275 lines
40 KiB
XML
275 lines
40 KiB
XML
<document id="0654291701587A47D460D655C0F47CBE" ID-DOI="10.11646/zootaxa.5562.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595752" ID-ZooBank="7344C9CA-BFE1-4239-9826-4A120D31B17E" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1735932195721" checkinUser="plazi" docAuthor="Álvarez-Parra, Sergio, Rasnitsyn, Alexandr P. & Azar, Dany" docDate="2024" docId="03E287B8FFEFFFF9DAFFF9ABCB1DF898" docLanguage="en" docName="zootaxa.5562.1.8.pdf" docOrigin="Zootaxa 5562 (1)" docSource="https://doi.org/10.11646/zootaxa.5562.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeromma phoenicium Álvarez-Parra, Rasnitsyn & Azar, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="50" masterDocId="FFDBFFC0FFEDFFFCDA68FF9CCA61FFC1" masterDocTitle="The oldest false fairy wasp (Hymenoptera: Mymarommatidae) from Lower Cretaceous Lebanese amber" masterLastPageNumber="55" masterPageNumber="45" pageNumber="47" updateTime="1736170905057" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:mods id="AF115841FAD47553DE8241959E9C3CE8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="541BB411050606D4B0EF899BA700D38A">
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<mods:title id="AA10F3747EB63ADB789B3E0D8A79691F">The oldest false fairy wasp (Hymenoptera: Mymarommatidae) from Lower Cretaceous Lebanese amber</mods:title>
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<mods:name id="AC0FC8621FBA641BB6CFE68268D08B14" type="personal">
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<mods:roleTerm id="AD56C36E062B683D662F6401704D06C0">Author</mods:roleTerm>
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<mods:namePart id="B2439977632AEAC2C1D96BBFEF6E4679">Álvarez-Parra, Sergio</mods:namePart>
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<mods:affiliation id="FB8756C44A0CBED3038F599F01CC9896">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:roleTerm id="54A5C6701AC7603EB519993CD271DA6E">Author</mods:roleTerm>
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<mods:namePart id="F472394C666CC57A231AAFD6C6D51343">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="B360C9999879E8A21397C7FF7A227C74">Palaeontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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<mods:roleTerm id="45B9933D8E0467955A4DBE95912B81B1">Author</mods:roleTerm>
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<mods:namePart id="D428C8F95074C9A94474F23210EE02CA">Azar, Dany</mods:namePart>
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<mods:affiliation id="A4E06BA6ED26598B5575E9C880F3192E">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Lebanese University, Faculty of Science II, Natural Sciences Department, Fanar - El-Matn, P. O. Box 90656 Jdeidet, Lebanon * Corresponding author</mods:affiliation>
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<mods:title id="6B1202AA06E382540A269FF95C39DBCC">Zootaxa</mods:title>
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<mods:part id="453ADFD0A51DB0B46F90995D47682B31">
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<mods:date id="E78A78CADF469248FE6044B2EBBD7C7D">2024</mods:date>
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<mods:number id="8977D02B0C988D46D771B755407EA071">2024-12-31</mods:number>
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<mods:number id="8B505A63B4CB413DA2755536D396368D">5562</mods:number>
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<mods:url id="9431164721ACB29277716708098C2C50">https://doi.org/10.11646/zootaxa.5562.1.8</mods:url>
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<subSubSection id="C3516525FFEFFFFEDAFFF9ABC820F990" box="[151,577,1591,1618]" pageId="2" pageNumber="47" type="nomenclature">
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<paragraph id="8BF436AEFFEFFFFEDAFFF9ABC820F990" blockId="2.[151,978,1591,1689]" box="[151,577,1591,1618]" pageId="2" pageNumber="47">
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<heading id="D0BC81C2FFEFFFFEDAFFF9ABC820F990" bold="true" box="[151,577,1591,1618]" fontSize="11" level="1" pageId="2" pageNumber="47" reason="1">
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<emphasis id="B93FEABCFFEFFFFEDAFFF9ABC820F990" bold="true" box="[151,577,1591,1618]" pageId="2" pageNumber="47">
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<taxonomicName id="4C4B4D2DFFEFFFFEDAFFF9ABCBBDF993" authorityName="Álvarez-Parra & Rasnitsyn & Azar" authorityYear="2024" box="[151,476,1591,1618]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="phoenicium" status="sp. nov.">
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<emphasis id="B93FEABCFFEFFFFEDAFFF9ABCBBDF993" bold="true" box="[151,476,1591,1618]" italics="true" pageId="2" pageNumber="47">Archaeromma phoenicium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57C7FFEFFFFEDB8CF9ABC820F990" box="[484,577,1591,1617]" pageId="2" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3516525FFEFFFFEDAFFF9C7CB6AF958" pageId="2" pageNumber="47" type="description">
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<paragraph id="8BF436AEFFEFFFFEDAFFF9C7C9B3F9B7" blockId="2.[151,978,1591,1689]" box="[151,978,1627,1654]" pageId="2" pageNumber="47">
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8BF436AEFFEFFFFEDAFFF9E3CB6AF958" blockId="2.[151,978,1591,1689]" box="[151,267,1663,1689]" pageId="2" pageNumber="47">
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(
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<figureCitation id="13702A2BFFEFFFFEDAF7F9E3CB63F958" box="[159,258,1663,1689]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1903,1927]" captionStartId-1="4.[151,250,815,839]" captionStartId-2="5.[151,250,1316,1340]" captionTargetBox-0="[151,1436,522,1856]" captionTargetBox-1="[151,1426,181,791]" captionTargetBox-2="[151,1436,181,1291]" captionTargetId-0="figure-166@3.[151,1436,522,1877]" captionTargetId-1="figure-20@4.[151,1436,181,791]" captionTargetId-2="figure-19@5.[151,1436,181,1291]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Archaeromma phoenicium sp. nov., holotype, female. A, Habitus, right lateral view. B, Habitus, left lateral view. C, Drawing of the specimen from left lateral view, head from ventral view. D, Interpretative drawing of antenna. Abbreviations: sc—scape, p—pedicel, f1–f7—funicular segments, cl1–cl4—claval segments." captionText-1="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." captionText-2="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14595754" figureDoi-1="http://doi.org/10.5281/zenodo.14595756" figureDoi-2="http://doi.org/10.5281/zenodo.14595758" httpUri-0="https://zenodo.org/record/14595754/files/figure.png" httpUri-1="https://zenodo.org/record/14595756/files/figure.png" httpUri-2="https://zenodo.org/record/14595758/files/figure.png" pageId="2" pageNumber="47">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516525FFEFFFFEDAFFF95BCF37F8E8" pageId="2" pageNumber="47" type="materials_examined">
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<paragraph id="8BF436AEFFEFFFFEDAFFF95BCF37F8E8" blockId="2.[151,1437,1735,2014]" pageId="2" pageNumber="47">
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<emphasis id="B93FEABCFFEFFFFEDAFFF95BCB28F920" bold="true" box="[151,329,1735,1761]" pageId="2" pageNumber="47">Type material.</emphasis>
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<materialsCitation id="3B233CF3FFEFFFFEDB3BF95BCF37F8E8" collectorName="Science II" country="Lebanon" location="Faculty of Science" pageId="2" pageNumber="47" specimenCode="1701-A" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F0880CFFEFFFFEDB3BF95BCBDAF920" box="[339,443,1735,1761]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBED9ED5FFEFFFFEDBAEF95BC841F920" box="[454,544,1735,1761]" pageId="2" pageNumber="47">1701-A</specimenCode>
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; a complete
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<specimenCount id="9D4DFD27FFEFFFFED8DFF95BC962F920" box="[695,771,1735,1761]" pageId="2" pageNumber="47" type="female">female</specimenCount>
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specimen; left antenna and left forewing are affected by fractures in the amber. Syninclusion with two spiders (Araneae) and two springtails (Collembola). The holotype is deposited at the Natural History Museum of the Lebanese University, Faculty of Science II, Fanar, Lebanon.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3516525FFEFFFFEDAAFF8AFCE3BF88C" box="[199,1114,1843,1870]" pageId="2" pageNumber="47" type="etymology">
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<paragraph id="8BF436AEFFEFFFFEDAAFF8AFCE3BF88C" blockId="2.[151,1437,1735,2014]" box="[199,1114,1843,1870]" pageId="2" pageNumber="47">
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<emphasis id="B93FEABCFFEFFFFEDAAFF8AFCB2FF88C" bold="true" box="[199,334,1843,1869]" pageId="2" pageNumber="47">Etymology.</emphasis>
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After Phoenicia, ancient name of the region of modern-day
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<collectingCountry id="F35C763EFFEFFFFED998F8AFCE37F88C" box="[1008,1110,1843,1869]" name="Lebanon" pageId="2" pageNumber="47">Lebanon</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516525FFEFFFFFDAAFF8CBC8E5FEA4" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47" type="diagnosis">
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<paragraph id="8BF436AEFFEFFFFFDAAFF8CBC8E5FEA4" blockId="2.[151,1437,1735,2014]" lastBlockId="3.[151,1437,151,501]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
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<emphasis id="B93FEABCFFEFFFFEDAAFF8CBCB23F8B0" bold="true" box="[199,322,1879,1905]" pageId="2" pageNumber="47">Diagnosis.</emphasis>
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Female. Compound eyes composed by about 30–40 ommatidia; no ocelli; mandibles exodont, bidentate; toruli not contiguous, inserted above compound eyes; antenna with funicle 7-segmented and clava 4- segmented; scape 2.8× longer than wide; pedicel longer than cumulative lengths of f1 and f2; clava slightly longer than cumulative lengths of scape and pedicel; forewing: submarginal and marginal veins separated by a bulla, stigmal vein present as a bulbous area, marginal fringe composed by: six short anterior setae, eight mid-sized anterior setae, 38 long erect setae deeply inserted in membrane, and four mid-sized posterior setae, no long posterobasal marginal seta; hind wing: stalk-like, with posterior membrane slender, distal pincer-like structure present; protibial calcar curved and simple; mesotibiae with two pseudospurs; metatibiae with four pseudospurs; basitarsi as long as cumulative lengths of second and third tarsomeres; petiole bi-segmented with first segment 1.2× longer than second; gaster flattened; dorsal part of gaster setose.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516525FFEEFFFFDAAFFEF2CFE9FE6C" pageId="3" pageNumber="48" type="distribution">
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<paragraph id="8BF436AEFFEEFFFFDAAFFEF2CFE9FE6C" blockId="3.[151,1437,151,501]" pageId="3" pageNumber="48">
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<emphasis id="B93FEABCFFEEFFFFDAAFFEF2CBA7FE49" bold="true" box="[199,454,366,393]" pageId="3" pageNumber="48">Locality and horizon.</emphasis>
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Hammana-Mdeyrij amber-bearing outcrop, Caza Baabda, Mouhafazet Mount
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<collectingCountry id="F35C763EFFEEFFFFDF5AFEF3CFF9FE48" box="[1330,1432,367,393]" name="Lebanon" pageId="3" pageNumber="48">Lebanon</collectingCountry>
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; lower Barremian (Lower Cretaceous), upper interval at the upper part of “Grès du
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<collectingCountry id="F35C763EFFEEFFFFDE5BFE0FCE16FE6C" box="[1075,1143,403,429]" name="Lebanon" pageId="3" pageNumber="48">Liban</collectingCountry>
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” (
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<bibRefCitation id="EFDA4B5FFFEEFFFFDEE7FE0FCF1DFE6C" author="Maksoud, S. & Azar, D. & Granier, B. & Geze, R." box="[1167,1404,403,429]" pageId="3" pageNumber="48" pagination="331 - 338" refId="ref8280" refString="Maksoud, S., Azar, D., Granier, B. & Geze, R. (2017) New data on the age of the Lower Cretaceous amber outcrops of Lebanon. Palaeoworld, 26 (2), 331 - 338. https: // doi. org / 10.1016 / j. palwor. 2016.03.003" type="journal article" year="2017">
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Maksoud
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<emphasis id="B93FEABCFFEEFFFFDF68FE0FCF50FE6C" box="[1280,1329,403,429]" italics="true" pageId="3" pageNumber="48">et al</emphasis>
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||
. 2017
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</bibRefCitation>
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||
).
|
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3516525FFEEFFF9DAAFFE2ACB1DF898" lastPageId="5" lastPageNumber="50" pageId="3" pageNumber="48" type="description">
|
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<paragraph id="8BF436AEFFEEFFFFDAAFFE2ACF74FE34" blockId="3.[151,1437,151,501]" pageId="3" pageNumber="48">
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<emphasis id="B93FEABCFFEEFFFFDAAFFE2ACB38FE11" bold="true" box="[199,345,438,464]" pageId="3" pageNumber="48">Description.</emphasis>
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Female. Body length
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<quantity id="4CB39B4BFFEEFFFFD81EFE2BC882FE10" box="[630,739,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="48" unit="mm" value="0.43">0.43 mm</quantity>
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(
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<figureCitation id="13702A2BFFEEFFFFD893FE2BC9E1FE10" box="[763,896,439,465]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1927]" captionTargetBox="[151,1436,522,1856]" captionTargetId="figure-166@3.[151,1436,522,1877]" captionTargetPageId="3" captionText="FIGURE 1. Archaeromma phoenicium sp. nov., holotype, female. A, Habitus, right lateral view. B, Habitus, left lateral view. C, Drawing of the specimen from left lateral view, head from ventral view. D, Interpretative drawing of antenna. Abbreviations: sc—scape, p—pedicel, f1–f7—funicular segments, cl1–cl4—claval segments." figureDoi="http://doi.org/10.5281/zenodo.14595754" httpUri="https://zenodo.org/record/14595754/files/figure.png" pageId="3" pageNumber="48">Fig. 1A–C</figureCitation>
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); macropterous (
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<figureCitation id="13702A2BFFEEFFFFDE26FE2BCEFAFE10" box="[1102,1179,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,815,839]" captionTargetBox="[151,1426,181,791]" captionTargetId="figure-20@4.[151,1436,181,791]" captionTargetPageId="4" captionText="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." figureDoi="http://doi.org/10.5281/zenodo.14595756" httpUri="https://zenodo.org/record/14595756/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
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); colour dark brown (preservational), without metallic shine; setae on antennae, head, wings, legs, and dorsal part of gaster.
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</paragraph>
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<caption id="DF346626FFEEFFFFDAFFF8F3C9D1F811" ID-DOI="http://doi.org/10.5281/zenodo.14595754" ID-Zenodo-Dep="14595754" httpUri="https://zenodo.org/record/14595754/files/figure.png" pageId="3" pageNumber="48" startId="3.[151,250,1903,1927]" targetBox="[151,1436,522,1856]" targetPageId="3" targetType="figure">
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<paragraph id="8BF436AEFFEEFFFFDAFFF8F3C9D1F811" blockId="3.[151,1437,1903,2000]" pageId="3" pageNumber="48">
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<emphasis id="B93FEABCFFEEFFFFDAFFF8F3CB75F846" bold="true" box="[151,276,1903,1927]" pageId="3" pageNumber="48">FIGURE 1.</emphasis>
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<taxonomicName id="4C4B4D2DFFEEFFFFDB73F8F3C846F846" authorityName="Álvarez-Parra & Rasnitsyn & Azar" authorityYear="2024" box="[283,551,1903,1927]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="phoenicium" status="sp. nov.">
|
||
<emphasis id="B93FEABCFFEEFFFFDB73F8F3C846F846" box="[283,551,1903,1927]" italics="true" pageId="3" pageNumber="48">Archaeromma phoenicium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEABCFFEEFFFFD846F8F3C81EF846" bold="true" box="[558,639,1903,1927]" pageId="3" pageNumber="48">
|
||
<taxonomicNameLabel id="A20C57C7FFEEFFFFD846F8F3C81EF846" box="[558,639,1903,1927]" pageId="3" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female.
|
||
<emphasis id="B93FEABCFFEEFFFFD92EF8F3C939F846" bold="true" box="[838,856,1903,1927]" pageId="3" pageNumber="48">A</emphasis>
|
||
, Habitus, right lateral view.
|
||
<emphasis id="B93FEABCFFEEFFFFDE14F8F3CEECF846" bold="true" box="[1148,1165,1903,1927]" pageId="3" pageNumber="48">B</emphasis>
|
||
, Habitus, left lateral view.
|
||
<emphasis id="B93FEABCFFEEFFFFDAFFF80FCAC8F86A" bold="true" box="[151,169,1939,1963]" pageId="3" pageNumber="48">C</emphasis>
|
||
, Drawing of the specimen from left lateral view, head from ventral view.
|
||
<emphasis id="B93FEABCFFEEFFFFD9E4F80FC9FFF86A" bold="true" box="[908,926,1939,1963]" pageId="3" pageNumber="48">D</emphasis>
|
||
, Interpretative drawing of antenna. Abbreviations: sc—scape, p—pedicel, f1–f7—funicular segments, cl1–cl4—claval segments.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF346626FFE9FFF8DAFFFCB3C910FC16" ID-DOI="http://doi.org/10.5281/zenodo.14595756" ID-Zenodo-Dep="14595756" httpUri="https://zenodo.org/record/14595756/files/figure.png" pageId="4" pageNumber="49" startId="4.[151,250,815,839]" targetBox="[151,1426,181,791]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BF436AEFFE9FFF8DAFFFCB3C910FC16" blockId="4.[151,1437,815,983]" pageId="4" pageNumber="49">
|
||
<emphasis id="B93FEABCFFE9FFF8DAFFFCB3CB77FC86" bold="true" box="[151,278,815,839]" pageId="4" pageNumber="49">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C4B4D2DFFE9FFF8DB77FCB3C84FFC86" authorityName="Álvarez-Parra & Rasnitsyn & Azar" authorityYear="2024" box="[287,558,815,839]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="phoenicium" status="sp. nov.">
|
||
<emphasis id="B93FEABCFFE9FFF8DB77FCB3C84FFC86" box="[287,558,815,839]" italics="true" pageId="4" pageNumber="49">Archaeromma phoenicium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEABCFFE9FFF8D85FFCACC8EAFC89" bold="true" box="[567,651,816,840]" pageId="4" pageNumber="49">
|
||
<taxonomicNameLabel id="A20C57C7FFE9FFF8D85FFCACC8EAFC89" box="[567,651,816,840]" pageId="4" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female.
|
||
<emphasis id="B93FEABCFFE9FFF8D93FFCB3C908FC86" bold="true" box="[855,873,815,839]" pageId="4" pageNumber="49">A</emphasis>
|
||
, Forewings in right lateral view.
|
||
<emphasis id="B93FEABCFFE9FFF8DEA4FCACCEBCFC89" bold="true" box="[1228,1245,816,840]" pageId="4" pageNumber="49">B</emphasis>
|
||
, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads).
|
||
<emphasis id="B93FEABCFFE9FFF8DEA1FCCFCEBAFCAA" bold="true" box="[1225,1243,851,875]" pageId="4" pageNumber="49">C</emphasis>
|
||
, Drawing of right forewing.
|
||
<emphasis id="B93FEABCFFE9FFF8DA97FCE4CB70FC51" bold="true" box="[255,273,888,912]" pageId="4" pageNumber="49">D</emphasis>
|
||
, Interpretative drawing of stigmal complex of left forewing.
|
||
<emphasis id="B93FEABCFFE9FFF8D91EFCE4C9E6FC51" bold="true" box="[886,903,888,912]" pageId="4" pageNumber="49">E</emphasis>
|
||
, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF436AEFFE9FFF8DAAFFBB6CA8AFA49" blockId="4.[151,1437,1066,1920]" pageId="4" pageNumber="49">
|
||
<emphasis id="B93FEABCFFE9FFF8DAAFFBB6CB65FB85" box="[199,260,1066,1092]" italics="true" pageId="4" pageNumber="49">Head</emphasis>
|
||
:
|
||
<quantity id="4CB39B4BFFE9FFF8DB7AFBB6CB17FB85" box="[274,374,1066,1092]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="49" unit="mm" value="0.1">0.10 mm</quantity>
|
||
long (
|
||
<figureCitation id="13702A2BFFE9FFF8DBD5FBB6C854FB85" box="[445,565,1066,1092]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="4" pageNumber="49">Fig. 3A, B</figureCitation>
|
||
); frontal plate convex; occipital plate seemingly flat; pleated membrane in the articulation between frontal and occipital plates not visible; compound eyes small,
|
||
<quantity id="4CB39B4BFFE9FFF8DE5DFBD2CEF8FBA9" box="[1077,1177,1102,1128]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="49" unit="mm" value="0.04">0.04 mm</quantity>
|
||
in diameter, composed of about 30–40 ommatidia; no ocelli; mandibles exodont, bidentate; labrum flap-like, with straight apex; toruli not contiguous, inserted above compound eyes; antenna clavate, 13-segmented (
|
||
<figureCitation id="13702A2BFFE9FFF8D99EFB0ACE2EFB71" box="[1014,1103,1174,1200]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1927]" captionTargetBox="[151,1436,522,1856]" captionTargetId="figure-166@3.[151,1436,522,1877]" captionTargetPageId="3" captionText="FIGURE 1. Archaeromma phoenicium sp. nov., holotype, female. A, Habitus, right lateral view. B, Habitus, left lateral view. C, Drawing of the specimen from left lateral view, head from ventral view. D, Interpretative drawing of antenna. Abbreviations: sc—scape, p—pedicel, f1–f7—funicular segments, cl1–cl4—claval segments." figureDoi="http://doi.org/10.5281/zenodo.14595754" httpUri="https://zenodo.org/record/14595754/files/figure.png" pageId="4" pageNumber="49">Fig. 1D</figureCitation>
|
||
); scape elongate, 2.8× longer than wide; pedicel wider and slightly shorter than scape, about 2× longer than wide; funicle 7-segmented; f1–f3 short and ovoid, f4–f7 mostly rectangular; pedicel longer than cumulative lengths of f1 and f2; each funicular segment with at least two trichoid sensillae; clava 4-segmented, segments closely compacted; clava slightly longer than cumulative lengths of scape and pedicel; distal claval segment elongate, with apex thin and rounded; lengths of antennal segments in μm: sc: 44, p: 36, f1: 17, f2: 15, f3: 18, f4: 17, f5: 19, f6: 24, f7: 23, cl1: 18, cl2: 18, cl3: 17, cl4: 23.
|
||
</paragraph>
|
||
<paragraph id="8BF436AEFFE9FFF8DAAFFA0EC97DFA11" blockId="4.[151,1437,1066,1920]" pageId="4" pageNumber="49">
|
||
<emphasis id="B93FEABCFFE9FFF8DAAFFA0ECB5EFA6A" box="[199,319,1426,1451]" italics="true" pageId="4" pageNumber="49">Mesosoma</emphasis>
|
||
:
|
||
<quantity id="4CB39B4BFFE9FFF8DB39FA0DCBD9FA6D" box="[337,440,1425,1452]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="49" unit="mm" value="0.15">0.15 mm</quantity>
|
||
long, narrower than head; mesoscutum convex, apparently granulose, lacking evident notauli (
|
||
<figureCitation id="13702A2BFFE9FFF8DA9CFA2ACB0FFA11" box="[244,366,1462,1488]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="4" pageNumber="49">Fig. 3A, B</figureCitation>
|
||
); other mesosomal structures obscure.
|
||
</paragraph>
|
||
<paragraph id="8BF436AEFFE9FFF8DAAFFA46C861F8D5" blockId="4.[151,1437,1066,1920]" pageId="4" pageNumber="49">
|
||
<emphasis id="B93FEABCFFE9FFF8DAAFFA46CB52FA32" box="[199,307,1498,1523]" italics="true" pageId="4" pageNumber="49">Forewing</emphasis>
|
||
: pedunculate with a broadly spatulate disc (
|
||
<figureCitation id="13702A2BFFE9FFF8D975FA46C9F6FA35" box="[797,919,1498,1524]" captionStart="FIGURE 2" captionStartId="4.[151,250,815,839]" captionTargetBox="[151,1426,181,791]" captionTargetId="figure-20@4.[151,1436,181,791]" captionTargetPageId="4" captionText="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." figureDoi="http://doi.org/10.5281/zenodo.14595756" httpUri="https://zenodo.org/record/14595756/files/figure.png" pageId="4" pageNumber="49">Fig. 2A, C</figureCitation>
|
||
);
|
||
<quantity id="4CB39B4BFFE9FFF8D9C3FA46CE6EFA35" box="[939,1039,1498,1524]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="49" unit="mm" value="0.27">0.27 mm</quantity>
|
||
long and
|
||
<quantity id="4CB39B4BFFE9FFF8DE15FA46CE81FA35" box="[1149,1248,1498,1524]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="49" unit="mm" value="0.13">0.13 mm</quantity>
|
||
maximum wide, 2× longer than wide; forewing stalk with venation extending
|
||
<quantity id="4CB39B4BFFE9FFF8D92DFA62C9C8F9D9" box="[837,937,1534,1560]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="49" unit="mm" value="0.08">0.08 mm</quantity>
|
||
from wing base; proximal part of stalk (
|
||
<figureCitation id="13702A2BFFE9FFF8DF07FA62CADBF9FD" captionStart="FIGURE 2" captionStartId="4.[151,250,815,839]" captionTargetBox="[151,1426,181,791]" captionTargetId="figure-20@4.[151,1436,181,791]" captionTargetPageId="4" captionText="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." figureDoi="http://doi.org/10.5281/zenodo.14595756" httpUri="https://zenodo.org/record/14595756/files/figure.png" pageId="4" pageNumber="49">Fig. 2D</figureCitation>
|
||
) with margin folded, corresponding to costal cell, with scattered spicules and lacking apparent campaniform sensillum; submarginal vein thin and short; marginal vein wide, extending to wing margin; submarginal and marginal veins separated by a bulla; macrochaeta present on distalmost part of marginal vein; stigmal vein present as a bulbous area at distal part of marginal vein; disc flat, without venation; discal setae combined as short and spinelike; a few hair-like discal setae close to anterior margin; marginal fringe composed by: six short anterior setae, eight mid-sized anterior setae, 38 long erect setae deeply inserted in membrane, and four mid-sized posterior setae; no long posterobasal marginal seta.
|
||
</paragraph>
|
||
<paragraph id="8BF436AEFFE9FFF8DAAFF882CE6BF841" blockId="4.[151,1437,1066,1920]" pageId="4" pageNumber="49">
|
||
<emphasis id="B93FEABCFFE9FFF8DAAFF882CB21F8F6" box="[199,320,1822,1848]" italics="true" pageId="4" pageNumber="49">Hind wing</emphasis>
|
||
: short and slender, stalk-like (
|
||
<figureCitation id="13702A2BFFE9FFF8D8CFF882C946F8F9" box="[679,807,1822,1848]" captionStart="FIGURE 2" captionStartId="4.[151,250,815,839]" captionTargetBox="[151,1426,181,791]" captionTargetId="figure-20@4.[151,1436,181,791]" captionTargetPageId="4" captionText="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." figureDoi="http://doi.org/10.5281/zenodo.14595756" httpUri="https://zenodo.org/record/14595756/files/figure.png" pageId="4" pageNumber="49">Fig. 2B, E</figureCitation>
|
||
);
|
||
<quantity id="4CB39B4BFFE9FFF8D928F882C9C9F8F9" box="[832,936,1822,1848]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="49" unit="mm" value="0.04">0.04 mm</quantity>
|
||
long; base formed by a tubular vein short reaching anterior margin and a posterior membranous section slender; two sub-basal setae on anterior margin; distal part with a pincer-like structure composed of hamulus and opposite projection.
|
||
</paragraph>
|
||
<caption id="DF346626FFE8FFF9DAFFFAB8C949FA69" ID-DOI="http://doi.org/10.5281/zenodo.14595758" ID-Zenodo-Dep="14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="5" pageNumber="50" startId="5.[151,250,1316,1340]" targetBox="[151,1436,181,1291]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BF436AEFFE8FFF9DAFFFAB8C949FA69" blockId="5.[151,1437,1316,1448]" pageId="5" pageNumber="50">
|
||
<emphasis id="B93FEABCFFE8FFF9DAFFFAB8CB72FAFD" bold="true" box="[151,275,1316,1340]" pageId="5" pageNumber="50">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C4B4D2DFFE8FFF9DB71FAB8C844FAFD" authorityName="Álvarez-Parra & Rasnitsyn & Azar" authorityYear="2024" box="[281,549,1316,1340]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="phoenicium" status="sp. nov.">
|
||
<emphasis id="B93FEABCFFE8FFF9DB71FAB8C844FAFD" box="[281,549,1316,1340]" italics="true" pageId="5" pageNumber="50">Archaeromma phoenicium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEABCFFE8FFF9D844FAB8C81CFAFD" bold="true" box="[556,637,1316,1340]" pageId="5" pageNumber="50">
|
||
<taxonomicNameLabel id="A20C57C7FFE8FFF9D844FAB8C81CFAFD" box="[556,637,1316,1340]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female.
|
||
<emphasis id="B93FEABCFFE8FFF9D929FAB8C932FAFD" bold="true" box="[833,851,1316,1340]" pageId="5" pageNumber="50">A</emphasis>
|
||
, Head in dorsal view and mesosoma in right lateral view.
|
||
<emphasis id="B93FEABCFFE8FFF9DAFFFAD4CAC9FAA1" bold="true" box="[151,168,1352,1376]" pageId="5" pageNumber="50">B</emphasis>
|
||
, Head in ventral view and mesosoma in left lateral view.
|
||
<emphasis id="B93FEABCFFE8FFF9D978FAD4C943FAA1" bold="true" box="[784,802,1352,1376]" pageId="5" pageNumber="50">C</emphasis>
|
||
, Protarsus and interpretative drawing of protibial calcar and basitarsus.
|
||
<emphasis id="B93FEABCFFE8FFF9DB61FAF0CB7AFA45" bold="true" box="[265,283,1388,1412]" pageId="5" pageNumber="50">D</emphasis>
|
||
, Mesotarsus with two pseudospurs (red arrowheads).
|
||
<emphasis id="B93FEABCFFE8FFF9D921FAF0C93BFA45" bold="true" box="[841,858,1388,1412]" pageId="5" pageNumber="50">E</emphasis>
|
||
, Metatarsus with four pseudospurs (red arrowheads).
|
||
<emphasis id="B93FEABCFFE8FFF9DFEEFAF0CFF4FA45" bold="true" box="[1414,1429,1388,1412]" pageId="5" pageNumber="50">F</emphasis>
|
||
, Metasoma in right lateral view.
|
||
<emphasis id="B93FEABCFFE8FFF9DBB1FA0CCB8CFA69" bold="true" box="[473,493,1424,1448]" pageId="5" pageNumber="50">G</emphasis>
|
||
, Metasoma in left lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF436AEFFE8FFF9DAAFFA67CBC8F964" blockId="5.[151,1437,1530,1881]" pageId="5" pageNumber="50">
|
||
<emphasis id="B93FEABCFFE8FFF9DAAFFA67CA9CF9D5" box="[199,253,1531,1556]" italics="true" pageId="5" pageNumber="50">Legs</emphasis>
|
||
: elongate and slender; tibiae and tarsi setose; mesofemorae with a few setae emerging from small protuberances; tibial spur formula 1:0:0; protibial calcar curved and simple (
|
||
<figureCitation id="13702A2BFFE8FFF9DE5AF982CEF0F9F9" box="[1074,1169,1566,1592]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="5" pageNumber="50">Fig. 3C</figureCitation>
|
||
); mesotibiae with two pseudospurs (
|
||
<figureCitation id="13702A2BFFE8FFF9DB52F9DECBFAF99D" box="[314,411,1602,1628]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="5" pageNumber="50">Fig. 3D</figureCitation>
|
||
); metatibiae with four pseudospurs (
|
||
<figureCitation id="13702A2BFFE8FFF9D93FF9DEC9D4F99D" box="[855,949,1602,1628]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="5" pageNumber="50">Fig. 3E</figureCitation>
|
||
); tarsi 5-segmented; basitarsi as long as cumulative lengths of second and third tarsomeres; two small pretarsal claws; arolium wide with a thin, short projection at distal part.
|
||
</paragraph>
|
||
<paragraph id="8BF436AEFFE8FFF9DAAFF933CB1DF898" blockId="5.[151,1437,1530,1881]" pageId="5" pageNumber="50">
|
||
<emphasis id="B93FEABCFFE8FFF9DAAFF933CB5FF909" box="[199,318,1711,1736]" italics="true" pageId="5" pageNumber="50">Metasoma</emphasis>
|
||
: composed of petiole bi-segmented and gaster globular (
|
||
<figureCitation id="13702A2BFFE8FFF9D9B0F932CE32F909" box="[984,1107,1710,1736]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1340]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-19@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14595758" httpUri="https://zenodo.org/record/14595758/files/figure.png" pageId="5" pageNumber="50">Fig. 3F, G</figureCitation>
|
||
); first petiolar segment
|
||
<quantity id="4CB39B4BFFE8FFF9DF03F932CAA2F92C" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="50" unit="mm" value="0.03">0.03 mm</quantity>
|
||
long, second petiolar segment
|
||
<quantity id="4CB39B4BFFE8FFF9D844F94EC8F3F92C" box="[556,658,1746,1773]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="50" unit="mm" value="0.02">0.02 mm</quantity>
|
||
long, first segment 1.2× longer than second, both segments narrow and same height, no sculpture; petiole inserted on the anterobasal part of gaster; gaster flattened,
|
||
<quantity id="4CB39B4BFFE8FFF9DE93F96ACF03F8D0" box="[1275,1378,1782,1809]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="50" unit="mm" value="0.16">0.16 mm</quantity>
|
||
long and
|
||
<quantity id="4CB39B4BFFE8FFF9DAAFF886CB4DF8F4" box="[199,300,1818,1845]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="50" unit="mm" value="0.05">0.05 mm</quantity>
|
||
heigh; tergites indiscernible; cerci and ovipositor sheath not visible; ovipositor thin,
|
||
<quantity id="4CB39B4BFFE8FFF9DE9EF886CF3AF8F4" box="[1270,1371,1818,1845]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="50" unit="mm" value="0.05">0.05 mm</quantity>
|
||
long; hypopygium setose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |