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<document id="C265215D272F83B53514D72A8320168F" ID-CLB-Dataset="170600" ID-DOI="10.11646/zootaxa.5196.1.1" ID-GBIF-Dataset="e1042177-7e89-413a-b10b-3803d000d20d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7224080" ID-ZooBank="1B12892E-D68F-4BFE-8E9A-7F19E220E73D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666165817387" checkinUser="plazi" docAuthor="Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo &amp; Templado, José" docDate="2022" docId="03D379474860FFFEA0B1F8FC466707DF" docLanguage="en" docName="zootaxa.5196.1.1.pdf" docOrigin="Zootaxa 5196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alvania wareni" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFEA013F486EFFF0A026FD52475B0043" masterDocTitle="Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="15" updateTime="1699408954272" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AE6F86B8E184D160A4F636E5777BA240">Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain)</mods:title>
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<mods:namePart id="0D0CB59644E7EBAA8C1B57CE91BEC438">Oliver, Joan Daniel</mods:namePart>
<mods:affiliation id="2C89192DBD394BFA021F97101C365FB4">Alcorisa 83 - 12 C, 28043 Madrid, Spain. joandanieloliver @ hotmail. com</mods:affiliation>
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<mods:namePart id="663F386813CF38258B0342E7334D8D9A">Gofas, Serge</mods:namePart>
<mods:affiliation id="348A781EAA7EC3A29947076494BF2B9C">Departamento de Biología Animal, Universidad de Málaga, E- 29071 Málaga, Spain.</mods:affiliation>
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<mods:namePart id="E11DB630BD8CF54633F759A464358858">Urgorri, Victoriano</mods:namePart>
<mods:affiliation id="7450B1208B6EDB46F1DDB0584206FC26">Estación de Bioloxía Mariña da Graña, REBUSC, Universidade de Santiago de Compostela, Spain.</mods:affiliation>
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<mods:namePart id="ADE72610B5D3A33556CCD7037E82F942">Díaz-Agras, Guillermo</mods:namePart>
<mods:affiliation id="4372291E5179BD0481D62C5C3BA7FA1C">Estación de Bioloxía Mariña da Graña, REBUSC, Universidade de Santiago de Compostela, Spain.</mods:affiliation>
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<mods:namePart id="2253A924D963DA747B50AF925C5092CE">Templado, José</mods:namePart>
<mods:affiliation id="BA6DE04A588F4DA3F1EB9710C633FB41">Museo Nacional de Ciencias Naturales (CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain.</mods:affiliation>
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<treatment id="03D379474860FFFEA0B1F8FC466707DF" ID-DOI="http://doi.org/10.5281/zenodo.7224155" ID-GBIF-Taxon="203121539" ID-Zenodo-Dep="7224155" LSID="urn:lsid:plazi:treatment:03D379474860FFFEA0B1F8FC466707DF" httpUri="http://treatment.plazi.org/id/03D379474860FFFEA0B1F8FC466707DF" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C3609BDA4860FFFEA0B1F8FC45F8058B" box="[151,675,1453,1481]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BC5C8514860FFFEA0B1F8FC45F8058B" blockId="14.[151,675,1453,1516]" box="[151,675,1453,1481]" pageId="14" pageNumber="15">
<heading id="D08D7F3D4860FFFEA0B1F8FC45F8058B" bold="true" box="[151,675,1453,1481]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C7AB3D24860FFFEA0B1F8FC45F8058B" ID-CoL="CBSM" authority="(Templado &amp; Rolan, 1986)" baseAuthorityName="Templado &amp; Rolan" baseAuthorityYear="1986" box="[151,675,1453,1481]" class="Gastropoda" family="Rissoidae" genus="Alvania" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="wareni">
<emphasis id="B90E14434860FFFEA0B1F8FC45F8058B" bold="true" box="[151,675,1453,1481]" pageId="14" pageNumber="15">
<emphasis id="B90E14434860FFFEA0B1F8FC460B058B" bold="true" box="[151,336,1454,1481]" italics="true" pageId="14" pageNumber="15">Alvania wareni</emphasis>
(
<bibRefCitation id="EFEBB5A04860FFFEA179F8FC45C1058B" author="Templado, J. &amp; Rolan, E." box="[351,666,1453,1480]" pageId="14" pageNumber="15" pagination="117 - 124" refId="ref31167" refString="Templado, J. &amp; Rolan, E. (1986) El genero Onoba H. &amp; A. Adams, 1854 (Gastropoda, Rissoidea) en las costas europeas (1). Iberus, 6, 117 - 124." type="journal article" year="1986">Templado &amp; Rolán, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3609BDA4860FFFEA0B1F88047B005AF" box="[151,235,1490,1516]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BC5C8514860FFFEA0B1F88047B005AF" blockId="14.[151,675,1453,1516]" box="[151,235,1490,1516]" pageId="14" pageNumber="15">
(
<figureCitation id="1341D4D44860FFFEA0B9F88047B905AF" box="[159,226,1490,1516]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3609BDA4860FFFEA0B1FB48442C061B" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BC5C8514860FFFEA0B1FB48442C061B" blockId="14.[151,1437,1562,1948]" pageId="14" pageNumber="15">
<emphasis id="B90E14434860FFFEA0B1FB4846C80677" bold="true" box="[151,403,1562,1588]" pageId="14" pageNumber="15">Material examined. (</emphasis>
5 empty shells in the same sample):
<emphasis id="B90E14434860FFFEA313FB4844640677" bold="true" box="[821,831,1562,1588]" pageId="14" pageNumber="15">•</emphasis>
5 sh;
<geoCoordinate id="EE4EAE964860FFFEA3A0FB48437A0674" box="[902,1057,1562,1591]" degrees="44" direction="north" minutes="09.896" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="44.164932">44° 09.896′N</geoCoordinate>
,
<geoCoordinate id="EE4EAE964860FFFEA40BFB48438C0674" box="[1069,1239,1562,1591]" degrees="008" direction="west" minutes="39.581" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-8.659683">008° 39.581′W</geoCoordinate>
to
<geoCoordinate id="EE4EAE964860FFFEA4D8FB4842C30674" box="[1278,1432,1562,1591]" degrees="44" direction="north" minutes="10.129" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="44.168816">44° 10.129′N</geoCoordinate>
,
<geoCoordinate id="EE4EAE964860FFFEA0B1FB6C46180618" box="[151,323,1598,1627]" degrees="008" direction="west" minutes="39.494" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-8.658234">008° 39.494′W</geoCoordinate>
;
<quantity id="4C8265B44860FFFEA169FB6C468A061A" box="[335,465,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="4.485" metricValueMax="4.59" metricValueMin="4.38" pageId="14" pageNumber="15" unit="m" value="448.5" valueMax="459.0" valueMin="438.0">438–459 m</quantity>
;
<date id="FFC4EE914860FFFEA1FAFB6C45C8061B" box="[476,659,1598,1625]" pageId="14" pageNumber="15" value="2008-07-15" valueMax="2008-07-24" valueMin="2008-07-15">15–24 Jul. 2008</date>
;
<materialsCitation id="3B12C20C4860FFFEA2BBFB6C4428061B" ID-GBIF-Occurrence="3940322525" box="[669,883,1598,1624]" collectionCode="SELVA" pageId="14" pageNumber="15" specimenCode="DRN-11" specimenCount="1">
A
<collectionCode id="ED6B50944860FFFEA291FB6C4452061B" box="[695,777,1598,1624]" pageId="14" pageNumber="15">SELVA</collectionCode>
<specimenCode id="DBDC602A4860FFFEA336FB6C4428061B" box="[784,883,1598,1624]" collectionCode="DRN-" pageId="14" pageNumber="15">DRN-11</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3609BDA4860FFFEA0E1FB30466707DF" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BC5C8514860FFFEA0E1FB30466707DF" blockId="14.[151,1437,1562,1948]" pageId="14" pageNumber="15">
<emphasis id="B90E14434860FFFEA0E1FB304660063F" bold="true" box="[199,315,1634,1660]" pageId="14" pageNumber="15">Remarks.</emphasis>
This species was only known previously from
<specimenCount id="9D7C03D84860FFFEA375FB3144A2063F" box="[851,1017,1634,1661]" pageId="14" pageNumber="15" type="generic">two specimens</specimenCount>
found off
<collectingCountry id="F36D88C14860FFFEA454FB30438F063F" box="[1138,1236,1634,1660]" name="Portugal" pageId="14" pageNumber="15">Portugal</collectingCountry>
, one the
<typeStatus id="54C176F34860FFFEA51CFB3042C7063F" box="[1338,1436,1634,1660]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(from PORCUPINE Expedition, at
<quantity id="4C8265B44860FFFEA204FBD4452C06E2" box="[546,631,1670,1697]" metricMagnitude="3" metricUnit="m" metricValue="1.809" pageId="14" pageNumber="15" unit="m" value="1809.0">1809 m</quantity>
depth) and the other from THALASSA expedition at
<quantity id="4C8265B44860FFFEA4EBFBD4424806E2" box="[1229,1299,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="14" pageNumber="15" unit="m" value="810.0">810 m</quantity>
(
<bibRefCitation id="EFEBB5A04860FFFEA507FBD446790687" author="Bouchet, P. &amp; Waren, A." pageId="14" pageNumber="15" pagination="579 - 840" refId="ref28417" refString="Bouchet, P. &amp; Waren, A. (1993) Revision of the Northeast Atlantic bathyal and abyssal Mesogastropoda. Bolletino Malacologico, Supplement 3, 579 - 840. https: // doi. org / 10.5962 / bhl. title. 140732" type="journal article" year="1993">Bouchet &amp; Warén 1993</bibRefCitation>
). Therefore, the five shells found in one of the samples studied at about
<quantity id="4C8265B44860FFFEA464FBF843D30686" box="[1090,1160,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="15" unit="m" value="450.0">450 m</quantity>
depth extend the knowledge on this rare species and expand its distribution area and bathymetric range. They are fusiform, up to 2.7 x
<quantity id="4C8265B44860FFFEA55FFB9C479E074E" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="15" unit="mm" value="1.4">1.4 mm</quantity>
, with a high spire, whose general shape resembles some European species of
<taxonomicName id="4C7AB3D24860FFFEA40CFBA0432D074F" authorityName="H. Adams &amp; A. Adams" authorityYear="1852" box="[1066,1142,1778,1804]" class="Gastropoda" family="Rissoidae" genus="Onoba" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="B90E14434860FFFEA40CFBA0432D074F" box="[1066,1142,1778,1804]" italics="true" pageId="14" pageNumber="15">Onoba</emphasis>
</taxonomicName>
(
<figureCitation id="1341D4D44860FFFEA4A0FBA043A4074F" box="[1158,1279,1778,1804]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Figs. 8a–b</figureCitation>
). Sculpture of weak spiral cords and axial ribs of similar width somewhat narrower than their respective interspaces. Microsculpture of fine striae (
<figureCitation id="1341D4D44860FFFEA1A6FA6846890717" box="[384,466,1850,1876]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Fig. 8c</figureCitation>
). Oval aperture with thin outer lip. Dome-shaped paucispiral protoconch (
<figureCitation id="1341D4D44860FFFEA53CFA6842CF0717" box="[1306,1428,1850,1876]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Figs. 8d, g</figureCitation>
) with fine equidistant spiral threads starting from the nucleus (
<figureCitation id="1341D4D44860FFFEA364FA0C44CE073B" box="[834,917,1886,1912]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Fig. 8h</figureCitation>
). Some details of its microsculpture are shown in
<figureCitation id="1341D4D44860FFFEA092FAD0466307DF" box="[180,312,1922,1948]" captionStart="FIGURE 8" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-20@15.[151,1436,181,1783]" captionTargetPageId="15" captionText="FIGURE 8. Alvania wareni (Templado &amp; Rolán, 1986) (SE: DRN-11). a Shell (2.5 mm); b Shell (2.7 mm); c Detail of the microsculpture of the teleoconch from b; d Lateral view of the protoconch; e–f Details of the microsculpture of the protoconch; g Apical view of another protoconch; h–i Detail of the microsculpture of the embryonic protoconch." figureDoi="http://doi.org/10.5281/zenodo.7224099" httpUri="https://zenodo.org/record/7224099/files/figure.png" pageId="14" pageNumber="15">Figure 8e–i</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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