190 lines
22 KiB
XML
190 lines
22 KiB
XML
<document id="40680F8CF384002A9B389B0320ABF204" ID-CLB-Dataset="50954" ID-DOI="10.5281/zenodo.170350" ID-GBIF-Dataset="01dcf5a1-d174-4270-907d-73a14d330404" ID-ISSN="11755326" ID-Zenodo-Dep="170350" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459747205460" checkinUser="plazi" docAuthor="Murányi, Dávid" docDate="2005" docId="03E287920721FFFE401DFD94FC4AEA7E" docLanguage="en" docName="zt01078p064.pdf" docOrigin="Zootaxa 1078" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Leuctra klapperichi Murányi, 2005, sp. n." docType="treatment" docVersion="10" lastPageNumber="62" masterDocId="FFDBFFEA0720FFFD4115FFC8FFEBEE50" masterDocTitle="Leuctra klapperichi, a new stonefly species (Plecoptera: Leuctridae) from Iran with geographic affinities of related species" masterLastPageNumber="64" masterPageNumber="59" pageNumber="60" updateTime="1698214200237" updateUser="ExternalLinkService">
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<mods:title id="1A80262433E420150F9C805712E27855">Leuctra klapperichi, a new stonefly species (Plecoptera: Leuctridae) from Iran with geographic affinities of related species</mods:title>
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<mods:namePart id="378FF8B85815375EB66E21A143326097">Murányi, Dávid</mods:namePart>
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<mods:title id="E491E1312C8156DF19805D7D0D6432E5">Zootaxa</mods:title>
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<mods:date id="17D4D7DC2CCFCD72F8D61E91213F0F47">2005</mods:date>
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<treatment id="03E287920721FFFE401DFD94FC4AEA7E" ID-DOI="http://doi.org/10.5281/zenodo.6266770" ID-GBIF-Taxon="119336707" ID-Zenodo-Dep="6266770" LSID="urn:lsid:plazi:treatment:03E287920721FFFE401DFD94FC4AEA7E" httpUri="http://treatment.plazi.org/id/03E287920721FFFE401DFD94FC4AEA7E" lastPageId="3" lastPageNumber="62" pageId="1" pageNumber="60">
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<subSubSection id="C351650F0721FFFC401DFD94FD21EC25" box="[264,714,603,630]" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="8BF436840721FFFC401DFD94FD21EC25" blockId="1.[264,714,603,630]" box="[264,714,603,630]" pageId="1" pageNumber="60">
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<heading id="D0BC81E80721FFFC401DFD94FD21EC25" bold="true" box="[264,714,603,630]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
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<emphasis id="B93FEA960721FFFC401DFD94FD21EC25" bold="true" box="[264,714,603,630]" pageId="1" pageNumber="60">
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<emphasis id="B93FEA960721FFFC401DFD94FE15EC26" bold="true" box="[264,510,604,630]" italics="true" pageId="1" pageNumber="60">
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<taxonomicName id="4C4B4D070721FFFC401DFD94FE13EC26" ID-CoL="727RQ" box="[264,504,604,630]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="klapperichi" status="sp. nov.">Leuctra klapperichi</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A20C57ED0721FFFC4310FD93FDACEC25" box="[517,583,603,629]" pageId="1" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="13702A010721FFFC4343FD93FD2AEC25" box="[598,705,603,629]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1652,1676]" captionTargetBox="[339,1263,315,1638]" captionTargetId="figure@2.[319,1268,296,1644]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male terminalia of Leuctra klapperichi sp. n. — 1: dorsal view; 2: lateral view; 3: paraprocts, caudal view; 4: ventral view of male terminalia — scale 0.5 mm" httpUri="https://zenodo.org/record/170351/files/figure.png" pageId="1" pageNumber="60">Figs 1–4</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351650F0721FFFC401DFD6EFD11ECB9" pageId="1" pageNumber="60" type="materials_examined">
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<paragraph id="8BF436840721FFFC401DFD6EFD11ECB9" blockId="1.[264,1324,678,1905]" pageId="1" pageNumber="60">
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<emphasis id="B93FEA960721FFFC401DFD6EFE29EC90" bold="true" box="[264,450,678,704]" pageId="1" pageNumber="60">
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<typeStatus id="54F088260721FFFC401DFD6EFEA8EC90" box="[264,323,678,704]" pageId="1" pageNumber="60">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="54F088260721FFFC40DBFD6FFDD0EC91" box="[462,571,679,705]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="F35C76140721FFFC4384FD6FFD37EC91" box="[657,732,679,705]" name="Iran" pageId="1" pageNumber="60">IRAN</collectingCountry>
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, Mts Elburs, Ewin (N of Teheran),
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<quantity id="4CB39B610721FFFC4584FD6FFB09EC91" box="[1169,1250,679,705]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="60" unit="m" value="1700.0">1700m</quantity>
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a.s.l.,
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<date id="FFF510440721FFFC401DFD07FE65ECB9" box="[264,398,719,745]" pageId="1" pageNumber="60" value="1960-10-14">14.10.1960</date>
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, leg. J. F. Klapperich (
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<collectionCode id="ED5AAE410721FFFC4380FD07FD18ECB9" LSID="urn:lsid:biocol.org:col:34991" box="[661,755,719,745]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="60">HNHM</collectionCode>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C351650F0721FFFC402DFD3EFEACEDE1" pageId="1" pageNumber="60" type="diagnosis">
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<paragraph id="8BF436840721FFFC402DFD3EFEACEDE1" blockId="1.[264,1324,678,1905]" pageId="1" pageNumber="60">
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<emphasis id="B93FEA960721FFFC402DFD3EFE51ED40" bold="true" box="[312,442,758,784]" pageId="1" pageNumber="60">Diagnosis:</emphasis>
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Member of the
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<taxonomicName id="4C4B4D070721FFFC438AFD3FFCE1ED40" box="[671,778,759,784]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="hippopus">
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<emphasis id="B93FEA960721FFFC438AFD3FFCE1ED40" box="[671,778,759,784]" italics="true" pageId="1" pageNumber="60">hippopus</emphasis>
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</taxonomicName>
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group,
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<taxonomicName id="4C4B4D070721FFFC426CFD3FFC0FED40" box="[889,996,759,784]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="hippopus">
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<emphasis id="B93FEA960721FFFC426CFD3FFC0FED40" box="[889,996,759,784]" italics="true" pageId="1" pageNumber="60">hippopus</emphasis>
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</taxonomicName>
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subgroup. Antecosta VII divided. Tergite VIII with a pigmented bridge between the sclerotized processes. Median sclerites of tergite IX subtriangular. Well developed vesicle on sternite IX. Posterior part of tergite X with a deep median notch. Styles and specilla long and gently curved near the apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="C351650F0721FFFE402DFC76FC4AEA7E" lastPageId="3" lastPageNumber="62" pageId="1" pageNumber="60" type="description">
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<paragraph id="8BF436840721FFFC402DFC76FCB7EA01" blockId="1.[264,1324,678,1905]" pageId="1" pageNumber="60">
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<emphasis id="B93FEA960721FFFC402DFC76FE39ED88" bold="true" box="[312,466,958,984]" pageId="1" pageNumber="60">Description:</emphasis>
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Medium sized species, macropterous, forewing length 7.7 mm. General colour brown. Head, antennae and pronotum uniformly brownish, pronotum as long as wide. Legs: femora brown; tibiae slightly lighter, distal part as brown as femora; tarsal segments brown. Body covered with short pilosity.
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</paragraph>
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<paragraph id="8BF436840721FFFC402DFB97FC91E921" blockId="1.[264,1324,678,1905]" pageId="1" pageNumber="60">Male abdomen: antecosta of tergites I–IX divided. Tergite I membranous anteriorly and medially, tergite II membranous anteriorly and mediolaterally, tergites III–VI weakly sclerotized medially and posteriorly. Tergites VII–X modified. All tergites covered with sparse pilosity. Transverse row of four pigmented spots on segments II–VIII. Antecosta VII divided for one fifth of the segment width. On tergite VII, bellshaped membranous area well delimited in its anterior half. Antecosta VIII divided for one third of the segment width. Medial sclerotized processes parallel, both inner and outer sides straight, apices rounded and raised in lateral view. The field between the sclerotized processes is lightly sclerotized, forming a pigmented bridge between the two processes. Lateral membranous fields reach down to one third of segment height in lateral view. Antecosta IX divided for about half the segment width. Median membranous area with two subtriangular sclerites separated by a space as wide as the width of one of them. Tergite X with a strongly bilobed expansion anteriorly, and a deep and wide median notch posteriorly. Epiproct large, rounded; its stalk having strongly sclerotized lateral edges. Cerci normal, covered with long thin setae. Paraprocts: Styles with wide short base, gently curved near the apex in lateral view, and ending in a sharp point. Base of the styles connected to a rectangular lateral expansion. Specilla long and thin, curved near the apex in lateral view, and ending in a rounded tip. Specilla a bit longer than the styles. Sternite IX: vesicle oval, well developed, with short peduncle; the sternite has a well delimited light area medially, beginning at the base of the vesicle and reaching the posterior margin.</paragraph>
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<caption id="DF34660C0722FFFF401DF9BCFBD9E8FE" httpUri="https://zenodo.org/record/170351/files/figure.png" pageId="2" pageNumber="61" targetBox="[339,1263,315,1638]" targetPageId="2">
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<paragraph id="8BF436840722FFFF401DF9BCFBD9E8FE" blockId="2.[264,1323,1652,1710]" pageId="2" pageNumber="61">
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<emphasis id="B93FEA960722FFFF401DF9BCFE5FE8DC" bold="true" box="[264,436,1652,1676]" pageId="2" pageNumber="61">FIGURES 1–4.</emphasis>
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Male terminalia of
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<taxonomicName id="4C4B4D070722FFFF439FF9BDFCB6E8DC" box="[650,861,1653,1676]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="klapperichi" status="sp. nov.">
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<emphasis id="B93FEA960722FFFF439FF9BDFCB6E8DC" box="[650,861,1653,1676]" italics="true" pageId="2" pageNumber="61">Leuctra klapperichi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ED0722FFFF4270F9BCFC75E8DB" box="[869,926,1652,1675]" pageId="2" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
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— 1: dorsal view; 2: lateral view; 3: paraprocts, caudal view; 4: ventral view of male terminalia — scale 0.5 mm
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</paragraph>
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</caption>
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<paragraph id="8BF436840722FFFF402DF92AFD7FE8AC" blockId="2.[264,1323,1762,1868]" box="[312,660,1762,1788]" pageId="2" pageNumber="61">Female and nymph unknown.</paragraph>
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<paragraph id="8BF436840722FFFE402DF8C1FCAEED8E" blockId="2.[264,1323,1762,1868]" lastBlockId="3.[264,1325,284,1070]" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
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<emphasis id="B93FEA960722FFFF402DF8C1FE5EE973" bold="true" box="[312,437,1801,1827]" pageId="2" pageNumber="61">Affinities:</emphasis>
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the species belongs to the
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<taxonomicName id="4C4B4D070722FFFF43FBF8C2FCB2E973" box="[750,857,1802,1827]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="hippopus">
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<emphasis id="B93FEA960722FFFF43FBF8C2FCB2E973" box="[750,857,1802,1827]" italics="true" pageId="2" pageNumber="61">hippopus</emphasis>
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</taxonomicName>
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group,
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<taxonomicName id="4C4B4D070722FFFF42A1F8C2FBCBE973" box="[948,1056,1802,1827]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="hippopus">
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<emphasis id="B93FEA960722FFFF42A1F8C2FBCBE973" box="[948,1056,1802,1827]" italics="true" pageId="2" pageNumber="61">hippopus</emphasis>
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</taxonomicName>
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subgroup, and is close to
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<taxonomicName id="4C4B4D070722FFFF4032F8FAFE4AE91B" box="[295,417,1842,1867]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="collaris">
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<emphasis id="B93FEA960722FFFF4032F8FAFE4AE91B" box="[295,417,1842,1867]" italics="true" pageId="2" pageNumber="61">L. collaris</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4B4D070722FFFF40CEF8FAFC8AE91C" authority="Vincon & Sivec. It" authorityName="Vincon & Sivec. It" box="[475,865,1842,1868]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="artvinensis">
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<emphasis id="B93FEA960722FFFF40CEF8FAFD90E91B" box="[475,635,1842,1867]" italics="true" pageId="2" pageNumber="61">L. artvinensis</emphasis>
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Vinçon & Sivec. It
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</taxonomicName>
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differs from both of them in the shape of the median sclerites of tergite IX, and by having a pigmented bridge between the sclerotized processes of tergite VIII. The dimensions of the animal is intermediate between the two related species. Moreover, it differs from
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<taxonomicName id="4C4B4D070723FFFE4208FEA4FC56EFD5" box="[797,957,364,389]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="artvinensis">
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<emphasis id="B93FEA960723FFFE4208FEA4FC56EFD5" box="[797,957,364,389]" italics="true" pageId="3" pageNumber="62">L. artvinensis</emphasis>
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</taxonomicName>
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with the divided antecosta VII, the straight inner sides of the sclerotized processes of tergite VIII, the smaller lateral membranous fields of tergite VIII, and the curvature of the styles. It differs from
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<taxonomicName id="4C4B4D070723FFFE4587FE74FAE0EF85" box="[1170,1291,444,469]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="collaris">
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<emphasis id="B93FEA960723FFFE4587FE74FAE0EF85" box="[1170,1291,444,469]" italics="true" pageId="3" pageNumber="62">L. collaris</emphasis>
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</taxonomicName>
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in the shape of the sclerotized processes of tergite VIII, deeper median notch of tergite X, longer styles, and the curvature of styles and specilla. The species also shows affinities with
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<taxonomicName id="4C4B4D070723FFFE4050FDFCFDBAEC1E" authority="Zhiltzova" authorityName="Zhiltzova" box="[325,593,564,590]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="svanetica">
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<emphasis id="B93FEA960723FFFE4050FDFCFE3FEC1D" box="[325,468,564,589]" italics="true" pageId="3" pageNumber="62">L. svanetica</emphasis>
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Zhiltzova
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</taxonomicName>
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, from which it differs in the parallel sclerotized processes of tergite VIII and in the shape of the median sclerites of tergite IX. Its small median sclerites also differ from the large undivided one of
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<taxonomicName id="4C4B4D070723FFFE4216FD4CFBE2ECCE" authority="Aubert. The" authorityName="Aubert. The" box="[771,1033,644,670]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="gallica">
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<emphasis id="B93FEA960723FFFE4216FD4CFC98ECCD" box="[771,883,644,669]" italics="true" pageId="3" pageNumber="62">L. gallica</emphasis>
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Aubert. The
|
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</taxonomicName>
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species differs from any other
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<taxonomicName id="4C4B4D070723FFFE4058FD64FE53EC95" box="[333,440,684,709]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="hippopus">
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||
<emphasis id="B93FEA960723FFFE4058FD64FE53EC95" box="[333,440,684,709]" italics="true" pageId="3" pageNumber="62">hippopus</emphasis>
|
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</taxonomicName>
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subgroup members that possess separated processes on tergite VIII by its divided antecosta VII and the pigmented bridge between the processes. The species also shows some resemblances with a few members of the
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<taxonomicName id="4C4B4D070723FFFE428EFD34FBEBED45" box="[923,1024,764,789]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="occitana">
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<emphasis id="B93FEA960723FFFE428EFD34FBEBED45" box="[923,1024,764,789]" italics="true" pageId="3" pageNumber="62">occitana</emphasis>
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</taxonomicName>
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subgroup, especially the following:
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<taxonomicName id="4C4B4D070723FFFE4098FCECFCDFED6E" authority="Vincon & Pardo" authorityName="Vincon & Pardo" box="[397,820,804,830]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="khroumiriensis">
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<emphasis id="B93FEA960723FFFE4098FCECFD8BED6D" box="[397,608,804,829]" italics="true" pageId="3" pageNumber="62">L. khroumiriensis</emphasis>
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Vinçon & Pardo
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</taxonomicName>
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,
|
||
<taxonomicName id="4C4B4D070723FFFE4251FCECFB7FED6E" authority="Vincon & Pardo" authorityName="Vincon & Pardo" box="[836,1172,804,830]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="sartorii">
|
||
<emphasis id="B93FEA960723FFFE4251FCECFC55ED6D" box="[836,958,804,829]" italics="true" pageId="3" pageNumber="62">L. sartorii</emphasis>
|
||
Vinçon & Pardo
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B4D070723FFFE45B6FCECFE88ED36" authority="Despax" authorityName="Despax" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="occitana">
|
||
<emphasis id="B93FEA960723FFFE45B6FCECFAC0ED6D" box="[1187,1323,804,829]" italics="true" pageId="3" pageNumber="62">L. occitana</emphasis>
|
||
Despax
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B4D070723FFFE4065FC84FD1CED36" authority="Vincon & Pardo" authorityName="Vincon & Pardo" box="[368,759,844,870]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="medjerdensis">
|
||
<emphasis id="B93FEA960723FFFE4065FC84FDC0ED35" box="[368,555,844,869]" italics="true" pageId="3" pageNumber="62">L. medjerdensis</emphasis>
|
||
Vinçon & Pardo
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B4D070723FFFE4210FC84FC0AED36" authority="Aubert" authorityName="Aubert" box="[773,993,844,870]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="vaillanti">
|
||
<emphasis id="B93FEA960723FFFE4210FC84FC63ED35" box="[773,904,844,869]" italics="true" pageId="3" pageNumber="62">L. vaillanti</emphasis>
|
||
Aubert
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D070723FFFE450BFC84FE03EDDE" authority="Consiglio & Giudicelli" authorityName="Consiglio & Giudicelli" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="subSpecies" species="cyrnea" subSpecies="cyrnea">
|
||
<emphasis id="B93FEA960723FFFE450BFC84FB0CED35" box="[1054,1255,844,869]" italics="true" pageId="3" pageNumber="62">L. cyrnea cyrnea</emphasis>
|
||
Consiglio & Giudicelli
|
||
</taxonomicName>
|
||
, from which it differs in the following features: divided antecosta VII, presence of a well developed vesicle on sternite IX and presence of a pigmented bridge between the sclerotized processes of tergite VIII.
|
||
</paragraph>
|
||
<paragraph id="8BF436840723FFFE402DFC24FC4AEA7E" blockId="3.[264,1325,284,1070]" pageId="3" pageNumber="62">
|
||
Etymology: The species is dedicated to the
|
||
<collectingCountry id="F35C76140723FFFE4244FC24FC5BEA56" box="[849,944,1004,1030]" name="Germany" pageId="3" pageNumber="62">German</collectingCountry>
|
||
entomologist, Johann Friedrich Klapperich (
|
||
<date id="FFF510440723FFFE408FFBDCFDCFEA7E" box="[410,548,1044,1070]" pageId="3" pageNumber="62" value="1913" valueMax="1987">1913–1987</date>
|
||
), collector of the
|
||
<typeStatus id="54F088260723FFFE43E5FBDCFCC8EA7E" box="[752,803,1044,1070]" pageId="3" pageNumber="62">type</typeStatus>
|
||
specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |