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<mods:title id="C37AF8ADBCC2D4BB7876026785A02D9A">A review of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) of Taiwan</mods:title>
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<emphasis id="B917EAD7FFB8FF8898808CB2FE43CE82" bold="true" box="[151,510,1158,1184]" pageId="1" pageNumber="522">
Genus
<taxonomicName id="4C634D46FFB8FF8898F98CB2FE43CE82" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[238,510,1158,1184]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898F98CB2FEE5CE82" bold="true" box="[238,344,1158,1184]" italics="true" pageId="1" pageNumber="522">Miridiba</emphasis>
<bibRefCitation id="EFF24B34FFB8FF8899488CB2FE43CE82" author="Reitter" box="[351,510,1158,1184]" pageId="1" pageNumber="522" refString="Reittter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Heft 50. Verhandlungen des Naturforschenden Vereins in Briinn, 40, 93 - 303." type="book chapter" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C379654EFFB8FF8898808CF9FB7FCF5E" pageId="1" pageNumber="522" type="reference_group">
<paragraph id="8BDC36C5FFB8FF8898808CF9FB3BCEC1" blockId="1.[151,1436,1229,1404]" box="[151,1158,1229,1251]" pageId="1" pageNumber="522">
<treatmentCitationGroup id="AB7311EBFFB8FF8898808CF9FB3BCEC1" box="[151,1158,1229,1251]" pageId="1" pageNumber="522">
<taxonomicName id="4C634D46FFB8FF8898808CF9FE0CCEC1" authority="Reitter, 1902: 170" authorityName="Reitter" authorityPageNumber="170" authorityYear="1902" box="[151,433,1229,1251]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898808CF9FF4CCEC1" box="[151,241,1229,1251]" italics="true" pageId="1" pageNumber="522">Miridiba</emphasis>
<treatmentCitation id="0AC210D4FFB8FF8898E08CF9FE0CCEC1" author="Reitter" box="[247,433,1229,1251]" page="170" pageId="1" pageNumber="522" year="1902">
<bibRefCitation id="EFF24B34FFB8FF8898E08CF9FEC3CEC1" author="Reitter" box="[247,382,1229,1251]" pageId="1" pageNumber="522" refString="Reittter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Heft 50. Verhandlungen des Naturforschenden Vereins in Briinn, 40, 93 - 303." type="book chapter" year="1902">Reitter, 1902</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D88867FFB8FF8899AD8CFAFE51CEC1" box="[442,492,1230,1251]" pageId="1" pageNumber="522">Type</typeStatus>
species:
<taxonomicName id="4C634D46FFB8FF889A5E8CF9FC49CEC1" authority="Fairmaire, 1891" authorityName="Fairmaire" authorityYear="1891" box="[585,1012,1229,1251]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="trichophorus">
<emphasis id="B917EAD7FFB8FF889A5E8CF9FCF1CEC1" box="[585,844,1229,1251]" italics="true" pageId="1" pageNumber="522">Rhizotrogus trichophorus</emphasis>
<bibRefCitation id="EFF24B34FFB8FF889B458CF9FC49CEC1" author="Fairmaire" box="[850,1012,1229,1251]" pageId="1" pageNumber="522" refString="Fairmaire, L. (1891) Coleopteres de l'interieur de la Chine (Suite: 7 e partie). Annales de la Societe entomologique de Belgique. Compte Rendus Bruxelles, 35, 187 - 219." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
</taxonomicName>
by monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36C5FFB8FF8898808CD8FCBFCF02" blockId="1.[151,1436,1229,1404]" pageId="1" pageNumber="522">
<treatmentCitationGroup id="AB7311EBFFB8FF8898808CD8FCBFCF02" pageId="1" pageNumber="522">
<taxonomicName id="4C634D46FFB8FF8898808CD8FE57CF20" ID-CoL="8VRQX" authority="Brenske, 1894: 75" authorityName="Brenske" authorityPageNumber="75" authorityYear="1894" box="[151,490,1260,1282]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898808CD8FEA8CF20" box="[151,277,1260,1282]" italics="true" pageId="1" pageNumber="522">Holotrochus</emphasis>
<treatmentCitation id="0AC210D4FFB8FF8899368CD8FE57CF20" author="Brenske" box="[289,490,1260,1282]" page="75" pageId="1" pageNumber="522" year="1894">
<bibRefCitation id="EFF24B34FFB8FF8899368CD8FE03CF20" author="Brenske" box="[289,446,1260,1282]" pageId="1" pageNumber="522" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Museum zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
: 75
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D88867FFB8FF8899EC8CD9FD90CF20" box="[507,557,1261,1282]" pageId="1" pageNumber="522">Type</typeStatus>
species:
<taxonomicName id="4C634D46FFB8FF889A8F8CD8FBAACF20" authority="Brenske, 1894" authorityName="Brenske" authorityYear="1894" box="[664,1047,1260,1282]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B917EAD7FFB8FF889A8F8CD8FCD2CF20" box="[664,879,1260,1282]" italics="true" pageId="1" pageNumber="522">Holotrochus vestitus</emphasis>
<bibRefCitation id="EFF24B34FFB8FF889B6B8CD8FBAACF20" author="Brenske" box="[892,1047,1260,1282]" pageId="1" pageNumber="522" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Museum zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
</taxonomicName>
, by monotypy. Junior homonym of
<taxonomicName id="4C634D46FFB8FF8898D08D3EFE59CF02" ID-CoL="8VRQX" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[199,484,1290,1312]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898D08D3EFEF8CF02" box="[199,325,1290,1312]" italics="true" pageId="1" pageNumber="522">Holotrochus</emphasis>
Erichson, 1840
</taxonomicName>
(
<taxonomicName id="4C634D46FFB8FF8899E68D3EFDDBCF02" ID-CoL="C2L" box="[497,614,1290,1312]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C634D46FFB8FF889A788D3EFD46CF02" ID-CoL="8VR4S" box="[623,763,1290,1312]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36C5FFB8FF8898808D1DFB81CF1D" blockId="1.[151,1436,1229,1404]" box="[151,1084,1321,1343]" pageId="1" pageNumber="522">
<treatmentCitationGroup id="AB7311EBFFB8FF8898808D1DFB81CF1D" box="[151,1084,1321,1343]" pageId="1" pageNumber="522">
<taxonomicName id="4C634D46FFB8FF8898808D1DFE12CF1D" ID-CoL="7PZLF" authority="Lucas, 1920: 592" authorityName="Lucas" authorityPageNumber="592" authorityYear="1920" box="[151,431,1321,1343]" class="Insecta" family="Scarabaeidae" genus="Shangaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898808D1DFF4ACF1D" box="[151,247,1321,1343]" italics="true" pageId="1" pageNumber="522">Shangaia</emphasis>
<treatmentCitation id="0AC210D4FFB8FF8898EA8D1EFE12CF1D" author="Lucas" box="[253,431,1321,1343]" page="592" pageId="1" pageNumber="522" year="1920">
<bibRefCitation id="EFF24B34FFB8FF8898EA8D1EFEC1CF1D" author="Lucas" box="[253,380,1321,1343]" pageId="1" pageNumber="522" refString="Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (jamil., trib., subtr., sect. incl.). Pars 1. Nicolaische Verlags-Buchhandlung R. Stricker, Berlin, 696 pp." type="book" year="1920">Lucas, 1920</bibRefCitation>
: 592
</treatmentCitation>
</taxonomicName>
, new name for the junior homonym
<taxonomicName id="4C634D46FFB8FF889B088D1DFB84CF1D" ID-CoL="8VRQX" authority="Brenske, 1894" authorityName="Brenske" authorityYear="1894" box="[799,1081,1321,1343]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF889B088D1DFC21CF1D" box="[799,924,1321,1343]" italics="true" pageId="1" pageNumber="522">Holotrochus</emphasis>
<bibRefCitation id="EFF24B34FFB8FF889BB48D1DFB84CF1D" author="Brenske" box="[931,1081,1321,1343]" pageId="1" pageNumber="522" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Museum zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36C5FFB8FF8898808D7CFADFCF7C" blockId="1.[151,1436,1229,1404]" box="[151,1378,1352,1374]" pageId="1" pageNumber="522">
<treatmentCitationGroup id="AB7311EBFFB8FF8898808D7CFADFCF7C" box="[151,1378,1352,1374]" pageId="1" pageNumber="522">
<taxonomicName id="4C634D46FFB8FF8898808D7CFE56CF7C" ID-CoL="7PQ25" authority="Arrow, 1948: 50" authorityName="Arrow" authorityPageNumber="50" authorityYear="1948" box="[151,491,1352,1374]" class="Insecta" family="Scarabaeidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898808D7CFE86CF7C" box="[151,315,1352,1374]" italics="true" pageId="1" pageNumber="522">Neodontocnema</emphasis>
<treatmentCitation id="0AC210D4FFB8FF8899558D7CFE56CF7C" author="Arrow" box="[322,491,1352,1374]" page="50" pageId="1" pageNumber="522" year="1948">
<bibRefCitation id="EFF24B34FFB8FF8899558D7CFE78CF7C" author="Arrow" box="[322,453,1352,1374]" pageId="1" pageNumber="522" refString="Arrow, G. J. (1948) Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. Proceedings of the Royal Entomological Society of London (B), 17, 49 - 54. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948</bibRefCitation>
: 50
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D88867FFB8FF8899E38D7DFD9BCF7C" box="[500,550,1353,1374]" pageId="1" pageNumber="522">Type</typeStatus>
species:
<taxonomicName id="4C634D46FFB8FF889A938D7CFC0FCF7C" box="[644,946,1352,1374]" class="Insecta" family="Scarabaeidae" genus="Ancylonycha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="coromandeliana">
<emphasis id="B917EAD7FFB8FF889A938D7CFC0FCF7C" box="[644,946,1352,1374]" italics="true" pageId="1" pageNumber="522">Ancylonycha coromandeliana</emphasis>
</taxonomicName>
B1anchard, 1851, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36C5FFB8FF8898808D52FB7FCF5E" blockId="1.[151,1436,1229,1404]" box="[151,1218,1382,1404]" pageId="1" pageNumber="522">
<treatmentCitationGroup id="AB7311EBFFB8FF8898808D52FB7FCF5E" box="[151,1218,1382,1404]" pageId="1" pageNumber="522">
<taxonomicName id="4C634D46FFB8FF8898808D52FE75CF5E" ID-CoL="7PKFL" authority="Arrow, 1948: 51" authorityName="Arrow" authorityPageNumber="51" authorityYear="1948" box="[151,456,1382,1404]" class="Insecta" family="Scarabaeidae" genus="Hippotrichia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8898808D52FEA7CF5E" box="[151,282,1382,1404]" italics="true" pageId="1" pageNumber="522">Hippotrichia</emphasis>
<treatmentCitation id="0AC210D4FFB8FF8899378D53FE75CF5E" author="Arrow" box="[288,456,1382,1404]" page="51" pageId="1" pageNumber="522" year="1948">
<bibRefCitation id="EFF24B34FFB8FF8899378D53FE1ECF5E" author="Arrow" box="[288,419,1382,1404]" pageId="1" pageNumber="522" refString="Arrow, G. J. (1948) Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. Proceedings of the Royal Entomological Society of London (B), 17, 49 - 54. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948</bibRefCitation>
: 51
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D88867FFB8FF8899C48D53FDB8CF5E" box="[467,517,1383,1404]" pageId="1" pageNumber="522">Type</typeStatus>
species:
<taxonomicName id="4C634D46FFB8FF889A758D52FC7BCF5E" authority="Arrow, 1948" authorityName="Arrow" authorityYear="1948" box="[610,966,1382,1404]" class="Insecta" family="Scarabaeidae" genus="Hippotrichia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="hainana">
<emphasis id="B917EAD7FFB8FF889A758D52FC83CF5E" box="[610,830,1382,1404]" italics="true" pageId="1" pageNumber="522">Hippotrichia hainana</emphasis>
<bibRefCitation id="EFF24B34FFB8FF889B538D53FC7BCF5E" author="Arrow" box="[836,966,1382,1404]" pageId="1" pageNumber="522" refString="Arrow, G. J. (1948) Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. Proceedings of the Royal Entomological Society of London (B), 17, 49 - 54. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948</bibRefCitation>
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C379654EFFB8FF8898808D9DFEDBCD50" pageId="1" pageNumber="522" type="description">
<paragraph id="8BDC36C5FFB8FF8898808D9DFEDBCD50" blockId="1.[151,1437,1449,2014]" pageId="1" pageNumber="522">
<emphasis id="B917EAD7FFB8FF8898808D9DFE38CFE0" bold="true" box="[151,389,1449,1474]" pageId="1" pageNumber="522">Description of male</emphasis>
. Body length 14.0
<quantity id="4C9B9B20FFB8FF889A7E8D9DFD6ECFE3" box="[617,723,1449,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="1" pageNumber="522" unit="mm" value="25.5">25.5 mm</quantity>
. Body oval-elongate; dark reddish brown to yellowish brown; weakly to moderately shiny. Dorsal surface densely punctate, punctures usually deep and rugose; dorsum glabrous or setiferous, setae fine (usually requiring high magnification for observation) or hair-like. Lateral margins of pronotum and elytra with fringe of long, robust setae. Antennae with 9 or 10 antennomeres; antennal club composed of 3 antennomeres, significantly shorter than basal antennomeres. Clypeus transversally narrow; frons flat, base of frons transversally carinate; lateral declivities with or without gibbosity behind middle of pronotum. Scutellum densely punctate. Elytral sutural costa well developed, discal costae absent. Pygidium convex centrally. Protarsi and mesotarsi with a tuft of short, dense setae on lower side of apex of tarsomeres 14. Parameres of male genitalia usually divided into upper and lower parts; inner margins of upper and lower parts with or without fringe of robust, long setae, tip of upper part with or without a beak-like small process.
<emphasis id="B917EAD7FFB8FF889C298ED9FB20CD24" bold="true" box="[1086,1181,1773,1798]" pageId="1" pageNumber="522">Female.</emphasis>
Antennal club shorter than in male and round; apex of clypeus more deeply emarginate at middle than in male; setae on dorsal surface (if any) thinner and shorter than in male; length of metatarsi subequal to metatibia; metatibia and metafemora more stout than in male.
</paragraph>
</subSubSection>
<subSubSection id="C379654EFFB8FF8B98D08F49FDAFCBAA" lastPageId="2" lastPageNumber="523" pageId="1" pageNumber="522" type="diagnosis">
<paragraph id="8BDC36C5FFB8FF8B98D08F49FDAFCBAA" blockId="1.[151,1437,1449,2014]" lastBlockId="2.[151,1436,151,608]" lastPageId="2" lastPageNumber="523" pageId="1" pageNumber="522">
<emphasis id="B917EAD7FFB8FF8898D08F49FEFCCDB4" bold="true" box="[199,321,1917,1942]" pageId="1" pageNumber="522">Diagnosis.</emphasis>
The genus
<taxonomicName id="4C634D46FFB8FF8899DD8F49FD93CDB4" box="[458,558,1917,1942]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF8899DD8F49FD93CDB4" box="[458,558,1917,1942]" italics="true" pageId="1" pageNumber="522">Miridiba</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C634D46FFB8FF889B298F49FC7CCDB4" box="[830,961,1917,1942]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF889B298F49FC7CCDB4" box="[830,961,1917,1942]" italics="true" pageId="1" pageNumber="522">Holotrichia</emphasis>
</taxonomicName>
(
<emphasis id="B917EAD7FFB8FF889BC38F4AFBAECDB7" box="[980,1043,1918,1941]" italics="true" pageId="1" pageNumber="522">sensu</emphasis>
<bibRefCitation id="EFF24B34FFB8FF889C0C8F4AFB69CDB4" author="Nomura" box="[1051,1236,1918,1942]" pageId="1" pageNumber="522" refString="Nomura, S. (1977) On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho, 27, 85 - 109." type="journal article" year="1977">Nomura (1977))</bibRefCitation>
by the following combination of characters: dorsal surface densely punctate (particularly on pronotum, elytra, and scutellum), punctures usually deep and rugose (usually sparsely and unevenly distributed in
<taxonomicName id="4C634D46FFB8FF889C228FF1FB05CDFC" box="[1077,1208,1989,2014]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFB8FF889C228FF1FB05CDFC" box="[1077,1208,1989,2014]" italics="true" pageId="1" pageNumber="522">Holotrichia</emphasis>
</taxonomicName>
); lateral margins of pronotum and elytra with fringe of long, robust setae (lacking in
<taxonomicName id="4C634D46FFBBFF8B9B6388A3FC4ACA92" box="[884,1015,151,176]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9B6388A3FC4ACA92" box="[884,1015,151,176]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
); clypeus transversally narrow; frons flat, base of frons transversally carinate (clypeus widened, frons convex and without carina in
<taxonomicName id="4C634D46FFBBFF8B9CAE8888FA81CAF7" box="[1209,1340,188,213]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9CAE8888FA81CAF7" box="[1209,1340,188,213]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
); elytral sutural costa well developed, discal costae absent (discal costae well developed in
<taxonomicName id="4C634D46FFBBFF8B9C6588EBFB48CADA" box="[1138,1269,223,248]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9C6588EBFB48CADA" box="[1138,1269,223,248]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
); protarsi and mesotarsi with a tuft of short, dense setae on lower side of apex of tarsomeres 14 (lacking in
<taxonomicName id="4C634D46FFBBFF8B9D1F8930FA36CB3F" box="[1288,1419,260,285]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9D1F8930FA36CB3F" box="[1288,1419,260,285]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
); parameres of male genitalia elongate (usually capsular in shape in
<taxonomicName id="4C634D46FFBBFF8B9B898913FB9CCB62" box="[926,1057,295,320]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9B898913FB9CCB62" box="[926,1057,295,320]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
), usually divided into upper and lower parts, inner margins of upper and lower parts with fringe of robust, long setae (not divided and lacking robust, long setae in
<taxonomicName id="4C634D46FFBBFF8B9968895BFDBFCBAA" box="[383,514,367,392]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B9968895BFDBFCBAA" box="[383,514,367,392]" italics="true" pageId="2" pageNumber="523">Holotrichia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C379654EFFBBFF8B98D089A0FB56CBF2" pageId="2" pageNumber="523" type="distribution">
<paragraph id="8BDC36C5FFBBFF8B98D089A0FB56CBF2" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="523">
<emphasis id="B917EAD7FFBBFF8B98D089A0FEE4CB8F" bold="true" box="[199,345,404,429]" pageId="2" pageNumber="523">Distribution</emphasis>
. Eastern to southwestern
<collectingCountry id="F3747655FFBBFF8B9A8289A0FD66CB8F" box="[661,731,404,429]" name="China" pageId="2" pageNumber="523">China</collectingCountry>
, Indochina, Sumatra, Java, Indian Subcontinent, Himalaya, Korean Peninsula,
<collectingCountry id="F3747655FFBBFF8B997C898CFE13CBED" box="[363,430,440,463]" name="Japan" pageId="2" pageNumber="523">Japan</collectingCountry>
, Ryukyu Archipelago (islands of Iriomotejima and Ishigakijima),
<collectingCountry id="F3747655FFBBFF8B9C868983FB5ACBF2" box="[1169,1255,439,464]" name="Taiwan" pageId="2" pageNumber="523">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C379654EFFBBFF8B98D089E8FE41C842" pageId="2" pageNumber="523" type="discussion">
<paragraph id="8BDC36C5FFBBFF8B98D089E8FE41C842" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="523">
<emphasis id="B917EAD7FFBBFF8B98D089E8FE86CBD7" bold="true" box="[199,315,476,501]" pageId="2" pageNumber="523">Remarks.</emphasis>
Adults of
<taxonomicName id="4C634D46FFBBFF8B99AE89E8FDA0CBD7" box="[441,541,476,501]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B99AE89E8FDA0CBD7" box="[441,541,476,501]" italics="true" pageId="2" pageNumber="523">Miridiba</emphasis>
</taxonomicName>
are active nocturnal and strongly or weakly attracted by lights.
<bibRefCitation id="EFF24B34FFBBFF8B9D1789E8FA21CBD6" author="Zhang" box="[1280,1436,476,501]" pageId="2" pageNumber="523" refString="Zhang, Z. - L. (1984) Economic Insect Fauna of China. Vol. 28. Coleoptera: Larvae of Scarabaeoidea. Science and Technology Press, Beijing, 107 pp. [in Chinese]" type="book" year="1984">Zhang (1984)</bibRefCitation>
reported that adults of
<taxonomicName id="4C634D46FFBBFF8B998389CBFE45C83A" box="[404,504,511,536]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAD7FFBBFF8B998389CBFE45C83A" box="[404,504,511,536]" italics="true" pageId="2" pageNumber="523">Miridiba</emphasis>
</taxonomicName>
do not feed and he also described four species larvae from
<collectingCountry id="F3747655FFBBFF8B9C8589CBFB65C83A" box="[1170,1240,511,536]" name="China" pageId="2" pageNumber="523">China</collectingCountry>
, which have oneyear life cycle and sometimes caused severe damages to corn, sweet potato, legumes, and peanuts by feeding on underground parts of the plants.
</paragraph>
</subSubSection>
</treatment>
</document>