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<document id="707A476C304586AF96FF442CD7690A7F" ID-CLB-Dataset="7153" ID-DOI="10.5852/ejt.2021.747.1333" ID-GBIF-Dataset="aca67cc8-96ca-4f24-b8af-2b532ecd355c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4836744" ID-ZooBank="3F0B6EAF-C616-4865-811A-414A094B590C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619605517524" checkinUser="plazi" docAuthor="Moctezuma, Victor &amp; Halffter, Gonzalo" docDate="2021" docId="03CA87F2FFE7FFDDFDF641E4BA453C24" docLanguage="en" docName="ejt-747_1-71.pdf" docOrigin="European Journal of Taxonomy 747" docSource="http://zoobank.org/3f0b6eaf-c616-4865-811a-414a094b590c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phanaeus malyi Arnaud 2002" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FFF3FF8AFFFEFFC1FFA8420BBF723A3A" masterDocTitle="Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="26" updateTime="1698937721231" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="17C2229315A1900DC55011CC7F3ACEBE">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="E9A98E02A7EF2C1EE6B5C17FD3989424">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="C9B7539E3A708956421F0CA55D76EA2B">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:namePart id="966F5A654FE887D1D5B68A24C8F2720D">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="4038F5FAC974AB2BF089F5AD8881445C">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:title id="D6796CCBBB243D153AD58C51DEEC8653">European Journal of Taxonomy</mods:title>
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<mods:date id="FB0012CCF241F8064B24F1E9C8B0CD89">2021</mods:date>
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<treatment id="03CA87F2FFE7FFDDFDF641E4BA453C24" ID-DOI="http://doi.org/10.5281/zenodo.4726229" ID-GBIF-Taxon="181753771" ID-Zenodo-Dep="4726229" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24" httpUri="http://treatment.plazi.org/id/03CA87F2FFE7FFDDFDF641E4BA453C24" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="C379656FFFE7FFD8FDF641E4BCA73E30" box="[606,981,1007,1034]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BDC36E4FFE7FFD8FDF641E4BCA73E30" blockId="25.[606,981,1007,1070]" box="[606,981,1007,1034]" pageId="25" pageNumber="26">
<heading id="D0948188FFE7FFD8FDF641E4BCA73E30" box="[606,981,1007,1034]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C634D67FFE7FFD8FDF641E4BCA73E30" ID-CoL="4FSJS" authority="Arnaud, 2002 a" authorityName="Arnaud" authorityYear="2002" box="[606,981,1007,1034]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE7FFD8FDF641E4BC533E30" bold="true" box="[606,801,1007,1034]" italics="true" pageId="25" pageNumber="26">Phanaeus malyi</emphasis>
<bibRefCitation id="EFF24B15FFE7FFD8FC8F41E4BCA73E30" author="Arnaud P." box="[807,981,1007,1034]" pageId="25" pageNumber="26" pagination="2 - 5" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." type="journal article" year="2002">Arnaud, 2002a</bibRefCitation>
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<subSubSection id="C379656FFFE7FFD8FDCF4618BCBE3E14" box="[615,972,1043,1070]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BDC36E4FFE7FFD8FDCF4618BCBE3E14" blockId="25.[606,981,1007,1070]" box="[615,972,1043,1070]" pageId="25" pageNumber="26">
<figureCitation id="13582A61FFE7FFD8FDCF4618BDC83E14" box="[615,698,1043,1070]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="25" pageNumber="26">Figs 1I</figureCitation>
,
<figureCitation id="13582A61FFE7FFD8FD60461FBD993E14" box="[712,747,1044,1070]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="25" pageNumber="26">2E</figureCitation>
,
<figureCitation id="13582A61FFE7FFD8FD5F461FBC7A3E14" box="[759,776,1044,1070]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="25" pageNumber="26">7</figureCitation>
,
<figureCitation id="13582A61FFE7FFD8FCBC4618BC123E17" box="[788,864,1043,1069]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="51.[189,232,1233,1259]" captionStartId-1="54.[191,234,1247,1273]" captionTargetBox-0="[193,1395,270,1197]" captionTargetBox-1="[193,1393,268,1195]" captionTargetId-0="figure-241@51.[189,1398,265,1199]" captionTargetId-1="figure-243@54.[189,1396,265,1198]" captionTargetPageId-0="51" captionTargetPageId-1="54" captionText-0="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." captionText-1="Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018)." figureDoi-0="http://doi.org/10.5281/zenodo.4836798" figureDoi-1="http://doi.org/10.5281/zenodo.4836800" httpUri-0="https://zenodo.org/record/4836798/files/figure.png" httpUri-1="https://zenodo.org/record/4836800/files/figure.png" pageId="25" pageNumber="26">16–17</figureCitation>
,
<figureCitation id="13582A61FFE7FFD8FCC4461FBCE53E14" box="[876,919,1044,1070]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="25" pageNumber="26">18J</figureCitation>
,
<figureCitation id="13582A61FFE7FFD8FC0C461FBCBE3E14" box="[932,972,1044,1070]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="25" pageNumber="26">19J</figureCitation>
</paragraph>
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<subSubSection id="C379656FFFE7FFDBFF15465DBB3D3BEE" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BDC36E4FFE7FFD8FF15465DBDD03E4A" blockId="25.[189,674,1110,1136]" box="[189,674,1110,1136]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF15465DBDD03E4A" box="[189,674,1110,1136]" pageId="25" pageNumber="26">
<treatmentCitation id="0AC210F5FFE7FFD8FF15465DBDEC3E4A" author="Arnaud P." box="[189,670,1110,1136]" page="4" pageId="25" pageNumber="26" year="2002">
<taxonomicName id="4C634D67FFE7FFD8FF15465DBDEC3E4A" ID-CoL="4FSJS" authority="Arnaud, 2002 a: 4" authorityName="Arnaud" authorityPageNumber="4" authorityYear="2002" box="[189,670,1110,1136]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="pyrois" subSpecies="malyi">
<emphasis id="B917EAF6FFE7FFD8FF15465DBEBB3E4A" box="[189,457,1110,1136]" italics="true" pageId="25" pageNumber="26">Phanaeus pyrois malyi</emphasis>
<bibRefCitation id="EFF24B15FFE7FFD8FE66465DBDEC3E4A" author="Arnaud P." box="[462,670,1110,1136]" pageId="25" pageNumber="26" pagination="2 - 5" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." type="journal article" year="2002">Arnaud, 2002a: 4</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154693BE593EED" blockId="25.[189,1399,1176,1742]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF154693BE593EED" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF154693BEF33E88" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[189,385,1176,1202]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FF154693BEF33E88" box="[189,385,1176,1202]" italics="true" pageId="25" pageNumber="26">Phanaeus pyrois</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC210F5FFE7FFD8FE094692BD263E89" author="Bates H. W." box="[417,596,1176,1203]" page="58" pageId="25" pageNumber="26" year="1887">
<bibRefCitation id="EFF24B15FFE7FFD8FE094692BD263E89" author="Bates H. W." box="[417,596,1176,1203]" pageId="25" pageNumber="26" refId="ref36135" refString="Bates H. W. 1887. Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor &amp; Francis, London." type="book" year="1887">Bates 1887: 58</bibRefCitation>
</treatmentCitation>
(in part, as a variety of
<taxonomicName id="4C634D67FFE7FFD8FCDF4692BCAE3E88" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[887,988,1177,1202]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FCDF4692BCAE3E88" box="[887,988,1177,1202]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FBB34693BA3F3E88" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[1051,1357,1176,1203]" page="78" pageId="25" pageNumber="26" year="2018">
<bibRefCitation id="EFF24B15FFE7FFD8FBB34693BA3F3E88" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[1051,1357,1176,1203]" pageId="25" pageNumber="26" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFE7FFD8FB094692BBA73E88" box="[1185,1237,1176,1202]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2018: 78
</bibRefCitation>
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(in part).
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</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF1546EBBC9D3F95" blockId="25.[189,1399,1176,1742]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF1546EBBC9D3F95" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF1546EBBEC83EC0" ID-CoL="856BJ" authorityName="Olsoufieff" authorityYear="1924" box="[189,442,1248,1274]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="B917EAF6FFE7FFD8FF1546EBBEC83EC0" box="[189,442,1248,1274]" italics="true" pageId="25" pageNumber="26">Phanaeus blanchardi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE7FFD8FD9A46EBBC693EC0" author="Olsoufieff G. d'" box="[562,795,1248,1274]" page="92" pageId="25" pageNumber="26" year="1924">
<bibRefCitation id="EFF24B15FFE7FFD8FD9A46EBBC693EC0" author="Olsoufieff G. d'" box="[562,795,1248,1274]" pageId="25" pageNumber="26" pagination="4 - 172" refId="ref41334" refString="Olsoufieff G. d'. 1924. Les Phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta, Revue illustree d'Entomologique 13: 4 - 172." type="journal article" year="1924">Olsoufieff 1924: 92</bibRefCitation>
</treatmentCitation>
(not
<treatmentCitation id="0AC210F5FFE7FFD8FCF646EABB913EC0" author="Harold E." box="[862,1251,1248,1274]" page="114" pageId="25" pageNumber="26" year="1871">
<taxonomicName id="4C634D67FFE7FFD8FCF646EABB913EC0" ID-CoL="856BJ" authority="Harold, 1871: 114" authorityName="Harold" authorityPageNumber="114" authorityYear="1871" box="[862,1251,1248,1274]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="B917EAF6FFE7FFD8FCF646EABC8F3EC0" box="[862,1021,1248,1274]" italics="true" pageId="25" pageNumber="26">P. blanchardi</emphasis>
<bibRefCitation id="EFF24B15FFE7FFD8FBAE46EBBB913EC0" author="Harold E." box="[1030,1251,1248,1274]" pageId="25" pageNumber="26" pagination="113 - 119" refId="ref38997" refString="Harold E. 1871. Berichtigungen und Zusatze zum Catalogus Coleopterorum synonymicus et systematicus. Coleopterologische Hefte 7: 113 - 119." type="journal article" year="1871">Harold, 1871: 114</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, in part). —
<treatmentCitation id="0AC210F5FFE7FFD8FF45470FBD3C3F25" author="Vulcano M. A. &amp; Pereira F. S." box="[237,590,1284,1311]" page="575" pageId="25" pageNumber="26" year="1967">
<bibRefCitation id="EFF24B15FFE7FFD8FF45470FBD3C3F25" author="Vulcano M. A. &amp; Pereira F. S." box="[237,590,1284,1311]" pageId="25" pageNumber="26" pagination="533 - 603" refId="ref42755" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao Amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 575</bibRefCitation>
</treatmentCitation>
(in part). —
<treatmentCitation id="0AC210F5FFE7FFD8FD5C470FBB233F24" author="Martinez A &amp; Pereira F. S." box="[756,1105,1284,1310]" page="68" pageId="25" pageNumber="26" year="1967">
<bibRefCitation id="EFF24B15FFE7FFD8FD5C470FBB233F24" author="Martinez A &amp; Pereira F. S." box="[756,1105,1284,1310]" pageId="25" pageNumber="26" pagination="53 - 69" refId="ref40118" refString="Martinez A &amp; Pereira F. S. 1967. Notas escarabeidologicas III (Col. Scarabaeidae-Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez &amp; Pereira 1967: 68</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FF424722BE1F3F78" ID-CoL="874G8" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[234,365,1320,1346]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE7FFD8FF424722BE1F3F78" box="[234,365,1320,1346]" italics="true" pageId="25" pageNumber="26">P. funereus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FE054723BDD23F78" author="Edmonds W. D." box="[429,672,1320,1346]" page="830" pageId="25" pageNumber="26" year="1972">
<bibRefCitation id="EFF24B15FFE7FFD8FE054723BDD23F78" author="Edmonds W. D." box="[429,672,1320,1346]" pageId="25" pageNumber="26" pagination="731 - 874" refId="ref37422" refString="Edmonds W. D. 1972. Comparative skeletal morphology, systematics and evolution of the Phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds 1972: 830</bibRefCitation>
</treatmentCitation>
, fig. 256 (in part); 1994: 3, 8, 45–46 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FF424746BE213F5C" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[234,339,1357,1382]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FF424746BE213F5C" box="[234,339,1357,1382]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FE3E4747BD033F5C" author="Arnaud P." box="[406,625,1356,1382]" page="116" pageId="25" pageNumber="26" year="1982">
<bibRefCitation id="EFF24B15FFE7FFD8FE3E4747BD033F5C" author="Arnaud P." box="[406,625,1356,1382]" pageId="25" pageNumber="26" pagination="113 - 118" refId="ref35745" refString="Arnaud P. 1982. Liste des types de Phanaeini du Museum National D'Histoire Naturelle de Paris [Coleoptera, Scarabaeidae]. Revue francaise d'Entomologie (N. S.) 4: 113 - 118." type="journal article" year="1982">Arnaud 1982: 116</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FC0C4746BB5B3F5C" ID-CoL="874G8" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[932,1065,1356,1382]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE7FFD8FC0C4746BB5B3F5C" box="[932,1065,1356,1382]" italics="true" pageId="25" pageNumber="26">P. funereus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FBC54747BA3B3F5D" author="Krajcik M." box="[1133,1353,1356,1383]" page="152" pageId="25" pageNumber="26" year="2006">
<bibRefCitation id="EFF24B15FFE7FFD8FBC54747BA3B3F5D" author="Krajcik M." box="[1133,1353,1356,1383]" pageId="25" pageNumber="26" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 152</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C634D67FFE7FFD8FE2B477ABE993FB0" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[387,491,1393,1418]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FE2B477ABE993FB0" box="[387,491,1393,1418]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FD87477BBC043FB1" author="Edmonds W. D. &amp; Zidek J." box="[559,886,1392,1419]" page="5" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FD87477BBC043FB1" author="Edmonds W. D. &amp; Zidek J." box="[559,886,1392,1419]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012: 5</bibRefCitation>
</treatmentCitation>
, 13 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FB7D477ABA4F3FB0" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1237,1341,1393,1418]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FB7D477ABA4F3FB0" box="[1237,1341,1393,1418]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FF42479FBD293F95" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A." box="[234,603,1428,1455]" page="220" pageId="25" pageNumber="26" year="2019">
<bibRefCitation id="EFF24B15FFE7FFD8FF42479FBD293F95" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A." box="[234,603,1428,1455]" pageId="25" pageNumber="26" pagination="1 - 343" refId="ref36626" refString="Chamorro W., Marin-Armijos D. &amp; Asenjo A. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="B917EAF6FFE7FFD8FEC1479EBEEE3F94" box="[361,412,1428,1454]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2019: 220–221
</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FCD1479EBCAC3F94" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[889,990,1429,1454]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FCD1479EBCAC3F94" box="[889,990,1429,1454]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF1547B3BB193C21" blockId="25.[189,1399,1176,1742]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF1547B3BB193C21" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF1547B3BEBB3FE8" ID-CoL="4FSJS" authorityName="Arnaud" authorityYear="2002" box="[189,457,1464,1490]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="pyrois" subSpecies="malyi">
<emphasis id="B917EAF6FFE7FFD8FF1547B3BEBB3FE8" box="[189,457,1464,1490]" italics="true" pageId="25" pageNumber="26">Phanaeus pyrois malyi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC210F5FFE7FFD8FE4D47B3BDC83FE9" author="Arnaud P." box="[485,698,1464,1491]" page="97" pageId="25" pageNumber="26" year="2002">
<bibRefCitation id="EFF24B15FFE7FFD8FE4D47B3BDC83FE9" author="Arnaud P." box="[485,698,1464,1491]" pageId="25" pageNumber="26" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 97</bibRefCitation>
</treatmentCitation>
(as subspecies of
<taxonomicName id="4C634D67FFE7FFD8FC3847B2BC873FE8" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[912,1013,1465,1490]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FC3847B2BC873FE8" box="[912,1013,1465,1490]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FB9B47B3BA1D3FE9" author="Solis A. &amp; Kohlmann B." box="[1075,1391,1464,1491]" page="1" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FB9B47B3BA1D3FE9" author="Solis A. &amp; Kohlmann B." box="[1075,1391,1464,1491]" pageId="25" pageNumber="26" pagination="1 - 32" refId="ref42322" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1 - 32." type="journal article" year="2012">Solís &amp; Kohlmann 2012: 1</bibRefCitation>
</treatmentCitation>
, 9–10 (as subspecies of
<taxonomicName id="4C634D67FFE7FFD8FE5147D6BD2F3FCC" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[505,605,1501,1526]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FE5147D6BD2F3FCC" box="[505,605,1501,1526]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FD3247D7BCA03FCD" author="Edmonds W. D. &amp; Zidek J." box="[666,978,1500,1527]" page="1" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FD3247D7BCA03FCD" author="Edmonds W. D. &amp; Zidek J." box="[666,978,1500,1527]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012: 1</bibRefCitation>
</treatmentCitation>
, 8, 13 (as synonym of
<taxonomicName id="4C634D67FFE7FFD8FB7447D6BA323FCC" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1244,1344,1501,1526]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FB7447D6BA323FCC" box="[1244,1344,1501,1526]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FF42440ABECD3C21" author="Krajcik M." box="[234,447,1536,1563]" page="152" pageId="25" pageNumber="26" year="2006">
<bibRefCitation id="EFF24B15FFE7FFD8FF42440ABECD3C21" author="Krajcik M." box="[234,447,1536,1563]" pageId="25" pageNumber="26" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 152</bibRefCitation>
</treatmentCitation>
(as subspecies of
<taxonomicName id="4C634D67FFE7FFD8FD3D440ABD883C20" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[661,762,1537,1562]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FD3D440ABD883C20" box="[661,762,1537,1562]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FC9F440ABB1A3C21" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[823,1128,1537,1563]" page="78" pageId="25" pageNumber="26" year="2018">
<bibRefCitation id="EFF24B15FFE7FFD8FC9F440ABB1A3C21" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[823,1128,1537,1563]" pageId="25" pageNumber="26" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFE7FFD8FC13440ABC9C3C21" box="[955,1006,1537,1563]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2018: 78
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF15442EBBA03C05" blockId="25.[189,1399,1176,1742]" box="[189,1234,1573,1599]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF15442EBBA03C05" box="[189,1234,1573,1599]" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF15442EBDE73C05" ID-CoL="4FSJS" authorityName="Arnaud" authorityYear="2002" box="[189,661,1573,1599]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="pyrois" subGenus="Notiophanaeus" subSpecies="malyi">
<emphasis id="B917EAF6FFE7FFD8FF15442EBDE73C05" box="[189,661,1573,1599]" italics="true" pageId="25" pageNumber="26">Phanaeus (Notiophanaeus) pyrois malyi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC210F5FFE7FFD8FD19442EBCF43C05" author="Arnaud P." box="[689,902,1573,1599]" page="96" pageId="25" pageNumber="26" year="2002">
<bibRefCitation id="EFF24B15FFE7FFD8FD19442EBCF43C05" author="Arnaud P." box="[689,902,1573,1599]" pageId="25" pageNumber="26" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 96</bibRefCitation>
</treatmentCitation>
(as subspecies of
<taxonomicName id="4C634D67FFE7FFD8FBF4442EBBB33C04" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1116,1217,1573,1598]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FBF4442EBBB33C04" box="[1116,1217,1573,1598]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154442BE0A3C90" blockId="25.[189,1399,1176,1742]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF154442BE0A3C90" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF154442BE0A3C59" ID-CoL="4FSJS" authorityName="Arnaud" authorityYear="2002" box="[189,376,1609,1635]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE7FFD8FF154442BE0A3C59" box="[189,376,1609,1635]" italics="true" pageId="25" pageNumber="26">Phanaeus malyi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC210F5FFE7FFD8FE3B4442BDB93C59" author="Solis A. &amp; Kohlmann B." box="[403,715,1609,1635]" page="1" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FE3B4442BDB93C59" author="Solis A. &amp; Kohlmann B." box="[403,715,1609,1635]" pageId="25" pageNumber="26" pagination="1 - 32" refId="ref42322" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1 - 32." type="journal article" year="2012">Solís &amp; Kohlmann 2012: 1</bibRefCitation>
</treatmentCitation>
, 7, 31, fig. 1. —
<treatmentCitation id="0AC210F5FFE7FFD8FC244442BBB33C59" author="Edmonds W. D. &amp; Zidek J." box="[908,1217,1609,1635]" page="3" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FC244442BBB33C59" author="Edmonds W. D. &amp; Zidek J." box="[908,1217,1609,1635]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012: 3</bibRefCitation>
</treatmentCitation>
, 6 (as synonym of
<taxonomicName id="4C634D67FFE7FFD8FEA24466BE1C3CBC" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[266,366,1645,1670]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FEA24466BE1C3CBC" box="[266,366,1645,1670]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FE024467BC763CBC" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[426,772,1644,1671]" httpUri="http://treatment.plazi.org/id/0D21E1174692236999186DBC687B22FA" page="67" pageId="25" pageNumber="26" year="2018">
<bibRefCitation id="EFF24B15FFE7FFD8FE024467BC763CBC" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[426,772,1644,1671]" pageId="25" pageNumber="26" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFE7FFD8FD844466BD2D3CBC" box="[556,607,1644,1670]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2018: 67–69
</bibRefCitation>
</treatmentCitation>
, 78–80, 82, 88–89, figs 8, S1b. —
<bibRefCitation id="EFF24B15FFE7FFD8FB304467BE5B3C90" author="Gillett C. P. D. T. &amp; Toussaint E. F. A." pageId="25" pageNumber="26" pagination="661 - 682" refId="ref38342" refString="Gillett C. P. D. T. &amp; Toussaint E. F. A. 2020. Macroevolution and shifts in the feeding biology of the New World scarab beetle tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae). Biological Journal of the Linnean Society 130: 661 - 682. https: // doi. org / 10.1093 / biolinnean / blaa 058" type="journal article" year="2020">Gillett &amp; Toussaint 2020</bibRefCitation>
: fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF1544BFBCE53CF4" blockId="25.[189,1399,1176,1742]" box="[189,919,1716,1742]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF1544BFBCE53CF4" box="[189,919,1716,1742]" pageId="25" pageNumber="26">
<taxonomicName id="4C634D67FFE7FFD8FF1544BFBD373CF4" ID-CoL="4FSJS" authorityName="Arnaud" authorityYear="2002" box="[189,581,1716,1742]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="malyi" subGenus="Notiophanaeus">
<emphasis id="B917EAF6FFE7FFD8FF1544BFBE433CF4" box="[189,305,1716,1742]" italics="true" pageId="25" pageNumber="26">Phanaeus</emphasis>
(
<emphasis id="B917EAF6FFE7FFD8FEEA44BFBE863CF4" box="[322,500,1716,1742]" italics="true" pageId="25" pageNumber="26">Notiophanaeus</emphasis>
)
<emphasis id="B917EAF6FFE7FFD8FDAC44BFBD373CF4" box="[516,581,1716,1742]" italics="true" pageId="25" pageNumber="26">malyi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC210F5FFE7FFD8FDCB44BFBCE13CF4" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[611,915,1716,1742]" httpUri="http://treatment.plazi.org/id/0D21E1174692236999186DBC687B22FA" page="78" pageId="25" pageNumber="26" year="2018">
<bibRefCitation id="EFF24B15FFE7FFD8FDCB44BFBCE13CF4" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[611,915,1716,1742]" pageId="25" pageNumber="26" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFE7FFD8FD4E44BEBC683CF4" box="[742,794,1716,1742]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2018: 78
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF1544FCBBE63D63" blockId="25.[189,1399,1782,2025]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF1544FCBBE63D63" pageId="25" pageNumber="26">
Non
<taxonomicName id="4C634D67FFE7FFD8FF5E44FDBE903D2A" ID-CoL="874GF" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[246,482,1782,1808]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFE7FFD8FF5E44FDBE903D2A" box="[246,482,1782,1808]" italics="true" pageId="25" pageNumber="26">Phanaeus olsoufieffi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE7FFD8FDF844FDBC493D2A" author="Balthasar V." box="[592,827,1782,1808]" page="242" pageId="25" pageNumber="26" year="1939">
<bibRefCitation id="EFF24B15FFE7FFD8FDF844FDBC493D2A" author="Balthasar V." box="[592,827,1782,1808]" pageId="25" pageNumber="26" pagination="238 - 247" refId="ref36059" refString="Balthasar V. 1939. Neue Phanaeus - Arten. Folia Zoologica et Hydrobiologica 9: 238 - 247." type="journal article" year="1939">Balthasar 1939: 242</bibRefCitation>
</treatmentCitation>
(in part). —
<treatmentCitation id="0AC210F5FFE7FFD8FC7944FDBBED3D2A" author="Edmonds W. D." box="[977,1183,1782,1808]" page="8" pageId="25" pageNumber="26" year="1994">
<bibRefCitation id="EFF24B15FFE7FFD8FC7944FDBBED3D2A" author="Edmonds W. D." box="[977,1183,1782,1808]" pageId="25" pageNumber="26" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994: 8</bibRefCitation>
</treatmentCitation>
, 45–46 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FED54510BE933D0E" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[381,481,1819,1844]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FED54510BE933D0E" box="[381,481,1819,1844]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FDB54511BD803D0E" author="Arnaud P." box="[541,754,1818,1844]" page="96" pageId="25" pageNumber="26" year="2002">
<bibRefCitation id="EFF24B15FFE7FFD8FDB54511BD803D0E" author="Arnaud P." box="[541,754,1818,1844]" pageId="25" pageNumber="26" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 96</bibRefCitation>
</treatmentCitation>
(in part). —
<treatmentCitation id="0AC210F5FFE7FFD8FC234511BB133D0F" author="Krajcik M." box="[907,1121,1818,1845]" page="152" pageId="25" pageNumber="26" year="2006">
<bibRefCitation id="EFF24B15FFE7FFD8FC234511BB133D0F" author="Krajcik M." box="[907,1121,1818,1845]" pageId="25" pageNumber="26" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 152</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FF424534BE3D3D62" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[234,335,1855,1880]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FF424534BE3D3D62" box="[234,335,1855,1880]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FE244535BDB53D62" author="Edmonds W. D. &amp; Zidek J." box="[396,711,1854,1880]" page="3" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FE244535BDB53D62" author="Edmonds W. D. &amp; Zidek J." box="[396,711,1854,1880]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012: 3</bibRefCitation>
</treatmentCitation>
, 5–6 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FBB64534BBF13D62" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1054,1155,1855,1880]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FBB64534BBF13D62" box="[1054,1155,1855,1880]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154568BBE43DD3" blockId="25.[189,1399,1782,2025]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB7311CAFFE7FFD8FF154568BBE43DD3" pageId="25" pageNumber="26">
Non
<taxonomicName id="4C634D67FFE7FFD8FF504569BEBC3D46" ID-CoL="856BK" authorityName="Blackwelder" authorityYear="1944" box="[248,462,1890,1916]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="bothrus">
<emphasis id="B917EAF6FFE7FFD8FF504569BEBC3D46" box="[248,462,1890,1916]" italics="true" pageId="25" pageNumber="26">Phanaeus bothrus</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE7FFD8FDED4569BC2B3D46" author="Blackwelder R. E." box="[581,857,1890,1916]" page="209" pageId="25" pageNumber="26" year="1944">
<bibRefCitation id="EFF24B15FFE7FFD8FDED4569BC2B3D46" author="Blackwelder R. E." box="[581,857,1890,1916]" pageId="25" pageNumber="26" pagination="1 - 1492" refId="ref36166" refString="Available from https: // www. biodiversitylibrary. org / item / 135502 # page / 7 / mode / 1 up [accessed Jun. 2020]. Blackwelder R. E. 1944 - 1957. Checklist of the coleopterous insects of Mexico, Central American, the West Indies, and South America. Parts 1 - 6. United States National Museum, Bulletin 185: 1 - 1492. https: // doi. org / 10.5479 / si. 03629236.185.927" type="journal article" year="1944">Blackwelder 1944: 209</bibRefCitation>
</treatmentCitation>
(in part). —
<treatmentCitation id="0AC210F5FFE7FFD8FC5F4569BA3F3D46" author="Martinez A &amp; Pereira F. S." box="[1015,1357,1890,1916]" page="68" pageId="25" pageNumber="26" year="1967">
<bibRefCitation id="EFF24B15FFE7FFD8FC5F4569BA3F3D46" author="Martinez A &amp; Pereira F. S." box="[1015,1357,1890,1916]" pageId="25" pageNumber="26" pagination="53 - 69" refId="ref40118" refString="Martinez A &amp; Pereira F. S. 1967. Notas escarabeidologicas III (Col. Scarabaeidae-Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez &amp; Pereira 1967: 68</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FE4D458CBD173D9A" ID-CoL="874G8" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[485,613,1926,1952]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE7FFD8FE4D458CBD173D9A" box="[485,613,1926,1952]" italics="true" pageId="25" pageNumber="26">P. funereus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FD00458DBC0F3D9A" author="Edmonds W. D." box="[680,893,1926,1952]" page="8" pageId="25" pageNumber="26" year="1994">
<bibRefCitation id="EFF24B15FFE7FFD8FD00458DBC0F3D9A" author="Edmonds W. D." box="[680,893,1926,1952]" pageId="25" pageNumber="26" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994: 8</bibRefCitation>
</treatmentCitation>
, 45 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FB71458CBA4C3D9A" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1241,1342,1927,1952]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FB71458CBA4C3D9A" box="[1241,1342,1927,1952]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FF4545A1BEB03DFE" author="Arnaud P." box="[237,450,1962,1988]" page="97" pageId="25" pageNumber="26" year="2002">
<bibRefCitation id="EFF24B15FFE7FFD8FF4545A1BEB03DFE" author="Arnaud P." box="[237,450,1962,1988]" pageId="25" pageNumber="26" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 97</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FD7645A0BC1F3DFE" ID-CoL="874GF" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[734,877,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFE7FFD8FD7645A0BC1F3DFE" box="[734,877,1962,1988]" italics="true" pageId="25" pageNumber="26">P. olsoufieffi</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FC0245A1BB0D3DFF" author="Krajcik M." box="[938,1151,1962,1989]" page="152" pageId="25" pageNumber="26" year="2006">
<bibRefCitation id="EFF24B15FFE7FFD8FC0245A1BB0D3DFF" author="Krajcik M." box="[938,1151,1962,1989]" pageId="25" pageNumber="26" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 152</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FEA245C4BE1D3DD2" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[266,367,1999,2024]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FEA245C4BE1D3DD2" box="[266,367,1999,2024]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0AC210F5FFE7FFD8FE0445C5BD943DD2" author="Edmonds W. D. &amp; Zidek J." box="[428,742,1998,2024]" page="3" pageId="25" pageNumber="26" year="2012">
<bibRefCitation id="EFF24B15FFE7FFD8FE0445C5BD943DD2" author="Edmonds W. D. &amp; Zidek J." box="[428,742,1998,2024]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012: 3</bibRefCitation>
</treatmentCitation>
, 5 (in part, as synonym of
<taxonomicName id="4C634D67FFE7FFD8FB8845C4BBF73DD2" ID-CoL="4FSJX" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1056,1157,1999,2024]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FB8845C4BBF73DD2" box="[1056,1157,1999,2024]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF154301BE0E3B7D" blockId="26.[189,1398,265,468]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB7311CAFFE4FFDBFF154301BE0E3B7D" pageId="26" pageNumber="27">
Non
<taxonomicName id="4C634D67FFE4FFDBFF514302BDF43B19" ID-CoL="856BJ" authorityName="Harold" authorityYear="1871" box="[249,646,265,292]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="blanchardi" subGenus="Phanaeus">
<emphasis id="B917EAF6FFE4FFDBFF514302BE1F3B19" box="[249,365,265,291]" italics="true" pageId="26" pageNumber="27">Phanaeus</emphasis>
(
<emphasis id="B917EAF6FFE4FFDBFE284302BE863B19" box="[384,500,265,291]" italics="true" pageId="26" pageNumber="27">Phanaeus</emphasis>
)
<emphasis id="B917EAF6FFE4FFDBFDAE4302BDF43B19" box="[518,646,265,291]" italics="true" pageId="26" pageNumber="27">blanchardi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE4FFDBFD554302BB243B19" author="Martinez A &amp; Pereira F. S." box="[765,1110,265,291]" page="68" pageId="26" pageNumber="27" year="1967">
<bibRefCitation id="EFF24B15FFE4FFDBFD554302BB243B19" author="Martinez A &amp; Pereira F. S." box="[765,1110,265,291]" pageId="26" pageNumber="27" pagination="53 - 69" refId="ref40118" refString="Martinez A &amp; Pereira F. S. 1967. Notas escarabeidologicas III (Col. Scarabaeidae-Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez &amp; Pereira 1967: 68</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE4FFDBFF444326BE193B7D" ID-CoL="874G8" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[236,363,301,327]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE4FFDBFF444326BE193B7D" box="[236,363,301,327]" italics="true" pageId="26" pageNumber="27">P. funereus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15435BBE0E3BB4" blockId="26.[189,1398,265,468]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB7311CAFFE4FFDBFF15435BBE0E3BB4" pageId="26" pageNumber="27">
Non
<taxonomicName id="4C634D67FFE4FFDBFF52435BBD093B50" ID-CoL="874GF" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[250,635,336,362]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="olsoufieffi" subGenus="Phanaeus">
<emphasis id="B917EAF6FFE4FFDBFF52435BBE1C3B50" box="[250,366,336,362]" italics="true" pageId="26" pageNumber="27">Phanaeus</emphasis>
(
<emphasis id="B917EAF6FFE4FFDBFE29435BBE873B50" box="[385,501,336,362]" italics="true" pageId="26" pageNumber="27">Phanaeus</emphasis>
)
<emphasis id="B917EAF6FFE4FFDBFDA1435BBD093B50" box="[521,635,336,362]" italics="true" pageId="26" pageNumber="27">olsoufieffi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE4FFDBFD5D435BBB233B50" author="Martinez A &amp; Pereira F. S." box="[757,1105,336,362]" page="68" pageId="26" pageNumber="27" year="1967">
<bibRefCitation id="EFF24B15FFE4FFDBFD5D435BBB233B50" author="Martinez A &amp; Pereira F. S." box="[757,1105,336,362]" pageId="26" pageNumber="27" pagination="53 - 69" refId="ref40118" refString="Martinez A &amp; Pereira F. S. 1967. Notas escarabeidologicas III (Col. Scarabaeidae-Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez &amp; Pereira 1967: 68</bibRefCitation>
</treatmentCitation>
(in part, as synonym of
<taxonomicName id="4C634D67FFE4FFDBFF42437FBE193BB7" ID-CoL="874G8" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[234,363,371,397]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE4FFDBFF42437FBE193BB7" box="[234,363,371,397]" italics="true" pageId="26" pageNumber="27">P. funereus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15439CBBC13B8B" blockId="26.[189,1398,265,468]" box="[189,1203,406,433]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB7311CAFFE4FFDBFF15439CBBC13B8B" box="[189,1203,406,433]" pageId="26" pageNumber="27">
Non
<taxonomicName id="4C634D67FFE4FFDBFF5F439DBC723B8A" ID-CoL="874GF" authorityName="Balthasar" authorityYear="1939" box="[247,768,406,433]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="pyrois" subGenus="Notiophanaeus" subSpecies="olsoufieffi">
<emphasis id="B917EAF6FFE4FFDBFF5F439DBE193B8A" box="[247,363,406,432]" italics="true" pageId="26" pageNumber="27">Phanaeus</emphasis>
(
<emphasis id="B917EAF6FFE4FFDBFED4439DBD5C3B8A" box="[380,558,406,432]" italics="true" pageId="26" pageNumber="27">Notiophanaeus</emphasis>
)
<emphasis id="B917EAF6FFE4FFDBFD96439CBC723B8A" box="[574,768,406,432]" italics="true" pageId="26" pageNumber="27">pyrois olsoufieffi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE4FFDBFCD8439DBB373B8A" author="Arnaud P." box="[880,1093,406,432]" page="96" pageId="26" pageNumber="27" year="2002">
<bibRefCitation id="EFF24B15FFE4FFDBFCD8439DBB373B8A" author="Arnaud P." box="[880,1093,406,432]" pageId="26" pageNumber="27" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 96</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF1543B1BB3D3BEE" blockId="26.[189,1398,265,468]" box="[189,1103,442,468]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB7311CAFFE4FFDBFF1543B1BB3D3BEE" box="[189,1103,442,468]" pageId="26" pageNumber="27">
Non
<taxonomicName id="4C634D67FFE4FFDBFF5F43B1BD413BEE" ID-CoL="874GF" authorityName="Balthasar" authorityYear="1939" box="[247,563,442,468]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="pyrois" subSpecies="olsoufieffi">
<emphasis id="B917EAF6FFE4FFDBFF5F43B1BD413BEE" box="[247,563,442,468]" italics="true" pageId="26" pageNumber="27">Phanaeus pyrois olsoufieffi</emphasis>
</taxonomicName>
(error) –
<treatmentCitation id="0AC210F5FFE4FFDBFD0B43B1BC0E3BEE" author="Arnaud P." box="[675,892,442,468]" page="98" pageId="26" pageNumber="27" year="2002">
<bibRefCitation id="EFF24B15FFE4FFDBFD0B43B1BC0E3BEE" author="Arnaud P." box="[675,892,442,468]" pageId="26" pageNumber="27" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 98</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210F5FFE4FFDBFC2043B1BC933BEE" author="Arnaud P." box="[904,993,442,468]" page="4" pageId="26" pageNumber="27" year="2018">
<bibRefCitation id="EFF24B15FFE4FFDBFC2043B1BC933BEE" author="Arnaud P." box="[904,993,442,468]" pageId="26" pageNumber="27" pagination="3 - 7" refId="ref35890" refString="Arnaud P. 2018. Description d'une nouvelle espece de Phanaeus d'Equateur et revalidation de l'espece Coprophanaeus (C.) edmondsi Arnaud (Coleoptera: Scarabaeidae, Scarabaeinae). Besoiro 26: 3 - 7." type="journal article" year="2018">2018: 4</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE4FFDBFF1543F0BD8038F5" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BDC36E4FFE4FFDBFF1543F0BE44382F" blockId="26.[189,310,507,533]" box="[189,310,507,533]" pageId="26" pageNumber="27">
<heading id="D0948188FFE4FFDBFF1543F0BE44382F" bold="true" box="[189,310,507,533]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B917EAF6FFE4FFDBFF1543F0BE44382F" bold="true" box="[189,310,507,533]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF154022BD8038F5" blockId="26.[188,1398,553,719]" pageId="26" pageNumber="27">
Easily diagnosed species by the bright black colour with red-green sheen on frontolateral angles of pronotum (
<figureCitation id="13582A61FFE4FFDBFE974047BEE9385C" box="[319,411,588,614]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="26" pageNumber="27">Fig. 2E</figureCitation>
); elytral striae fine, smooth, impressed basally as distinct fossae (
<figureCitation id="13582A61FFE4FFDBFB314047BB93385D" box="[1177,1249,588,615]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="26" pageNumber="27">Fig. 7</figureCitation>
). The rest of black species within the
<taxonomicName id="4C634D67FFE4FFDBFE4B407BBD0038B3" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[483,626,623,649]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B917EAF6FFE4FFDBFE4B407BBD0038B3" box="[483,626,623,649]" italics="true" pageId="26" pageNumber="27">P. endymion</emphasis>
</taxonomicName>
species group (
<taxonomicName id="4C634D67FFE4FFDBFC87407BBCC338B3" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[815,945,623,649]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="funereus">
<emphasis id="B917EAF6FFE4FFDBFC87407BBCC338B3" box="[815,945,623,649]" italics="true" pageId="26" pageNumber="27">P. funereus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D67FFE4FFDBFC68407BBB2238B3" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[960,1104,623,649]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFE4FFDBFC68407BBB2238B3" box="[960,1104,623,649]" italics="true" pageId="26" pageNumber="27">P. olsoufieffi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D67FFE4FFDBFBC8407BBA7B38B3" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[1120,1289,624,649]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFE4FFDBFBC8407BBA7B38B3" box="[1120,1289,624,649]" italics="true" pageId="26" pageNumber="27">P. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE4FFDBFAB9407BBA1C38B0" box="[1297,1390,624,650]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
) are differentiated from
<taxonomicName id="4C634D67FFE4FFDBFE634098BD5A3896" authorityName="Arnaud" authorityYear="2002" box="[459,552,658,684]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE4FFDBFE634098BD5A3896" box="[459,552,658,684]" italics="true" pageId="26" pageNumber="27">P. malyi</emphasis>
</taxonomicName>
by the elytral striae not strongly impressed basally as distinct fossae and the shape of the endophallite copulatrix (
<figureCitation id="13582A61FFE4FFDBFD3440BEBD9338F5" box="[668,737,693,719]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="26" pageNumber="27">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE4FFDBFF1540FDBB6F3CB2" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BDC36E4FFE4FFDBFF1540FDBE1E392A" blockId="26.[189,364,758,784]" box="[189,364,758,784]" pageId="26" pageNumber="27">
<heading id="D0948188FFE4FFDBFF1540FDBE1E392A" bold="true" box="[189,364,758,784]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B917EAF6FFE4FFDBFF1540FDBE1E392A" bold="true" box="[189,364,758,784]" pageId="26" pageNumber="27">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF42412FBE8B3905" blockId="26.[189,1398,804,901]" box="[234,505,804,831]" pageId="26" pageNumber="27">
<emphasis id="B917EAF6FFE4FFDBFF42412FBE2E3904" bold="true" box="[234,348,804,830]" pageId="26" pageNumber="27">
<typeStatus id="54D88846FFE4FFDBFF42412FBE2E3904" box="[234,348,804,830]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
(not studied)
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15414CBDF839BE" blockId="26.[189,1398,804,901]" pageId="26" pageNumber="27">
<materialsCitation id="3B0B3CB9FFE4FFDBFF15414CBDF839BE" ID-GBIF-Occurrence="3093980471" collectionCode="INBIO, MNCR" country="Costa Rica" location="Estacion Quebrada Bonita" municipality="Carara National Park" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
<collectingCountry id="F3747674FFE4FFDBFF15414CBE18395B" box="[189,362,839,865]" name="Costa Rica" pageId="26" pageNumber="27">COSTA RICA</collectingCountry>
•
<specimenCount id="9D65FD6DFFE4FFDBFE214143BED33958" box="[393,417,840,866]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
,
<bibRefCitation id="EFF24B15FFE4FFDBFE07414CBD0C395B" author="Arnaud P." box="[431,638,839,865]" pageId="26" pageNumber="27" pagination="2 - 5" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." type="journal article" year="2002">Arnaud 2002a: 3</bibRefCitation>
;
<collectingRegion id="49A7F806FFE4FFDBFD274143BC613958" box="[655,787,840,866]" country="Costa Rica" name="Puntarenas" pageId="26" pageNumber="27">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6BB8AC9EFFE4FFDBFC8A414CBB56395B" box="[802,1060,839,865]" pageId="26" pageNumber="27">Carara National Park</collectingMunicipality>
,
<location id="8EBC603FFFE4FFDBFB9B414CBA00395B" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFB9B414CBA00395B" box="[1075,1394,839,865]" country="Costa Rica" municipality="Carara National Park" name="Estacion Quebrada Bonita" pageId="26" pageNumber="27" stateProvince="Puntarenas">Estación Quebrada Bonita</location>
; originally deposited at
<collectionCode id="ED72AE21FFE4FFDBFE634161BD5239BE" box="[459,544,874,900]" pageId="26" pageNumber="27">INBIO</collectionCode>
;
<collectionCode id="ED72AE21FFE4FFDBFD844161BDF439BE" box="[556,646,874,900]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="26" pageNumber="27">MNCR</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF1541A0BDCD39FC" blockId="26.[189,703,939,966]" box="[189,703,939,966]" pageId="26" pageNumber="27">
<emphasis id="B917EAF6FFE4FFDBFF1541A0BD7039FF" bold="true" box="[189,514,939,965]" pageId="26" pageNumber="27">Non-type material revised</emphasis>
(
<specimenCount id="9D65FD6DFFE4FFDBFDBB41A0BD1439FC" box="[531,614,939,966]" pageId="26" pageNumber="27" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D65FD6DFFE4FFDBFDDB41A0BDC139FC" box="[627,691,939,966]" pageId="26" pageNumber="27" type="female">6 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF1541D1BDA83E00" blockId="26.[189,1399,985,1082]" pageId="26" pageNumber="27">
<materialsCitation id="3B0B3CB9FFE4FFDBFF1541D1BDA43E00" ID-GBIF-Occurrence="3093980462" collectionCode="MNHN" collectorName="Coll. R. Oberthur" country="Colombia" location="COLOMBIA" pageId="26" pageNumber="27" specimenCode="MNHN EC10569" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="F3747674FFE4FFDBFF1541D1BE2B39CE" box="[189,345,986,1012]" name="Colombia" pageId="26" pageNumber="27">COLOMBIA</collectingCountry>
•
<specimenCount id="9D65FD6DFFE4FFDBFED441D1BEC239CE" box="[380,432,986,1012]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; unknown locality; “[Illegible data]/ Ex-mus ӕo D. Sharp 1890/ Museum Paris ex
<collectorName id="26965332FFE4FFDBFF4141F7BEB53E2C" box="[233,455,1020,1047]" pageId="26" pageNumber="27">Coll. R. Oberthur</collectorName>
/ PARALECTOTYPE/ PARALECTOTYPE
<taxonomicName id="4C634D67FFE4FFDBFC7041F7BE0F3E00" authority="Olsoufieff, 1924" authorityName="Olsoufieff" authorityYear="1924" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="blanchardi" subGenus="Phanaeus">
<emphasis id="B917EAF6FFE4FFDBFC7041F7BA053E2C" box="[984,1399,1020,1046]" italics="true" pageId="26" pageNumber="27">Phanaeus (Phanaeus) blanchardi</emphasis>
<bibRefCitation id="EFF24B15FFE4FFDBFF154614BE0F3E00" author="Olsoufieff G. d'" box="[189,381,1055,1082]" pageId="26" pageNumber="27" pagination="4 - 172" refId="ref41334" refString="Olsoufieff G. d'. 1924. Les Phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta, Revue illustree d'Entomologique 13: 4 - 172." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
</taxonomicName>
/
<specimenCode id="DBC59E9FFFE4FFDBFE24462BBD103E00" box="[396,610,1055,1082]" pageId="26" pageNumber="27">MNHN EC10569</specimenCode>
”;
<collectionCode id="ED72AE21FFE4FFDBFDD0462BBDA43E00" box="[632,726,1056,1082]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15466ABECD3F8F" blockId="26.[189,1399,1121,1462]" pageId="26" pageNumber="27">
<materialsCitation id="3B0B3CB9FFE4FFDBFF15466ABE723EA4" ID-GBIF-Occurrence="3093980513" collectingDate="1996-07-29" collectionCode="VMC" collectorName="Coll. Louis La Pierre" country="Costa Rica" location="San Luis Vly." municipality="San Luis" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<collectingCountry id="F3747674FFE4FFDBFF15466ABE183E41" box="[189,362,1121,1147]" name="Costa Rica" pageId="26" pageNumber="27">COSTA RICA</collectingCountry>
–
<emphasis id="B917EAF6FFE4FFDBFE24466ABD693E41" bold="true" box="[396,539,1121,1147]" pageId="26" pageNumber="27">
<collectingRegion id="49A7F806FFE4FFDBFE24466ABD693E41" box="[396,539,1121,1147]" country="Costa Rica" name="Puntarenas" pageId="26" pageNumber="27">Puntarenas</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE4FFDBFD92466ABD1E3E41" box="[570,620,1121,1147]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFE4FFDBFD2E466ABD843E41" box="[646,758,1121,1147]" pageId="26" pageNumber="27">San Luis</collectingMunicipality>
,
<location id="8EBC603FFFE4FFDBFCAD466ABCDE3E41" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFCAD466ABCDE3E41" box="[773,940,1121,1147]" country="Costa Rica" municipality="San Luis" name="San Luis Vly." pageId="26" pageNumber="27" stateProvince="Puntarenas">San Luis Vly.</location>
<date id="FFDD1024FFE4FFDBFC1D466ABB203E41" box="[949,1106,1121,1147]" pageId="26" pageNumber="27" value="1996-07-29">
<collectingDate id="EF99E9CCFFE4FFDBFC1D466ABB203E41" box="[949,1106,1121,1147]" pageId="26" pageNumber="27" value="1996-07-29">VII-29–1996</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFBC8466ABA1A3E41" box="[1120,1384,1121,1147]" pageId="26" pageNumber="27">Coll. Louis La Pierre</collectorName>
”;
<collectionCode id="ED72AE21FFE4FFDBFF15468FBE723EA4" box="[189,256,1156,1182]" pageId="26" pageNumber="27">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFEB4468FBA573EA4" ID-GBIF-Occurrence="3093980371" box="[284,1317,1155,1182]" collectingDate="1927-04" collectingDateMax="1967-05-07" collectingDateMin="1927-04" collectingMethod="human feces?" collectionCode="TAMU" collectorName="D. F. Viers" country="Costa Rica" location="San Vito." pageId="26" pageNumber="27" specimenCount="101" specimenCount-female="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFEB4468FBE383EA4" box="[284,330,1156,1182]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; “
<locationDeviation id="9A6E2FC6FFE4FFDBFECC4688BD5D3EA7" box="[356,559,1155,1182]" pageId="26" pageNumber="27">
<quantity id="4C9B9B01FFE4FFDBFECC4688BEE93EA7" box="[356,411,1155,1181]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="27" unit="km" value="6.0">6km</quantity>
S. San Vito
</locationDeviation>
.
<date id="FFDD1024FFE4FFDBFD93468FBDF33EA4" box="[571,641,1156,1182]" bridgedPair="-" pageId="26" pageNumber="27" value="1927-04">
<collectingDate id="EF99E9CCFFE4FFDBFD93468FBDF33EA4" box="[571,641,1156,1182]" bridgedPair="-" pageId="26" pageNumber="27" value="1927-04">27-IV</collectingDate>
</date>
/
<date id="FFDD1024FFE4FFDBFD23468FBD8F3EA7" box="[651,765,1155,1182]" pageId="26" pageNumber="27" value="1967-05-07">
<collectingDate id="EF99E9CCFFE4FFDBFD23468FBD8F3EA7" box="[651,765,1155,1182]" pageId="26" pageNumber="27" value="1967-05-07">
7-V
<quantity id="4C9B9B01FFE4FFDBFD174688BD8F3EA7" box="[703,765,1155,1181]" metricMagnitude="3" metricUnit="m" metricValue="1.9670999999999998" pageId="26" pageNumber="27" unit="m" value="1967.1">1967</quantity>
</collectingDate>
</date>
.
<specimenCount id="9D65FD6DFFE4FFDBFCA2468FBC273EA4" box="[778,853,1156,1182]" pageId="26" pageNumber="27" type="generic">100m.</specimenCount>
<collectorName id="26965332FFE4FFDBFCF5468FBCA73EA7" box="[861,981,1155,1182]" pageId="26" pageNumber="27">D.F. Viers</collectorName>
col.
<collectingMethod id="52224EF3FFE4FFDBFBBA4688BBC83EA7" box="[1042,1210,1155,1181]" pageId="26" pageNumber="27">human feces?</collectingMethod>
”;
<collectionCode id="ED72AE21FFE4FFDBFB79468FBA573EA4" box="[1233,1317,1156,1182]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="26" pageNumber="27">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFAE9468FBC4F3EFB" ID-GBIF-Occurrence="3093980327" collectingDate="1982-07" collectionCode="IEXA" collectorName="B. Gill." country="Costa Rica" elevation="1200" location="S. Vito" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFAE9468FBA033EA4" box="[1345,1393,1156,1182]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFF6046ADBE643EFA" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFF6046ADBE643EFA" box="[200,278,1190,1216]" country="Costa Rica" name="S. Vito" pageId="26" pageNumber="27" stateProvince="Puntarenas">S.Vito</location>
,
<location id="8EBC603FFFE4FFDBFE8946ACBED93EFB" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFE8946ACBED93EFB" box="[289,427,1191,1217]" country="Costa Rica" name="Las Cruces." pageId="26" pageNumber="27" stateProvince="Puntarenas">Las Cruces.</location>
<date id="FFDD1024FFE4FFDBFE1846ADBD573EFB" box="[432,549,1190,1217]" pageId="26" pageNumber="27" value="1982-07">
<collectingDate id="EF99E9CCFFE4FFDBFE1846ADBD573EFB" box="[432,549,1190,1217]" pageId="26" pageNumber="27" value="1982-07">July 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFD8746ADBD0D3EFA" box="[559,639,1190,1216]" pageId="26" pageNumber="27">B.Gill.</collectorName>
<quantity id="4C9B9B01FFE4FFDBFD2D46ACBD923EFB" box="[645,736,1191,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">
<elevation id="004ED1D7FFE4FFDBFD2D46ACBD923EFB" box="[645,736,1191,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">1200 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFE4FFDBFD5F46ACBC4F3EFB" box="[759,829,1191,1217]" pageId="26" pageNumber="27">IEXA</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFCF946ACBE633EDE" ID-GBIF-Occurrence="3093980559" collectingDate="1982-07" collectionCode="TAMU" collectorName="S. Vito &amp; B. Gill." country="Costa Rica" elevation="1200" location="Las Cruces." pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFCF946ACBC0C3EFB" box="[849,894,1191,1217]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D65FD6DFFE4FFDBFC2146ACBCC73EFB" box="[905,949,1191,1217]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFE4FFDBFF1546C1BE633EDE" box="[189,273,1226,1252]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="26" pageNumber="27">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFE9846C1BBA83EDE" ID-GBIF-Occurrence="3093980437" box="[304,1242,1225,1252]" collectingDate="1982-08-17" collectingDateMax="1982-09-12" collectingDateMin="1982-08-17" collectionCode="TAMU" collectorName="B. Gill." country="Costa Rica" elevation="1200" location="S. Vito" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFE9846C1BE133EDE" box="[304,353,1226,1252]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D65FD6DFFE4FFDBFED846C1BED23EDE" box="[368,416,1226,1252]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFE1446C2BD793ED9" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFE1446C2BD793ED9" box="[444,523,1225,1251]" country="Costa Rica" name="S. Vito" pageId="26" pageNumber="27" stateProvince="Puntarenas">S.Vito</location>
,
<location id="8EBC603FFFE4FFDBFDB246C1BDDA3EDE" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFDB246C1BDDA3EDE" box="[538,680,1226,1252]" country="Costa Rica" name="Las Cruces." pageId="26" pageNumber="27" stateProvince="Puntarenas">Las Cruces.</location>
<date id="FFDD1024FFE4FFDBFD1946C1BCD13EDE" box="[689,931,1226,1252]" pageId="26" pageNumber="27" value="1982-08-17" valueMax="1982-09-12" valueMin="1982-08-17">
<collectingDate id="EF99E9CCFFE4FFDBFD1946C1BCD13EDE" box="[689,931,1226,1252]" pageId="26" pageNumber="27" value="1982-08-17" valueMax="1982-09-12" valueMin="1982-08-17">17 VIII-12 IX 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFC1A46C2BB703ED9" box="[946,1026,1225,1251]" pageId="26" pageNumber="27">B.Gill.</collectorName>
<quantity id="4C9B9B01FFE4FFDBFBA446C1BB1E3EDE" box="[1036,1132,1226,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">
<elevation id="004ED1D7FFE4FFDBFBA446C1BB1E3EDE" box="[1036,1132,1226,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">1200 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFE4FFDBFB2E46C1BBA83EDE" box="[1158,1242,1226,1252]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="26" pageNumber="27">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFB5146C1BDE23F3C" ID-GBIF-Occurrence="3093980567" collectingDate="1982-08-17" collectingDateMax="1982-09-12" collectingDateMin="1982-08-17" collectionCode="UVGC" collectorName="S. Vito &amp; B. Gill." country="Costa Rica" elevation="1200" location="Las Cruces." pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFB5146C1BA593EDE" box="[1273,1323,1226,1252]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFE4FFDBFD9546E7BDE23F3C" box="[573,656,1260,1286]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="26" pageNumber="27">UVGC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFD0246E7BE723F13" ID-GBIF-Occurrence="3093980429" collectingDate="1982-08-15" collectingDateMax="1982-08-18" collectingDateMin="1982-08-15" collectionCode="VMC" collectorName="B. Gill." country="Costa Rica" elevation="1200" location="S. Vito" pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFD0246E7BDA53F3D" box="[682,727,1260,1287]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFD5846E7BC4D3F3C" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFD5846E7BC4D3F3C" box="[752,831,1260,1286]" country="Costa Rica" name="S. Vito" pageId="26" pageNumber="27" stateProvince="Puntarenas">S.Vito</location>
,
<location id="8EBC603FFFE4FFDBFCE346E6BCA43F3C" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFCE346E6BCA43F3C" box="[843,982,1260,1287]" country="Costa Rica" name="Las Cruces." pageId="26" pageNumber="27" stateProvince="Puntarenas">Las Cruces.</location>
<date id="FFDD1024FFE4FFDBFC7546E7BBD43F3C" box="[989,1190,1260,1287]" pageId="26" pageNumber="27" value="1982-08-15" valueMax="1982-08-18" valueMin="1982-08-15">
<collectingDate id="EF99E9CCFFE4FFDBFC7546E7BBD43F3C" box="[989,1190,1260,1287]" pageId="26" pageNumber="27" value="1982-08-15" valueMax="1982-08-18" valueMin="1982-08-15">15–18 VIII 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFB1946E7BA733F3C" box="[1201,1281,1260,1286]" pageId="26" pageNumber="27">B.Gill.</collectorName>
<quantity id="4C9B9B01FFE4FFDBFAA046E7BA173F3C" box="[1288,1381,1260,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">
<elevation id="004ED1D7FFE4FFDBFAA046E7BA173F3C" box="[1288,1381,1260,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">1200 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFE4FFDBFF154704BE723F13" box="[189,256,1295,1321]" pageId="26" pageNumber="27">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFE884704BEAE3F76" ID-GBIF-Occurrence="3093980406" collectedFrom="tropical lluviosa" collectingDate="1969-10-03" collectingMethod="Cebo excremento" collectionCode="GHC" collectorName="G. Halffter &amp; P. Reyes C. &amp; Selva" country="Costa Rica" location="Rincon de Osa." pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFE884704BE203F10" box="[288,338,1295,1322]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFEC44704BD593F13" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFEC44704BD593F13" box="[364,555,1295,1321]" country="Costa Rica" name="Rincon de Osa." pageId="26" pageNumber="27" stateProvince="Puntarenas">Rincón de Osa.</location>
<date id="FFDD1024FFE4FFDBFD9D4704BDFF3F13" box="[565,653,1295,1321]" pageId="26" pageNumber="27" value="1969-10-03">
<collectingDate id="EF99E9CCFFE4FFDBFD9D4704BDFF3F13" box="[565,653,1295,1321]" pageId="26" pageNumber="27" value="1969-10-03">3-X-69</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFD344704BC6D3F13" box="[668,799,1295,1321]" pageId="26" pageNumber="27">G. Halffter</collectorName>
y
<collectorName id="26965332FFE4FFDBFCEC471BBCA03F13" box="[836,978,1295,1322]" pageId="26" pageNumber="27">P. Reyes C.</collectorName>
, col.
<collectorName id="26965332FFE4FFDBFBB14704BB283F13" box="[1049,1114,1295,1321]" pageId="26" pageNumber="27">Selva</collectorName>
<collectedFrom id="387F7519FFE4FFDBFBCD4704BA583F13" box="[1125,1322,1295,1321]" pageId="26" pageNumber="27">tropical lluviosa</collectedFrom>
.
<collectingMethod id="52224EF3FFE4FFDBFA914704BE383F77" pageId="26" pageNumber="27">Cebo excremento</collectingMethod>
. Día”;
<collectionCode id="ED72AE21FFE4FFDBFE364739BEAE3F76" box="[414,476,1330,1356]" pageId="26" pageNumber="27">GHC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFE5F4739BDFA3F55" ID-GBIF-Occurrence="3093980538" collectedFrom="tropical lluviosa" collectingDate="1969-10-05" collectingMethod="Cebo excremento" collectionCode="GHC" collectorName="G. Halffter &amp; P. Reyes C. &amp; Selva" country="Costa Rica" location="Rincon de Osa." pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFE5F4739BD573F77" box="[503,549,1330,1357]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFD974739BD8B3F76" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFD974739BD8B3F76" box="[575,761,1330,1356]" country="Costa Rica" name="Rincon de Osa." pageId="26" pageNumber="27" stateProvince="Puntarenas">Rincón de Osa.</location>
<date id="FFDD1024FFE4FFDBFCA94739BC283F77" box="[769,858,1330,1357]" pageId="26" pageNumber="27" value="1969-10-05">
<collectingDate id="EF99E9CCFFE4FFDBFCA94739BC283F77" box="[769,858,1330,1357]" pageId="26" pageNumber="27" value="1969-10-05">5-X-69</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFCCE4739BC953F76" box="[870,999,1330,1356]" pageId="26" pageNumber="27">G. Halffter</collectorName>
y
<collectorName id="26965332FFE4FFDBFBAF4738BBE33F76" box="[1031,1169,1330,1357]" pageId="26" pageNumber="27">P. Reyes C.</collectorName>
, col.
<collectorName id="26965332FFE4FFDBFB7B4739BA663F76" box="[1235,1300,1330,1356]" pageId="26" pageNumber="27">Selva</collectorName>
<collectedFrom id="387F7519FFE4FFDBFAB54739BE6C3F55" pageId="26" pageNumber="27">tropical lluviosa</collectedFrom>
.
<collectingMethod id="52224EF3FFE4FFDBFE80475EBE883F4A" box="[296,506,1365,1392]" pageId="26" pageNumber="27">Cebo excremento</collectingMethod>
. Día”;
<collectionCode id="ED72AE21FFE4FFDBFDE2475EBDFA3F55" box="[586,648,1365,1391]" pageId="26" pageNumber="27">GHC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFD08475EBCBC3FA8" ID-GBIF-Occurrence="3093980550" collectedFrom="Selva tropical lluviosa" collectingDate="1969-09-30" collectingMethod="Cebo excremento" collectionCode="VMC" collectorName="G. Halffter &amp; P. Reyes" country="Costa Rica" location="Rincon de Osa." pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFD08475EBDBE3F4A" box="[672,716,1365,1392]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFD4C475EBCE93F55" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFD4C475EBCE93F55" box="[740,923,1365,1391]" country="Costa Rica" name="Rincon de Osa." pageId="26" pageNumber="27" stateProvince="Puntarenas">Rincón de Osa.</location>
<date id="FFDD1024FFE4FFDBFC09475EBB603F55" box="[929,1042,1365,1391]" pageId="26" pageNumber="27" value="1969-09-30">
<collectingDate id="EF99E9CCFFE4FFDBFC09475EBB603F55" box="[929,1042,1365,1391]" pageId="26" pageNumber="27" value="1969-09-30">30-IX-69</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFBB5475EBBEE3F55" box="[1053,1180,1365,1391]" pageId="26" pageNumber="27">G. Halffter</collectorName>
y
<collectorName id="26965332FFE4FFDBFB10475DBA6E3F4A" box="[1208,1308,1366,1392]" pageId="26" pageNumber="27">P. Reyes</collectorName>
C., col.
<collectedFrom id="387F7519FFE4FFDBFF154773BEBA3FA8" box="[189,456,1400,1426]" pageId="26" pageNumber="27">Selva tropical lluviosa</collectedFrom>
.
<collectingMethod id="52224EF3FFE4FFDBFE7A4773BDD63FA9" box="[466,676,1400,1427]" pageId="26" pageNumber="27">Cebo excremento</collectingMethod>
. Día (11–17 hrs.)”;
<collectionCode id="ED72AE21FFE4FFDBFC234773BCBC3FA8" box="[907,974,1400,1426]" pageId="26" pageNumber="27">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFC4F4773BEC93F8F" ID-GBIF-Occurrence="3093980544" collectingDate="1965-05-22" collectionCode="GHC" collectorName="Col. P. Kazan" country="Costa Rica" location="Rincon de Osa." pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<specimenCount id="9D65FD6DFFE4FFDBFC4F4773BB643FA9" box="[999,1046,1400,1427]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE4FFDBFB854773BB963FA8" LSID="urn:lsid:plazi:treatment:03CA87F2FFE7FFDDFDF641E4BA453C24:8EBC603FFFE4FFDBFB854773BB963FA8" box="[1069,1252,1400,1426]" country="Costa Rica" name="Rincon de Osa." pageId="26" pageNumber="27" stateProvince="Puntarenas">Rincón de Osa.</location>
<date id="FFDD1024FFE4FFDBFB434773BA003FA9" box="[1259,1394,1400,1427]" pageId="26" pageNumber="27" value="1965-05-22">
<collectingDate id="EF99E9CCFFE4FFDBFB434773BA003FA9" box="[1259,1394,1400,1427]" pageId="26" pageNumber="27" value="1965-05-22">22–5–1965</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFF154790BE113F8F" box="[189,355,1435,1462]" pageId="26" pageNumber="27">Col. P. Kazan</collectorName>
”;
<collectionCode id="ED72AE21FFE4FFDBFED14790BEC93F8F" box="[377,443,1435,1461]" pageId="26" pageNumber="27">GHC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF1547D7BBB83C23" blockId="26.[189,1398,1499,1562]" pageId="26" pageNumber="27">
<materialsCitation id="3B0B3CB9FFE4FFDBFF1547D7BE633C23" ID-GBIF-Occurrence="3093980584" collectingDate="1982-06-17" collectingDateMax="1982-06-21" collectingDateMin="1982-06-17" collectionCode="TAMU" collectorName="B. Gill." country="Panama" elevation="1200" location="Chiriqui" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui">
<collectingCountry id="F3747674FFE4FFDBFF1547D7BE4A3FCC" box="[189,312,1500,1526]" name="Panama" pageId="26" pageNumber="27">PANAMA</collectingCountry>
–
<emphasis id="B917EAF6FFE4FFDBFEF247D0BEB43FCC" bold="true" box="[346,454,1499,1526]" pageId="26" pageNumber="27">
<collectingRegion id="49A7F806FFE4FFDBFEF247D0BEB43FCC" box="[346,454,1499,1526]" country="Panama" name="Chiriqui" pageId="26" pageNumber="27">Chiriquí</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE4FFDBFE4C47D7BD643FCD" box="[484,534,1500,1527]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<locationDeviation id="9A6E2FC6FFE4FFDBFD8747D7BCA23FCC" box="[559,976,1500,1527]" pageId="26" pageNumber="27">
Cerro Hornito
<quantity id="4C9B9B01FFE4FFDBFD4C47D7BC423FCC" box="[740,816,1500,1527]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="km" value="15.0">15 km</quantity>
NE Gualaca
</locationDeviation>
.
<date id="FFDD1024FFE4FFDBFC7547D7BBE53FCC" box="[989,1175,1500,1527]" pageId="26" pageNumber="27" value="1982-06-17" valueMax="1982-06-21" valueMin="1982-06-17">
<collectingDate id="EF99E9CCFFE4FFDBFC7547D7BBE53FCC" box="[989,1175,1500,1527]" pageId="26" pageNumber="27" value="1982-06-17" valueMax="1982-06-21" valueMin="1982-06-17">17–21 VI 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFB0D47D7BB873FCC" box="[1189,1269,1500,1526]" pageId="26" pageNumber="27">B.Gill.</collectorName>
<quantity id="4C9B9B01FFE4FFDBFB5647D7BA123FCC" box="[1278,1376,1500,1526]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">
<elevation id="004ED1D7FFE4FFDBFB5647D7BA123FCC" box="[1278,1376,1500,1526]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="m" value="1200.0">1200 m</elevation>
</quantity>
.”;
<collectionCode id="ED72AE21FFE4FFDBFF1547F4BE633C23" box="[189,273,1535,1561]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="26" pageNumber="27">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE4FFDBFE8247F4BBB43C23" ID-GBIF-Occurrence="3093980527" box="[298,1222,1535,1562]" collectingDate="1982-08-09" collectingDateMax="1982-08-18" collectingDateMin="1982-08-09" collectionCode="VMC" collectorName="B. Gill." country="Panama" elevation="1500" location="Sta. Clara." pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui">
<specimenCount id="9D65FD6DFFE4FFDBFE8247F4BE2B3C20" box="[298,345,1535,1562]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
; “
<locationDeviation id="9A6E2FC6FFE4FFDBFEC747F4BDAF3C23" box="[367,733,1535,1562]" pageId="26" pageNumber="27">
Cerro Pelota
<quantity id="4C9B9B01FFE4FFDBFDA547F4BD3A3C23" box="[525,584,1535,1561]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="km" value="4.0">4 km</quantity>
N Sta.Clara
</locationDeviation>
.
<date id="FFDD1024FFE4FFDBFD4047F4BCD33C23" box="[744,929,1535,1562]" pageId="26" pageNumber="27" value="1982-08-09" valueMax="1982-08-18" valueMin="1982-08-09">
<collectingDate id="EF99E9CCFFE4FFDBFD4047F4BCD33C23" box="[744,929,1535,1562]" pageId="26" pageNumber="27" value="1982-08-09" valueMax="1982-08-18" valueMin="1982-08-09">9–18 VIII 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE4FFDBFC0447F4BC8E3C23" box="[940,1020,1535,1561]" pageId="26" pageNumber="27">B.Gill.</collectorName>
<quantity id="4C9B9B01FFE4FFDBFBAB47F4BB103C23" box="[1027,1122,1535,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">
<elevation id="004ED1D7FFE4FFDBFBAB47F4BB103C23" box="[1027,1122,1535,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">1500 m</elevation>
</quantity>
.”;
<collectionCode id="ED72AE21FFE4FFDBFBD747F4BBB43C23" box="[1151,1222,1535,1561]" pageId="26" pageNumber="27">VMC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15444BBE2F3C60" blockId="26.[189,349,1600,1626]" box="[189,349,1600,1626]" pageId="26" pageNumber="27">
<heading id="D0948188FFE4FFDBFF15444BBE2F3C60" bold="true" box="[189,349,1600,1626]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B917EAF6FFE4FFDBFF15444BBE2F3C60" bold="true" box="[189,349,1600,1626]" pageId="26" pageNumber="27">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF154464BB6F3CB2" blockId="26.[189,1053,1646,1673]" box="[189,1053,1646,1673]" pageId="26" pageNumber="27">
<collectingCountry id="F3747674FFE4FFDBFF154464BE323CB2" box="[189,320,1646,1673]" name="Costa Rica" pageId="26" pageNumber="27">Costa Rica</collectingCountry>
,
<collectingRegion id="49A7F806FFE4FFDBFEE44464BEA23CB3" box="[332,464,1647,1673]" country="Costa Rica" name="Puntarenas" pageId="26" pageNumber="27">Puntarenas</collectingRegion>
, Carara National Park, Estación Quebrada Bonita.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE4FFDDFF1544A4BBEA39FF" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="8BDC36E4FFE4FFDBFF1544A4BE193CF3" blockId="26.[189,363,1711,1737]" box="[189,363,1711,1737]" pageId="26" pageNumber="27">
<heading id="D0948188FFE4FFDBFF1544A4BE193CF3" bold="true" box="[189,363,1711,1737]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B917EAF6FFE4FFDBFF1544A4BE193CF3" bold="true" box="[189,363,1711,1737]" pageId="26" pageNumber="27">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF4244D5BE0F3CC2" blockId="26.[189,1399,1758,1854]" box="[234,381,1758,1784]" pageId="26" pageNumber="27">
<emphasis id="B917EAF6FFE4FFDBFF4244D5BE0F3CC2" bold="true" box="[234,381,1758,1784]" pageId="26" pageNumber="27">Major male</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15450ABB6C3D04" blockId="26.[189,1399,1758,1854]" pageId="26" pageNumber="27">
HEAD. Completely bright black. Clypeus bidentate, with roughened sculpture. Genae with roughened sculpture. Cephalic horn curved posteriorly over pronotum (
<figureCitation id="13582A61FFE4FFDBFCD7452FBCAF3D04" box="[895,989,1828,1854]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="26" pageNumber="27">Figs 2E</figureCitation>
,
<figureCitation id="13582A61FFE4FFDBFC41452FBB603D04" box="[1001,1042,1828,1854]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="26" pageNumber="27">7A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE4FFDBFF15456DBA043DD2" blockId="26.[187,1399,1893,2025]" pageId="26" pageNumber="27">PRONOTUM. Keel absent in the middle of anterior pronotal margin. Disc triangular, flat; with two weakly developed, elongate tubercles on anterior portion. Triangle bright black, smooth, scabriculous, impunctate. Sides bright black, becoming bright metallic red-green; scabriculous, with smooth sculpture, almost effaced punctures. Lateral lines of triangle straight. Posterolateral angles short, widened. Lateral</paragraph>
<paragraph id="8BDC36E4FFE5FFDAFF154302BCFA3B7C" blockId="27.[189,1398,265,327]" pageId="27" pageNumber="28">
fossae distinctly impressed. Basal fossae distinctly to superficially impressed. Posterior margin with distinctly to superficially impressed punctures (
<figureCitation id="13582A61FFE5FFDAFD404327BC353B7C" box="[744,839,300,326]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="27" pageNumber="28">Figs 2E</figureCitation>
,
<figureCitation id="13582A61FFE5FFDAFCFB4327BC093B7C" box="[851,891,300,326]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="27" pageNumber="28">7A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE5FFDAFF154365BA043BF4" blockId="27.[189,1398,366,463]" pageId="27" pageNumber="28">
ELYTRA. Striae fine, smooth, completely bright black, with superficially impressed to effaced punctures, scabriculous, impressed basally as distinct fossae. Interstriae bright black, with smooth surface, scabriculous, impunctate or with almost effaced punctures. Sutural margin without apical tooth (
<figureCitation id="13582A61FFE5FFDAFAA143BFBA183BF4" box="[1289,1386,436,462]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="27" pageNumber="28">Figs 7A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE5FFDAFF1543FDBDCA3835" blockId="27.[189,696,501,528]" box="[189,696,501,528]" pageId="27" pageNumber="28">PROTIBIAE. Quadridentate with apical spine.</paragraph>
<caption id="DF1C666CFFE5FFDAFF15458EBDBB3DDF" ID-DOI="http://doi.org/10.5281/zenodo.4836774" ID-Zenodo-Dep="4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1925,1951]" targetBox="[219,1380,610,1882]" targetPageId="27">
<paragraph id="8BDC36E4FFE5FFDAFF15458EBDBB3DDF" blockId="27.[187,1399,1925,2022]" pageId="27" pageNumber="28">
<emphasis id="B917EAF6FFE5FFDAFF15458EBE7B3DA5" bold="true" box="[189,265,1925,1951]" pageId="27" pageNumber="28">Fig. 7.</emphasis>
<taxonomicName id="4C634D67FFE5FFDAFEA5458EBD023DA5" authority="Arnaud, 2002" authorityName="Arnaud" authorityYear="2002" box="[269,624,1925,1951]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE5FFDAFEA5458EBEBB3DA5" box="[269,457,1925,1951]" italics="true" pageId="27" pageNumber="28">Phanaeus malyi</emphasis>
Arnaud, 2002
</taxonomicName>
.
<emphasis id="B917EAF6FFE5FFDAFDDE458EBDF93DA5" bold="true" box="[630,651,1925,1951]" pageId="27" pageNumber="28">A</emphasis>
. ♂ (TAMU).
<emphasis id="B917EAF6FFE5FFDAFC84458EBC323DA5" bold="true" box="[812,832,1925,1951]" pageId="27" pageNumber="28">B</emphasis>
. ♀ (TAMU).
<emphasis id="B917EAF6FFE5FFDAFC74458EBC833DA5" bold="true" box="[988,1009,1925,1951]" pageId="27" pageNumber="28">C</emphasis>
. Colombian specimen previously labelled as a paralectotype of
<taxonomicName id="4C634D67FFE5FFDAFD8D45A2BCFF3DF8" authority="Olsoufieff, 1924" authorityName="Olsoufieff" authorityYear="1924" box="[549,909,1960,1986]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="B917EAF6FFE5FFDAFD8D45A2BDB63DF8" box="[549,708,1960,1986]" italics="true" pageId="27" pageNumber="28">P. blanchardi</emphasis>
<bibRefCitation id="EFF24B15FFE5FFDAFD6345A3BCFF3DF8" author="Olsoufieff G. d'" box="[715,909,1960,1986]" pageId="27" pageNumber="28" pagination="4 - 172" refId="ref41334" refString="Olsoufieff G. d'. 1924. Les Phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta, Revue illustree d'Entomologique 13: 4 - 172." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
</taxonomicName>
(by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN).
</paragraph>
</caption>
<paragraph id="8BDC36E4FFE2FFDDFF154302BC653B7F" blockId="28.[189,1398,265,326]" pageId="28" pageNumber="29">TERGITE VIII. Bright metallic red-green, scabriculous sculpture; with rough, superficially impressed punctures. Basal margin with setae variable in size.</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF154367BE573BF1" blockId="28.[188,1398,364,459]" pageId="28" pageNumber="29">
GENITALIA. Right and left lobes of endophallite copulatrix similar in size. Right lobe obtusely triangular in shape, rounded superiorly. Left lobe obtusely lobed. Central ridge less developed than central column (
<figureCitation id="13582A61FFE2FFDDFF6D43BABE673BF1" box="[197,277,433,459]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="28" pageNumber="29">Fig. 1I</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF4243F9BE0F3836" blockId="28.[189,1398,498,594]" box="[234,381,498,524]" pageId="28" pageNumber="29">
<emphasis id="B917EAF6FFE2FFDDFF4243F9BE0F3836" bold="true" box="[234,381,498,524]" pageId="28" pageNumber="29">Minor male</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF15401FBCA03868" blockId="28.[189,1398,498,594]" pageId="28" pageNumber="29">Like the major male, except for the reduction of the secondary sexual characters (i.e., cephalic horn, pronotal triangle and tubercles, and pronotal posterolateral angles).</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF424073BE3738A8" blockId="28.[189,1400,632,865]" box="[234,325,632,658]" pageId="28" pageNumber="29">
<emphasis id="B917EAF6FFE2FFDDFF424073BE3738A8" bold="true" box="[234,325,632,658]" pageId="28" pageNumber="29">Female</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF154091BC3A395B" blockId="28.[189,1400,632,865]" pageId="28" pageNumber="29">
Similar to the male, except for the head showing a cephalic trituberculate carina, with nearly aligned, rounded or carinate, weakly developed tubercles; middle tubercle slightly more developed than lateral tubercles; frons with superficially impressed punctures; pronotum with almost effaced punctures; pronotal process trituberculate, lacking concavity; middle pronotal tubercle more developed, slightly more projected posteriorly than lateral tubercles; all tubercles rounded in shape; posterior pronotal midline completely effaced to almost effaced (
<figureCitation id="13582A61FFE2FFDDFD76414CBC49395B" box="[734,827,839,865]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="28" pageNumber="29">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF42418CBE12399B" blockId="28.[189,1176,903,965]" box="[234,352,903,929]" pageId="28" pageNumber="29">
<emphasis id="B917EAF6FFE2FFDDFF42418CBE12399B" bold="true" box="[234,352,903,929]" pageId="28" pageNumber="29">Variation</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF1541A0BBEA39FF" blockId="28.[189,1176,903,965]" box="[189,1176,938,965]" pageId="28" pageNumber="29">
Mean length
<quantity id="4C9B9B01FFE2FFDDFEF041A1BEB239FF" box="[344,448,938,965]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="28" pageNumber="29" unit="mm" value="17.8">17.8 mm</quantity>
(
<quantity id="4C9B9B01FFE2FFDDFE7841A1BDF339FF" box="[464,641,938,965]" metricMagnitude="-2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="2.05" metricValueMin="1.28" pageId="28" pageNumber="29" unit="mm" value="16.65" valueMax="20.5" valueMin="12.8">12.8–20.5 mm</quantity>
). Colour variation was not found for
<taxonomicName id="4C634D67FFE2FFDDFB9B41A0BBE339FE" authorityName="Arnaud" authorityYear="2002" box="[1075,1169,938,964]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFB9B41A0BBE339FE" box="[1075,1169,938,964]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE2FFDDFF1541E0BCB33F1E" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BDC36E4FFE2FFDDFF1541E0BE243E3F" blockId="28.[189,342,1003,1029]" box="[189,342,1003,1029]" pageId="28" pageNumber="29">
<heading id="D0948188FFE2FFDDFF1541E0BE243E3F" bold="true" box="[189,342,1003,1029]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B917EAF6FFE2FFDDFF1541E0BE243E3F" bold="true" box="[189,342,1003,1029]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF154612BCB33F1E" blockId="28.[188,1399,1049,1317]" pageId="28" pageNumber="29">
Southern Pacific costal area,
<collectingCountry id="F3747674FFE2FFDDFDA54612BDFD3E09" box="[525,655,1049,1075]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
,
<collectingCountry id="F3747674FFE2FFDDFD314612BD843E09" box="[665,758,1049,1075]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
and
<collectingCountry id="F3747674FFE2FFDDFC854612BCD33E09" box="[813,929,1049,1075]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
(
<figureCitation id="13582A61FFE2FFDDFC074612BB453E0E" box="[943,1079,1049,1076]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="51.[189,232,1233,1259]" captionStartId-1="54.[191,234,1247,1273]" captionTargetBox-0="[193,1395,270,1197]" captionTargetBox-1="[193,1393,268,1195]" captionTargetId-0="figure-241@51.[189,1398,265,1199]" captionTargetId-1="figure-243@54.[189,1396,265,1198]" captionTargetPageId-0="51" captionTargetPageId-1="54" captionText-0="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." captionText-1="Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018)." figureDoi-0="http://doi.org/10.5281/zenodo.4836798" figureDoi-1="http://doi.org/10.5281/zenodo.4836800" httpUri-0="https://zenodo.org/record/4836798/files/figure.png" httpUri-1="https://zenodo.org/record/4836800/files/figure.png" pageId="28" pageNumber="29">Figs 16–17</figureCitation>
). The specimens revised by us represent the first accurate records of
<taxonomicName id="4C634D67FFE2FFDDFD3E4637BD813E6F" authorityName="Arnaud" authorityYear="2002" box="[662,755,1083,1109]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFD3E4637BD813E6F" box="[662,755,1083,1109]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
from
<collectingCountry id="F3747674FFE2FFDDFC934637BCEE3E6C" box="[827,924,1084,1110]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
. Apparently, the distributions of
<taxonomicName id="4C634D67FFE2FFDDFAB04637BA043E6F" authorityName="Arnaud" authorityYear="2002" box="[1304,1398,1083,1109]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFAB04637BA043E6F" box="[1304,1398,1083,1109]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D67FFE2FFDDFF594655BEE83E4D" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[241,410,1118,1143]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFE2FFDDFF594655BEE83E4D" box="[241,410,1118,1143]" italics="true" pageId="28" pageNumber="29">P. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE2FFDDFE0B4655BD723E42" box="[419,512,1118,1144]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
show a significant area of sympatry and both species may be collected in the same locality (e.g., Cerro Hornito,
<collectingCountry id="F3747674FFE2FFDDFD20468ABD983EA1" box="[648,746,1153,1179]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
). Nevertheless,
<taxonomicName id="4C634D67FFE2FFDDFC0E468ABB773EA0" authorityName="Arnaud" authorityYear="2002" box="[934,1029,1152,1178]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFC0E468ABB773EA0" box="[934,1029,1152,1178]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D67FFE2FFDDFBE8468ABB9A3EA0" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[1088,1256,1153,1178]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFE2FFDDFBE8468ABB9A3EA0" box="[1088,1256,1153,1178]" italics="true" pageId="28" pageNumber="29">P. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE2FFDDFB47468ABA393EA1" box="[1263,1355,1153,1179]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
are confidently identified by the diagnosis provided herein and putative hybrid specimens were not found by us. The MNHN EC10569 specimen represents the first record for
<taxonomicName id="4C634D67FFE2FFDDFC6546CDBB5E3EE5" authorityName="Arnaud" authorityYear="2002" box="[973,1068,1221,1247]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFC6546CDBB5E3EE5" box="[973,1068,1221,1247]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
in
<collectingCountry id="F3747674FFE2FFDDFBFC46CEBBBE3EE5" box="[1108,1228,1221,1247]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
. Nevertheless, the illegible label data prevents us to provide an accurate locality (
<figureCitation id="13582A61FFE2FFDDFC1146E3BB663F38" box="[953,1044,1256,1282]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="28" pageNumber="29">Fig. 7C</figureCitation>
). The extent of the distribution of
<taxonomicName id="4C634D67FFE2FFDDFF754700BE493F1E" authorityName="Arnaud" authorityYear="2002" box="[221,315,1290,1316]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFF754700BE493F1E" box="[221,315,1290,1316]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
in
<collectingCountry id="F3747674FFE2FFDDFEC84701BEA63F1E" box="[352,468,1290,1316]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
needs to be confirmed by future research.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE2FFDDFF154740BA453C24" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BDC36E4FFE2FFDDFF154740BE5D3F5F" blockId="28.[189,303,1355,1381]" box="[189,303,1355,1381]" pageId="28" pageNumber="29">
<heading id="D0948188FFE2FFDDFF154740BE5D3F5F" bold="true" box="[189,303,1355,1381]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B917EAF6FFE2FFDDFF154740BE5D3F5F" bold="true" box="[189,303,1355,1381]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE2FFDDFF154772BA453C24" blockId="28.[187,1401,1401,1566]" pageId="28" pageNumber="29">
<taxonomicName id="4C634D67FFE2FFDDFF154772BE053FA9" authorityName="Arnaud" authorityYear="2002" box="[189,375,1401,1427]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFF154772BE053FA9" box="[189,375,1401,1427]" italics="true" pageId="28" pageNumber="29">Phanaeus malyi</emphasis>
</taxonomicName>
was considered by previous authors as a synonym of
<taxonomicName id="4C634D67FFE2FFDDFC744771BA003FA9" authority="(Edmonds &amp; Zidek 2012)" baseAuthorityName="Edmonds &amp; Zidek" baseAuthorityYear="2012" box="[988,1394,1401,1427]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE2FFDDFC744771BB323FA9" box="[988,1088,1402,1427]" italics="true" pageId="28" pageNumber="29">P. pyrois</emphasis>
(
<bibRefCitation id="EFF24B15FFE2FFDDFBE64772BA193FA9" author="Edmonds W. D. &amp; Zidek J." box="[1102,1387,1401,1427]" pageId="28" pageNumber="29" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012</bibRefCitation>
)
</taxonomicName>
. Nevertheless, clear differences in the external and genital morphology of
<taxonomicName id="4C634D67FFE2FFDDFB9D4797BBE43F8F" authorityName="Arnaud" authorityYear="2002" box="[1077,1174,1435,1461]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFB9D4797BBE43F8F" box="[1077,1174,1435,1461]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
were found by us. These differences support the full species status suggested by
<bibRefCitation id="EFF24B15FFE2FFDDFC3C47B5BBB63FE2" author="Solis A. &amp; Kohlmann B." box="[916,1220,1470,1496]" pageId="28" pageNumber="29" pagination="1 - 32" refId="ref42322" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1 - 32." type="journal article" year="2012">Solís &amp; Kohlmann (2012)</bibRefCitation>
and
<bibRefCitation id="EFF24B15FFE2FFDDFB5547B5BE233FC0" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." pageId="28" pageNumber="29" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFE2FFDDFF1547EABF823FC0" box="[189,240,1504,1530]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. (2018)
</bibRefCitation>
. When revising the type material housed at MNHM, a Colombian minor male specimen of
<taxonomicName id="4C634D67FFE2FFDDFF15440FBE693C27" authorityName="Arnaud" authorityYear="2002" box="[189,283,1539,1565]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFE2FFDDFF15440FBE693C27" box="[189,283,1539,1565]" italics="true" pageId="28" pageNumber="29">P. malyi</emphasis>
</taxonomicName>
was found to be labelled as a
<typeStatus id="54D88846FFE2FFDDFDD54408BC693C27" box="[637,795,1539,1565]" pageId="28" pageNumber="29" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C634D67FFE2FFDDFCEA440FBCAC3C27" authorityName="Olsoufieff" authorityYear="1924" box="[834,990,1539,1565]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="B917EAF6FFE2FFDDFCEA440FBCAC3C27" box="[834,990,1539,1565]" italics="true" pageId="28" pageNumber="29">P. blanchardi</emphasis>
</taxonomicName>
(
<figureCitation id="13582A61FFE2FFDDFC464408BB3B3C27" box="[1006,1097,1539,1565]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="28" pageNumber="29">Fig. 7C</figureCitation>
, MNHN EC10569).
</paragraph>
</subSubSection>
</treatment>
</document>
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