669 lines
108 KiB
XML
669 lines
108 KiB
XML
<document id="F8A325E124C0E099197FEAA7F80FFA51" ID-CLB-Dataset="20048" ID-DOI="10.11646/zootaxa.5120.4.7" ID-GBIF-Dataset="886176b1-83b0-4af0-9c87-fed4c1f2060e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6392944" ID-ZooBank="8EB14962-1EC2-49BA-AB42-8627E7571301" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648538407559" checkinUser="plazi" docAuthor="Onishko, Vladimir V. & Kosterin, Oleg E." docDate="2022" docId="03CA87C468338B1A73E3FB2250B24AC3" docLanguage="en" docName="zootaxa.5120.4.7.pdf" docOrigin="Zootaxa 5120 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ischnura elegans subsp. malikovae Onishko & Kosterin 2022, subspecies nova" docType="treatment" docVersion="8" lastPageNumber="579" masterDocId="FFF3FFBC68328B1C7374FF88522B4D73" masterDocTitle="Ischnura elegans malikovae subspecies nova (Odonata, Coenagrionidae) from the Far East of Eurasia, with discussion of other possible subspecies" masterLastPageNumber="585" masterPageNumber="573" pageNumber="574" updateTime="1699318824062" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="2505143217813275A149EA6E6FF94BAC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="57497924F2C0E75BE2A445336E232E08">
|
||
<mods:title id="30A0325182C87B2738926CD231789E4E">Ischnura elegans malikovae subspecies nova (Odonata, Coenagrionidae) from the Far East of Eurasia, with discussion of other possible subspecies</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="EFB621FD95F8ECE42A0F9175765DAF64" type="personal">
|
||
<mods:role id="1721C44B3BBF67CACF1A879C55677244">
|
||
<mods:roleTerm id="D0B74AA09AC2B9E6E4FF27CFE6E8BF5E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0122B78952854B6AFB9AAE0352B398BA">Onishko, Vladimir V.</mods:namePart>
|
||
<mods:nameIdentifier id="2D98A6C6D51BFCC3F759A3DFAFD083BD" type="ORCID">0000-0002-6469-6778</mods:nameIdentifier>
|
||
<mods:affiliation id="0BA2864E65BE2CE3FE374E691C198883">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia. https: // orcid. org / 0000 - 0002 - 6469 - 6778</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="D13B066B71532C67C8096CC3BFE37C32" type="personal">
|
||
<mods:role id="21CE7042C63C8D8E34B753060631DAB0">
|
||
<mods:roleTerm id="C2CF6369C8FA1412FD5211692E21BBAB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="18EE34592378836E4F20652A7E7A5DDA">Kosterin, Oleg E.</mods:namePart>
|
||
<mods:affiliation id="CD32C6E795192D1B21FCA600D0A65ADA">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DF72490979C6EBC825BBC00BDC66D525">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D9F2A39756A82BA534668F72AA1BD8EF" type="host">
|
||
<mods:titleInfo id="D029CFD13B4DBB355178A036D570B7F5">
|
||
<mods:title id="50F181CC0E4C634B607D69F3D6AF5F85">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7A99C11EECC9B022E7B49C805D318751">
|
||
<mods:date id="EDB11A649C55E8B76598F7AEE18D1EC7">2022</mods:date>
|
||
<mods:detail id="71780D79AF0DECCBFA6C75A0FA828E07" type="pubDate">
|
||
<mods:number id="BB423D806203953B69384478ABDA36C2">2022-03-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F04FB10E6E473DA1BDC9294BDA942076" type="volume">
|
||
<mods:number id="27D4221CFD14515137A86BA24E38E2F4">5120</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="693EF68BB00E1132741655F22B18B116" type="issue">
|
||
<mods:number id="60ECFB6D36874B153386ACD99DCEDD09">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4BE2664B392F6671811C3C0884634EAC" unit="page">
|
||
<mods:start id="693DF1217DB6FDB47530BD1D3DC92BCA">573</mods:start>
|
||
<mods:end id="96621ED9043F368945DBE9E66C9F613E">585</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="992CD76388A52ECDDB0E80C96674D34A">journal article</mods:classification>
|
||
<mods:identifier id="01A293C841F31F0EA5AF5F3E3CF6BAAB" type="CLB-Dataset">20048</mods:identifier>
|
||
<mods:identifier id="7AEB43C4F29A46C40BCA9BB1CAC31B37" type="DOI">10.11646/zootaxa.5120.4.7</mods:identifier>
|
||
<mods:identifier id="D991EFDD5C3851B355D1B521F1C75004" type="GBIF-Dataset">886176b1-83b0-4af0-9c87-fed4c1f2060e</mods:identifier>
|
||
<mods:identifier id="3342DBEBF1FFA910AB65845845686071" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="D2A6979035150E6C5842979DAEDD3093" type="Zenodo-Dep">6392944</mods:identifier>
|
||
<mods:identifier id="E7FAFB1F7E7BF1734431072F3674A6CA" type="ZooBank">8EB14962-1EC2-49BA-AB42-8627E7571301</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CA87C468338B1A73E3FB2250B24AC3" ID-DOI="http://doi.org/10.5281/zenodo.6402209" ID-GBIF-Taxon="194317688" ID-Zenodo-Dep="6402209" LSID="urn:lsid:plazi:treatment:03CA87C468338B1A73E3FB2250B24AC3" httpUri="http://treatment.plazi.org/id/03CA87C468338B1A73E3FB2250B24AC3" lastPageId="6" lastPageNumber="579" pageId="1" pageNumber="574">
|
||
<subSubSection id="C379655968338B1D73E3FB22509249B6" box="[151,697,1194,1221]" pageId="1" pageNumber="574" type="nomenclature">
|
||
<paragraph id="8BDC36D268338B1D73E3FB22509249B6" blockId="1.[151,697,1194,1257]" box="[151,697,1194,1221]" pageId="1" pageNumber="574">
|
||
<heading id="D09481BE68338B1D73E3FB22509249B6" bold="true" box="[151,697,1194,1221]" fontSize="11" level="1" pageId="1" pageNumber="574" reason="1">
|
||
<emphasis id="B917EAC068338B1D73E3FB22509249B6" bold="true" box="[151,697,1194,1221]" pageId="1" pageNumber="574">
|
||
<taxonomicName id="4C634D5168338B1D73E3FB2253C749B6" authority="Onishko & Kosterin, 2022" authorityName="Onishko & Kosterin" authorityYear="2022" box="[151,492,1194,1221]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="574" phylum="Arthropoda" rank="subSpecies" species="elegans" status="subsp. nov." subSpecies="malikovae">
|
||
<emphasis id="B917EAC068338B1D73E3FB2253C749B6" bold="true" box="[151,492,1194,1221]" italics="true" pageId="1" pageNumber="574">Ischnura elegans malikovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457BB68338B1D7287FB22509249B6" box="[499,697,1194,1221]" pageId="1" pageNumber="574" rank="subSpecies">subspecies nova</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968338B1D73E3FB475364499A" box="[151,335,1231,1257]" pageId="1" pageNumber="574" type="description">
|
||
<paragraph id="8BDC36D268338B1D73E3FB475364499A" blockId="1.[151,697,1194,1257]" box="[151,335,1231,1257]" pageId="1" pageNumber="574">
|
||
(
|
||
<figureCitation id="13582A5768338B1D73EBFB47532F499A" box="[159,260,1231,1257]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1943,1967]" captionStartId-1="5.[151,250,1943,1967]" captionTargetBox-0="[389,1199,181,1920]" captionTargetBox-1="[394,1193,181,1897]" captionTargetId-0="figure-21@4.[389,1199,181,1920]" captionTargetId-1="figure-17@5.[394,1193,181,1920]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." captionText-1="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6392946" figureDoi-1="http://doi.org/10.5281/zenodo.6392948" httpUri-0="https://zenodo.org/record/6392946/files/figure.png" httpUri-1="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">Figs 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D7264FB475306499A" box="[272,301,1231,1257]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="1" pageNumber="574">3a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D724CFB47536C499A" box="[312,327,1231,1257]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="1" pageNumber="574">4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968338B1D73E3FA9F51ED4B7A" pageId="1" pageNumber="574" type="materials_examined">
|
||
<paragraph id="8BDC36D268338B1D73E3FA9F56D248B2" blockId="1.[151,1437,1302,2013]" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73E3FA9F53634843" bold="true" box="[151,328,1302,1329]" pageId="1" pageNumber="574">Type material.</emphasis>
|
||
<materialsCitation id="3B0B3C8F68338B1D7226FA9F5355480A" ID-GBIF-Occurrence="3744208302" collectingDate="2020-07-28" collectionCode="V, RMNH" collectorName="Onishko" country="Russia" elevation="68" latitude="46.4625" location="Luchegorsk Reservoir" longLatPrecision="6" longitude="134.3009" municipality="Luchegorsk Town" pageId="1" pageNumber="574" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
|
||
<emphasis id="B917EAC068338B1D7226FA9F53E34842" bold="true" box="[338,456,1303,1329]" pageId="1" pageNumber="574">
|
||
<typeStatus id="54D8887068338B1D7226FA9F53E84842" box="[338,451,1303,1329]" pageId="1" pageNumber="574" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D65FD5B68338B1D72A6FA9F53CC4842" box="[466,487,1303,1329]" pageId="1" pageNumber="574" type="male">♂</specimenCount>
|
||
(
|
||
<figureCitation id="13582A5768338B1D728DFA9F507B4842" box="[505,592,1303,1329]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="1" pageNumber="574">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D7114FA9F505B4842" box="[608,624,1303,1329]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">2</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D710BFA9F50B74842" box="[639,668,1303,1329]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="1" pageNumber="574">3a</figureCitation>
|
||
),
|
||
<collectingCountry id="F374764268338B1D71C0FA9F512A4842" box="[692,769,1303,1329]" name="Russia" pageId="1" pageNumber="574">Russia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A7F83068338B1D707BFA9F51E44842" box="[783,975,1303,1329]" country="Russia" name="Primorskiy" pageId="1" pageNumber="574">Primorskiy Kray</collectingRegion>
|
||
, Pozharskiy District,
|
||
<collectingMunicipality id="6BB8ACA868338B1D77BCFA9F57B34842" box="[1224,1432,1303,1329]" pageId="1" pageNumber="574">Luchegorsk Town</collectingMunicipality>
|
||
, the
|
||
<location id="8EBC600968338B1D73B6FAB353974826" LSID="urn:lsid:plazi:treatment:03CA87C468338B1A73E3FB2250B24AC3:8EBC600968338B1D73B6FAB353974826" box="[194,444,1339,1365]" country="Russia" latitude="46.4625" longLatPrecision="6" longitude="134.3009" municipality="Luchegorsk Town" name="Luchegorsk Reservoir" pageId="1" pageNumber="574" stateProvince="Primorskiy">Luchegorsk Reservoir</location>
|
||
bank at the outlet,
|
||
<geoCoordinate id="EE57501568338B1D71EFFAB2510A4826" box="[667,801,1338,1365]" degrees="46.4625" direction="north" orientation="latitude" pageId="1" pageNumber="574" precision="5" value="46.4625">46.4625° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE57501568338B1D7059FAB351964826" box="[813,957,1339,1365]" degrees="134.3009" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="5" value="134.3009">134.3009° E</geoCoordinate>
|
||
,
|
||
<elevation id="004ED1E168338B1D70BFFAB3566F4826" box="[971,1092,1339,1365]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="574" unit="m" value="68.0">
|
||
<quantity id="4C9B9B3768338B1D70BFFAB3562D4826" box="[971,1030,1339,1365]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="574" unit="m" value="68.0">68 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFDD101268338B1D7726FAB356F34826" box="[1106,1240,1339,1365]" pageId="1" pageNumber="574" value="2020-07-28">
|
||
<collectingDate id="EF99E9FA68338B1D7726FAB356F34826" box="[1106,1240,1339,1365]" pageId="1" pageNumber="574" value="2020-07-28">28 vii 2020</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED72AE1768338B1D7790FAB356D34826" box="[1252,1272,1339,1365]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="574" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="2696530468338B1D7670FAB357484826" box="[1284,1379,1339,1365]" pageId="1" pageNumber="574">Onishko</collectorName>
|
||
leg., deposited in
|
||
<collectionCode id="ED72AE1768338B1D7252FAD75355480A" box="[294,382,1375,1401]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="574" type="Museum">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0B3C8F68338B1D72F3FAD75156480A" ID-GBIF-Occurrence="3744208303" box="[391,893,1375,1401]" collectingDate="2020-07-28" collectionCode="V, RMNH" collectorName="Onishko" country="Russia" elevation="68" latitude="46.4625" location="Luchegorsk Reservoir" longLatPrecision="6" longitude="134.3009" municipality="Luchegorsk Town" pageId="1" pageNumber="574" specimenCount="2" specimenCount-female="2" stateProvince="Primorskiy" typeStatus="paratype">
|
||
<emphasis id="B917EAC068338B1D72F3FAD75022480A" bold="true" box="[391,521,1375,1401]" pageId="1" pageNumber="574">
|
||
<typeStatus id="54D8887068338B1D72F3FAD75028480A" box="[391,515,1375,1401]" pageId="1" pageNumber="574" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D65FD5B68338B1D717BFAD75065480A" box="[527,590,1375,1401]" pageId="1" pageNumber="574" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<figureCitation id="13582A5768338B1D7129FAD7509B480A" box="[605,688,1375,1401]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="1" pageNumber="574">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D71C9FAD750E7480A" box="[701,716,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="1" pageNumber="574">3</figureCitation>
|
||
), the same data
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0B3C8F68338B1D70FCFAD756DE48B2" ID-GBIF-Occurrence="3744208301" collectingDate="2020-07-25" collectionCode="V, RMNH" collectorName="Onishko & O. Kosterin" country="Russia" elevation="105" latitude="48.2226" location="Korfovskiy Settlement" longLatPrecision="6" longitude="135.0632" pageId="1" pageNumber="574" specimenCount="4" specimenCount-male="4" stateProvince="Khabarovsk" typeStatus="paratype">
|
||
<specimenCount id="9D65FD5B68338B1D70FCFAD751E2480A" box="[904,969,1375,1401]" pageId="1" pageNumber="574" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<collectingCountry id="F374764268338B1D70A1FAD75609480A" box="[981,1058,1375,1401]" name="Russia" pageId="1" pageNumber="574">Russia</collectingCountry>
|
||
, Khabarovskiy Kray,
|
||
<collectingRegion id="49A7F83068338B1D7661FAD752C548EE" country="Russia" name="Khabarovsk" pageId="1" pageNumber="574">Khabarovsk District</collectingRegion>
|
||
, the pond in
|
||
<location id="8EBC600968338B1D720AFA0B50AB48EE" LSID="urn:lsid:plazi:treatment:03CA87C468338B1A73E3FB2250B24AC3:8EBC600968338B1D720AFA0B50AB48EE" box="[382,640,1411,1437]" country="Russia" latitude="48.2226" longLatPrecision="6" longitude="135.0632" name="Korfovskiy Settlement" pageId="1" pageNumber="574" stateProvince="Khabarovsk">Korfovskiy Settlement</location>
|
||
, 48.2226–
|
||
<geoCoordinate id="EE57501568338B1D7181FA0B515C48EE" box="[757,887,1411,1437]" degrees="48.2240" direction="west" orientation="longitude" pageId="1" pageNumber="574" precision="5" value="-48.224">48.2240° N</geoCoordinate>
|
||
, 135.0619–
|
||
<geoCoordinate id="EE57501568338B1D708FFA0A56A348EE" box="[1019,1160,1410,1437]" degrees="135.0632" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="5" value="135.0632">135.0632° E</geoCoordinate>
|
||
,
|
||
<elevation id="004ED1E168338B1D77E0FA0B576448EE" box="[1172,1359,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.06" metricValueMin="1.04" pageId="1" pageNumber="574" unit="m" value="105.0" valueMax="106.0" valueMin="104.0">
|
||
<quantity id="4C9B9B3768338B1D77E0FA0B573848EE" box="[1172,1299,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.06" metricValueMin="1.04" pageId="1" pageNumber="574" unit="m" value="105.0" valueMax="106.0" valueMin="104.0">104–106 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFDD101268338B1D7628FA0A52F948B2" pageId="1" pageNumber="574" value="2020-07-25">
|
||
<collectingDate id="EF99E9FA68338B1D7628FA0A52F948B2" pageId="1" pageNumber="574" value="2020-07-25">25 vii 2020</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED72AE1768338B1D73A9FA2F52DA48B2" box="[221,241,1447,1473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="574" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="2696530468338B1D738FFA2F537148B2" box="[251,346,1447,1473]" pageId="1" pageNumber="574">Onishko</collectorName>
|
||
and
|
||
<collectorName id="2696530468338B1D72E5FA2F503848B2" box="[401,531,1447,1473]" pageId="1" pageNumber="574">O. Kosterin</collectorName>
|
||
leg.; deposited in
|
||
<collectionCode id="ED72AE1768338B1D7195FA2F511E48B2" box="[737,821,1447,1473]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="574" type="Museum">RMNH</collectionCode>
|
||
and the collection by the second author
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268338B1D73B3FA4251ED4B7A" blockId="1.[151,1437,1302,2013]" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73B3FA4253A74896" bold="true" box="[199,396,1482,1509]" pageId="1" pageNumber="574">Additional data:</emphasis>
|
||
<materialsCitation id="3B0B3C8F68338B1D72E7FA4351E94B7A" ID-GBIF-Occurrence="3744208304" collectingDate="2020-07-31" country="Russia" elevation="83" latitude="44.513" location="Prokhory village" longLatPrecision="67" longitude="132.696" municipality="Spasskiy District" pageId="1" pageNumber="574" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
|
||
<specimenCount id="9D65FD5B68338B1D72E7FA4353834896" box="[403,424,1483,1509]" pageId="1" pageNumber="574" type="male">♂</specimenCount>
|
||
photographed,
|
||
<collectingCountry id="F374764268338B1D7122FA4350884896" box="[598,675,1483,1509]" name="Russia" pageId="1" pageNumber="574">Russia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A7F83068338B1D71DBFA4351404896" box="[687,875,1483,1509]" country="Russia" name="Primorskiy" pageId="1" pageNumber="574">Primorskiy Kray</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BB8ACA868338B1D7002FA43561C4896" box="[886,1079,1483,1509]" pageId="1" pageNumber="574">Spasskiy District</collectingMunicipality>
|
||
,
|
||
<location id="8EBC600968338B1D7736FA43572B4896" LSID="urn:lsid:plazi:treatment:03CA87C468338B1A73E3FB2250B24AC3:8EBC600968338B1D7736FA43572B4896" box="[1090,1280,1483,1509]" country="Russia" latitude="44.513" longLatPrecision="67" longitude="132.696" municipality="Spasskiy District" name="Prokhory village" pageId="1" pageNumber="574" stateProvince="Primorskiy">Prokhory village</location>
|
||
, the bank of a big reservoir at the dam,
|
||
<geoCoordinate id="EE57501568338B1D72DBFA66500E4B7A" box="[431,549,1518,1545]" degrees="44.513" direction="north" orientation="latitude" pageId="1" pageNumber="574" precision="55" value="44.513">44.513° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE57501568338B1D7145FA67509A4B7A" box="[561,689,1519,1545]" degrees="132.696" direction="east" orientation="longitude" pageId="1" pageNumber="574" precision="55" value="132.696">132.696° E</geoCoordinate>
|
||
,
|
||
<elevation id="004ED1E168338B1D71C9FA6751194B7A" box="[701,818,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="574" unit="m" value="83.0">
|
||
<quantity id="4C9B9B3768338B1D71C9FA6750DD4B7A" box="[701,758,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="574" unit="m" value="83.0">83 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFDD101268338B1D704BFA6751E94B7A" box="[831,962,1519,1545]" pageId="1" pageNumber="574" value="2020-07-31">
|
||
<collectingDate id="EF99E9FA68338B1D704BFA6751E94B7A" box="[831,962,1519,1545]" pageId="1" pageNumber="574" value="2020-07-31">31 vii 2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968338B1D73B3F99B56B44B22" pageId="1" pageNumber="574" type="etymology">
|
||
<paragraph id="8BDC36D268338B1D73B3F99B56B44B22" blockId="1.[151,1437,1302,2013]" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73B3F99B53654B5E" bold="true" box="[199,334,1555,1581]" pageId="1" pageNumber="574">Etymology.</emphasis>
|
||
The subspecies is named after Dr. Elena Ivanovna Malikova, a prominent expert of
|
||
<taxonomicName id="4C634D5168338B1D7783F99B577D4B5E" authorityName="Fabricius" authorityYear="1793" box="[1271,1366,1555,1581]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="1" pageNumber="574" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
|
||
of the Far East of
|
||
<collectingCountry id="F374764268338B1D726DF9BF534D4B22" box="[281,358,1591,1617]" name="Russia" pageId="1" pageNumber="574">Russia</collectingCountry>
|
||
, who for the first time reported this taxon (as
|
||
<taxonomicName id="4C634D5168338B1D7013F9BF51FE4B22" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[871,981,1591,1617]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="574" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068338B1D7013F9BF51FE4B22" box="[871,981,1591,1617]" italics="true" pageId="1" pageNumber="574">I. elegans</emphasis>
|
||
</taxonomicName>
|
||
) for that territory.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968338B1E73B3F9D356B34B06" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574" type="materials_examined">
|
||
<paragraph id="8BDC36D268338B1D73B3F9D3510F4BEA" blockId="1.[151,1437,1302,2013]" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73B3F9D353104B06" bold="true" box="[199,315,1627,1653]" pageId="1" pageNumber="574">
|
||
<typeStatus id="54D8887068338B1D73B3F9D3531D4B06" box="[199,310,1627,1653]" pageId="1" pageNumber="574" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Mostly bicolorous blue and black, but head with pale greenish ground colour and the ventral part of tergites in the middle part of abdomen yellowish (
|
||
<figureCitation id="13582A5768338B1D71B2F9F7513E4BEA" box="[710,789,1663,1689]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="1" pageNumber="574">Fig. 1a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268338B1D73B3F92B501B4A3E" blockId="1.[151,1437,1302,2013]" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73B3F92B532F4BCE" box="[199,260,1699,1725]" italics="true" pageId="1" pageNumber="574">Head</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768338B1D7262F92B53464BCD" box="[278,365,1699,1726]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">Fig. 2b</figureCitation>
|
||
). Labium pale greenish. Labrum pale green with a black stripe along base with a gentle and rounded central projection. Mandibles greenish. Anteclypeus greenish. Postclypeus black. Antefrons largely pale green; postfrons black but black colour somewhat extends from the former to the latter between antennae, its border uneven. Vertex and occiput black, with two large nearly triangular (with rounded corners) azure blue postocular spots. Rear head surface pale bluish.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268338B1E73B3F8DF51D44CDE" blockId="1.[151,1437,1302,2013]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574">
|
||
<emphasis id="B917EAC068338B1D73B3F8DF53334A02" box="[199,280,1879,1905]" italics="true" pageId="1" pageNumber="574">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768338B1D725CF8DF53564A02" box="[296,381,1879,1905]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="1" pageNumber="574">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768338B1D72FFF8DF53ED4A01" box="[395,454,1879,1906]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">2a–b</figureCitation>
|
||
): Most of anterior lobe of prothorax (
|
||
<figureCitation id="13582A5768338B1D7004F8DF51F64A01" box="[880,989,1879,1906]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">Fig. 2d–e</figureCitation>
|
||
) occupied by a broad azure blue stripe. Median prothorax lobe black above, azure blue below, with black on either side making a projection at its hind margin (
|
||
<figureCitation id="13582A5768338B1D7382F81753604AC9" box="[246,331,1951,1978]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">Fig. 2d</figureCitation>
|
||
). Posterior lobe azure blue laterally, black below; in dorsal view (
|
||
<figureCitation id="13582A5768338B1D7741F81756AD4ACA" box="[1077,1158,1951,1977]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="1" pageNumber="574">Fig. 2e</figureCitation>
|
||
) has a bluntly triangular margin overlaid by a strong process protruding posteriorly. This process has a longitudinally concave surface, straight, slightly converging margins and a rounded tip bearing a large pale-blue spot with brownish margins (
|
||
<figureCitation id="13582A5768308B1E761BFF1F529E4DA6" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">Fig. 2e</figureCitation>
|
||
). In lateral view (
|
||
<figureCitation id="13582A5768308B1E72F0FF3353F34DA5" box="[388,472,187,214]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">Fig. 2d</figureCitation>
|
||
) posterior lobe fore margin convex and hind margin concave, process attenuated in dorso-caudal direction. Mesostigmal plate (
|
||
<figureCitation id="13582A5768308B1E71F1FF5750FD4D8A" box="[645,726,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">Fig. 2e</figureCitation>
|
||
) rather simple, its margin forming two gentle waves on either side. Either side of mesostigmal plate with azure blue outer half (but suture remains black), these blue stripes intruding into otherwise black pterothorax dorsum (
|
||
<figureCitation id="13582A5768308B1E71A9FEAF511B4C32" box="[733,816,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">Fig. 2e</figureCitation>
|
||
). Antehumeral stripes well developed, azure blue (
|
||
<figureCitation id="13582A5768308B1E761FFEAF52984C16" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="2" pageNumber="575">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768308B1E73B4FEC352F54C16" box="[192,222,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">2a</figureCitation>
|
||
). Black humeral stripes tapering posteriorly. Sides of pterothorax (including entire mesinfraepisternum) and coxae azure blue, with short black streak at tops of interpleural and metapleural sutures, the former very narrow, the latter expanding as a rounded diamond. (
|
||
<figureCitation id="13582A5768308B1E7114FE1B509F4CDE" box="[608,692,403,429]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="2" pageNumber="575">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768308B1E71B5FE1B50F44CDE" box="[705,735,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">2a</figureCitation>
|
||
). Poststernum azure blue.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268308B1E73B3FE3F50A34F6A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="575">
|
||
Legs (
|
||
<figureCitation id="13582A5768308B1E7279FE3F53494CA2" box="[269,354,439,465]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="2" pageNumber="575">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768308B1E721BFE3F53A64CA2" box="[367,397,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">2a</figureCitation>
|
||
): ground colour of femora pale bluish, of tibia and tarsi yellowish-white; femora with broad black stripes along dorsal side and tibia with black stripes along ventral and posterior sides; tarsal segments pale with dark apices; all spines and claws black.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268308B1E73B3FDAB519B4F82" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="575">
|
||
Wings hyaline; main longitudinal veins yellowish, narrow veins blackish. Ten postnodals on fore wing, eight on hind wing. Pterostigmata diamond-shaped, with slightly convex costal side, those on hindwing slightly smaller. Fore wing pterostigma tricolorous above—inner half black, outer half blue but becomes white at apex, and bicolorous below—inner ca 2/3 black, outer 1/3 pale blue; black parts with narrow paler rims; bordering veins blackish at its dark part but become yellowish at pale part. Hind wing pterostigma dark-grey inside, with whitish margins at bordering veins but blue at outer tip; bordering veins as above (
|
||
<figureCitation id="13582A5768308B1E702AFD5F51B44F82" box="[862,927,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="2" pageNumber="575">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268308B1E73B3FD7356C7487E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="575">
|
||
<emphasis id="B917EAC068308B1E73B3FD73531A4E66" box="[199,305,763,789]" italics="true" pageId="2" pageNumber="575">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768308B1E724BFD7353B94E66" box="[319,402,763,789]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="2" pageNumber="575">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768308B1E72EAFD7353914E66" box="[414,442,763,789]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[394,1193,181,1897]" captionTargetId="figure-17@5.[394,1193,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Morphological details of the holotype male, of Ischnura elegans malikovae: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="2" pageNumber="575">2c</figureCitation>
|
||
): S1 azure blue at sides, black dorsally. S2 azure blue with a large black wine-cup-shaped black dorsal spot, with a ‘base’ as dorsal trasnversal stripe at hind margin of segment. S2–S5 pale with broad dorsal black stripes, indented from anterior margin of segment, convex before their posterior margins and contacting to black stripes along the latter; ground colour changing from azure blue to bluish-yellow on S3, yellowish on S3–4 and bluish-yellow on S5. S6 ground colour azure blue, black dorsal spot broader but near posterior margin of segment on either side with roundish incisions filled with ground colour. S8 azure blue with a black streak along posterior margin going for half of its height in lateral view. S9 and S10 azure blue below and black above, with black occupying slightly less than half of S9 tergite height on S9 but about 2/3 of S10 tergite height in lateral view. S9 posterior margin with a dorsal blue dot; S10 posterior margin brownish. Secondary genitalia yellowish, hamuli with pale-brown margins, ligula brownish-black. Sternites of S1–S6 barely seen, black, of S7 bluish-yellow with a black mid-streak, on S8 pale blue with a black mid-streak, on S9–S10 pale blue, black. S10 raised dorsally towards apical tubercle, its dorsal margin in dorsal view oblique but straight. Apical tubercle has a rectangular outline with rounded corners in caudal view, its lateral margins thick and black, somewhat extending to dorsal side where they acquire brownish colour and form rounded, laterad inclined projections; tubercle inner area azure blue. In dorsal view S10 posterior margin, as well as tubercle posterior margin, shallowly incised as a blunt angle.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268308B1E73B3FA9F52D44B22" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="575">
|
||
Cerci brown but their dorsal side and processes blackish brown; short and rounded in lateral view and nearly rectangular, disposed at a blunt angle to each other in dorsal view (
|
||
<figureCitation id="13582A5768308B1E70F1FAB351FC4826" box="[901,983,1339,1365]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="2" pageNumber="575">Fig. 3a</figureCitation>
|
||
). In caudal view they are roundish, their inner processes long, blunt and deeply crossed with each other (
|
||
<figureCitation id="13582A5768308B1E700BFAD751FF480A" box="[895,980,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="2" pageNumber="575">Fig. 3a</figureCitation>
|
||
). Paraprocts yellowish basally but their dorsal side and protruding distal part brownish-black. They are twice as long as the cerci and twice as short than S
|
||
<quantity id="4C9B9B3768308B1E73D2FA2F52F548B2" box="[166,222,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="575" unit="in" value="10.0">10 in</quantity>
|
||
lateral view, directed obliquely upward, the left one slightly curved, the right one very slightly so; their apices blunt; dorsal side and apices coarse because of scarcely defined blunt teeth, about three of which disposed on apex (
|
||
<figureCitation id="13582A5768308B1E73D4FA6752DB4B7A" box="[160,240,1519,1545]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="2" pageNumber="575">Fig. 3a</figureCitation>
|
||
). In dorsal view they are very slightly diverging, almost parallel, slightly hooked at base (
|
||
<figureCitation id="13582A5768308B1E77A5FA6757094B7A" box="[1233,1314,1519,1545]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="2" pageNumber="575">Fig. 3a</figureCitation>
|
||
). In caudal view the tips of paraprocts project onto the hind margin of S10, their inner processes yellowish with rounded tips (
|
||
<figureCitation id="13582A5768308B1E73D4F9BF52DA4B22" box="[160,241,1591,1617]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="2" pageNumber="575">Fig. 3a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268308B1E73B3F9D356B34B06" blockId="2.[151,1437,151,2013]" box="[199,1176,1626,1653]" pageId="2" pageNumber="575">
|
||
<emphasis id="B917EAC068308B1E73B3F9D353984B06" box="[199,435,1627,1653]" italics="true" pageId="2" pageNumber="575">Measurements (mm).</emphasis>
|
||
Hindwing 18.5; abdomen without appendages 29; total length 37.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968308B1F73B3F9F653AF4EF2" lastPageId="3" lastPageNumber="576" pageId="2" pageNumber="575" type="description">
|
||
<paragraph id="8BDC36D268308B1F73B3F9F653414C6E" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1437,151,1977]" lastPageId="3" lastPageNumber="576" pageId="2" pageNumber="575">
|
||
<emphasis id="B917EAC068308B1E73B3F9F650314BEA" bold="true" box="[199,538,1662,1689]" pageId="2" pageNumber="575">
|
||
Variation in male
|
||
<typeStatus id="54D8887068308B1E72E9F9F7503E4BEA" box="[413,533,1663,1689]" pageId="2" pageNumber="575" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The angle at which the paraprocts are directed obliquely upward varies between ca 30° and 40°. In
|
||
<specimenCount id="9D65FD5B68308B1E721FF92B53D04BCE" box="[363,507,1699,1725]" pageId="2" pageNumber="575" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
the paraprocts are slightly curved, whereas in others their shape is nearly straight. The number of irregular teeth on the apices of the paraprocts vary: in
|
||
<specimenCount id="9D65FD5B68308B1E70ECF94F56184B92" box="[920,1075,1735,1761]" pageId="2" pageNumber="575" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
the paraproct ends with a robust blunt tooth and a larger tubercle on dorsal surface just before it; in another
|
||
<typeStatus id="54D8887068308B1E7098F96356674A76" box="[1004,1100,1771,1797]" pageId="2" pageNumber="575" type="paratype">paratype</typeStatus>
|
||
the left paraproct is the same while the right one ends with three irregular teeth, as in the
|
||
<typeStatus id="54D8887068308B1E704EF887518B4A5A" box="[826,928,1807,1833]" pageId="2" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
; in the fourth
|
||
<typeStatus id="54D8887068308B1E774BF88756B44A5A" box="[1087,1183,1807,1833]" pageId="2" pageNumber="575" type="paratype">paratype</typeStatus>
|
||
the left paraproct also bears two and the right one five blunt teeth at apex, which is slightly expanded after a slight subapical constriction. The blue spot at the apex of the prothoracic process is as large as in the
|
||
<typeStatus id="54D8887068308B1E70C9F8DF56344A02" box="[957,1055,1879,1905]" pageId="2" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
in
|
||
<specimenCount id="9D65FD5B68308B1E7730F8DF56FD4A02" box="[1092,1238,1879,1905]" pageId="2" pageNumber="575" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
, but is reduced to occupy only its hind margin in
|
||
<specimenCount id="9D65FD5B68308B1E728AF8F350B54AE6" box="[510,670,1915,1941]" pageId="2" pageNumber="575" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
, and the blue colour is completely missing from the process in
|
||
<specimenCount id="9D65FD5B68308B1E7600F8F352D04ACA" pageId="2" pageNumber="575" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
. The pterostigma coloration varies considerably: in
|
||
<specimenCount id="9D65FD5B68308B1E7036F81751FA4ACA" box="[834,977,1951,1977]" pageId="2" pageNumber="575" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
the fore wing pterostigma lacks the blue colour; in
|
||
<specimenCount id="9D65FD5B68308B1E7278F84B53B54AAE" box="[268,414,1987,2013]" pageId="2" pageNumber="575" type="generic" typeStatus="paratype">two paratype</specimenCount>
|
||
it lacks the blue colour only its upper side while the black colour expands to ca 4/5 of the pterostigma or more on both sides; the hind wing pterostigma varies in the degree of darkening of its main area and the breadth of the pale rims. There are nine rather than ten postnodals in both fore wings of
|
||
<specimenCount id="9D65FD5B68318B1F77E6FF33570A4DA6" box="[1170,1313,187,213]" pageId="3" pageNumber="576" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
and in one of those in another one. Ranges of measurements (mm): hindwing 17–19; abdomen without appendages 26.5–29; total length 33–36.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FEAF512F4C16" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FEAF530E4C32" bold="true" box="[199,293,295,321]" pageId="3" pageNumber="576">Female.</emphasis>
|
||
The
|
||
<specimenCount id="9D65FD5B68318B1F7215FEAF53F44C32" box="[353,479,295,321]" pageId="3" pageNumber="576" type="female">two female</specimenCount>
|
||
<typeStatus id="54D8887068318B1F7293FEAF50794C32" box="[487,594,295,321]" pageId="3" pageNumber="576" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="13582A5768318B1F7116FEAF509C4C32" box="[610,695,295,321]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="3" pageNumber="576">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768318B1F71B1FEAF50FF4C32" box="[709,724,295,321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">4</figureCitation>
|
||
) resemble males in coloration; they slightly differ in size from each other; the smaller individual is illustrated in
|
||
<figureCitation id="13582A5768318B1F71B6FEC3512B4C16" box="[706,768,331,357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig 4</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FEE750694CDE" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FEE7532F4CFA" box="[199,260,367,393]" italics="true" pageId="3" pageNumber="576">Head</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768318B1F7267FEE7534F4CF9" box="[275,356,367,394]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4b</figureCitation>
|
||
) As in male but in the smaller female the postocular spots also smaller (about twice as small with respect to area), with uneven margins.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FE3F515E4FD9" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FE3F53334CA2" box="[199,280,439,465]" italics="true" pageId="3" pageNumber="576">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768318B1F725DFE3F53AB4CA2" box="[297,384,439,465]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="3" pageNumber="576">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768318B1F72E4FE3F53804CA2" box="[400,427,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">4a</figureCitation>
|
||
): Pterothorax as in male. In the larger female
|
||
<typeStatus id="54D8887068318B1F70B2FE3F560D4CA2" box="[966,1062,439,465]" pageId="3" pageNumber="576" type="paratype">paratype</typeStatus>
|
||
the posterior process of its hind margin is as in the male
|
||
<typeStatus id="54D8887068318B1F72C5FE5350384C86" box="[433,531,475,501]" pageId="3" pageNumber="576" type="holotype">holotype</typeStatus>
|
||
but the blue spot is larger and there is a blue dot on the left side of the median lobe. In the smaller female
|
||
<typeStatus id="54D8887068318B1F72A1FE77501E4F6A" box="[469,565,511,537]" pageId="3" pageNumber="576" type="paratype">paratype</typeStatus>
|
||
the process is flat, slightly shorter and much broader, trapezoid with barely converging sides, almost rectangular, with rounded angles and a small and shallow incision at apex (
|
||
<figureCitation id="13582A5768318B1F7781FDAB57624F4D" box="[1269,1353,547,574]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4d</figureCitation>
|
||
). In the larger female venation is dark and there is one more postnodal on each wing than in the
|
||
<typeStatus id="54D8887068318B1F77FAFDCF56DF4F12" box="[1166,1268,583,609]" pageId="3" pageNumber="576" type="holotype">holotype</typeStatus>
|
||
; in the smaller female venation is as in the
|
||
<typeStatus id="54D8887068318B1F72A1FDE3501C4FF6" box="[469,567,619,645]" pageId="3" pageNumber="576" type="holotype">holotype</typeStatus>
|
||
but the longitudinal veins are paler. All pterostigmata pale grey with whitish margins, three bordering veins dark but costal one pale. (
|
||
<figureCitation id="13582A5768318B1F7060FD0751434FD9" box="[788,872,655,682]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FD3B50E44E4A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FD3B531A4FBE" box="[199,305,691,717]" italics="true" pageId="3" pageNumber="576">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768318B1F7234FD3B53B84FBE" box="[320,403,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[389,1199,181,1920]" captionTargetId="figure-21@4.[389,1199,181,1920]" captionTargetPageId="4" captionText="FIGURE 1. Ischnura elegans malikovae. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia." figureDoi="http://doi.org/10.5281/zenodo.6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="3" pageNumber="576">Figs 1a</figureCitation>
|
||
,
|
||
<figureCitation id="13582A5768318B1F72D4FD3B53974FBE" box="[416,444,691,717]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">4c</figureCitation>
|
||
) as in male but somewhat thicker, dorsal black spot of S2 somewhat narrower, that of S7 with shallower posterior incisions and that of S9 with deeply incised margins (
|
||
<figureCitation id="13582A5768318B1F7776FD5F56794F82" box="[1026,1106,727,753]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4c</figureCitation>
|
||
) (which are straight in male). Sternite of S8 with blue spots, vulvar spine well developed (
|
||
<figureCitation id="13582A5768318B1F704DFD7351A04E66" box="[825,907,763,789]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4c</figureCitation>
|
||
). Ovipositor yellowish but style brownish (
|
||
<figureCitation id="13582A5768318B1F761BFD73529E4E4A" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4c</figureCitation>
|
||
). Cerci blackish, paraprocts yellowish (
|
||
<figureCitation id="13582A5768318B1F7104FC9750E94E4A" box="[624,706,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[483,1105,181,1916]" captionTargetId="figure-17@7.[483,1105,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Morphological details of the smaller female paratype of Ischnura elegans malikovae: a—head and thorax, lateral view; b—head (dorsal view) and prothorax (lateral view); c—end of abdomen, lateral view; d—prothorax, anteriodorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392952" httpUri="https://zenodo.org/record/6392952/files/figure.png" pageId="3" pageNumber="576">Fig. 4c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FCCB53AF4EF2" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FCCB50424E2E" box="[199,617,835,861]" italics="true" pageId="3" pageNumber="576">
|
||
Measurements (mm;
|
||
<specimenCount id="9D65FD5B68318B1F72C1FCCB50774E2E" box="[437,604,835,861]" pageId="3" pageNumber="576" type="generic">two specimens</specimenCount>
|
||
).
|
||
</emphasis>
|
||
Hindwing 18 and 19.5; abdomen without appendages 25 and 27.5; total length 32.5 and 35.5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968318B1F73B3FC0256D14B92" pageId="3" pageNumber="576" type="diagnosis">
|
||
<paragraph id="8BDC36D268318B1F73B3FC0250B848EE" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3FC0253694ED7" bold="true" box="[199,322,906,932]" pageId="3" pageNumber="576">Diagnosis.</emphasis>
|
||
While currently
|
||
<taxonomicName id="4C634D5168318B1F728EFC03504E4ED6" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[506,613,907,933]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068318B1F728EFC03504E4ED6" box="[506,613,907,933]" italics="true" pageId="3" pageNumber="576">I. elegans</emphasis>
|
||
</taxonomicName>
|
||
is considered as a monotypical species (
|
||
<bibRefCitation id="EFF24B2368318B1F7769FC0357184ED6" author="Boudot, J. - P. & Salamun, A." box="[1053,1331,906,933]" pageId="3" pageNumber="576" pagination="125 - 126" refId="ref8282" refString="Boudot, J. - P. & Salamun, A. (2015) In: Boudot, J. - P. & Kalkman, V. J. (Eds.), Atlas of the European Dragonflies and Damselflies. KKNV publishing, Zeist, pp. 125 - 126." type="book chapter" year="2015">Boudot & Salamun 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF24B2368318B1F7649FC03533B4EBA" author="Schroter, A. & Seehausen, M. & Kunz, B. & Gunther, A. & Schneider, T. & Jodicke, R." pageId="3" pageNumber="576" pagination="279 - 342" refId="ref9838" refString="Schroter, A., Seehausen, M., Kunz, B., Gunther, A., Schneider, T. & Jodicke, R. (2015) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Odonatologica, 44, 279 - 342." type="journal article" year="2015">
|
||
Schröter
|
||
<emphasis id="B917EAC068318B1F73E3FC2752E44EBA" box="[151,207,943,969]" italics="true" pageId="3" pageNumber="576">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
; Schneider
|
||
<emphasis id="B917EAC068318B1F72E6FC2753E24EBA" box="[402,457,943,969]" italics="true" pageId="3" pageNumber="576">et al.</emphasis>
|
||
2018;
|
||
<bibRefCitation id="EFF24B2368318B1F7162FC27511F4EBA" author="Kosterin, O. E. & Ahmadi, A." box="[534,820,943,969]" pageId="3" pageNumber="576" pagination="1 - 65" refId="ref9163" refString="Kosterin, O. E. & Ahmadi, A. (2018) Odonata observed in Central Zagros, Iran, in late May 2017. International Dragonfly Fund Report, 117, 1 - 65." type="journal article" year="2018">Kosterin & Ahmadi 2018</bibRefCitation>
|
||
), a new subspecies described for it should be compared to the nominotypical subspecies which is to be redefined. Its
|
||
<typeStatus id="54D8887068318B1F7025FC5B51AA4E9E" box="[849,897,979,1005]" pageId="3" pageNumber="576">type</typeStatus>
|
||
locality is Europe, from where the species was described. We find specimens of
|
||
<taxonomicName id="4C634D5168318B1F717BFC7F50564962" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[527,637,1015,1041]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068318B1F717BFC7F50564962" box="[527,637,1015,1041]" italics="true" pageId="3" pageNumber="576">I. elegans</emphasis>
|
||
</taxonomicName>
|
||
from Europe, the Caucasus, Ural and West Siberia homogeneous with respect to the characters considered as diagnostic below, so we assume
|
||
<emphasis id="B917EAC068318B1F70B9FB9356734946" box="[973,1112,1051,1077]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7776FB9356734946" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[1026,1112,1051,1077]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
to range in these territories. The new subspecies is very similar to
|
||
<emphasis id="B917EAC068318B1F714BFBB750ED492A" box="[575,710,1087,1113]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7104FBB750ED492A" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[624,710,1087,1113]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
but the males strongly differ from it in their paraprocts. In dorsal view, the male paraprocts of
|
||
<emphasis id="B917EAC068318B1F7296FBEB50A1490E" box="[482,650,1123,1149]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7163FBEB50A1490E" authorityName="Onishko & Kosterin" authorityYear="2022" box="[535,650,1123,1149]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
are nearly parallel (
|
||
<figureCitation id="13582A5768318B1F7005FBEB51E9490E" box="[881,962,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3a</figureCitation>
|
||
) while in
|
||
<emphasis id="B917EAC068318B1F774CFBEB56E8490E" box="[1080,1219,1123,1149]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7719FBEB56E8490E" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[1133,1219,1123,1149]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
from Europe, Ural and West Siberia they are strongly divaricate and strikingly (1.5 times) longer (
|
||
<figureCitation id="13582A5768318B1F776AFB0F565849D1" box="[1054,1139,1159,1186]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3b</figureCitation>
|
||
). In lateral view, the male paraprocts of
|
||
<emphasis id="B917EAC068318B1F7247FB2353F149B6" box="[307,474,1195,1221]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7213FB2353F149B6" authorityName="Onishko & Kosterin" authorityYear="2022" box="[359,474,1195,1221]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
are directed obliquely upward (
|
||
<figureCitation id="13582A5768318B1F7032FB2351B349B6" box="[838,920,1195,1221]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3a</figureCitation>
|
||
), as in
|
||
<taxonomicName id="4C634D5168318B1F709FFB23577A49B6" authority="(Rambur, 1842)" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1003,1361,1195,1221]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="genei">
|
||
<emphasis id="B917EAC068318B1F709FFB2356BF49B7" box="[1003,1172,1195,1221]" italics="true" pageId="3" pageNumber="576">Ischnura genei</emphasis>
|
||
(Rambur, 1842)
|
||
</taxonomicName>
|
||
, while in
|
||
<emphasis id="B917EAC068318B1F73C1FB475314499A" box="[181,319,1231,1257]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F739DFB475314499A" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[233,319,1231,1257]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
they are directed strictly posteriorly (
|
||
<figureCitation id="13582A5768318B1F7198FB47516A4999" box="[748,833,1231,1258]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3b</figureCitation>
|
||
). In specimens from
|
||
<collectingCountry id="F374764268318B1F7745FB475675499A" box="[1073,1118,1231,1257]" name="Iran" pageId="3" pageNumber="576">Iran</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A7F83068318B1F771AFB475731499A" box="[1134,1306,1231,1257]" country="Iran" name="Markazi" pageId="3" pageNumber="576">Markazi Ostan</collectingRegion>
|
||
), which we consider to represent the subspecies
|
||
<emphasis id="B917EAC068318B1F7140FB7B5084487E" box="[564,687,1267,1293]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7113FB7B5084487E" authorityName="Schmidt" authorityYear="1938" box="[615,687,1267,1293]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="ebneri">ebneri</taxonomicName>
|
||
</emphasis>
|
||
(see below), the paraprocts are scarcely or not at all divergent but of the same length as in
|
||
<emphasis id="B917EAC068318B1F72C1FA9F506A4842" box="[437,577,1303,1329]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F729FFA9F506A4842" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[491,577,1303,1329]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
in dorsal view and strictly caudally directed in lateral view (
|
||
<figureCitation id="13582A5768318B1F7674FA9F575B4841" box="[1280,1392,1303,1330]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3c–d</figureCitation>
|
||
), in this view being 0.71–0.73 as long as S10 (
|
||
<figureCitation id="13582A5768318B1F7102FAB350C94825" box="[630,738,1339,1366]" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3c–d</figureCitation>
|
||
) versus ca.
|
||
<quantity id="4C9B9B3768318B1F701DFAB251824826" box="[873,937,1338,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="576" unit="in" value="0.5">0.5 in</quantity>
|
||
<emphasis id="B917EAC068318B1F70C5FAB3567C4826" box="[945,1111,1339,1365]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7090FAB3567C4826" authorityName="Onishko & Kosterin" authorityYear="2022" box="[996,1111,1339,1365]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<quantity id="4C9B9B3768318B1F77FAFAB356E44826" box="[1166,1231,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="576" unit="in" value="0.6">0.6 in</quantity>
|
||
<emphasis id="B917EAC068318B1F77A2FAB357744826" box="[1238,1375,1339,1365]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F767DFAB357744826" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[1289,1375,1339,1365]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13582A5768318B1F761BFAB352F94809" captionStart="FIGURE 3" captionStartId="6.[151,250,1547,1571]" captionTargetBox="[182,1436,181,1515]" captionTargetId="figure-21@6.[151,1436,181,1523]" captionTargetPageId="6" captionText="FIGURE 3. Male anal appendages of Ischnura elegans sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—I. elegans malikovae, the holotype; b—I. elegans elegans elegans from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (Kosterin 2017); c–d—I. elegans elegans ebneri from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (Kosterin & Ahmadi 2018); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="3" pageNumber="576">Fig. 3c–d</figureCitation>
|
||
). The prothoracic process in males of
|
||
<emphasis id="B917EAC068318B1F710CFAD75137480A" box="[632,796,1375,1401]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F71DDFAD75137480A" authority="Onishko & Kosterin, 2022" authorityName="Onishko & Kosterin" authorityYear="2022" box="[681,796,1375,1401]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae" status="sp. nov.">malikovae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A22457BB68318B1F7057FAD75152480A" box="[803,889,1375,1401]" pageId="3" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is longer than broad and parallel-sided, which is, however, within the variation of
|
||
<emphasis id="B917EAC068318B1F7171FA0B50A648EE" box="[517,653,1411,1437]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7143FA0B50A648EE" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[567,653,1411,1437]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FA2F577148B2" blockId="3.[151,1437,151,1977]" box="[199,1370,1447,1473]" pageId="3" pageNumber="576">
|
||
We failed to trace any diagnostic difference of females of the new subspecies from those of
|
||
<emphasis id="B917EAC068318B1F77BFFA2F577848B2" box="[1227,1363,1447,1473]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F7789FA2F577848B2" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[1277,1363,1447,1473]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3FA43539D4B06" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
From the superficially similar
|
||
<taxonomicName id="4C634D5168318B1F715EFA43519F4895" authority="(Ris, 1916)" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[554,948,1483,1510]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="senegalensis">
|
||
<emphasis id="B917EAC068318B1F715EFA43510D4896" box="[554,806,1483,1509]" italics="true" pageId="3" pageNumber="576">Ischnura senegalensis</emphasis>
|
||
(Ris, 1916)
|
||
</taxonomicName>
|
||
, with which the new species co-occurs in
|
||
<collectingCountry id="F374764268318B1F73E3FA6752F04B7A" box="[151,219,1519,1545]" name="South Korea" pageId="3" pageNumber="576">Korea</collectingCountry>
|
||
and
|
||
<collectingCountry id="F374764268318B1F7266FA6753734B7A" box="[274,344,1519,1545]" name="China" pageId="3" pageNumber="576">China</collectingCountry>
|
||
,
|
||
<emphasis id="B917EAC068318B1F7217FA6750234B7A" box="[355,520,1519,1545]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F72E1FA6750234B7A" authorityName="Onishko & Kosterin" authorityYear="2022" box="[405,520,1519,1545]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
differs by the presence of a strong process of the prothorax posterior lobe in both sexes, by the black dorsal spot on S
|
||
<quantity id="4C9B9B3768318B1F7155F99B50664B5E" box="[545,589,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="576" unit="in" value="2.0">2 in</quantity>
|
||
males not glittering green and the ventral parts of the S7 tergite largely blue rather than black. In addition, the male paraprocts in
|
||
<taxonomicName id="4C634D5168318B1F7187F9BF51B24B22" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[755,921,1591,1617]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="senegalensis">
|
||
<emphasis id="B917EAC068318B1F7187F9BF51B24B22" box="[755,921,1591,1617]" italics="true" pageId="3" pageNumber="576">I. senegalensis</emphasis>
|
||
</taxonomicName>
|
||
are longer, pointed and more curved towards each other in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="8BDC36D268318B1F73B3F9F756D14B92" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
One more species of
|
||
<taxonomicName id="4C634D5168318B1F72CBF9F750084BEA" authorityName="Onishko & Kosterin" authorityYear="2022" box="[447,547,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EAC068318B1F72CBF9F750084BEA" box="[447,547,1663,1689]" italics="true" pageId="3" pageNumber="576">Ischnura</emphasis>
|
||
</taxonomicName>
|
||
co-occurring with
|
||
<emphasis id="B917EAC068318B1F7074F9F751804BEA" box="[768,939,1663,1689]" italics="true" pageId="3" pageNumber="576">
|
||
I. e.
|
||
<taxonomicName id="4C634D5168318B1F704CF9F751804BEA" authorityName="Onishko & Kosterin" authorityYear="2022" box="[824,939,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
is
|
||
<taxonomicName id="4C634D5168318B1F70A5F9F757624BEA" authority="(Brauer, 1865)" baseAuthorityName="Brauer" baseAuthorityYear="1865" box="[977,1353,1662,1689]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B917EAC068318B1F70A5F9F756BC4BEB" box="[977,1175,1663,1689]" italics="true" pageId="3" pageNumber="576">Ischnura asiatica</emphasis>
|
||
(Brauer, 1865)
|
||
</taxonomicName>
|
||
, which belongs to the
|
||
<taxonomicName id="4C634D5168318B1F724CF92B53A54BCE" authorityName="Charpentier" authorityYear="1825" box="[312,398,1699,1725]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B917EAC068318B1F724CF92B53A54BCE" box="[312,398,1699,1725]" italics="true" pageId="3" pageNumber="576">pumilio</emphasis>
|
||
</taxonomicName>
|
||
-group and well differs by the pterostigmata substantially smaller in the hind wings than in the fore wings, S8 black but S9 pale (blue or green) in males (vice versa in
|
||
<taxonomicName id="4C634D5168318B1F70CEF94F560C4B92" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[954,1063,1735,1761]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068318B1F70CEF94F560C4B92" box="[954,1063,1735,1761]" italics="true" pageId="3" pageNumber="576">I. elegans</emphasis>
|
||
</taxonomicName>
|
||
), and smaller size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379655968318B1A73B3F96250B24AC3" lastPageId="6" lastPageNumber="579" pageId="3" pageNumber="576" type="distribution">
|
||
<paragraph id="8BDC36D268318B1F73B3F96256BA4ACA" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="576">
|
||
<emphasis id="B917EAC068318B1F73B3F962534A4A77" bold="true" box="[199,353,1770,1796]" pageId="3" pageNumber="576">Distribution.</emphasis>
|
||
<emphasis id="B917EAC068318B1F721CF96350494A76" box="[360,610,1771,1797]" italics="true" pageId="3" pageNumber="576">
|
||
<taxonomicName id="4C634D5168318B1F721CF96353E74A76" authorityName="Onishko & Kosterin" authorityYear="2022" box="[360,460,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">Ischnura</taxonomicName>
|
||
e.
|
||
<taxonomicName id="4C634D5168318B1F729BF96350494A76" authorityName="Onishko & Kosterin" authorityYear="2022" box="[495,610,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="malikovae">malikovae</taxonomicName>
|
||
</emphasis>
|
||
occurs for sure in the southern Khabarovskiy Kray and
|
||
<collectingRegion id="49A7F83068318B1F7794F96357B74A76" box="[1248,1436,1771,1797]" country="Russia" name="Primorskiy" pageId="3" pageNumber="576">Primorskiy Kray</collectingRegion>
|
||
of
|
||
<collectingCountry id="F374764268318B1F73CFF88753224A5A" box="[187,265,1807,1833]" name="Russia" pageId="3" pageNumber="576">Russia</collectingCountry>
|
||
. However, based on biogeographical considerations and available illustrations in the literature (see Discussion below), it appears to enjoy a broad East Asian distribution. We believe that it likely occupies the eastern Zabaikalskiy Kray,
|
||
<collectingRegion id="49A7F83068318B1F7205F8DF50354A02" box="[369,542,1879,1905]" country="Russia" name="Amur" pageId="3" pageNumber="576">Amur Province</collectingRegion>
|
||
and
|
||
<collectingRegion id="49A7F83068318B1F7120F8DF508B4A02" box="[596,672,1879,1905]" country="Russia" name="Yevrey" pageId="3" pageNumber="576">Jewish</collectingRegion>
|
||
Autonomous Region in
|
||
<collectingCountry id="F374764268318B1F70C6F8DF51D64A02" box="[946,1021,1879,1905]" name="Russia" pageId="3" pageNumber="576">Russia</collectingCountry>
|
||
and broadly ranges across N and NE
|
||
<collectingCountry id="F374764268318B1F73E3F8F352F64AE6" box="[151,221,1915,1941]" name="China" pageId="3" pageNumber="576">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F374764268318B1F739CF8F353044AE6" box="[232,303,1915,1941]" name="South Korea" pageId="3" pageNumber="576">Korea</collectingCountry>
|
||
, and
|
||
<collectingRegion id="49A7F83068318B1F721EF8F353F34AE6" box="[362,472,1915,1941]" country="Japan" name="Hokkaido" pageId="3" pageNumber="576">Hokkaido</collectingRegion>
|
||
and the northernmost Honshu in
|
||
<collectingCountry id="F374764268318B1F703BF8F351B94AE6" box="[847,914,1915,1941]" name="Japan" pageId="3" pageNumber="576">Japan</collectingCountry>
|
||
. In
|
||
<collectingCountry id="F374764268318B1F70CFF8F351D04AE6" box="[955,1019,1915,1941]" name="Japan" pageId="3" pageNumber="576">Japan</collectingCountry>
|
||
it seems to have been mistaken for
|
||
<taxonomicName id="4C634D5168318B1F76F8F8F353614ACA" authority="elegans" authorityName="elegans (Vander Linden" authorityYear="1820" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="576" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="elegans">
|
||
<emphasis id="B917EAC068318B1F76F8F8F353614ACA" italics="true" pageId="3" pageNumber="576">I. elegans elegans</emphasis>
|
||
</taxonomicName>
|
||
, while specimens from
|
||
<collectingCountry id="F374764268318B1F7122F81750B24ACA" box="[598,665,1951,1977]" name="China" pageId="3" pageNumber="576">China</collectingCountry>
|
||
and
|
||
<collectingCountry id="F374764268318B1F71BBF81751384ACA" box="[719,787,1951,1977]" name="South Korea" pageId="3" pageNumber="576">Korea</collectingCountry>
|
||
were not identified to subspecies.
|
||
</paragraph>
|
||
<caption id="DF1C665A68368B1873E3F81F530B4A8B" ID-DOI="http://doi.org/10.5281/zenodo.6392946" ID-Zenodo-Dep="6392946" httpUri="https://zenodo.org/record/6392946/files/figure.png" pageId="4" pageNumber="577" startId="4.[151,250,1943,1967]" targetBox="[389,1199,181,1920]" targetPageId="4">
|
||
<paragraph id="8BDC36D268368B1873E3F81F530B4A8B" blockId="4.[151,1436,1943,2040]" pageId="4" pageNumber="577">
|
||
<emphasis id="B917EAC068368B1873E3F81F533F4ADC" bold="true" box="[151,276,1943,1967]" pageId="4" pageNumber="577">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C634D5168368B18726EF810501C4AC3" authorityName="Onishko & Kosterin" authorityYear="2022" box="[282,567,1944,1968]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="577" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="malikovae">
|
||
<emphasis id="B917EAC068368B18726EF810501C4AC3" box="[282,567,1944,1968]" italics="true" pageId="4" pageNumber="577">Ischnura elegans malikovae</emphasis>
|
||
</taxonomicName>
|
||
. in nature: a—in copula (the holotype male and paratype female) in the type locality, the reservoir in Luchegorsk Town, Primorskiy Kray, Russia; b–c—males at the pond in Korfovskiy settlement, Khabarovskiy Kray, Russia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1C665A68378B1973E3F81F57A34AA0" ID-DOI="http://doi.org/10.5281/zenodo.6392948" ID-Zenodo-Dep="6392948" httpUri="https://zenodo.org/record/6392948/files/figure.png" pageId="5" pageNumber="578" startId="5.[151,250,1943,1967]" targetBox="[394,1193,181,1897]" targetPageId="5">
|
||
<paragraph id="8BDC36D268378B1973E3F81F57A34AA0" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="578">
|
||
<emphasis id="B917EAC068378B1973E3F81F53384ADC" bold="true" box="[151,275,1943,1967]" pageId="5" pageNumber="578">FIGURE 2.</emphasis>
|
||
Morphological details of the holotype male, of
|
||
<taxonomicName id="4C634D5168378B197182F81056394AC3" authorityName="Onishko & Kosterin" authorityYear="2022" box="[758,1042,1944,1968]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="578" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="malikovae">
|
||
<emphasis id="B917EAC068378B197182F81056394AC3" box="[758,1042,1944,1968]" italics="true" pageId="5" pageNumber="578">Ischnura elegans malikovae</emphasis>
|
||
</taxonomicName>
|
||
: a—head and thorax, lateral view; b— head, frontal view; c—end of abdomen, lateral view; d—prothorax, lateral view; e—prothorax, dorsal view. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1C665A68348B1A73E3F98357644BC0" ID-DOI="http://doi.org/10.5281/zenodo.6392950" ID-Zenodo-Dep="6392950" httpUri="https://zenodo.org/record/6392950/files/figure.png" pageId="6" pageNumber="579" startId="6.[151,250,1547,1571]" targetBox="[182,1436,181,1515]" targetPageId="6">
|
||
<paragraph id="8BDC36D268348B1A73E3F98357644BC0" blockId="6.[151,1437,1547,1716]" pageId="6" pageNumber="579">
|
||
<emphasis id="B917EAC068348B1A73E3F98353384B50" bold="true" box="[151,275,1547,1571]" pageId="6" pageNumber="579">FIGURE 3.</emphasis>
|
||
Male anal appendages of
|
||
<taxonomicName id="4C634D5168348B1A716FF98350E34B50" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[539,712,1547,1571]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068348B1A716FF98350E34B50" box="[539,712,1547,1571]" italics="true" pageId="6" pageNumber="579">Ischnura elegans</emphasis>
|
||
</taxonomicName>
|
||
sspp. (left—dorsal view; middle—lateral view; right—posterior view): a—
|
||
<taxonomicName id="4C634D5168348B1A73C8F9B853A04B34" authorityName="Onishko & Kosterin" authorityYear="2022" box="[188,395,1583,1607]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="malikovae">
|
||
<emphasis id="B917EAC068348B1A73C8F9B853A04B34" box="[188,395,1583,1607]" italics="true" pageId="6" pageNumber="579">I. elegans malikovae</emphasis>
|
||
</taxonomicName>
|
||
, the holotype; b—
|
||
<emphasis id="B917EAC068348B1A7133F9B851654B34" box="[583,846,1583,1607]" italics="true" pageId="6" pageNumber="579">
|
||
<taxonomicName id="4C634D5168348B1A7133F9B850D04B34" authority="elegans" authorityName="elegans (Vander Linden" authorityYear="1820" box="[583,763,1583,1607]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="elegans">I. elegans elegans</taxonomicName>
|
||
<taxonomicName id="4C634D5168348B1A7075F9A751654B34" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[769,846,1583,1607]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
||
</emphasis>
|
||
from Russia, Krasnodarskiy Kray Province, Novorossiysk Municipality, Lake Krugloe, 44.677° N 37.597° E, 10 vii 2016 (
|
||
<bibRefCitation id="EFF24B2368348B1A704EF9DB51F84B1F" author="Kosterin, O. E." box="[826,979,1619,1644]" pageId="6" pageNumber="579" pagination="1 - 23" refId="ref9128" refString="Kosterin, O. E. (2017) Update to Odonata of the Black Sea coast of the western Caucasus, Russia. International Dragonfly Fund Report, 110, 1 - 23." type="journal article" year="2017">Kosterin 2017</bibRefCitation>
|
||
); c–d—
|
||
<emphasis id="B917EAC068348B1A7751F9DC57014B18" box="[1061,1322,1619,1643]" italics="true" pageId="6" pageNumber="579">
|
||
<taxonomicName id="4C634D5168348B1A7751F9DC56CB4B18" authority="elegans" authorityName="elegans (Vander Linden" authorityYear="1820" box="[1061,1248,1619,1643]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="elegans">I. elegans elegans</taxonomicName>
|
||
<taxonomicName id="4C634D5168348B1A779DF9DB57014B18" authorityName="Schmidt" authorityYear="1938" box="[1257,1322,1619,1643]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="ebneri">ebneri</taxonomicName>
|
||
</emphasis>
|
||
from Iran, Markazi Province, Garehchay River at Gavigodar village 34.109° N 49.363° E, 19 v 2017 (
|
||
<bibRefCitation id="EFF24B2368348B1A775CF9FF57074BE3" author="Kosterin, O. E. & Ahmadi, A." box="[1064,1324,1655,1680]" pageId="6" pageNumber="579" pagination="1 - 65" refId="ref9163" refString="Kosterin, O. E. & Ahmadi, A. (2018) Odonata observed in Central Zagros, Iran, in late May 2017. International Dragonfly Fund Report, 117, 1 - 65." type="journal article" year="2018">Kosterin & Ahmadi 2018</bibRefCitation>
|
||
); these two specimens exhibit the extremities of the slight variation in the paraproct orientation in the same series. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDC36D268348B1A73B3F96950B24AC3" blockId="6.[151,1436,1761,1968]" pageId="6" pageNumber="579">
|
||
<emphasis id="B917EAC068348B1A73B3F96953014B88" bold="true" box="[199,298,1761,1787]" pageId="6" pageNumber="579">Habitat.</emphasis>
|
||
The new subspecies was found in sedge (
|
||
<taxonomicName id="4C634D5168348B1A7073F96A51584B8F" box="[775,883,1762,1788]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="579" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B917EAC068348B1A7073F96A51614B8F" box="[775,842,1762,1788]" italics="true" pageId="6" pageNumber="579">Carex</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
) at the bank of a small and clear pond inside the town of Korfovskiy (southern Khabarovskiy Kray), of a large reservoir quite polluted with cattle dung inside the village of Prokhory (southern
|
||
<collectingRegion id="49A7F83068348B1A729EF8A250834A37" box="[490,680,1834,1860]" country="Russia" name="Primorskiy" pageId="6" pageNumber="579">Primorskiy Kray</collectingRegion>
|
||
) and of a very large, clear and warm reservoir at Luchegorsk Town (northern
|
||
<collectingRegion id="49A7F83068348B1A727CF8C653E04A1B" box="[264,459,1870,1896]" country="Russia" name="Primorskiy" pageId="6" pageNumber="579">Primorskiy Kray</collectingRegion>
|
||
). So, it inhabits a variety of lentic habitats and prefers sedge bordering open water. It is noteworthy that in the southern Far East of
|
||
<collectingCountry id="F374764268348B1A71DAF8FA50D24AFF" box="[686,761,1906,1932]" name="Russia" pageId="6" pageNumber="579">Russia</collectingCountry>
|
||
it appears to be highly local and never so widespread and numerous as
|
||
<taxonomicName id="4C634D5168348B1A725FF81E53B34AC3" baseAuthorityName="Vander Linden" baseAuthorityYear="1820" box="[299,408,1942,1968]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B917EAC068348B1A725FF81E53B34AC3" box="[299,408,1942,1968]" italics="true" pageId="6" pageNumber="579">I. elegans</emphasis>
|
||
</taxonomicName>
|
||
in the West Palearctic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |