330 lines
36 KiB
XML
330 lines
36 KiB
XML
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<mods:title id="798B5719B0F9C16AC29F04A278E797A7">From the mud to the tree: phylogeny of Austrolebias killifishes, new generic structure and description of a new species (Cyprinodontiformes: Rivulidae)</mods:title>
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<mods:namePart id="8A06CCE5905000A3A0EE6B5A51AA3072">Alonso, Felipe</mods:namePart>
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<mods:affiliation id="583BBC3C9F2765F88EF3150FF0E2482D">Instituto de Bio y Geociencias del NOA (IBIGEO), National Scientific and Technical Research Council (CONICET) - UNSa, Av. 9 de Julio 14, A 4405 BBA Rosario de Lerma, Salta, Argentina & Killifish Foundation, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="A8481D4A27777959179C051F2470ADEC">Terán, Guillermo Enrique</mods:namePart>
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<mods:affiliation id="BCDE7BDBC8B305C249B1F1A63068F715">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Fundación Miguel Lillo - Unidad Ejecutora Lillo (CONICET), San Miguel de Tucumán, Tucumán, Tucuman Province, Argentina</mods:affiliation>
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<mods:affiliation id="0ABC02BE4F9747E23FCE28E4774B992F">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Sección Ictiología, Departamento de Zoología, Museo Nacional de Historia Natural, Montevideo, Uruguay & Centro Universitario Regional del Este (CURE) Sede Rocha, Rocha, Uruguay</mods:affiliation>
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<mods:namePart id="CC259EB49EF0F6AC2432E22D0C535B3A">Calviño, Pablo</mods:namePart>
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<mods:affiliation id="B4796144EA7C41C869EE00DFDF68D259">Killifish Foundation, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="BDE7F33FFABF34D8F955DB0E2AF008D9">Montes, Martin Miguel</mods:namePart>
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<mods:affiliation id="D7A63D813058209816901A87A0AEC3C2">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Centro de Estudios Parasitológicos y de Vectores - [CCT-CONICET - La Plata] Centro Científico Tecnológico Conicet, La Plata - [Conicet] Consejo Nacional De Investigaciones Científicas y Técnicas, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="6FBBA98FA12FAD3196EF8749FF684170">García, Ignacio Daniel</mods:namePart>
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<mods:affiliation id="BFBD58255B48F3CDC4DAD796BB6F3F1F">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Instituto de Limnología ‘ Dr. Raúl Ringuelet’ (ILPLA) - UNLP-CONICET, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:affiliation id="615BDF62A2EDD4BA525FFF73A3EE907A">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Centro de Estudios Parasitológicos y de Vectores - [CCT-CONICET - La Plata] Centro Científico Tecnológico Conicet, La Plata - [Conicet] Consejo Nacional De Investigaciones Científicas y Técnicas, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:affiliation id="0091E043DADADD8FA48B48A39A5F7B77">Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:affiliation id="4663B1FE3E684472F38E16901E31C690">Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:title id="98BD23942EAC3FAF54A1838EBA10A393">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="8809F4E75C73AE410F9B937AA7F1D917">2023</mods:date>
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<taxonomicName id="4C634D0FFFA2FFF2FBC4F9ECFBD1F966" authority="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa, 2023" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1024,1149,1639,1666]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="286" phylum="Chordata" rank="genus" status="gen. nov.">
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urn:lsid:zoobank.org:act:
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species:
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</emphasis>
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<taxonomicName id="4C634D0FFFA2FFF2FC6FF96CFAB0F91B" authority="Alonso et al., 2018" authorityName="Alonso" authorityYear="2018" box="[939,1308,1767,1791]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="286" phylum="Chordata" rank="species" species="wichi">
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<emphasis id="B917EA9EFFA2FFF2FC6FF96CFBF6F91B" box="[939,1114,1767,1791]" italics="true" pageId="6" pageNumber="286">Austrolebias wichi</emphasis>
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<bibRefCitation id="EFF24B7DFFA2FFF2FB9BF96CFAB0F91B" author="Alonso F & Teran GE & Calvino P" box="[1119,1308,1767,1791]" pageId="6" pageNumber="286" pagination="0196261" refId="ref21540" refString="Alonso F, Teran GE, Calvino P, et al. An endangered new species of seasonal killifish of the genus Austrolebias (Cyprinodontiformes: Aplocheiloidei) from the Bermejo river basin in the Western Chacoan Region. PLoS One 2018; 13: e 0196261." type="journal article" year="2018">
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Alonso
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<emphasis id="B917EA9EFFA2FFF2FB6AF963FB70F91B" box="[1198,1244,1767,1791]" italics="true" pageId="6" pageNumber="286">et al.</emphasis>
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, 2018
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</taxonomicName>
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.
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<subSubSection id="C3796507FFA2FFF3FCEEF8AAFC3EF8AF" lastPageId="7" lastPageNumber="287" pageId="6" pageNumber="286" type="diagnosis">
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Diagnosis:
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<taxonomicName id="4C634D0FFFA2FFF2FC5BF8ABFBB4F8DC" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[927,1048,1824,1848]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="286" phylum="Chordata" rank="genus">Amatolebias</taxonomicName>
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is distinguished from the remaining genera of the
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<taxonomicName id="4C634D0FFFA2FFF2FC7CF8B4FB83F8B3" authorityName="Costa" authorityYear="1998" box="[952,1071,1855,1879]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="286" phylum="Chordata" rank="genus">
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<emphasis id="B917EA9EFFA2FFF2FC7CF8B4FB83F8B3" box="[952,1071,1855,1879]" italics="true" pageId="6" pageNumber="286">Austrolebias</emphasis>
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genus group by the following unique combination of characters in mature dominant males: (1) well-defined and regular vertical dark grey bars only on the anterior portion of the flank (anterior to the anal fin origin); (2) scales on flanks, above anal fin and below dorsal fin leaden bluish to greenish, without wide dark grey border; (3) anal fin with a black band at the distal margin; (4) thin most exterior border of scales on dorsal half portion of flank, below dorsal fin, light grey to brownish grey; and (5) lateral margins of basihyal cartilage laterally expanded.
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</paragraph>
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|
||
<emphasis id="B917EA9EFFA3FFF3FF45FAF0FF74FA77" bold="true" box="[129,216,1403,1427]" pageId="7" pageNumber="287">Figure 3.</emphasis>
|
||
<taxonomicName id="4C634D0FFFA3FFF3FF19FAF0FE97FA77" authorityName="sensu Costa" authorityYear="2008" box="[221,315,1403,1427]" family="Rivulidae" genus="Argolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FF19FAF0FE97FA77" box="[221,315,1403,1427]" italics="true" pageId="7" pageNumber="287">Argolebias</emphasis>
|
||
</taxonomicName>
|
||
diagnostic characters: black blotch on most anterior region of dorsal fin (A, B); fused iridescent marks on distal portion of dorsal fin (C, D); a wide dark grey to black border on most scales on ventral half portion of flank, above anal fin, in mature dominant males (not considering bands; E, F); and iridescent spots on medial distal portion of pectoral fin (G, H).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDC368CFFA3FFF3FF45F95AFC3EF8AF" blockId="7.[129,778,1745,1988]" lastBlockId="7.[825,1475,1530,1867]" pageId="7" pageNumber="287">
|
||
<emphasis id="B917EA9EFFA3FFF3FF45F95AFEF0F90D" box="[129,348,1745,1769]" italics="true" pageId="7" pageNumber="287">Differential diagnosis:</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C634D0FFFA3FFF3FE2DF95AFDCEF90D" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[489,610,1745,1769]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FE2DF95AFDCEF90D" box="[489,610,1745,1769]" italics="true" pageId="7" pageNumber="287">Amatolebias</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other genera of the genus group
|
||
<taxonomicName id="4C634D0FFFA3FFF3FD56F97BFCA5F8EC" authorityName="Costa" authorityYear="1998" box="[658,777,1776,1800]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FD56F97BFCA5F8EC" box="[658,777,1776,1800]" italics="true" pageId="7" pageNumber="287">Austrolebias</emphasis>
|
||
</taxonomicName>
|
||
by having <40 scales in the longitudinal series (vs.> 40 scales in
|
||
<taxonomicName id="4C634D0FFFA3FFF3FF5AF8A4FEBAF8A3" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[158,278,1839,1863]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FF5AF8A4FEBAF8A3" box="[158,278,1839,1863]" italics="true" pageId="7" pageNumber="287">Titanolebias</emphasis>
|
||
</taxonomicName>
|
||
); by the presence of scales in the preopercular region (vs. absent in
|
||
<taxonomicName id="4C634D0FFFA3FFF3FE9EF8C5FE74F882" authorityName="Costa" authorityYear="2008" box="[346,472,1870,1894]" family="Rivulidae" genus="Gymnolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FE9EF8C5FE74F882" box="[346,472,1870,1894]" italics="true" pageId="7" pageNumber="287">Gymnolebias</emphasis>
|
||
</taxonomicName>
|
||
); by the basal and medial region of pectoral fins hyaline in adult males (vs. non-hyaline in
|
||
<taxonomicName id="4C634D0FFFA3FFF3FF45F807FF4AF840" authorityName="sensu Costa" authorityYear="2008" box="[129,230,1932,1956]" family="Rivulidae" genus="Argolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FF45F807FF4AF840" box="[129,230,1932,1956]" italics="true" pageId="7" pageNumber="287">Argolebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FF3FF807FE2FF840" authorityName="Costa" authorityYear="2008" box="[251,387,1932,1956]" family="Rivulidae" genus="Acantholebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FF3FF807FE2FF840" box="[251,387,1932,1956]" italics="true" pageId="7" pageNumber="287">Acantholebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FE5CF807FDA8F840" authorityName="Costa" authorityYear="1998" box="[408,516,1932,1956]" class="Actinopterygii" family="Rivulidae" genus="Megalebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FE5CF807FDA8F840" box="[408,516,1932,1956]" italics="true" pageId="7" pageNumber="287">Megalebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FDDEF807FDD2F840" authorityName="Costa" authorityYear="2008" box="[538,638,1932,1956]" family="Rivulidae" genus="Acrolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FDDEF807FDD2F840" box="[538,638,1932,1956]" italics="true" pageId="7" pageNumber="287">Acrolebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FD57F807FCA5F840" authorityName="Costa" authorityYear="2008" box="[659,777,1932,1956]" family="Rivulidae" genus="Cypholebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FD57F807FCA5F840" box="[659,777,1932,1956]" italics="true" pageId="7" pageNumber="287">Cypholebias</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0FFFA3FFF3FF69F827FEBFF820" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[173,275,1964,1988]" family="Rivulidae" genus="Matilebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FF69F827FEBFF820" box="[173,275,1964,1988]" italics="true" pageId="7" pageNumber="287">Matilebias</emphasis>
|
||
</taxonomicName>
|
||
); by presenting a black band at the distal margin of the anal fin and posterior region of the flanks with white spots vertically aligned in adult males (vs. without black band and white markings in
|
||
<taxonomicName id="4C634D0FFFA3FFF3FBEBF9B2FB04F9B5" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1071,1192,1593,1617]" family="Rivulidae" genus="Garcialebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FBEBF9B2FB04F9B5" box="[1071,1192,1593,1617]" italics="true" pageId="7" pageNumber="287">Garcialebias</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="4C634D0FFFA3FFF3FB3BF9B2FADAF9B5" authorityName="Costa" authorityYear="1998" box="[1279,1398,1593,1617]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFA3FFF3FB3BF9B2FADAF9B5" box="[1279,1398,1593,1617]" italics="true" pageId="7" pageNumber="287">Austrolebias</emphasis>
|
||
</taxonomicName>
|
||
by the urogenital papillae not aưached to the anal fin and no contact organs on anal fin in males [vs. urogenital papillae aưached in
|
||
<taxonomicName id="4C634D0FFFA3FFF3FCFDF91CFB1DF94B" authority="Amato, 1986" authorityName="Amato" authorityYear="1986" box="[825,1201,1687,1711]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="melanoorus">
|
||
<emphasis id="B917EA9EFFA3FFF3FCFDF91CFB8AF94B" box="[825,1062,1687,1711]" italics="true" pageId="7" pageNumber="287">Austrolebias melanoorus</emphasis>
|
||
Amato, 1986
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FB7EF91CFC5FF92B" authority="Serra & Loureiro, 2018" authorityName="Serra & Loureiro" authorityYear="2018" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="queguay">
|
||
<emphasis id="B917EA9EFFA3FFF3FB7EF91CFA2BF94B" box="[1210,1415,1687,1711]" italics="true" pageId="7" pageNumber="287">Austrolebias queguay</emphasis>
|
||
<bibRefCitation id="EFF24B7DFFA3FFF3FA49F91CFC5FF92B" author="Serra WS & Loureiro M" pageId="7" pageNumber="287" pagination="547 - 56" refId="ref25328" refString="Serra WS, Loureiro M. Austrolebias queguay (Cyprinodontiformes, Rivulidae), a new species of annual killifish endemic to the lower Uruguay river basin. Zoosystematics and Evolution 2018; 94: 547 - 56." type="journal article" year="2018">Serra & Loureiro, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FC3BF93DFCC3F90A" authority="Costa & Cheffe, 2005" authorityName="Costa & Cheffe" authorityYear="2005" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="univentripinnis">
|
||
<emphasis id="B917EA9EFFA3FFF3FC3BF93DFAA3F92A" box="[1023,1295,1718,1742]" italics="true" pageId="7" pageNumber="287">Austrolebias univentripinnis</emphasis>
|
||
Costa & Cheffe, 2005
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFA3FFF3FCBFF95DFAB8F90A" authority="Steindachner, 1881" authorityName="Steindachner" authorityYear="1881" box="[891,1300,1750,1774]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="bellottii">
|
||
<emphasis id="B917EA9EFFA3FFF3FCBFF95DFBEFF90A" box="[891,1091,1750,1774]" italics="true" pageId="7" pageNumber="287">Austrolebias bellottii</emphasis>
|
||
Steindachner, 1881
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0FFFA3FFF3FA8FF95DFB48F8E9" authority="Volcan & Severo-Neto, 2019" authorityName="Volcan & Severo-Neto" authorityYear="2019" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="ephemerus">
|
||
<emphasis id="B917EA9EFFA3FFF3FA8FF95DFC0CF8E9" italics="true" pageId="7" pageNumber="287">Austrolebias ephemerus</emphasis>
|
||
<bibRefCitation id="EFF24B7DFFA3FFF3FC68F97EFB48F8E9" author="Volcan MV & Severo-Neto F" box="[940,1252,1781,1805]" pageId="7" pageNumber="287" pagination="541 - 53" refId="ref26068" refString="Volcan MV, Severo-Neto F. Austrolebias ephemerus (Cyprinodontiformes: Rivulidae), a new annual fish from the upper Rio Paraguai basin, Brazilian Chaco. Zootaxa 2019; 4560: 541 - 53." type="journal article" year="2019">Volcan & Severo-Neto, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, and contact organs present on anal fin of
|
||
<taxonomicName id="4C634D0FFFA3FFF3FBE5F89FFAB9F8C8" baseAuthorityName="Huber" baseAuthorityYear="1995" box="[1057,1301,1812,1836]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="vandenbergi">
|
||
<emphasis id="B917EA9EFFA3FFF3FBE5F89FFAB9F8C8" box="[1057,1301,1812,1836]" italics="true" pageId="7" pageNumber="287">Austrolebias vandenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0FFFA3FFF3FA8FF89FFC2DF8AF" authorityName="Nielsen & Pillet" authorityYear="2015" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="species" species="accorsii">
|
||
<emphasis id="B917EA9EFFA3FFF3FA8FF89FFC2DF8AF" italics="true" pageId="7" pageNumber="287">Austrolebias accorsii</emphasis>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFA3FFFCFCFDF8E6FEABFDDB" lastPageId="8" lastPageNumber="288" pageId="7" pageNumber="287" type="discussion">
|
||
<paragraph id="8BDC368CFFA3FFFCFCFDF8E6FEABFDDB" blockId="7.[825,1474,1901,1988]" lastBlockId="8.[113,763,144,575]" lastPageId="8" lastPageNumber="288" pageId="7" pageNumber="287">
|
||
<emphasis id="B917EA9EFFA3FFF3FCFDF8E6FBCCF861" box="[825,1120,1901,1925]" italics="true" pageId="7" pageNumber="287">
|
||
Synapomorphies:
|
||
<taxonomicName id="4C634D0FFFA3FFF3FC23F8E6FBCCF861" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[999,1120,1901,1925]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="287" phylum="Chordata" rank="genus">Amatolebias</taxonomicName>
|
||
</emphasis>
|
||
is recovered as monophyletic, supported by eight molecular synapomorphies and the following six morphological synapomorphies (see Supporting Information, Appendix S5): centre of most scales on ventral half of flank, above anal fin, in dominant mature males (excluding bands, bars and iridescent lines) bluish to greenish leaden (1:7); wide border of most scales on ventral half of flank, above anal fin, in dominant mature males (not considering bands, bars and iridescent lines) absent (2:0); centre of most scales on dorsal half of flank, below dorsal fin, in mature dominant males (not considering bands, bars and iridescent lines) bluish to greenish leaden (6:7); thin outermost edge of scales on dorsal half of flank, below dorsal fin, in dominant mature males (not considering bands, bars and iridescent lines) light grey to brownish grey (10:1); flanks with regular, well-defined vertical dark grey bars (16:1); and dark grey bars only on anterior portion of flank (anterior to anal fin origin) (17:1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFACFFFCFFB5FDEAFE6CFD53" pageId="8" pageNumber="288" type="taxon_list">
|
||
<paragraph id="8BDC368CFFACFFFCFFB5FDEAFE6CFD53" blockId="8.[113,763,609,695]" pageId="8" pageNumber="288">
|
||
<emphasis id="B917EA9EFFACFFFCFFB5FDEAFEBCFD9C" box="[113,272,609,633]" italics="true" pageId="8" pageNumber="288">Included species:</emphasis>
|
||
<taxonomicName id="4C634D0FFFACFFFCFED1FDEAFD2EFD9D" authority="Alonso et al., 2018" authorityName="Alonso" authorityYear="2018" box="[277,642,609,633]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="288" phylum="Chordata" rank="species" species="wichi" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFACFFFCFED1FDEAFE69FD9D" box="[277,453,609,633]" italics="true" pageId="8" pageNumber="288">Amatolebias wichi</emphasis>
|
||
<bibRefCitation id="EFF24B7DFFACFFFCFE0DFDEAFD2EFD9D" author="Alonso F & Teran GE & Calvino P" box="[457,642,609,633]" pageId="8" pageNumber="288" pagination="0196261" refId="ref21540" refString="Alonso F, Teran GE, Calvino P, et al. An endangered new species of seasonal killifish of the genus Austrolebias (Cyprinodontiformes: Aplocheiloidei) from the Bermejo river basin in the Western Chacoan Region. PLoS One 2018; 13: e 0196261." type="journal article" year="2018">
|
||
Alonso
|
||
<emphasis id="B917EA9EFFACFFFCFDD3FDEAFDE9FD9D" box="[535,581,609,633]" italics="true" pageId="8" pageNumber="288">et al.</emphasis>
|
||
, 2018
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFACFFFCFD43FDEAFD5BFD9C" box="[647,759,609,633]" pageId="8" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C634D0FFFACFFFCFFB5FD0BFE48FD7C" authority="Costa et al., 2004" authorityName="Costa" authorityYear="2004" box="[113,484,640,664]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="288" phylum="Chordata" rank="species" species="varzeae" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFACFFFCFFB5FD0BFE94FD7C" box="[113,312,640,664]" italics="true" pageId="8" pageNumber="288">Amatolebias varzeae</emphasis>
|
||
Costa
|
||
<emphasis id="B917EA9EFFACFFFCFEBFFD0AFE04FD7C" box="[379,424,640,664]" italics="true" pageId="8" pageNumber="288">et al.</emphasis>
|
||
, 2004
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFACFFFCFE2CFD0BFDF9FD73" box="[488,597,640,664]" pageId="8" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C634D0FFFACFFFCFD46FD0BFEE0FD53" authority="Huber, 1995" authorityName="Huber" authorityYear="1995" baseAuthorityName="Huber" baseAuthorityYear="1995" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="288" phylum="Chordata" rank="species" species="patriciae" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFACFFFCFD46FD0BFF6AFD53" italics="true" pageId="8" pageNumber="288">Amatolebias patriciae</emphasis>
|
||
Huber, 1995
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFACFFFCFE95FD14FE6CFD53" box="[337,448,671,695]" pageId="8" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFACFFFCFFB5FD53FF17FC69" pageId="8" pageNumber="288" type="etymology">
|
||
<paragraph id="8BDC368CFFACFFFCFFB5FD53FF17FC69" blockId="8.[113,763,728,909]" pageId="8" pageNumber="288">
|
||
<emphasis id="B917EA9EFFACFFFCFFB5FD53FF4DFD14" box="[113,225,728,752]" italics="true" pageId="8" pageNumber="288">Etymology:</emphasis>
|
||
Genus dedicated to Luis H. Amato, for his important contribution to the knowledge of the
|
||
<taxonomicName id="4C634D0FFFACFFFCFDD1FD73FDDBFCF4" box="[533,631,760,784]" family="Rivulidae" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="288" phylum="Chordata" rank="family">Rivulidae</taxonomicName>
|
||
diversity of
|
||
<collectingCountry id="F374761CFFACFFFCFFB5FC93FF66FCD4" box="[113,202,792,816]" name="Uruguay" pageId="8" pageNumber="288">Uruguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F374761CFFACFFFCFF11FC93FE9DFCD4" box="[213,305,792,816]" name="Paraguay" pageId="8" pageNumber="288">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="F374761CFFACFFFCFEA2FC9CFE0DFCCB" box="[358,417,791,815]" name="Brazil" pageId="8" pageNumber="288">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A7F86EFFACFFFCFE74FC9CFDD0FCCB" box="[432,636,791,815]" country="Brazil" name="Rio Grande do Sul" pageId="8" pageNumber="288">Rio Grande do Sul</collectingRegion>
|
||
); composed of his last name and from the Greek
|
||
<emphasis id="B917EA9EFFACFFFCFE32FCBDFD80FCAA" box="[502,556,822,846]" italics="true" pageId="8" pageNumber="288">lebias</emphasis>
|
||
(small fish, a name commonly used to compose generic names of cyprinodontiform fishes).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFACFFFCFFB5FC25FF1FFBE1" pageId="8" pageNumber="288" type="distribution">
|
||
<paragraph id="8BDC368CFFACFFFCFFB5FC25FF1FFBE1" blockId="8.[113,762,942,1029]" pageId="8" pageNumber="288">
|
||
<emphasis id="B917EA9EFFACFFFCFFB5FC25FF43FC22" box="[113,239,942,966]" italics="true" pageId="8" pageNumber="288">Distribution:</emphasis>
|
||
Chacoan region of the
|
||
<collectingCountry id="F374761CFFACFFFCFE3FFC24FDFBFC23" box="[507,599,943,967]" name="Paraguay" pageId="8" pageNumber="288">Paraguay</collectingCountry>
|
||
River basin in
|
||
<collectingCountry id="F374761CFFACFFFCFFB5FC45FF7AFC02" box="[113,214,974,998]" name="Argentina" pageId="8" pageNumber="288">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="F374761CFFACFFFCFEC8FC45FEC4FC02" box="[268,360,974,998]" name="Paraguay" pageId="8" pageNumber="288">Paraguay</collectingCountry>
|
||
and the upper
|
||
<collectingCountry id="F374761CFFACFFFCFDCEFC45FDCFFC02" box="[522,611,974,998]" name="Uruguay" pageId="8" pageNumber="288">Uruguay</collectingCountry>
|
||
River basin in
|
||
<collectingCountry id="F374761CFFACFFFCFFB5FC66FF03FBE1" box="[113,175,1005,1029]" name="Brazil" pageId="8" pageNumber="288">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |