219 lines
40 KiB
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219 lines
40 KiB
XML
<document id="7349466903299D7E8B0F20A799B0219A" ID-CLB-Dataset="31373" ID-DOI="10.11646/zootaxa.4350.3.5" ID-GBIF-Dataset="adb428f5-289d-43f0-91d6-a5da8981da8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1059327" ID-ZooBank="7B3B84E2-4521-4591-8F53-71F56018A403" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511168099826" checkinUser="plazi" docAuthor="Qiu, Jian-Yue, Xu, Hao & Chen, Li" docDate="2017" docId="03CD6B0AFFD0FC4E2E81F934F7A4F842" docLanguage="en" docName="zootaxa.4350.3.5.pdf" docOrigin="Zootaxa 4350 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coilodera penicillata Hope 1831" docType="key" docVersion="8" lastPageNumber="515" masterDocId="FFF41372FFD4FC4A2E16FC2CF23FFFE3" masterDocTitle="Taxonomic review of the Oriental flower beetle Coilodera penicillata species complex (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="537" masterPageNumber="511" pageNumber="515" updateTime="1698501641652" updateUser="ExternalLinkService">
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<mods:title id="0FED480EF0C7EAD716BF50A5DA6572DA">Taxonomic review of the Oriental flower beetle Coilodera penicillata species complex (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<mods:namePart id="F53DB372BA1105F982C9F23668B0D63C">Qiu, Jian-Yue</mods:namePart>
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<mods:namePart id="B63C78857490AE8D3F17BAB13897F901">Xu, Hao</mods:namePart>
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<mods:namePart id="C469091CC32848652A676780FF2151B5">Chen, Li</mods:namePart>
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<mods:title id="6BE58BEA9C748EA11DB828A349229C73">Zootaxa</mods:title>
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<treatment id="03CD6B0AFFD0FC4E2E81F934F7A4F842" ID-DOI="http://doi.org/10.5281/zenodo.6040971" ID-GBIF-Taxon="137453749" ID-Zenodo-Dep="6040971" LSID="urn:lsid:plazi:treatment:03CD6B0AFFD0FC4E2E81F934F7A4F842" httpUri="http://treatment.plazi.org/id/03CD6B0AFFD0FC4E2E81F934F7A4F842" lastPageNumber="515" pageId="4" pageNumber="515">
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<emphasis id="B910060EFFD0FC4E2E81F934F15AFAD1" bold="true" box="[151,869,1303,1330]" pageId="4" pageNumber="515">
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Key to species of the
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<taxonomicName id="4C64A19FFFD0FC4E2F8AF93BF0A7FAD2" authorityName="Hope" authorityYear="1831" box="[412,664,1303,1329]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="penicillata">
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<emphasis id="B910060EFFD0FC4E2F8AF93BF0A7FAD2" bold="true" box="[412,664,1303,1329]" italics="true" pageId="4" pageNumber="515">Coilodera penicillata</emphasis>
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</taxonomicName>
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species complex
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</emphasis>
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</paragraph>
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<subSubSection id="C37E8997FFD0FC4E2E81F971F7A4F842" pageId="4" pageNumber="515" type="key">
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<paragraph id="8BDBDA1CFFD0FC4E2E81F971F7A3FA90" blockId="4.[151,1436,1373,1953]" box="[151,1436,1373,1395]" pageId="4" pageNumber="515">
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1 Mesometasternal suture not engraved (
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<figureCitation id="135FC699FFD0FC4E2C42F971F0D4FA90" box="[596,747,1373,1395]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 63–64, 66</figureCitation>
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).......................................................2
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</keyLead>
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</paragraph>
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<paragraph id="8BDBDA1CFFD0FC4E2E81F950F7A4FA72" blockId="4.[151,1436,1373,1953]" box="[151,1435,1403,1425]" pageId="4" pageNumber="515">
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<keyLead id="30957929FFD0FC4E2E81F950F7A4FA72" box="[151,1435,1403,1425]" pageId="4" pageNumber="515">
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- Mesometasternal suture engraved (
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<figureCitation id="135FC699FFD0FC4E2C26F957F0F6FA72" box="[560,713,1403,1425]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 59–62, 65</figureCitation>
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)...........................................................3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="3090C2B9FFD0FC4E2E81F9B4F7A3F9E5" pageId="4" pageNumber="515">
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<paragraph id="8BDBDA1CFFD0FC4E2E81F9B4F7A3FA28" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
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<keyLead id="30957929FFD0FC4E2E81F9B4F7A3FA28" pageId="4" pageNumber="515">
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2 Scutellum partly clad with tomentum (rarely glabrous) (
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<figureCitation id="135FC699FFD0FC4E2CEFF9B4F150FA4D" box="[761,879,1432,1454]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 52–53</figureCitation>
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); basal piece of male genitalia distinctly convex (
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<figureCitation id="135FC699FFD0FC4E2B5EF9B4F7B1FA4D" box="[1352,1422,1432,1454]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="4" pageNumber="515">Fig. 80</figureCitation>
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)................................................................................
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<taxonomicName id="4C64A19FFFD0FC4E2AB5F999F7A3FA28" authority="Hope, 1831" authorityName="Hope" authorityYear="1831" box="[1187,1436,1461,1483]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="penicillata">
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<emphasis id="B910060EFFD0FC4E2AB5F999F719FA28" box="[1187,1318,1461,1483]" italics="true" pageId="4" pageNumber="515">C. penicillata</emphasis>
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<bibRefCitation id="EFF5A7EDFFD0FC4E2B3AF99AF7A3FA28" author="Hope" box="[1324,1436,1461,1483]" pageId="4" pageNumber="515" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope, 1831</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BDBDA1CFFD0FC4E2E81F9F8F7A3F9E5" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
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<keyLead id="30957929FFD0FC4E2E81F9F8F7A3F9E5" pageId="4" pageNumber="515">
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- Scutellum glabrous (
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<figureCitation id="135FC699FFD0FC4E2FBEF9FFF3D0FA0A" box="[424,495,1491,1513]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Fig. 58</figureCitation>
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); basal piece of male genitalia large and flat in lateral view (
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<figureCitation id="135FC699FFD0FC4E2A24F9FFF647FA0A" box="[1074,1144,1491,1513]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="4" pageNumber="515">Fig. 84</figureCitation>
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)..................................................................................................
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<taxonomicName id="4C64A19FFFD0FC4E2A7CF9DCF7A3F9E5" authority="Moser, 1902" authorityName="Moser" authorityYear="1902" box="[1130,1436,1520,1542]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="nigroscutellaris">
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<emphasis id="B910060EFFD0FC4E2A7CF9DCF722F9E5" box="[1130,1309,1520,1542]" italics="true" pageId="4" pageNumber="515">C. nigroscutellaris</emphasis>
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<bibRefCitation id="EFF5A7EDFFD0FC4E2B32F9DDF7A3F9E5" author="Moser" box="[1316,1436,1520,1542]" pageId="4" pageNumber="515" refString="Moser, J. (1902) Neue Cetoniden-Arten aus Tonkin, gesammelt von H. Fruhstorfer. Berliner entomologische Zeitschrift, 46, 525 - 538." type="journal article" year="1902">Moser, 1902</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<keyStep id="3090C2B9FFD0FC4E2E81FA21F7A3F955" pageId="4" pageNumber="515">
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<paragraph id="8BDBDA1CFFD0FC4E2E81FA21F7A8F9BD" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
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<keyLead id="30957929FFD0FC4E2E81FA21F7A8F9BD" pageId="4" pageNumber="515">
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3 Scutellum glabrous (
|
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<figureCitation id="135FC699FFD0FC4E2FBAFA21F3CAF9C0" box="[428,501,1549,1571]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Fig. 57</figureCitation>
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); posterior margin of median tomentose area of pronotum distinctly notched (
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<figureCitation id="135FC699FFD0FC4E2AE5FA21F756F9C0" box="[1267,1385,1549,1571]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 & 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 & 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="4" pageNumber="515">Figs. 25–26</figureCitation>
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,
|
||
<figureCitation id="135FC699FFD0FC4E2B63FA21F7AFF9C0" box="[1397,1424,1549,1571]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1679,1702]" captionTargetBox="[155,1440,274,1648]" captionTargetId="figure@13.[155,1440,267,1648]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Types and labels of Coilodera. 34 – 36, C. dives, female lectotype (photograph by Giulio Cuccodoro). 37 – 41, C. miksici: 37 – 39, male holotype; 40 – 41, female paratype. 42 – 44, C. mearesii, female syntype (provided by Amoret Spooner). 45 – 46, C. formosana, female syntype (photograph by Joachim Willers). 47 – 48, C. nigroscutellaris, male syntype (photograph by Joachim Willers)." httpUri="https://zenodo.org/record/1059345/files/figure.png" pageId="4" pageNumber="515">45</figureCitation>
|
||
); outer teeth of male metatibia longer than ventral teeth (
|
||
<figureCitation id="135FC699FFD0FC4E2CE5FA07F104F9A2" box="[755,827,1579,1601]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Fig. 71</figureCitation>
|
||
); the length of male antennal club about 1.5 times as the length of antennomeres 2–7 combined......................................................
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A8CFA64F7A8F9BD" authority="Moser, 1910" authorityName="Moser" authorityYear="1910" box="[1178,1431,1608,1630]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="formosana">
|
||
<emphasis id="B910060EFFD0FC4E2A8CFA64F724F9BD" box="[1178,1307,1608,1630]" italics="true" pageId="4" pageNumber="515">C. formosana</emphasis>
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2B36FA65F7A8F9BD" author="Moser" box="[1312,1431,1608,1630]" pageId="4" pageNumber="515" refString="Moser, I. (1910) Beitrag zur Kenntnis der Cetoniden VII. Deutsche Entomologische Zeitschrift, 1910, 293 - 301." type="journal article" year="1910">Moser, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD0FC4E2E81FA4AF7A3F955" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
|
||
<keyLead id="30957929FFD0FC4E2E81FA4AF7A3F955" pageId="4" pageNumber="515">
|
||
- Scutellum covered with tomentum (
|
||
<figureCitation id="135FC699FFD0FC4E2C28FA49F08AF998" box="[574,693,1637,1659]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 49–51</figureCitation>
|
||
); posterior margin of median tomentose area of pronotum curved or slightly notched (
|
||
<figureCitation id="135FC699FFD0FC4E2F2CFAAFF3C9F97A" box="[314,502,1667,1689]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Figs. 4, 6, 8, 10, 13</figureCitation>
|
||
,
|
||
<figureCitation id="135FC699FFD0FC4E2C16FAAFF001F97A" box="[512,574,1667,1689]" captionStart="FIGURES 15 – 20" captionStartId="8.[151,264,1844,1867]" captionTargetBox="[158,1409,224,1802]" captionTargetId="figure@8.[151,1436,195,1823]" captionTargetPageId="8" captionText="FIGURES 15 – 20. Habitus of Coilodera grandimaculata: 15 – 17, male holotype; 18 – 20, female paratype." httpUri="https://zenodo.org/record/1059339/files/figure.png" pageId="4" pageNumber="515">15, 18</figureCitation>
|
||
); outer teeth of male metatibia shorter than ventral teeth (
|
||
<figureCitation id="135FC699FFD0FC4E2A71FAAFF6E2F97A" box="[1127,1245,1667,1689]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 67–69</figureCitation>
|
||
); the length of male antennal club about twice as the length of antennomeres 2–7 combined...........................................4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="3090C2B9FFD0FC4E2E81FA91F7A4F8C8" pageId="4" pageNumber="515">
|
||
<paragraph id="8BDBDA1CFFD0FC4E2E81FA91F7A3F912" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
|
||
<keyLead id="30957929FFD0FC4E2E81FA91F7A3F912" pageId="4" pageNumber="515">
|
||
4 Body surface with green metallic sheen and tomentum lemon-yellow (
|
||
<figureCitation id="135FC699FFD0FC4E2D9FFA91F63FF930" box="[905,1024,1725,1747]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Figs. 10–13</figureCitation>
|
||
); abdomen with long setae around midline in some individuals (
|
||
<figureCitation id="135FC699FFD0FC4E2FBCFAF7F3C6F912" box="[426,505,1755,1777]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="135FC699FFD0FC4E2C15FAF7F023F912" box="[515,540,1755,1777]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1679,1702]" captionTargetBox="[155,1440,274,1648]" captionTargetId="figure@13.[155,1440,267,1648]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Types and labels of Coilodera. 34 – 36, C. dives, female lectotype (photograph by Giulio Cuccodoro). 37 – 41, C. miksici: 37 – 39, male holotype; 40 – 41, female paratype. 42 – 44, C. mearesii, female syntype (provided by Amoret Spooner). 45 – 46, C. formosana, female syntype (photograph by Joachim Willers). 47 – 48, C. nigroscutellaris, male syntype (photograph by Joachim Willers)." httpUri="https://zenodo.org/record/1059345/files/figure.png" pageId="4" pageNumber="515">43</figureCitation>
|
||
).................................................
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A60FAF7F7A3F912" authority="(Westwood, 1842)" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[1142,1436,1755,1777]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="mearesii">
|
||
<emphasis id="B910060EFFD0FC4E2A60FAF7F6DBF912" box="[1142,1252,1755,1777]" italics="true" pageId="4" pageNumber="515">C. mearesii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2AE4FAF7F7AAF912" author="Westwood" box="[1266,1429,1755,1777]" pageId="4" pageNumber="515" refString="Westwood, J. O. (1842) Descriptions of some new species of Cetoniidae from Australia, Asia, and the Asiatic Islands. Arcana Entomologica; or Illustrations of new, rare, and interesting Insects, 1, 103 - 104." type="journal article" year="1842">Westwood, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD0FC4E2E81FAD5F7A4F8C8" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
|
||
<keyLead id="30957929FFD0FC4E2E81FAD5F7A4F8C8" pageId="4" pageNumber="515">
|
||
- Body surface with wine-red and/or green metallic sheen and tomentum orange or orange-red (
|
||
<figureCitation id="135FC699FFD0FC4E2A67FAD4F6F0F8ED" box="[1137,1231,1784,1806]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Figs. 4–9</figureCitation>
|
||
,
|
||
<figureCitation id="135FC699FFD0FC4E2ACFFAD4F727F8ED" box="[1241,1304,1784,1806]" captionStart="FIGURES 15 – 20" captionStartId="8.[151,264,1844,1867]" captionTargetBox="[158,1409,224,1802]" captionTargetId="figure@8.[151,1436,195,1823]" captionTargetPageId="8" captionText="FIGURES 15 – 20. Habitus of Coilodera grandimaculata: 15 – 17, male holotype; 18 – 20, female paratype." httpUri="https://zenodo.org/record/1059339/files/figure.png" pageId="4" pageNumber="515">15–20</figureCitation>
|
||
); middle portion of abdomen without setae, or with a few short setae (
|
||
<figureCitation id="135FC699FFD0FC4E2D11FB39F144F8C8" box="[775,891,1813,1835]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Figs. 5, 7, 9</figureCitation>
|
||
,
|
||
<figureCitation id="135FC699FFD0FC4E2D93FB39F19EF8C8" box="[901,929,1813,1835]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1679,1702]" captionTargetBox="[155,1440,274,1648]" captionTargetId="figure@13.[155,1440,267,1648]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Types and labels of Coilodera. 34 – 36, C. dives, female lectotype (photograph by Giulio Cuccodoro). 37 – 41, C. miksici: 37 – 39, male holotype; 40 – 41, female paratype. 42 – 44, C. mearesii, female syntype (provided by Amoret Spooner). 45 – 46, C. formosana, female syntype (photograph by Joachim Willers). 47 – 48, C. nigroscutellaris, male syntype (photograph by Joachim Willers)." httpUri="https://zenodo.org/record/1059345/files/figure.png" pageId="4" pageNumber="515">38</figureCitation>
|
||
).........................................5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="3090C2B9FFD0FC4E2E81FB1FF7A4F842" pageId="4" pageNumber="515">
|
||
<paragraph id="8BDBDA1CFFD0FC4E2E81FB1FF7A3F885" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
|
||
<keyLead id="30957929FFD0FC4E2E81FB1FF7A3F885" pageId="4" pageNumber="515">
|
||
5 Base of mesometasternal process distinctly constricted (
|
||
<figureCitation id="135FC699FFD0FC4E2D0EFB1FF15BF8AA" box="[792,868,1843,1865]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Fig. 59</figureCitation>
|
||
); tomentum orange-red; posthumeral maculae in female smaller in size (
|
||
<figureCitation id="135FC699FFD0FC4E2F6EFB7CF38CF885" box="[376,435,1872,1894]" captionStart="FIGURES 4 – 14" captionStartId="6.[151,264,1076,1099]" captionTargetBox="[157,1436,198,1050]" captionTargetId="figure@6.[151,1436,193,1056]" captionTargetPageId="6" captionText="FIGURES 4 – 14. Habitus of Coilodera. 4 – 9, C. dives: 4 – 5, northern Kachin, Myanmar (male); 6 – 7, Mêdog Xizang (female); 8 – 9, Mêdog, Xizang (male). 8, Mêdog, Xizang (male). 10 – 14, C. mearesii (male): 10 – 11, Tiger Hill, northern India; 12, Kurseong, northern India; 13 – 14, female paratype of C. miksici." httpUri="https://zenodo.org/record/1059335/files/figure.png" pageId="4" pageNumber="515">Fig. 6</figureCitation>
|
||
,
|
||
<figureCitation id="135FC699FFD0FC4E2FABFB7CF3C3F885" box="[445,508,1872,1894]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1679,1702]" captionTargetBox="[155,1440,274,1648]" captionTargetId="figure@13.[155,1440,267,1648]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Types and labels of Coilodera. 34 – 36, C. dives, female lectotype (photograph by Giulio Cuccodoro). 37 – 41, C. miksici: 37 – 39, male holotype; 40 – 41, female paratype. 42 – 44, C. mearesii, female syntype (provided by Amoret Spooner). 45 – 46, C. formosana, female syntype (photograph by Joachim Willers). 47 – 48, C. nigroscutellaris, male syntype (photograph by Joachim Willers)." httpUri="https://zenodo.org/record/1059345/files/figure.png" pageId="4" pageNumber="515">34, 40</figureCitation>
|
||
); parameres short (
|
||
<figureCitation id="135FC699FFD0FC4E2CA5FB7CF118F885" box="[691,807,1872,1894]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="4" pageNumber="515">Figs. 73–74</figureCitation>
|
||
)........................
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A50FB7CF7A3F885" authority="(Gory & Percheron, 1833)" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[1094,1436,1872,1894]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="dives">
|
||
<emphasis id="B910060EFFD0FC4E2A50FB7CF6A7F885" box="[1094,1176,1872,1894]" italics="true" pageId="4" pageNumber="515">C. dives</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2AB3FB7CF7AAF885" author="Gory" box="[1189,1429,1872,1894]" pageId="4" pageNumber="515" refString="Gory, M. H. & Percheron, M. A. (1833) Monographie des Cetoines et genres voisins, formant dans les familles naturelles de Latreille, le division des Scarabees Melitophiles. J. B. Baillieres, Paris, France, 410 pp." type="book" year="1833">Gory & Percheron, 1833</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD0FC4E2E81FB42F7A4F842" blockId="4.[151,1436,1373,1953]" pageId="4" pageNumber="515">
|
||
<keyLead id="30957929FFD0FC4E2E81FB42F7A4F842" pageId="4" pageNumber="515">
|
||
- Base of mesometasternal process distinctly slightly constricted (
|
||
<figureCitation id="135FC699FFD0FC4E2D75FB41F1E2F860" box="[867,989,1901,1923]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="4" pageNumber="515">Figs. 60–61</figureCitation>
|
||
); tomentum orange; posthumeral maculae in female larger in size (
|
||
<figureCitation id="135FC699FFD0FC4E2FA6FBA7F3C8F842" box="[432,503,1931,1953]" captionStart="FIGURES 15 – 20" captionStartId="8.[151,264,1844,1867]" captionTargetBox="[158,1409,224,1802]" captionTargetId="figure@8.[151,1436,195,1823]" captionTargetPageId="8" captionText="FIGURES 15 – 20. Habitus of Coilodera grandimaculata: 15 – 17, male holotype; 18 – 20, female paratype." httpUri="https://zenodo.org/record/1059339/files/figure.png" pageId="4" pageNumber="515">Fig. 18</figureCitation>
|
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,
|
||
<figureCitation id="135FC699FFD0FC4E2C17FBA7F023F842" box="[513,540,1931,1953]" captionStart="FIGURES 91 – 94" captionStartId="18.[151,264,1972,1995]" captionTargetBox="[167,1395,293,1927]" captionTargetId="figure@18.[167,1395,193,1927]" captionTargetPageId="18" captionText="FIGURES 91 – 94. Habitat of Coilodera. 91 – 93, C. dives: 91, habitat (Tongbiguan Nature Reserve, Yingjiang, Yunnan); 92, closer view of habitat; 93, larva in rotten wood. 94, Host plant (Zanthoxylum sp.) of C. grandimaculata in Mêdog, Xizang." httpUri="https://zenodo.org/record/1059359/files/figure.png" pageId="4" pageNumber="515">94</figureCitation>
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); parameres elongate (
|
||
<figureCitation id="135FC699FFD0FC4E2CE2FBA7F157F842" box="[756,872,1931,1953]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="4" pageNumber="515">Figs. 75–76</figureCitation>
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)....................
|
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<taxonomicName id="4C64A19FFFD0FC4E2A7CFBA7F720F842" authority="Qiu & Xu & Chen, 2017" authorityName="Qiu & Xu & Chen" authorityYear="2017" box="[1130,1311,1931,1953]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="grandimaculata" status="sp. nov.">
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<emphasis id="B910060EFFD0FC4E2A7CFBA7F720F842" box="[1130,1311,1931,1953]" italics="true" pageId="4" pageNumber="515">C. grandimaculata</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A223BB75FFD0FC4E2B30FBA7F7A4F842" box="[1318,1435,1931,1953]" pageId="4" pageNumber="515" rank="species">
|
||
<emphasis id="B910060EFFD0FC4E2B30FBA7F7A4F842" bold="true" box="[1318,1435,1931,1953]" pageId="4" pageNumber="515">new species</emphasis>
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</taxonomicNameLabel>
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||
</keyLead>
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||
</paragraph>
|
||
</keyStep>
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||
</subSubSection>
|
||
</treatment>
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</document> |