698 lines
92 KiB
XML
698 lines
92 KiB
XML
<document id="EDF9B521235EE92DF6FD7A14515DE6F7" ID-CLB-Dataset="57492" ID-DOI="10.5852/ejt.2023.885.2183" ID-GBIF-Dataset="f1935cd8-83c6-45fd-aac5-8c6659a934aa" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177540" ID-ZooBank="A928AB6A-0029-4CEA-B941-C2238643835A" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201302250" checkinUser="felipe" docAuthor="Arriaga-Jiménez, Alfonsina, Zunino, Mario, Halffter, Gonzalo, Escobar-Hernández, Fernando & Rossini, Michele" docDate="2023" docId="03CF324D047E1F44FDC7F71DFCDEFB6D" docLanguage="en" docName="ejt-885_1-20.pdf" docOrigin="European Journal of Taxonomy 885" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratotrupes sturmi Jekel" docType="treatment" docVersion="7" lastPageNumber="12" masterDocId="FFF64A3504761F4FFF97F328FF93FFF7" masterDocTitle="A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur" masterLastPageNumber="20" masterPageNumber="1" pageNumber="9" updateTime="1699544954118" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9CC4C3AE1F16031F1F96869BE72504BF">A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur</mods:title>
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<mods:namePart id="CBB1375C2E6AF620895C1875BB1CB926">Arriaga-Jiménez, Alfonsina</mods:namePart>
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<mods:affiliation id="6DEB7A0B00A3DEEA2421625A3D5603C7">Insect Ecology Lab, School of Environmental and Rural Sciences, University of New England, Armidale NSW 2351, Australia.</mods:affiliation>
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<mods:nameIdentifier id="FE230A7CB8413D72B944E99BFE58CAD0" type="email">aarriaga@une.edu.au</mods:nameIdentifier>
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<mods:namePart id="8C2E5017F0DE037613E08A1A5B64BC96">Zunino, Mario</mods:namePart>
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<mods:affiliation id="BC7BAEAB7A9D2B7A63DEA61EE069363F">Instituto de Ecología A. C., Invited researcher, Carretera antigua a Coatepec 351, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="B1B376C5574CE4FCFE2436E6AD3EFF02" type="email">mariozunino@hotmail.it</mods:nameIdentifier>
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<mods:namePart id="5672F592C671593D102F6BC4672A217B">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="2353BCB77AB7A09840CAA051767CAB63">Deceased 11 March 2022. Former address: Aldama Street 17, Centro, 91500 Coatepec, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="FF6AAC329AC6B855DDD699B4A9BE06AD">Escobar-Hernández, Fernando</mods:namePart>
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<mods:affiliation id="81CC5F2765C34667174A52948FFD13EC">Red de Ecoetología, Instituto de Ecología, INECOL A. C. Xalapa, México, Carretera antigua a Coatepec 351, El Haya, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="E66FCA6871692D7CE58F74B350BE9818">Rossini, Michele</mods:namePart>
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<mods:affiliation id="CCF2875E0993547165DEB8EB6D11117A">Finnish Museum of Natural History (LUOMUS), University of Helsinki, Pohjoinen Rautatiekatu 13, Helsinki, 00014, Finland. & Department of Agronomy, Food, Natural resources, Animals and Environment (DAFNAE), University of Padova, Via dell'Università 16, 35020, Legnaro (PD), Italy.</mods:affiliation>
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<mods:title id="32A72D084AA28F183A7EF3C14E88FD1F">European Journal of Taxonomy</mods:title>
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<mods:date id="7997CD84C78EFF6D3241CCF5879F13FB">2023</mods:date>
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<treatment id="03CF324D047E1F44FDC7F71DFCDEFB6D" ID-DOI="http://doi.org/10.5281/zenodo.8182485" ID-GBIF-Taxon="212419183" ID-Zenodo-Dep="8182485" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D" httpUri="http://treatment.plazi.org/id/03CF324D047E1F44FDC7F71DFCDEFB6D" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<taxonomicName id="4C66F8D8047E1F47FDC7F71DFC70FBA7" ID-CoL="SQJ6" authority="(Jekel, 1865)" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[592,995,1077,1104]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi">
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<emphasis id="B9125F49047E1F47FDC7F71DFCD1FBB8" bold="true" box="[592,834,1077,1103]" italics="true" pageId="8" pageNumber="9">Ceratotrupes sturmi</emphasis>
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(
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<bibRefCitation id="EFF7FEAA047E1F47FCC5F71DFC48FBA7" author="Jekel H." box="[850,987,1077,1104]" pageId="8" pageNumber="9" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865</bibRefCitation>
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)
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<paragraph id="8BD9835B047E1F47FD5CF770FCFAFB84" blockId="8.[592,995,1077,1139]" box="[715,873,1112,1139]" pageId="8" pageNumber="9">
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<figureCitation id="135D9FDE047E1F47FD5CF770FC86FB84" box="[715,789,1112,1139]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Figs 3</figureCitation>
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,
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<figureCitation id="135D9FDE047E1F47FCB5F770FCFAFB84" box="[802,873,1112,1139]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[209,1382,582,1879]" captionTargetId="figure-82@14.[189,1398,565,1888]" captionTargetPageId="14" captionText="Fig. 5. Type specimens and original labels. A–B. Ceratotrupes sturmi (Jekel, 1865), holotype, ♂ (NHML). A. Dorsal habitus. B. Labels. C–D. Ceratotrupes mniszechi (Jekel, 1865), holotype, ♂ (MNHN). C. Dorsal habitus. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.8177552" httpUri="https://zenodo.org/record/8177552/files/figure.png" pageId="8" pageNumber="9">5A–B</figureCitation>
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<paragraph id="8BD9835B047E1F47FF2AF7B2FC84FB42" blockId="8.[189,975,1178,1240]" box="[189,791,1178,1205]" pageId="8" pageNumber="9">
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<taxonomicName id="4C66F8D8047E1F47FF2AF7B2FC87FB42" ID-CoL="6KD8P" authority="Jekel, 1865: 543" authorityName="Jekel" authorityPageNumber="543" authorityYear="1865" box="[189,788,1178,1205]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmii" subGenus="Ceratotrupes">
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<emphasis id="B9125F49047E1F47FF2AF7B2FDD4FB43" box="[189,583,1178,1204]" italics="true" pageId="8" pageNumber="9">Geotrupes (Ceratotrupes) sturmii</emphasis>
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<bibRefCitation id="EFF7FEAA047E1F47FDD8F7B2FC87FB42" author="Jekel H." box="[591,788,1178,1205]" pageId="8" pageNumber="9" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865: 543</bibRefCitation>
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.
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<treatmentCitation id="0AC7A54A047E1F47FF2AF796FCA3FB2F" author="Jekel H." box="[189,816,1213,1240]" page="544" pageId="8" pageNumber="9" year="1865">
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<taxonomicName id="4C66F8D8047E1F47FF2AF796FCA3FB2F" ID-CoL="3FTDX" authority="Jekel, 1865: 544" authorityName="Jekel" authorityPageNumber="544" authorityYear="1865" box="[189,816,1213,1240]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mniszechi" subGenus="Ceratotrupes">
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<emphasis id="B9125F49047E1F47FF2AF796FDFBFB20" box="[189,616,1213,1240]" italics="true" pageId="8" pageNumber="9">Geotrupes (Ceratotrupes) mniszechi</emphasis>
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<bibRefCitation id="EFF7FEAA047E1F47FDF8F795FCA3FB2F" author="Jekel H." box="[623,816,1213,1240]" pageId="8" pageNumber="9" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865: 544</bibRefCitation>
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(
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<figureCitation id="135D9FDE047E1F47FCD7F795FC51FB2F" box="[832,962,1213,1240]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[209,1382,582,1879]" captionTargetId="figure-82@14.[189,1398,565,1888]" captionTargetPageId="14" captionText="Fig. 5. Type specimens and original labels. A–B. Ceratotrupes sturmi (Jekel, 1865), holotype, ♂ (NHML). A. Dorsal habitus. B. Labels. C–D. Ceratotrupes mniszechi (Jekel, 1865), holotype, ♂ (MNHN). C. Dorsal habitus. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.8177552" httpUri="https://zenodo.org/record/8177552/files/figure.png" pageId="8" pageNumber="9">Fig. 5C–D</figureCitation>
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).
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<paragraph id="8BD9835B047E1F47FF2AF7D7FBBDFAEE" blockId="8.[189,1399,1278,1587]" box="[189,1070,1278,1305]" pageId="8" pageNumber="9">
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<taxonomicName id="4C66F8D8047E1F47FF2AF7D7FE19FAEF" ID-CoL="6KD8P" authorityName="Jekel" authorityYear="1865" box="[189,394,1279,1305]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi">
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<emphasis id="B9125F49047E1F47FF2AF7D7FE19FAEF" box="[189,394,1279,1305]" italics="true" pageId="8" pageNumber="9">Geotrupes sturmi</emphasis>
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–
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<treatmentCitation id="0AC7A54A047E1F47FE30F7D7FCAAFAEE" author="Gemminger M. & Harold E. von" box="[423,825,1278,1305]" page="1085" pageId="8" pageNumber="9" year="1869">
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<bibRefCitation id="EFF7FEAA047E1F47FE30F7D7FCAAFAEE" author="Gemminger M. & Harold E. von" box="[423,825,1278,1305]" pageId="8" pageNumber="9" refId="ref9661" refString="Gemminger M. & Harold E. von 1869. Catalogous Coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus IV. Scarabaeidae. E. H. Gummi, Monachii [Munich]. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Gemminger & Harold 1869: 1085</bibRefCitation>
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. —
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<treatmentCitation id="0AC7A54A047E1F47FCFEF7D7FBB9FAEE" author="Bates H. W." box="[873,1066,1279,1305]" page="112" pageId="8" pageNumber="9" year="1887">
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<bibRefCitation id="EFF7FEAA047E1F47FCFEF7D7FBB9FAEE" author="Bates H. W." box="[873,1066,1279,1305]" pageId="8" pageNumber="9" refId="ref9012" refString="Bates H. W. 1887. Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730" type="book" year="1887">Bates 1887: 112</bibRefCitation>
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.
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<paragraph id="8BD9835B047E1F47FF2AF60AFBC4FACB" blockId="8.[189,1399,1278,1587]" box="[189,1111,1314,1340]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup id="AB76A475047E1F47FF2AF60AFBC4FACB" box="[189,1111,1314,1340]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C66F8D8047E1F47FF2AF60AFE20FACB" ID-CoL="3FTDX" authorityName="Jekel" authorityYear="1865" box="[189,435,1314,1340]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F49047E1F47FF2AF60AFE20FACB" box="[189,435,1314,1340]" italics="true" pageId="8" pageNumber="9">Geotrupes mniszechi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC7A54A047E1F47FE47F60AFCF1FACB" author="Gemminger M. & Harold E. von" box="[464,866,1314,1340]" page="1083" pageId="8" pageNumber="9" year="1869">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FE47F60AFCF1FACB" author="Gemminger M. & Harold E. von" box="[464,866,1314,1340]" pageId="8" pageNumber="9" refId="ref9661" refString="Gemminger M. & Harold E. von 1869. Catalogous Coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus IV. Scarabaeidae. E. H. Gummi, Monachii [Munich]. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Gemminger & Harold 1869: 1083</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FC05F60AFBC0FACB" author="Bates H. W." box="[914,1107,1314,1340]" page="113" pageId="8" pageNumber="9" year="1887">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FC05F60AFBC0FACB" author="Bates H. W." box="[914,1107,1314,1340]" pageId="8" pageNumber="9" refId="ref9012" refString="Bates H. W. 1887. Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730" type="book" year="1887">Bates 1887: 113</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047E1F47FF2AF66DFCE5FAA8" blockId="8.[189,1399,1278,1587]" box="[189,886,1349,1376]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup id="AB76A475047E1F47FF2AF66DFCE5FAA8" box="[189,886,1349,1376]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C66F8D8047E1F47FF2AF66DFDC8FAA8" ID-CoL="SQJ6" authority="(Jekel, 1865)" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[189,603,1349,1375]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi" subGenus="Ceratotrupes">
|
||
<emphasis id="B9125F49047E1F47FF2AF66DFDC8FAA8" box="[189,603,1349,1375]" italics="true" pageId="8" pageNumber="9">Ceratophyus (Ceratotrupes) sturmi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC7A54A047E1F47FDEFF66EFCE1FAA8" author="Boucomont A." box="[632,882,1349,1376]" page="11" pageId="8" pageNumber="9" year="1902">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FDEFF66EFCE1FAA8" author="Boucomont A." box="[632,882,1349,1376]" pageId="8" pageNumber="9" pagination="1 - 20" refId="ref9252" refString="Boucomont A. 1902. Coleoptera Lamellicornia Fam. Geotrupidae. Genera Insectorum. Proceedings of the United States National Museum 7: 1 - 20." type="journal article" year="1902">Boucomont 1902: 11</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047E1F47FF2AF640FC0DFA75" blockId="8.[189,1399,1278,1587]" box="[189,926,1384,1411]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup id="AB76A475047E1F47FF2AF640FC0DFA75" box="[189,926,1384,1411]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C66F8D8047E1F47FF2AF640FD17FA75" ID-CoL="3FTDX" authority="Jekel, 1865: 544" authorityName="Jekel" authorityPageNumber="544" authorityYear="1865" box="[189,644,1384,1411]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mniszechi" subGenus="Ceratotrupes">
|
||
<emphasis id="B9125F49047E1F47FF2AF640FEC0FA75" box="[189,339,1384,1410]" italics="true" pageId="8" pageNumber="9">Ceratophyus</emphasis>
|
||
(
|
||
<emphasis id="B9125F49047E1F47FEF3F641FE6CFA74" box="[356,511,1385,1411]" italics="true" pageId="8" pageNumber="9">Ceratotrupes</emphasis>
|
||
)
|
||
<emphasis id="B9125F49047E1F47FD87F640FD17FA75" box="[528,644,1384,1410]" italics="true" pageId="8" pageNumber="9">mniszechi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC7A54A047E1F47FD36F641FC09FA75" author="Boucomont A." box="[673,922,1384,1411]" page="11" pageId="8" pageNumber="9" year="1902">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FD36F641FC09FA75" author="Boucomont A." box="[673,922,1384,1411]" pageId="8" pageNumber="9" pagination="1 - 20" refId="ref9252" refString="Boucomont A. 1902. Coleoptera Lamellicornia Fam. Geotrupidae. Genera Insectorum. Proceedings of the United States National Museum 7: 1 - 20." type="journal article" year="1902">Boucomont 1902: 11</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047E1F47FF2AF6A4FCA5FA1B" blockId="8.[189,1399,1278,1587]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup id="AB76A475047E1F47FF2AF6A4FCA5FA1B" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C66F8D8047E1F47FF2AF6A4FE38FA52" ID-CoL="SQJ6" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[189,427,1420,1446]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047E1F47FF2AF6A4FE38FA52" box="[189,427,1420,1446]" italics="true" pageId="8" pageNumber="9">Ceratotrupes sturmi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC7A54A047E1F47FE5DF6A4FD54FA51" author="Boucomont A." box="[458,711,1420,1446]" page="20" pageId="8" pageNumber="9" year="1912">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FE5DF6A4FD54FA51" author="Boucomont A." box="[458,711,1420,1446]" pageId="8" pageNumber="9" pagination="47" refId="ref9279" refString="Boucomont A. 1912. Scarabaeidae: Taurocerastinae, Geotrupinae. In: Junk W. & Schenkling S. (eds) Coleopterum Catalogus, Pars 46: 47. W. Junk, Berlin." type="journal article" year="1912">Boucomont 1912: 20</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FD6FF6A3FB9CFA51" author="Blackwelder R. E." box="[760,1039,1419,1446]" page="220" pageId="8" pageNumber="9" year="1944">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FD6FF6A3FB9CFA51" author="Blackwelder R. E." box="[760,1039,1419,1446]" pageId="8" pageNumber="9" pagination="189 - 341" refId="ref9097" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185 (2): 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 220</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FBA8F6A3FE89FA3E" author="Halffter G. & Martinez A." page="155" pageId="8" pageNumber="9" year="1962">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FBA8F6A3FE89FA3E" author="Halffter G. & Martinez A." pageId="8" pageNumber="9" pagination="145 - 159" refId="ref9786" refString="Halffter G. & Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter & Martínez 1962: 155</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FEDFF687FD88FA3E" author="Howden H. F." box="[328,539,1455,1481]" page="47" pageId="8" pageNumber="9" year="1964">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FEDFF687FD88FA3E" author="Howden H. F." box="[328,539,1455,1481]" pageId="8" pageNumber="9" pagination="5 - 91" refId="ref9941" refString="Howden H. F. 1964. The Geotrupinae of North and Central America. The Memoirs of the Entomological Society of Canada 96 (S 39): 5 - 91. https: // doi. org / 10.4039 / entm 9639 fv" type="journal article" year="1964">Howden 1964: 47</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC7A54A047E1F47FDB1F687FD0CFA3E" author="Howden H. F." box="[550,671,1454,1481]" page="569" pageId="8" pageNumber="9" year="1974">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FDB1F687FD0CFA3E" author="Howden H. F." box="[550,671,1454,1481]" pageId="8" pageNumber="9" pagination="567 - 575" refId="ref9986" refString="Howden H. F. 1974. Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 52 (5): 567 - 575. https: // doi. org / 10.1139 / z 74 - 073" type="journal article" year="1974">1974: 569</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC7A54A047E1F47FD3EF687FCB2FA3E" author="Howden H. F." box="[681,801,1455,1481]" page="100" pageId="8" pageNumber="9" year="2003">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FD3EF687FCB2FA3E" author="Howden H. F." box="[681,801,1455,1481]" pageId="8" pageNumber="9" pagination="95 - 108" refId="ref10098" refString="Howden H. F. 2003. Subfamilia Geotrupinae. In: Moron M. A. (ed.) Atlas de los Escarabajos de Mexico. Coleoptera: Lamellicornia Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 95 - 108. Argania, Barcelona." type="book chapter" year="2003">2003: 100</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FCD8F687FBB1FA3E" author="Zunino M." box="[847,1058,1455,1481]" page="109" pageId="8" pageNumber="9" year="1984">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FCD8F687FBB1FA3E" author="Zunino M." box="[847,1058,1455,1481]" pageId="8" pageNumber="9" pagination="9 - 162" refId="ref11069" refString="Zunino M. 1984. Sistematica generica dei Geotrupinae filogenesi della sottofamiglia e considerazioni biogeografiche. Bollettino del Museo regionale di Scienze naturali 2 (1): 9 - 162." type="journal article" year="1984">Zunino 1984: 109</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FBC7F687FEA6FA1B" author="Trotta-Moreu N. & Lobo J. M. & Cabrero-Sanudo F. J." page="42" pageId="8" pageNumber="9" year="2008">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FBC7F687FEA6FA1B" author="Trotta-Moreu N. & Lobo J. M. & Cabrero-Sanudo F. J." pageId="8" pageNumber="9" pagination="39 - 65" refId="ref10890" refString="Trotta-Moreu N., Lobo J. M. & Cabrero-Sanudo F. J. 2008. Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico. Acta zoologica mexicana 24 (2): 39 - 65. https: // doi. org / 10.21829 / azm. 2008.242702" type="journal article" year="2008">
|
||
Trotta-Moreu
|
||
<emphasis id="B9125F49047E1F47FB64F687FABFFA3E" box="[1267,1324,1455,1481]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2008: 42–43
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 48, 54. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FE2AF6FAFCA1FA1B" author="Trotta-Moreu N. & Lobo J. M." box="[445,818,1490,1516]" page="46" pageId="8" pageNumber="9" year="2010">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FE2AF6FAFCA1FA1B" author="Trotta-Moreu N. & Lobo J. M." box="[445,818,1490,1516]" pageId="8" pageNumber="9" pagination="42 - 49" refId="ref10835" refString="Trotta-Moreu N. & Lobo J. M. 2010. Deriving the species richness distribution of Geotrupinae (Coleoptera: Scarabaeoidea) in Mexico from the overlap of individual model predictions. Environmental Entomology 39 (1): 42 - 49. https: // doi. org / 10.1603 / EN 08179" type="journal article" year="2010">Trotta-Moreu & Lobo 2010: 46</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047E1F47FF2AF6DDFBFDF9C5" blockId="8.[189,1399,1278,1587]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup id="AB76A475047E1F47FF2AF6DDFBFDF9C5" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C66F8D8047E1F47FF2AF6DDFE45F9F8" ID-CoL="3FTDX" authority="Jekel, 1865: 544" authorityName="Jekel" authorityPageNumber="544" authorityYear="1865" box="[189,470,1525,1551]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F49047E1F47FF2AF6DDFE45F9F8" box="[189,470,1525,1551]" italics="true" pageId="8" pageNumber="9">Ceratotrupes mniszechi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC7A54A047E1F47FE6DF6DEFD6FF9F8" author="Boucomont A." box="[506,764,1525,1552]" page="20" pageId="8" pageNumber="9" year="1912">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FE6DF6DEFD6FF9F8" author="Boucomont A." box="[506,764,1525,1552]" pageId="8" pageNumber="9" pagination="47" refId="ref9279" refString="Boucomont A. 1912. Scarabaeidae: Taurocerastinae, Geotrupinae. In: Junk W. & Schenkling S. (eds) Coleopterum Catalogus, Pars 46: 47. W. Junk, Berlin." type="journal article" year="1912">Boucomont 1912: 20</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FCA5F6DDFBDDF9F8" author="Blackwelder R. E." box="[818,1102,1525,1551]" page="220" pageId="8" pageNumber="9" year="1944">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FCA5F6DDFBDDF9F8" author="Blackwelder R. E." box="[818,1102,1525,1551]" pageId="8" pageNumber="9" pagination="189 - 341" refId="ref9097" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185 (2): 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 220</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FB13F6DDFEF1F9C4" author="Halffter G. & Martinez A." page="153" pageId="8" pageNumber="9" year="1962">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FB13F6DDFEF1F9C4" author="Halffter G. & Martinez A." pageId="8" pageNumber="9" pagination="145 - 159" refId="ref9786" refString="Halffter G. & Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter & Martínez 1962: 153</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 155, 157. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FD9EF530FD4CF9C5" author="Howden H. F." box="[521,735,1560,1586]" page="47" pageId="8" pageNumber="9" year="1964">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FD9EF530FD4CF9C5" author="Howden H. F." box="[521,735,1560,1586]" pageId="8" pageNumber="9" pagination="5 - 91" refId="ref9941" refString="Howden H. F. 1964. The Geotrupinae of North and Central America. The Memoirs of the Entomological Society of Canada 96 (S 39): 5 - 91. https: // doi. org / 10.4039 / entm 9639 fv" type="journal article" year="1964">Howden 1964: 47</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC7A54A047E1F47FD7CF530FCF6F9C4" author="Howden H. F." box="[747,869,1560,1587]" page="569" pageId="8" pageNumber="9" year="1974">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FD7CF530FCF6F9C4" author="Howden H. F." box="[747,869,1560,1587]" pageId="8" pageNumber="9" pagination="567 - 575" refId="ref9986" refString="Howden H. F. 1974. Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 52 (5): 567 - 575. https: // doi. org / 10.1139 / z 74 - 073" type="journal article" year="1974">1974: 569</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AC7A54A047E1F47FC03F530FBF9F9C5" author="Zunino M." box="[916,1130,1560,1586]" page="109" pageId="8" pageNumber="9" year="1984">
|
||
<bibRefCitation id="EFF7FEAA047E1F47FC03F530FBF9F9C5" author="Zunino M." box="[916,1130,1560,1586]" pageId="8" pageNumber="9" pagination="9 - 162" refId="ref11069" refString="Zunino M. 1984. Sistematica generica dei Geotrupinae filogenesi della sottofamiglia e considerazioni biogeografiche. Bollettino del Museo regionale di Scienze naturali 2 (1): 9 - 162." type="journal article" year="1984">Zunino 1984: 109</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047E1F47FF2AF571FA95F84B" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="8BD9835B047E1F47FF2AF571FEA5F984" blockId="8.[189,310,1625,1651]" box="[189,310,1625,1651]" pageId="8" pageNumber="9">
|
||
<heading id="D0913437047E1F47FF2AF571FEA5F984" bold="true" box="[189,310,1625,1651]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B9125F49047E1F47FF2AF571FEA5F984" bold="true" box="[189,310,1625,1651]" pageId="8" pageNumber="9">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047E1F47FF2AF5A0FA95F84B" blockId="8.[187,1399,1672,1980]" pageId="8" pageNumber="9">
|
||
Within the genus,
|
||
<taxonomicName id="4C66F8D8047E1F47FE1CF5A1FE64F955" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[395,503,1673,1699]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047E1F47FE1CF5A1FE64F955" box="[395,503,1673,1699]" italics="true" pageId="8" pageNumber="9">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
is here considered to be related to
|
||
<taxonomicName id="4C66F8D8047E1F47FC16F5A1FB95F955" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[897,1030,1672,1699]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047E1F47FC16F5A1FB95F955" box="[897,1030,1672,1699]" italics="true" pageId="8" pageNumber="9">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047E1F47FB9BF5A1FBF4F954" box="[1036,1127,1673,1699]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and it can be separated by the following diagnostic characters: posterior edge of pronotum with a thin, medial margin (
|
||
<figureCitation id="135D9FDE047E1F47FA84F583FF47F91E" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, C</figureCitation>
|
||
); clypeal horn slender, laterally narrower, tip obtuse (
|
||
<figureCitation id="135D9FDE047E1F47FCDEF5E7FC5FF91E" box="[841,972,1743,1769]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, D</figureCitation>
|
||
); pronotal projection narrower, with convergent sides (
|
||
<figureCitation id="135D9FDE047E1F47FE06F5DAFD80F8FB" box="[401,531,1778,1804]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, D</figureCitation>
|
||
); elytral interstriae unpunctured. Furthermore, the male genitalia exhibit a series of clear morphological differences: distal tip of right ventral lobe of phallobase curved medially (
|
||
<figureCitation id="135D9FDE047E1F47FF51F410FE89F8A5" box="[198,282,1848,1874]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3J</figureCitation>
|
||
); shorter branch of the bifurcation of the right lobe of the phallobase longer than in
|
||
<taxonomicName id="4C66F8D8047E1F47FB78F411FAE5F8A5" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[1263,1398,1848,1875]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gonzaloi">
|
||
<emphasis id="B9125F49047E1F47FB78F411FAE5F8A5" box="[1263,1398,1848,1875]" italics="true" pageId="8" pageNumber="9">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135D9FDE047E1F47FF51F473FEB0F881" box="[198,291,1883,1910]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3L</figureCitation>
|
||
); different shape of the parameres (
|
||
<figureCitation id="135D9FDE047E1F47FD45F473FCBEF881" box="[722,813,1883,1910]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3F</figureCitation>
|
||
; arrows indicate the main differences); in lateral view, left paramere with a distinct lobe beneath the apical spur (
|
||
<figureCitation id="135D9FDE047E1F47FC32F457FB90F86E" box="[933,1027,1919,1945]" captionStart="Fig" captionStartId="10.[189,232,1752,1778]" captionTargetBox="[303,1294,293,1672]" captionTargetId="figure-19@10.[283,1305,265,1711]" captionTargetPageId="10" captionText="Fig. 3. Ceratotrupes sturmi (Jekel, 1865) (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. sturmi and C. gonzaloi sp. nov. G. Right paramere in lateral view; arrow indicates lobe beneath apical spur. H. Tergite. I. Pleurotergites. J. Phallobase in ventral view; arrow indicates curved margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates long bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="8" pageNumber="9">Fig. 3G</figureCitation>
|
||
). It is noteworthy that
|
||
<taxonomicName id="4C66F8D8047E1F47FA9EF457FAE5F86F" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1289,1398,1919,1945]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047E1F47FA9EF457FAE5F86F" box="[1289,1398,1919,1945]" italics="true" pageId="8" pageNumber="9">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C66F8D8047E1F47FF78F48AFEE5F84B" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[239,374,1954,1980]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gonzaloi">
|
||
<emphasis id="B9125F49047E1F47FF78F48AFEE5F84B" box="[239,374,1954,1980]" italics="true" pageId="8" pageNumber="9">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
share with
|
||
<taxonomicName id="4C66F8D8047E1F47FE69F48AFDEEF84B" authorityName="Halffter & Martinez" authorityYear="1962" box="[510,637,1954,1980]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bolivari">
|
||
<emphasis id="B9125F49047E1F47FE69F48AFDEEF84B" box="[510,637,1954,1980]" italics="true" pageId="8" pageNumber="9">C. bolivari</emphasis>
|
||
</taxonomicName>
|
||
the presence of an apical, acute tooth on left paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047F1F46FF2AF221FAFBFB8B" pageId="9" pageNumber="10" type="materials_examined">
|
||
<paragraph id="8BD9835B047F1F46FF2AF221FE7FFED5" blockId="9.[189,492,264,291]" box="[189,492,264,291]" pageId="9" pageNumber="10">
|
||
<heading id="D0913437047F1F46FF2AF221FE7FFED5" bold="true" box="[189,492,264,291]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B9125F49047F1F46FF2AF221FE7FFED5" bold="true" box="[189,492,264,291]" pageId="9" pageNumber="10">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF7DF21FFE7DFE29" blockId="9.[188,1399,311,478]" pageId="9" pageNumber="10">
|
||
<materialsCitation id="3B0E8906047F1F46FF7DF21FFE7DFE29" ID-GBIF-Occurrence="4164095348" collectionCode="NHML" country="Mexico" location="Juquila" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B9125F49047F1F46FF7DF21FFE67FEA6" bold="true" box="[234,500,311,337]" pageId="9" pageNumber="10">
|
||
<typeStatus id="54DD3DF9047F1F46FF7DF21FFECFFEA6" box="[234,348,311,337]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C66F8D8047F1F46FE13F21FFE67FEA6" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[388,500,311,337]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047F1F46FE13F21FFE67FEA6" bold="true" box="[388,500,311,337]" italics="true" pageId="9" pageNumber="10">
|
||
<originalDetermination id="A8783BA7047F1F46FE13F21FFE67FEA6" box="[388,500,311,337]" pageId="9" pageNumber="10">C. sturmi</originalDetermination>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<collectingCountry id="F371C3CB047F1F46FF2AF272FEA2FE83" box="[189,305,346,372]" name="Mexico" pageId="9" pageNumber="10">MEXICO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D6048D2047F1F46FED0F273FEF3FE82" box="[327,352,347,373]" pageId="9" pageNumber="10" type="male">♂</specimenCount>
|
||
; “
|
||
<location id="8EB9D580047F1F46FEE1F272FE5EFE83" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FEE1F272FE5EFE83" box="[374,461,346,372]" country="Mexico" name="Juquila" pageId="9" pageNumber="10">Juquila</location>
|
||
[printed] /
|
||
<collectingRegion id="49A24DB9047F1F46FDD8F272FD3FFE83" box="[591,684,346,372]" country="Mexico" name="Mexico" pageId="9" pageNumber="10">Mexico</collectingRegion>
|
||
, Salle Coll. [printed] / 844 [green label, printed] / Sp. figured. [printed] /
|
||
<taxonomicName id="4C66F8D8047F1F46FEADF256FDFFFE60" authority="Jekel" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[314,620,381,408]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">Ceratotrupes Sturmi Jekel</taxonomicName>
|
||
, male symbol [handwritten] /
|
||
<taxonomicName id="4C66F8D8047F1F46FC5CF256FB6BFE60" authority="Jekel" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[971,1272,381,408]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">Ceratotrupes sturmi Jekel</taxonomicName>
|
||
, type apud Sallé [handwritten] / B.C.A. Col., II(2).,
|
||
<taxonomicName id="4C66F8D8047F1F46FD01F289FCF2FE4C" authorityName="Jekel" authorityYear="1865" box="[662,865,417,443]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">Geotrupes sturmi</taxonomicName>
|
||
[printed] Jek / Holotype [white label with red border, printed]”;
|
||
<collectionCode id="ED771B9E047F1F46FE07F2ECFE79FE29" box="[400,490,452,478]" collectionName="Libya, Tripoli, Natural History Museum" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="9" pageNumber="10">NHML</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF7DF12DFCF9FD7E" blockId="9.[189,1399,517,649]" pageId="9" pageNumber="10">
|
||
<materialsCitation id="3B0E8906047F1F46FF7DF12DFCF9FD7E" ID-GBIF-Occurrence="4164095342" collectionCode="MNHN" country="Mexico" location="MEXICO" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1">
|
||
<emphasis id="B9125F49047F1F46FF7DF12DFD8CFDD7" bold="true" box="[234,543,517,544]" pageId="9" pageNumber="10">
|
||
Holotype of
|
||
<taxonomicName id="4C66F8D8047F1F46FE13F12DFD8CFDD7" box="[388,543,517,544]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F49047F1F46FE13F12DFD8CFDD7" bold="true" box="[388,543,517,544]" italics="true" pageId="9" pageNumber="10">
|
||
<originalDetermination id="A8783BA7047F1F46FE13F12DFD8CFDD7" box="[388,543,517,544]" pageId="9" pageNumber="10">C. mniszechi</originalDetermination>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<collectingCountry id="F371C3CB047F1F46FF2AF101FEA2FDB4" box="[189,305,553,579]" name="Mexico" pageId="9" pageNumber="10">MEXICO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D6048D2047F1F46FEC4F101FEFFFDB4" box="[339,364,553,579]" pageId="9" pageNumber="10" type="male">♂</specimenCount>
|
||
; “Collection Mniszech [printed] / Ex-Musaeo VAN LANSBERGE [printed] / Geotr.
|
||
<taxonomicName id="4C66F8D8047F1F46FF2AF164FDBCFD91" authority="Jekel" authorityName="Jekel" authorityYear="1865" box="[189,559,588,614]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mniszechi" subGenus="Ceratotrupes">(Ceratotrupes) Mniszechi Jekel</taxonomicName>
|
||
[green label, handwritten] / Holo TYPUS
|
||
<taxonomicName id="4C66F8D8047F1F46FBA7F164FAE5FD91" authority="Jek" authorityName="Jekel" authorityYear="1865" box="[1072,1398,588,614]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mniszechi">Ceratotrupes mniszechi Jek</taxonomicName>
|
||
[red label, handwritten except TYPUS, printed]”;
|
||
<collectionCode id="ED771B9E047F1F46FC9EF147FCF4FD7E" box="[777,871,623,649]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF198FDA7FD3D" blockId="9.[189,564,688,714]" box="[189,564,688,714]" pageId="9" pageNumber="10">
|
||
<heading id="D0913437047F1F46FF2AF198FDA7FD3D" bold="true" box="[189,564,688,714]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B9125F49047F1F46FF2AF198FDA7FD3D" bold="true" box="[189,564,688,714]" pageId="9" pageNumber="10">Additional material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF1F7FAFBFB8B" blockId="9.[186,1399,735,1149]" pageId="9" pageNumber="10">
|
||
<materialsCitation id="3B0E8906047F1F46FF2AF1F7FE45FCEB" ID-GBIF-Occurrence="4164095318" collectingDate="1971-05-27" collectingDateMax="1971-05-30" collectingDateMin="1971-05-27" collectionCode="CMNC" collectorName="H. F. Howden" country="Mexico" location="Rt. 131" pageId="9" pageNumber="10" specimenCode="CMNC 00031072" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
|
||
<collectingCountry id="F371C3CB047F1F46FF2AF1F7FEA2FD0E" box="[189,305,735,761]" name="Mexico" pageId="9" pageNumber="10">MEXICO</collectingCountry>
|
||
–
|
||
<emphasis id="B9125F49047F1F46FED8F1F7FE3DFD0E" bold="true" box="[335,430,735,761]" pageId="9" pageNumber="10">
|
||
<collectingRegion id="49A24DB9047F1F46FED8F1F7FE3DFD0E" box="[335,430,735,761]" country="Mexico" name="Oaxaca" pageId="9" pageNumber="10">Oaxaca</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D6048D2047F1F46FE50F1F7FE65FD0E" box="[455,502,735,761]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<locationDeviation id="9A6B9A79047F1F46FD99F1F7FC89FD0E" box="[526,794,735,761]" location="Juchatengo" pageId="9" pageNumber="10">
|
||
<quantity id="4C9E2EBE047F1F46FD99F1F7FDC7FD0E" box="[526,596,735,761]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="9" pageNumber="10" unit="mi" value="20.0">20 mi</quantity>
|
||
S of Juchatengo
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EB9D580047F1F46FCB1F1F7FC13FD0E" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FCB1F1F7FC13FD0E" box="[806,896,735,761]" country="Mexico" name="Rt. 131" pageId="9" pageNumber="10" stateProvince="Oaxaca">Rt. 131</location>
|
||
. 6000’ alt.
|
||
<date id="FFD8A59B047F1F46FB95F1F7FB5DFD0E" box="[1026,1230,735,761]" pageId="9" pageNumber="10" value="1971-05-27" valueMax="1971-05-30" valueMin="1971-05-27">
|
||
<collectingDate id="EF9C5C73047F1F46FB95F1F7FB5DFD0E" box="[1026,1230,735,761]" pageId="9" pageNumber="10" value="1971-05-27" valueMax="1971-05-30" valueMin="1971-05-27">27–30 May 1971</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="2693E68D047F1F46FB4DF1F7FAE5FD0E" box="[1242,1398,735,761]" pageId="9" pageNumber="10">H.F. Howden</collectorName>
|
||
leg.”;
|
||
<specimenCode id="DBC02B20047F1F46FE93F02AFE45FCEB" box="[260,470,770,796]" collectionCode="CMNC" pageId="9" pageNumber="10">CMNC 00031072</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FE79F02AFB2BFCEB" ID-GBIF-Occurrence="4164095315" box="[494,1208,770,797]" collectingDate="1971-05-27" collectingDateMax="1971-05-30" collectingDateMin="1971-05-27" collectionCode="CMNC" collectorName="H. F. Howden" country="Mexico" location="Rt. 131" pageId="9" pageNumber="10" specimenCode="CMNC 00031071" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
|
||
<specimenCount id="9D6048D2047F1F46FE79F02AFD89FCEA" box="[494,538,770,797]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="DBC02B20047F1F46FC71F02AFB2BFCEB" box="[998,1208,770,796]" collectionCode="CMNC" pageId="9" pageNumber="10">CMNC 00031071</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FB47F02AFAF6FC94" ID-GBIF-Occurrence="4164095350" collectingDate="1995-06-24" collectingDateMax="1995-06-26" collectingDateMin="1995-06-24" collectionCode="FMNH" collectorName="A. Perez & J. Blackaller" elevation="1900" location="km 164.5, carr. 131" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
|
||
<specimenCount id="9D6048D2047F1F46FB47F02AFB6DFCEA" box="[1232,1278,770,797]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047F1F46FA82F02AFAE1FCEB" box="[1301,1394,770,796]" pageId="9" pageNumber="10">Oaxaca</collectingRegion>
|
||
,
|
||
<location id="8EB9D580047F1F46FF2AF00DFE0FFCC8" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FF2AF00DFE0FFCC8" box="[189,412,805,832]" name="km 164.5, carr. 131" pageId="9" pageNumber="10" stateProvince="Oaxaca">km 164.5, carr. 131</location>
|
||
,
|
||
<location id="8EB9D580047F1F46FE31F00DFD33FCC8" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FE31F00DFD33FCC8" box="[422,672,805,831]" name="Oax. - Pto. Escondido" pageId="9" pageNumber="10" stateProvince="Oaxaca">Oax.–Pto. Escondido</location>
|
||
.
|
||
<elevation id="004B6468047F1F46FD3FF00DFC10FCC8" box="[680,899,805,832]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="9" pageNumber="10" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
|
||
<quantity id="4C9E2EBE047F1F46FD3FF00DFCDBFCB7" box="[680,840,805,832]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="9" pageNumber="10" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<date id="FFD8A59B047F1F46FC1FF00DFBDAFCB7" box="[904,1097,805,832]" pageId="9" pageNumber="10" value="1995-06-24" valueMax="1995-06-26" valueMin="1995-06-24">
|
||
<collectingDate id="EF9C5C73047F1F46FC1FF00DFBDAFCB7" box="[904,1097,805,832]" pageId="9" pageNumber="10" value="1995-06-24" valueMax="1995-06-26" valueMin="1995-06-24">24–26 Jun. 1995</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="2693E68D047F1F46FBC7F00DFB21FCB7" box="[1104,1202,805,832]" pageId="9" pageNumber="10">A. Perez</collectorName>
|
||
and
|
||
<collectorName id="2693E68D047F1F46FB71F00EFAE4FCC8" box="[1254,1399,805,832]" pageId="9" pageNumber="10">J. Blackaller</collectorName>
|
||
leg. Bosque de Encino, en excremento [oak forest, in excrement]”; http://id.luomus.fi/NC.04;
|
||
<collectionCode id="ED771B9E047F1F46FA87F061FAF6FC94" box="[1296,1381,841,867]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10" type="Museum">FMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FF2AF044FB7EFC5E" ID-GBIF-Occurrence="4164095330" collectingDate="2004-05-26" collectionCode="FMNH" collectorName="G. Nogueira" determinerName="M. A. Moron" elevation="1800" location="Cerro del Vidrio" municipality="Cerro del Vidrio" pageId="9" pageNumber="10" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="9D6048D2047F1F46FF2AF044FF7BFC71" box="[189,232,876,902]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D6048D2047F1F46FF65F044FE8FFC71" box="[242,284,876,902]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
; “
|
||
<collectingMunicipality id="6BBD1921047F1F46FEA5F044FE61FC71" box="[306,498,876,902]" country="Mexico" name="Cerro del Vidrio" pageId="9" pageNumber="10" stateProvince="Oaxaca">Cerro del Vidrio</collectingMunicipality>
|
||
,
|
||
<elevation id="004B6468047F1F46FE6CF044FD1FFC71" box="[507,652,876,902]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="m" value="1800.0">
|
||
<quantity id="4C9E2EBE047F1F46FE6CF044FDC2FC71" box="[507,593,876,902]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="m" value="1800.0">1800 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<date id="FFD8A59B047F1F46FD06F043FCB8FC71" box="[657,811,875,902]" pageId="9" pageNumber="10" value="2004-05-26">
|
||
<collectingDate id="EF9C5C73047F1F46FD06F043FCB8FC71" box="[657,811,875,902]" pageId="9" pageNumber="10" value="2004-05-26">26 May 2004</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="2693E68D047F1F46FCA4F044FC52FC71" box="[819,961,876,902]" pageId="9" pageNumber="10">G. Nogueira</collectorName>
|
||
leg. /
|
||
<taxonomicName id="4C66F8D8047F1F46FB95F044FAE2FC72" authority="Jekel, 1866" authorityName="Jekel" authorityYear="1866" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1026,1393,875,902]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<originalDetermination id="A8783BA7047F1F46FB95F044FAE2FC72" box="[1026,1393,875,902]" pageId="9" pageNumber="10">Ceratotrupes sturmi Jekel, 1866</originalDetermination>
|
||
</taxonomicName>
|
||
,
|
||
<determinerName id="7E056F9F047F1F46FF2AF0A7FEC1FC5E" box="[189,338,911,937]" pageId="9" pageNumber="10">M.A. Morón</determinerName>
|
||
det.
|
||
<date id="FFD8A59B047F1F46FE1DF0A7FD86FC5E" box="[394,533,911,937]" pageId="9" pageNumber="10" value="2007-05">Mayo 2007</date>
|
||
”; http://id.luomus.fi/NC.05, http://id.luomus.fi/NC.06;
|
||
<collectionCode id="ED771B9E047F1F46FB0FF0A7FB7EFC5E" box="[1176,1261,911,937]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10" type="Museum">FMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FA92F0A7FE4CFC18" ID-GBIF-Occurrence="4164095312" collectingDate="1993-06-17" collectingDateMax="1993-06-20" collectingDateMin="1993-06-17" collectionCode="CEMT" collectorName="L. Delgado" elevation="1575" location="Oaxaca - Puerto Escondido" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D6048D2047F1F46FA92F0A7FAA0FC5E" box="[1285,1331,911,937]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
|
||
; “Rm 164.5, carr.
|
||
<location id="8EB9D580047F1F46FEDDF09AFD16FC3B" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FEDDF09AFD16FC3B" box="[330,645,946,972]" name="Oaxaca - Puerto Escondido" pageId="9" pageNumber="10">Oaxaca–Puerto Escondido</location>
|
||
.
|
||
<elevation id="004B6468047F1F46FD06F09AFCE0FC3B" box="[657,883,946,973]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="9" pageNumber="10" unit="m" value="1575.0" valueMax="1650.0" valueMin="1500.0">
|
||
<quantity id="4C9E2EBE047F1F46FD06F09AFCA7FC3A" box="[657,820,946,973]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="9" pageNumber="10" unit="m" value="1575.0" valueMax="1650.0" valueMin="1500.0">1500–1650 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<date id="FFD8A59B047F1F46FCECF09AFBD0FC3B" box="[891,1091,946,973]" pageId="9" pageNumber="10" value="1993-06-17" valueMax="1993-06-20" valueMin="1993-06-17">
|
||
<collectingDate id="EF9C5C73047F1F46FCECF09AFBD0FC3B" box="[891,1091,946,973]" pageId="9" pageNumber="10" value="1993-06-17" valueMax="1993-06-20" valueMin="1993-06-17">17–20 Jun. 1993</collectingDate>
|
||
</date>
|
||
. coprotrampa [coprotrap].
|
||
<collectorName id="2693E68D047F1F46FF29F0FEFED3FC18" box="[190,320,981,1008]" pageId="9" pageNumber="10">L. Delgado</collectorName>
|
||
leg.”;
|
||
<collectionCode id="ED771B9E047F1F46FE19F0FDFE4CFC18" box="[398,479,981,1007]" pageId="9" pageNumber="10">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FE61F0FDFBADFC18" ID-GBIF-Occurrence="4164095349" box="[502,1086,981,1008]" collectingDate="1993-06-17" collectingDateMax="1993-06-20" collectingDateMin="1993-06-17" collectionCode="CEMT" collectorName="L. Delgado" elevation="1575" location="Oaxaca - Puerto Escondido" pageId="9" pageNumber="10" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D6048D2047F1F46FE61F0FDFDB1FC07" box="[502,546,981,1008]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED771B9E047F1F46FC7AF0FDFBADFC18" box="[1005,1086,981,1007]" pageId="9" pageNumber="10">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FBC2F0FDFB40FBE4" ID-GBIF-Occurrence="4164095322" collectingDate="2006-06-01" collectingDateMax="2006-06-24" collectingDateMin="2006-06-01" collectionCode="CEMT" collectorName="D. Curoe" elevation="1710" latitude="16.201666" location="Oaxaca" longLatPrecision="92" longitude="-97.13617" municipality="Oaxaca" pageId="9" pageNumber="10" specimenCount="15" specimenCount-female="15">
|
||
<specimenCount id="9D6048D2047F1F46FBC2F0FDFB35FC07" box="[1109,1190,981,1008]" pageId="9" pageNumber="10" type="female">15 ♀♀</specimenCount>
|
||
; “
|
||
<collectingMunicipality id="6BBD1921047F1F46FB29F0FDFA88FC18" box="[1214,1307,981,1007]" country="Mexico" latitude="16.201666" longLatPrecision="92" longitude="-97.13617" name="Oaxaca" pageId="9" pageNumber="10" stateProvince="Oaxaca">Oaxaca</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="9A6B9A79047F1F46FAB1F0FDFEA9FBE5" location="Lachao." pageId="9" pageNumber="10">
|
||
<quantity id="4C9E2EBE047F1F46FAB1F0FDFAF3FC18" box="[1318,1376,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="km" value="4.0">4 km</quantity>
|
||
S of Lachao.
|
||
</locationDeviation>
|
||
<geoCoordinate id="EE52E59C047F1F46FED7F0D0FE56FBE4" box="[320,453,1016,1043]" degrees="16" direction="north" minutes="12.1" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="16.201666">16°12.1´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE52E59C047F1F46FE47F0DFFDF7FBE4" box="[464,612,1015,1043]" degrees="97" direction="west" minutes="08.17" orientation="longitude" pageId="9" pageNumber="10" precision="9" value="-97.13617">97°08.17′ W</geoCoordinate>
|
||
.
|
||
<elevation id="004B6468047F1F46FDF9F0D1FC90FBE5" box="[622,771,1016,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.71" pageId="9" pageNumber="10" unit="m" value="1710.0">
|
||
<quantity id="4C9E2EBE047F1F46FDF9F0D1FD55FBE4" box="[622,710,1017,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.71" pageId="9" pageNumber="10" unit="m" value="1710.0">1710 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<date id="FFD8A59B047F1F46FC9DF0D1FC2CFBE5" box="[778,959,1016,1043]" pageId="9" pageNumber="10" value="2006-06-01" valueMax="2006-06-24" valueMin="2006-06-01">
|
||
<collectingDate id="EF9C5C73047F1F46FC9DF0D1FC2CFBE5" box="[778,959,1016,1043]" pageId="9" pageNumber="10" value="2006-06-01" valueMax="2006-06-24" valueMin="2006-06-01">1–24 Jun. 2006</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="2693E68D047F1F46FC5DF0D1FBA6FBE4" box="[970,1077,1017,1043]" pageId="9" pageNumber="10">D. Curoe</collectorName>
|
||
leg.”;
|
||
<collectionCode id="ED771B9E047F1F46FB15F0D1FB40FBE4" box="[1154,1235,1017,1043]" pageId="9" pageNumber="10">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FB7DF0D1FD00FBC1" ID-GBIF-Occurrence="4164095351" collectingDate="2006-06-01" collectingDateMax="2006-06-24" collectingDateMin="2006-06-01" collectionCode="CEMT" collectorName="D. Curoe" elevation="1710" latitude="16.201666" location="Oaxaca" longLatPrecision="92" longitude="-97.13617" municipality="Oaxaca" pageId="9" pageNumber="10" specimenCount="3" specimenCount-male="3">
|
||
<specimenCount id="9D6048D2047F1F46FB7DF0D1FABCFBE4" box="[1258,1327,1017,1043]" pageId="9" pageNumber="10" type="male">3 ♂♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED771B9E047F1F46FDD5F734FD00FBC1" box="[578,659,1052,1078]" pageId="9" pageNumber="10">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FD39F734FDFBFB8B" ID-GBIF-Occurrence="4164095339" collectingDate="1993-06-17" collectingDateMax="1993-06-20" collectingDateMin="1993-06-17" collectionCode="CMNC" collectorName="L. Delgado" determinerName="H. F. HOWDEN" elevation="1600" location="Oaxaca - Puerto Escondido." pageId="9" pageNumber="10" specimenCode="CMNC 00031062" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D6048D2047F1F46FD39F734FD4DFBC1" box="[686,734,1052,1078]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
|
||
; “km 164.5 carr.
|
||
<location id="8EB9D580047F1F46FC3AF734FB7EFBC1" LSID="urn:lsid:plazi:treatment:03CF324D047E1F44FDC7F71DFCDEFB6D:8EB9D580047F1F46FC3AF734FB7EFBC1" box="[941,1261,1052,1078]" country="Mexico" name="Oaxaca - Puerto Escondido." pageId="9" pageNumber="10" stateProvince="Oaxaca">Oaxaca–Puerto Escondido.</location>
|
||
<date id="FFD8A59B047F1F46FB62F734FF68FBAE" pageId="9" pageNumber="10" value="1993-06-17" valueMax="1993-06-20" valueMin="1993-06-17">
|
||
<collectingDate id="EF9C5C73047F1F46FB62F734FF68FBAE" pageId="9" pageNumber="10" value="1993-06-17" valueMax="1993-06-20" valueMin="1993-06-17">17–20 Jun. 1993</collectingDate>
|
||
</date>
|
||
.
|
||
<elevation id="004B6468047F1F46FE90F717FE7BFBAE" box="[263,488,1087,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="9" pageNumber="10" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">
|
||
<quantity id="4C9E2EBE047F1F46FE90F717FE39FBAE" box="[263,426,1087,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="9" pageNumber="10" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">1550–1650 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
coprotrampa [coprotrap].
|
||
<collectorName id="2693E68D047F1F46FC88F717FC30FBAE" box="[799,931,1087,1113]" pageId="9" pageNumber="10">L. Delgado</collectorName>
|
||
leg. /
|
||
<taxonomicName id="4C66F8D8047F1F46FC79F717FAB2FBAE" authority="(Jek.)" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1006,1313,1087,1113]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<originalDetermination id="A8783BA7047F1F46FC79F717FAB2FBAE" box="[1006,1313,1087,1113]" pageId="9" pageNumber="10">Ceratotrupes sturmi (Jek.)</originalDetermination>
|
||
</taxonomicName>
|
||
/ DET.
|
||
<determinerName id="7E056F9F047F1F46FF2AF74BFE13FB8B" box="[189,384,1122,1149]" pageId="9" pageNumber="10">H.F. HOWDEN</determinerName>
|
||
”;
|
||
<specimenCode id="DBC02B20047F1F46FE02F74AFDFBFB8B" box="[405,616,1122,1148]" collectionCode="CMNC" pageId="9" pageNumber="10">CMNC 00031062</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047F1F46FD16F74AFAFBFB8B" ID-GBIF-Occurrence="4164095306" box="[641,1384,1122,1149]" collectingDate="1993-06-17" collectingDateMax="1993-06-20" collectingDateMin="1993-06-17" collectionCode="CMNC" collectorName="L. Delgado" determinerName="H. F. HOWDEN" elevation="1600" location="Oaxaca - Puerto Escondido." pageId="9" pageNumber="10" specimenCode="CMNC 00031067" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D6048D2047F1F46FD16F74AFD3DFB8A" box="[641,686,1122,1149]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding”;
|
||
<specimenCode id="DBC02B20047F1F46FB1AF74AFAF7FB8B" box="[1165,1380,1122,1148]" collectionCode="CMNC" pageId="9" pageNumber="10">CMNC 00031067</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047F1F44FF2AF78BFAE0FE60" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BD9835B047F1F46FF2AF78BFE6EFB49" blockId="9.[189,509,1187,1214]" box="[189,509,1187,1214]" pageId="9" pageNumber="10">
|
||
<heading id="D0913437047F1F46FF2AF78BFE6EFB49" bold="true" box="[189,509,1187,1214]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B9125F49047F1F46FF2AF78BFE6EFB49" bold="true" box="[189,509,1187,1214]" pageId="9" pageNumber="10">Redescription of the male</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF7FAFE80FAC5" blockId="9.[189,1399,1234,1330]" pageId="9" pageNumber="10">
|
||
<smallCapsWord id="8D3F1587047F1F46FF2AF7FAFE8BFB1C" baselines="1253,1254" box="[189,280,1234,1260]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="9" pageNumber="10">COLOUR</smallCapsWord>
|
||
. Body black with purple iridescence, especially on both dorsal and ventral sides; dorsally, tegument shining, pronotal disc occasionally weakly silky; antennal segments brown, antennal club yellow.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF672FC09FA29" blockId="9.[187,1398,1370,1502]" pageId="9" pageNumber="10">
|
||
<smallCapsWord id="8D3F1587047F1F46FF2AF672FF6FFA85" baselines="1389,1389" box="[189,252,1370,1396]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="9" pageNumber="10">HEAD</smallCapsWord>
|
||
. Clypeus obtusely triangular and elongated, with a medial, high and triangular horn; from posterior side, horn narrow and longitudinally sulcate, tip narrowly rounded; clypeal punctation dense, coarse and superficial; genae squared, with elevated and acuminate angle, genal punctation obsolete; frons smooth, unpunctured, with two oblique ridges along inner side of eyes.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF52DFD33F8E1" blockId="9.[188,1400,1541,1814]" pageId="9" pageNumber="10">
|
||
<smallCapsWord id="8D3F1587047F1F46FF2AF52DFE8AF9EA" baselines="1560,1560" box="[189,281,1541,1567]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="9" pageNumber="10">THORAX</smallCapsWord>
|
||
. In lateral view, pronotum weakly convex; in dorsal view, posterior edge of pronotum finely margined at middle, lateral edges weakly curved from posterior angles to midline, straight and convergent from midline to anterior angles; anterior angles obtusely squared; pronotal disc unpunctured, sides with strong and coarse punctures sparsely distributed from posterior to anterior angles; in lateral view, anteromedial projection curved upward apically; in dorsal view, projection with sides convergent apically, tip narrowly emarginated at middle. Elytral interstriae slightly convex, unpunctured, apex of elytra with distinct but shallow punctation. Ventral side of thoracic sclerites (i.e., pro-, meso- and metasternum) with long, yellowish setae.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047F1F46FF2AF415FC60F836" blockId="9.[189,1398,1853,1985]" pageId="9" pageNumber="10">
|
||
<smallCapsWord id="8D3F1587047F1F46FF2AF415FF65F8A2" baselines="1872,1872" box="[189,246,1853,1879]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="9" pageNumber="10">LEGS</smallCapsWord>
|
||
. Protibiae robust, with six external teeth, ventral side with a longitudinal, crenulated carina; apical spur long and slender; meso- and metatibiae respectively with 3 and 4 entire, transversal carinae on posterior side (among the examined material, only
|
||
<specimenCount id="9D6048D2047F1F46FCBAF4ACFC42F86A" box="[813,977,1923,1950]" pageId="9" pageNumber="10" type="generic">one specimen</specimenCount>
|
||
shows four entire carinae on both meso- and metatibiae); femora unmodified, with long yellowish setae.
|
||
</paragraph>
|
||
<caption id="DF19D3D3047C1F45FF2AF5F0FCA9F81F" ID-DOI="http://doi.org/10.5281/zenodo.8177548" ID-Zenodo-Dep="8177548" httpUri="https://zenodo.org/record/8177548/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1752,1778]" targetBox="[303,1294,293,1672]" targetPageId="10">
|
||
<paragraph id="8BD9835B047C1F45FF2AF5F0FCA9F81F" blockId="10.[188,1399,1752,2025]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9125F49047C1F45FF2AF5F0FE9EF905" bold="true" box="[189,269,1752,1778]" pageId="10" pageNumber="11">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C66F8D8047C1F45FE81F5F0FC88F905" authority="(Jekel, 1865) (CMNC)" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[278,795,1752,1779]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047C1F45FE81F5F0FD97F905" box="[278,516,1752,1778]" italics="true" pageId="10" pageNumber="11">Ceratotrupes sturmi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF7FEAA047C1F45FD80F5F0FD32F904" author="Jekel H." box="[535,673,1752,1779]" pageId="10" pageNumber="11" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865</bibRefCitation>
|
||
) (CMNC)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9125F49047C1F45FCBCF5F0FCF0F905" bold="true" box="[811,867,1752,1778]" pageId="10" pageNumber="11">A–B</emphasis>
|
||
. Dorsal habitus of male (A) and female (B).
|
||
<emphasis id="B9125F49047C1F45FF28F5D3FF40F8E2" bold="true" box="[191,211,1787,1813]" pageId="10" pageNumber="11">C</emphasis>
|
||
. Details of head and prothorax; arrows indicate posteromedial edge of pronotum margined.
|
||
<emphasis id="B9125F49047C1F45FB68F5D3FA87F8E2" bold="true" box="[1279,1300,1787,1813]" pageId="10" pageNumber="11">D</emphasis>
|
||
. Details of cephalic horn and pronotal projection of male.
|
||
<emphasis id="B9125F49047C1F45FC84F436FCB6F8CF" bold="true" box="[787,805,1822,1848]" pageId="10" pageNumber="11">E</emphasis>
|
||
. Aedeagus in dorsal view.
|
||
<emphasis id="B9125F49047C1F45FBFCF436FBE8F8CF" bold="true" box="[1131,1147,1822,1848]" pageId="10" pageNumber="11">F</emphasis>
|
||
. Parameres in dorsal view; arrows indicate main differences between
|
||
<taxonomicName id="4C66F8D8047C1F45FD6BF46AFCF8F8AC" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[764,875,1858,1884]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047C1F45FD6BF46AFCF8F8AC" box="[764,875,1858,1884]" italics="true" pageId="10" pageNumber="11">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C66F8D8047C1F45FC32F46AFBBEF8AC" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[933,1069,1857,1884]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047C1F45FC32F46AFBBEF8AC" box="[933,1069,1857,1884]" italics="true" pageId="10" pageNumber="11">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047C1F45FBA0F46AFB01F8AB" box="[1079,1170,1858,1884]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B9125F49047C1F45FB0EF469FB23F8AC" bold="true" box="[1177,1200,1857,1883]" pageId="10" pageNumber="11">G</emphasis>
|
||
. Right paramere in lateral view; arrow indicates lobe beneath apical spur.
|
||
<emphasis id="B9125F49047C1F45FCF2F44DFCE8F888" bold="true" box="[869,891,1893,1919]" pageId="10" pageNumber="11">H</emphasis>
|
||
. Tergite.
|
||
<emphasis id="B9125F49047C1F45FC7AF44DFC6BF888" bold="true" box="[1005,1016,1893,1919]" pageId="10" pageNumber="11">I</emphasis>
|
||
. Pleurotergites.
|
||
<emphasis id="B9125F49047C1F45FB2CF44DFB5BF888" bold="true" box="[1211,1224,1893,1919]" pageId="10" pageNumber="11">J</emphasis>
|
||
. Phallobase in ventral view; arrow indicates curved margin of ventral lobe.
|
||
<emphasis id="B9125F49047C1F45FCECF4A0FC01F855" bold="true" box="[891,914,1928,1954]" pageId="10" pageNumber="11">K</emphasis>
|
||
. Phallobase in dorsal view.
|
||
<emphasis id="B9125F49047C1F45FB42F4A0FB7AF855" bold="true" box="[1237,1257,1928,1954]" pageId="10" pageNumber="11">L</emphasis>
|
||
. Phallobase in lateral view; arrow indicates long bifurcation.
|
||
<emphasis id="B9125F49047C1F45FC9FF483FCB1F832" bold="true" box="[776,802,1963,1989]" pageId="10" pageNumber="11">M</emphasis>
|
||
. Internal sac (is) and median struts (ms) in dorsal view.
|
||
<emphasis id="B9125F49047C1F45FE94F4E6FE8BF81F" bold="true" box="[259,280,1998,2024]" pageId="10" pageNumber="11">N</emphasis>
|
||
. Internal sac and median struts in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD9835B047D1F44FF2AF220FE23FED5" blockId="11.[189,432,264,290]" box="[189,432,264,290]" pageId="11" pageNumber="12">
|
||
<heading id="D0913437047D1F44FF2AF220FE23FED5" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B9125F49047D1F44FF2AF220FE23FED5" bold="true" box="[189,432,264,290]" pageId="11" pageNumber="12">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF21FFAE0FE60" blockId="11.[189,1399,311,408]" pageId="11" pageNumber="12">Female specimens have the clypeal horn distinctly shorter and clearly depressed posteriorly; from above, the clypeal horn is triangular-shaped, apex almost reaching clypeal edge; anteromedial pronotal protuberance callus-like and transverse, depressed on superior side and close to anterior pronotal edge.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047D1F44FF2AF297FCDEFB6D" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8BD9835B047D1F44FF2AF297FEBCFE2E" blockId="11.[189,303,447,473]" box="[189,303,447,473]" pageId="11" pageNumber="12">
|
||
<heading id="D0913437047D1F44FF2AF297FEBCFE2E" bold="true" box="[189,303,447,473]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B9125F49047D1F44FF2AF297FEBCFE2E" bold="true" box="[189,303,447,473]" pageId="11" pageNumber="12">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF2C5FCB1FD4F" blockId="11.[186,1400,493,696]" pageId="11" pageNumber="12">
|
||
This species was originally described using the specific epithet “
|
||
<taxonomicName id="4C66F8D8047D1F44FC3AF2C6FB93FDF0" authorityName="Jekel" authorityYear="1865" box="[941,1024,494,519]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmii">
|
||
<emphasis id="B9125F49047D1F44FC3AF2C6FB93FDF0" box="[941,1024,494,519]" italics="true" pageId="11" pageNumber="12">sturmii</emphasis>
|
||
</taxonomicName>
|
||
”, which is probably the derived genitive form of the Latinization “Sturmius” (
|
||
<bibRefCitation id="EFF7FEAA047D1F44FD51F139FCDBFDDC" author="Jekel H." box="[710,840,528,555]" pageId="11" pageNumber="12" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel 1865</bibRefCitation>
|
||
). Nonetheless, later authors, from
|
||
<bibRefCitation id="EFF7FEAA047D1F44FB59F139FEF0FDB9" author="Gemminger M. & Harold E. von" pageId="11" pageNumber="12" refId="ref9661" refString="Gemminger M. & Harold E. von 1869. Catalogous Coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus IV. Scarabaeidae. E. H. Gummi, Monachii [Munich]. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Gemminger & Harold (1869)</bibRefCitation>
|
||
to today’s publications, indicated the same species as “
|
||
<taxonomicName id="4C66F8D8047D1F44FB96F11DFBDFFDB9" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1025,1100,565,590]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FB96F11DFBDFFDB9" box="[1025,1100,565,590]" italics="true" pageId="11" pageNumber="12">sturmi</emphasis>
|
||
</taxonomicName>
|
||
”. According to Art. 31.1 of the
|
||
<bibRefCitation id="EFF7FEAA047D1F44FE9DF17FFE3BFD86" author="ICZN & International Commission on Zoological Nomenclature" box="[266,424,599,625]" pageId="11" pageNumber="12" refId="ref10150" refString="ICZN (International Commission on Zoological Nomenclature) 1999. International Code of Zoological Nomenclature. Fourth edition. International Trust For Zoological Nomenclature, London. https: // doi. org / 10.5962 / bhl. title. 50608" type="book" year="1999">ICZN (1999)</bibRefCitation>
|
||
, the original spelling of the species-group name must be preserved. However, Art. 33.3.1 suggests that an “incorrect subsequent spelling” must be maintained if in prevailing use in the literature, which is the case for
|
||
<taxonomicName id="4C66F8D8047D1F44FDB9F1B6FC88FD40" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[558,795,670,696]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FDB9F1B6FC88FD40" box="[558,795,670,696]" italics="true" pageId="11" pageNumber="12">Ceratotrupes sturmi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF1F7FEC6FC5D" blockId="11.[187,1400,735,938]" pageId="11" pageNumber="12">
|
||
<bibRefCitation id="EFF7FEAA047D1F44FF2AF1F7FEC0FD0D" author="Jekel H." box="[189,339,735,762]" pageId="11" pageNumber="12" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel (1865)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C66F8D8047D1F44FE4DF1F7FDD8FD0E" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[474,587,735,761]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FE4DF1F7FDD8FD0E" box="[474,587,735,761]" italics="true" pageId="11" pageNumber="12">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C66F8D8047D1F44FD1CF1F7FCB5FD0E" authorityName="Jekel" authorityYear="1865" box="[651,806,735,761]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F49047D1F44FD1CF1F7FCB5FD0E" box="[651,806,735,761]" italics="true" pageId="11" pageNumber="12">C. mniszechi</emphasis>
|
||
</taxonomicName>
|
||
on
|
||
<specimenCount id="9D6048D2047D1F44FCCEF1F7FB99FD0E" box="[857,1034,735,761]" pageId="11" pageNumber="12" type="generic">two specimens</specimenCount>
|
||
from the Sallé and Mniszech collections and today deposited in the NHML and MNHN, respectively. In addition, the author mentioned the generic “
|
||
<collectingCountry id="F371C3CB047D1F44FEC6F00DFE3CFCC8" box="[337,431,805,831]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
|
||
” as type locality of the two species. The
|
||
<typeStatus id="54DD3DF9047D1F44FC09F00DFB96FCC8" box="[926,1029,805,831]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C66F8D8047D1F44FBA4F00EFB58FCC8" authorityName="Jekel" authorityYear="1865" box="[1075,1227,805,832]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F49047D1F44FBA4F00EFB58FCC8" box="[1075,1227,805,832]" italics="true" pageId="11" pageNumber="12">C. mniszechi</emphasis>
|
||
</taxonomicName>
|
||
does not have any label saying “
|
||
<collectingCountry id="F371C3CB047D1F44FE18F060FE62FC95" box="[399,497,840,866]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
|
||
”. However, Jekel’s handwritten label (“Collection Mniszech”) and the match between the morphology of the specimen and its original description remove any doubt to its identity (
|
||
<figureCitation id="135D9FDE047D1F44FF50F0A7FEDAFC5D" box="[199,329,911,938]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[209,1382,582,1879]" captionTargetId="figure-82@14.[189,1398,565,1888]" captionTargetPageId="14" captionText="Fig. 5. Type specimens and original labels. A–B. Ceratotrupes sturmi (Jekel, 1865), holotype, ♂ (NHML). A. Dorsal habitus. B. Labels. C–D. Ceratotrupes mniszechi (Jekel, 1865), holotype, ♂ (MNHN). C. Dorsal habitus. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.8177552" httpUri="https://zenodo.org/record/8177552/files/figure.png" pageId="11" pageNumber="12">Fig. 5C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF0F8FCDEFB6D" blockId="11.[187,1400,976,1179]" pageId="11" pageNumber="12">
|
||
At the same time, the
|
||
<typeStatus id="54DD3DF9047D1F44FE52F0F8FDBFFC1D" box="[453,556,976,1002]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C66F8D8047D1F44FDC0F0F9FD55FC1D" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[599,710,977,1003]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FDC0F0F9FD55FC1D" box="[599,710,977,1003]" italics="true" pageId="11" pageNumber="12">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
has a label indicating “
|
||
<collectingCountry id="F371C3CB047D1F44FC4AF0F8FBAEFC1D" box="[989,1085,976,1002]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
|
||
. Salle Coll.” and a second saying “Juquila”, a locality near
|
||
<collectingRegion id="49A24DB9047D1F44FDACF0DCFD00FBF9" box="[571,659,1012,1038]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="12">Oaxaca</collectingRegion>
|
||
(
|
||
<figureCitation id="135D9FDE047D1F44FD35F0DBFCB0FBF9" box="[674,803,1011,1038]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[209,1382,582,1879]" captionTargetId="figure-82@14.[189,1398,565,1888]" captionTargetPageId="14" captionText="Fig. 5. Type specimens and original labels. A–B. Ceratotrupes sturmi (Jekel, 1865), holotype, ♂ (NHML). A. Dorsal habitus. B. Labels. C–D. Ceratotrupes mniszechi (Jekel, 1865), holotype, ♂ (MNHN). C. Dorsal habitus. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.8177552" httpUri="https://zenodo.org/record/8177552/files/figure.png" pageId="11" pageNumber="12">Fig. 5A–B</figureCitation>
|
||
). Probably the latter was added subsequently to the description of the species. At the moment, there is no evidence that indicates who might have attached this label to the
|
||
<typeStatus id="54DD3DF9047D1F44FEE6F712FE4FFBA3" box="[369,476,1082,1108]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
. Nonetheless, the morphology of the
|
||
<typeStatus id="54DD3DF9047D1F44FC12F712FC7FFBA3" box="[901,1004,1082,1108]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
matches perfectly with specimens recently collected in the mountains around
|
||
<collectingRegion id="49A24DB9047D1F44FD57F775FC8BFB80" box="[704,792,1117,1143]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="12">Oaxaca</collectingRegion>
|
||
(cephalic horn slender; pronotal projection clearly narrower distally; pronotal posterior edge with margin).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |