to setae on posterior margin of male sternite VIII.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |