152 lines
19 KiB
XML
152 lines
19 KiB
XML
<document id="AA1D405BEA47ED8DDA1CD7ACD58A8AE1" ID-DOI="10.5281/zenodo.186646" ID-GBIF-Dataset="e80d7f6c-b367-43d7-80a8-1403082fb549" ID-ISSN="1175-5326" ID-Zenodo-Dep="186646" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054379412" checkinUser="plazi" docAuthor="Wu, Min & Zheng, Wei" docDate="2009" docId="03E18793FF87FF95FF16F8B371CE77D1" docLanguage="en" docName="zt02053p031.pdf" docOrigin="Zootaxa 2053" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupinidius pupinella subsp. altispirus Moellendorff 1901" docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="FFD8FFEBFF95FF86FF81FF937359701A" masterDocTitle="A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="19" updateTime="1698580698258" updateUser="plazi">
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<mods:titleInfo id="B6086BD8818564C1FA789DE6A6DA3C31">
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<mods:title id="3BD5B19BE6769056D82937636ED3D323">A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species</mods:title>
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<mods:name id="BE57DCC3ACE7CC39B3B0DEE4724A645B" type="personal">
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<mods:namePart id="1B63F30B3D1C4F0E51E1DC789EA1E84A">Wu, Min</mods:namePart>
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<mods:roleTerm id="396FF8D5DE0DEC1D16C90FF15ABBCEBF">Author</mods:roleTerm>
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<mods:namePart id="C581E24081EC528B238A28DC472B8D42">Zheng, Wei</mods:namePart>
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<mods:typeOfResource id="ADA00CBAAE560D96E85AD2692BB9AA55">text</mods:typeOfResource>
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<mods:title id="EC65ADD558A0136C38DB42F9A3A9FF2D">Zootaxa</mods:title>
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<mods:part id="8C7219BC3051844AD3583DA2616A8AAC">
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<mods:date id="F7A781D247C71EFCE833FFDA9255A905">2009</mods:date>
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<mods:detail id="58E3FDA516056630CA0DAC50BF631336" type="volume">
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<mods:number id="417004163349CAF508A659427B4CD8FB">2053</mods:number>
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<mods:start id="FAC5DF3002A42D25DDECD28ECA24A47C">1</mods:start>
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<mods:end id="8A9BF2865C826F2BA33EC6554585E7AF">31</mods:end>
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<mods:identifier id="204156B78A3D411251549DAAAC5B53CF" type="DOI">10.5281/zenodo.186646</mods:identifier>
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<mods:identifier id="3FDFF3B92FC72ADA72CBF907DB54BAE3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="334C3A8BF6CCDF87AA296787F0B704FA" type="Zenodo-Dep">186646</mods:identifier>
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<treatment id="03E18793FF87FF95FF16F8B371CE77D1" ID-DOI="http://doi.org/10.5281/zenodo.6212440" ID-GBIF-Taxon="119612781" ID-Zenodo-Dep="6212440" LSID="urn:lsid:plazi:treatment:03E18793FF87FF95FF16F8B371CE77D1" httpUri="http://treatment.plazi.org/id/03E18793FF87FF95FF16F8B371CE77D1" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="C352650EFF87FF94FF16F8B370197720" box="[151,832,1824,1851]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BF73685FF87FF94FF16F8B370197720" blockId="18.[151,832,1824,1886]" box="[151,832,1824,1851]" pageId="18" pageNumber="19">
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<heading id="D0BF81E9FF87FF94FF16F8B370197720" bold="true" box="[151,832,1824,1851]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="B93CEA97FF87FF94FF16F8B370197720" bold="true" box="[151,832,1824,1851]" pageId="18" pageNumber="19">
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14.
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<taxonomicName id="4C484D06FF87FF94FF42F8B270197720" ID-CoL="7V2BH" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[195,832,1824,1851]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="subSpecies" species="pupinella" subSpecies="altispirus">
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<emphasis id="B93CEA97FF87FF94FF42F8B2716E7721" bold="true" box="[195,567,1825,1851]" italics="true" pageId="18" pageNumber="19">Pupinidius pupinella altispirus</emphasis>
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(
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<bibRefCitation id="EFD94B74FF87FF94FDC6F8B370617720" author="Moellendorff" box="[583,824,1824,1850]" pageId="18" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF87FF94FF16F8D772C97744" box="[151,400,1860,1886]" pageId="18" pageNumber="19" type="description">
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<paragraph id="8BF73685FF87FF94FF16F8D772C97744" blockId="18.[151,832,1824,1886]" box="[151,400,1860,1886]" pageId="18" pageNumber="19">
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<figureCitation id="13732A00FF87FF94FF16F8D773B97744" box="[151,224,1860,1886]" captionStart="FIGURE 1" captionStartId="3.[151,255,1609,1633]" captionTargetBox="[161,1429,545,1583]" captionTargetId="figure@3.[151,1436,527,1584]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of Pupinidius spp. Red dot, Pupinidius anocamptus, P. nanpingensis nanpingensis and P. nanpingensis ambigua; red circle, P. porrectus; black dot, P. pupinella altispirus; black circle, P. chrysalis; blue dot, P. obrutschewi contractus and P. pupinella pupinella; blue circle: P. latilabrum; green dot, P. melinostoma, P. nanpingensis nanpingensis and P. wenxian sp. nov.; yellow dot, P. obrutschewi obrutschewi and P. streptaxis; yellow circle, P. porrecta pygmaea. P. pupinidius and P. gregorii have not been mapped here due to the unrecognizable locality names. Although the original description of the type locality of P. chrysalis is not accurate and geographically wrong, mapping of the species is possible here based only on the recognized information therein." httpUri="https://zenodo.org/record/186647/files/figure.png" pageId="18" pageNumber="19">Figs 1</figureCitation>
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,
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<figureCitation id="13732A00FF87FF94FF6FF8D773A57744" box="[238,252,1860,1886]" captionStart="FIGURE 6. A – C" captionStartId="17.[151,261,1762,1786]" captionTargetBox="[211,1375,197,1736]" captionTargetId="figure@17.[211,1376,197,1738]" captionTargetPageId="17" captionText="FIGURE 6. A – C, Pupinidius pupinella pupinella (Moellendorff, 1901). A, Lectotype, SMF- 41988; B – C, type specimens, ZI RAS-N 498. D – H, Pupinidius pupinella altispirus (Moellendorff, 1901). D, Lectotype, SMF- 41990; E – F, type specimens, ZI RAS-N 602; G, 1 paratype, SMF- 41991; H, HBUMM- 05723 - specimen 2." httpUri="https://zenodo.org/record/186652/files/figure.png" pageId="18" pageNumber="19">6</figureCitation>
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D–H, 8G, 15
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF87FF94FF16F802704D77D2" pageId="18" pageNumber="19" type="reference_group">
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<paragraph id="8BF73685FF87FF94FF16F802704D77D2" blockId="18.[151,1271,1937,1992]" pageId="18" pageNumber="19">
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<treatmentCitationGroup id="AB5811ABFF87FF94FF16F802704D77D2" pageId="18" pageNumber="19">
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<taxonomicName id="4C484D06FF87FF94FF16F802702F77B2" ID-CoL="7TR79" authority="Moellendorff, 1901: 344" authorityName="Moellendorff" authorityPageNumber="344" authorityYear="1901" box="[151,886,1937,1960]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="subSpecies" species="pupinella" subGenus="Pupinidius" subSpecies="altispirus">
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<emphasis id="B93CEA97FF87FF94FF16F802725A77B2" box="[151,259,1937,1960]" italics="true" pageId="18" pageNumber="19">Buliminus</emphasis>
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(
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<emphasis id="B93CEA97FF87FF94FE93F80272DC77B2" box="[274,389,1937,1960]" italics="true" pageId="18" pageNumber="19">Pupinidius</emphasis>
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)
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<emphasis id="B93CEA97FF87FF94FE14F802713C77B2" box="[405,613,1937,1960]" italics="true" pageId="18" pageNumber="19">pupinella altispirus</emphasis>
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<treatmentCitation id="0AE91094FF87FF94FDEDF802702F77B2" author="Moellendorff" box="[620,886,1937,1960]" page="344" pageId="18" pageNumber="19" year="1901">
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<bibRefCitation id="EFD94B74FF87FF94FDEDF802701977B2" author="Moellendorff" box="[620,832,1937,1960]" pageId="18" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
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: 344
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</treatmentCitation>
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</taxonomicName>
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, pl. 14, figs 4, 5;
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<treatmentCitation id="0AE91094FF87FF94FBAEF80277AA77B2" author="Kobelt" box="[1071,1267,1937,1960]" page="848" pageId="18" pageNumber="19" year="1902">
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<bibRefCitation id="EFD94B74FF87FF94FBAEF80277E777B2" author="Kobelt" box="[1071,1214,1937,1960]" pageId="18" pageNumber="31" refString="Kobelt, W. (1902) Die Familie Buliminidae. Systematisches Conchylien-Cabinet, 397 - 1051, 133 pls." type="book chapter" year="1902">Kobelt, 1902</bibRefCitation>
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: 848
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</treatmentCitation>
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.
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<taxonomicName id="4C484D06FF87FF94FF16F822712677D2" ID-CoL="7V2BH" authority="Yen, 1939: 84" authorityName="Yen" authorityPageNumber="84" authorityYear="1939" box="[151,639,1969,1992]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="subSpecies" species="pupinella" subSpecies="altispirus">
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<emphasis id="B93CEA97FF87FF94FF16F82272B877D2" box="[151,481,1969,1992]" italics="true" pageId="18" pageNumber="19">Pupinidius pupinella altispirus</emphasis>
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<treatmentCitation id="0AE91094FF87FF94FE69F822712677D2" author="Yen" box="[488,639,1969,1992]" page="84" pageId="18" pageNumber="19" year="1939">
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<bibRefCitation id="EFD94B74FF87FF94FE69F822710177D2" author="Yen" box="[488,600,1969,1992]" pageId="18" pageNumber="31" refString="Yen, T. C. (1939) Die chinesischen Land-und Sußwasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 444, 1 - 234, pls. 1 - 16." type="journal article" year="1939">Yen, 1939</bibRefCitation>
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: 84
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</treatmentCitation>
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</taxonomicName>
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, pl. 7, fig. 50.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C352650EFF86FF95FF16FF0470837180" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="8BF73685FF86FF95FF16FF0470837180" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">
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<emphasis id="B93CEA97FF86FF95FF16FF0472D770AB" bold="true" box="[151,398,151,177]" pageId="19" pageNumber="20">Material examined.</emphasis>
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HBUMM-05723 (20 shell measured, 3 with broken apex; specimen-1 for dissection; 1 young):
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<quantity id="4CB09B60FF86FF95FF78FF2D720B70C2" box="[249,338,190,216]" metricMagnitude="3" metricUnit="m" metricValue="1.661" pageId="19" pageNumber="20" unit="m" value="1661.0">1661 m</quantity>
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a.s.l., 34°0΄127.6΄ N, 104°25΄16.4΄ E, Dangchang, Gansu, loessial area,
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<date id="FFF61045FF86FF95FB24FF2D764370C2" box="[1189,1306,190,216]" pageId="19" pageNumber="20" value="2006-10-04">2006-X-4</date>
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, coll. J.Liu & W.Zheng;
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<collectingCountry id="F35F7615FF86FF95FEB8FF76720F70E5" box="[313,342,229,255]" name="Serbia" pageId="19" pageNumber="20">ZI</collectingCountry>
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RAS-N602:
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<typeStatus id="54F38827FF86FF95FE78FF76717770E5" box="[505,558,229,255]" pageId="19" pageNumber="20">type</typeStatus>
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material, det. Moellendorff,
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<date id="FFF61045FF86FF95FC18FF76777470E5" box="[921,1069,229,255]" pageId="19" pageNumber="20" value="1885-06-19">1885-VI-19</date>
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, coll.? Potanin. SMF-41990:
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<typeStatus id="54F38827FF86FF95FF16FE9F724C713C" box="[151,277,268,294]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
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. Tan-tshang, Kansu,
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<collectingCountry id="F35F7615FF86FF95FD94FE9F7138713C" box="[533,609,268,294]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
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. Slg.
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<collectingCountry id="F35F7615FF86FF95FD26FE9F71E5713C" box="[679,700,268,294]" name="Iceland" pageId="19" pageNumber="20">O</collectingCountry>
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.v.Moellendorff. SMF-41991:
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<typeStatus id="54F38827FF86FF95FBAFFE9F77F1713C" box="[1070,1192,268,294]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
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. 3 fmss. Tan-tshang (Dangchang of new spelling), Kansu,
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<collectingCountry id="F35F7615FF86FF95FD15FEA171BE7156" box="[660,743,306,332]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
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. Slg.
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<collectingCountry id="F35F7615FF86FF95FCB5FEA170127156" box="[820,843,306,332]" name="Iceland" pageId="19" pageNumber="20">O</collectingCountry>
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.v.Moellendorff (measurement of
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<typeStatus id="54F38827FF86FF95FA8FFEA176037156" box="[1294,1370,306,332]" pageId="19" pageNumber="20">types</typeStatus>
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: ewh 1.500–1.656–1.750, wh 6.375–6.500–6.625, h 15.4–16.8–17.9, diam. maj. 11.4–11.8–12.0, aph 8.7–9.3–9.7, apw 7.2–7.8–8.6, h/diam. maj. 1.35–1.40–1.49, h/aph 1.68–1.77–1.87).
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF86FF95FF44FE35724671FD" pageId="19" pageNumber="20" type="distribution">
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<paragraph id="8BF73685FF86FF95FF44FE35724671FD" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">
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<emphasis id="B93CEA97FF86FF95FF44FE35723271DA" bold="true" box="[197,363,422,448]" pageId="19" pageNumber="20">Distribution.</emphasis>
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Gansu (
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<typeStatus id="54F38827FF86FF95FE50FE35715E71DA" box="[465,519,422,448]" pageId="19" pageNumber="20">type</typeStatus>
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locality, between Li-dshia-pu and His-gu-tsheng, between Li-dshia-pu and Guan-ting).
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF86FF95FF44FE6072A67214" box="[197,511,499,526]" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="8BF73685FF86FF95FF44FE6072A67214" blockId="19.[151,1438,151,1995]" box="[197,511,499,526]" pageId="19" pageNumber="20">
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<emphasis id="B93CEA97FF86FF95FF44FE6072337217" bold="true" box="[197,362,499,525]" pageId="19" pageNumber="20">
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<typeStatus id="54F38827FF86FF95FF44FE60725A7217" box="[197,259,499,525]" pageId="19" pageNumber="20">Type</typeStatus>
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locality.
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</emphasis>
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Dangchang.
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF86FF95FF44FD8971CE77D1" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="8BF73685FF86FF95FF44FD89729372D5" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">
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<emphasis id="B93CEA97FF86FF95FF44FD89721E722E" bold="true" box="[197,327,538,564]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
|
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Shell bandless; not or faintly striate. Aperture more or less distinctly with angular tubercle. Most swollen part at penultimate and body whorls. With 6.375–7.000 whorls; height 15.4–18.7, diam. maj. 11.4–14.0. Penis without penial caecum; internally with more than 2 fine longitudinal pilasters. Papilla developed by fused proximal pilasters forming V-shaped structure absent. Epiphallus forming a few loops. Epiphallic caecum absent.
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</paragraph>
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<paragraph id="8BF73685FF86FF95FF44FD4F718274F6" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">Re-description of shell. Ovate; apex protruded. Most swollen part at penultimate and body whorls. Shell dextral; thin; solid; opaque; glossy; not speckled; with spiral grooves (only at umbilical region); with 6.375–6.588–7.000 whorls. Whorls not shouldered; flattened. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.375–1.529–1.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl gradually ascending towards aperture; rounded at periphery; without smoothed spiral peripheral depression nor rugate region. Shell 16.4–17.5–18.7 high; 12.0–12.7–14.0 in diam. maj. Height/width ratio 1.30–1.38–1.48. Aperture in a plane; completely attached to body whorl; subcircular or truncate-ovate; not adnate; oblique; not armed; more or less distictly with angular tubercle; 9.0–10.0–10.9 high; 7.2–8.4–9.3 wide. Ratio of shell height to aperture height 1.60–1.75–1.92. Peristome reflexed, without distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical or slightly arched. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured to differently tinted. Shell multicoloured; and umbilical region. in light greyish brown. From about the third or the fourth whorl on, with a broad whitish spiral band which occupies nearly the whole whorl. Aperture white.</paragraph>
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<paragraph id="8BF73685FF86FF95FF44FB6A76377764" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">Genitalia. Vas deferens long; of even thickness; entering epiphallus apically with distinct demarcation. Epiphallus long; cylindrical; of uniform thickness; forming a few loops; externally smooth; without caecum. Flagellum short; tubular; proximally normal; with tip blunt. Penis with terminal entrance of epiphallus; clavate; evenly thick; without penial caecum; thin-walled; internally with more than 2 fine longitudinal pilasters; with 1 V-shaped pilaster. V-shaped pilaster with proximal free end approaching at penial retractor insertion; distally not developing into a papilla. Penial process nipple-shaped; situated at boundary between epiphallus and penis. Penial verge absent. Penial appendix long; branched off from penis at some distance from atrium; divided into sections sections including A-1+A-2, A-3 and A-4+A-5. A-1 and A-2 fused; interiorly with more than 2 fine longitudinal pilasters. A-2 near A-3 internally without a ring of papilla. A-2 and A-3 not fused. A-3 opening into A-2 by no papilla. Boundary between A-4 and A-5 indistinct. A-5 long; convoluted. Penial retractor biramous; attaching to penis distally and to A-1+ A-2; with penial retractor arms arising from diaphragm closed to each other. Muscular band connecting vagina and epiphallus absent.. Additional retractor other than penial or appendicular absent. Atrium short; with weak atrial retractor. Free oviduct short; much shorter than vagina. Vagina long; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct rather long; proximally twisted. Bursa copulatrix with stalk; without apical ligament; normal in size; with long neck; well defined. Diverticle normally present; longer than reservoir; expanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base.</paragraph>
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<paragraph id="8BF73685FF86FF95FF44F81871CE77D1" blockId="19.[151,1438,151,1995]" pageId="19" pageNumber="20">Measurement of genitalia. P: 4.4; Ep: 19.0; Fl: 0.7; VD: 13.0; Va: 3.9; FO: 2.2; BCD: 26.1; BC: 1.6; D: 4.5; A-1+A-2: 6.0; A3: 1.6. A-4+A-5: 20.2.</paragraph>
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</subSubSection>
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</treatment>
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</document> |