treatments-xml/data/03/CA/87/03CA8795FFB5FFA3FF3F689BFB2D9584.xml

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<mods:title id="A93E2127840DC631B682474A6BC259C7">Two new species of Syzygium (Myrtaceae) from Wawonii Island, Southeastern Sulawesi, Indonesia</mods:title>
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<mods:namePart id="BD49EEADA17085D7219B1903B19D39E1">Sunarti, Siti</mods:namePart>
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<mods:affiliation id="66C41C6AD9FD7E9AE553FA71091E1AF6">Herbarium Bogoriense ”, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Jl. Raya Jakarta-Bogor KM 46, Cibinong, Bogor, Indonesia</mods:affiliation>
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<treatment id="03CA8795FFB5FFA3FF3F689BFB2D9584" ID-DOI="http://doi.org/10.5281/zenodo.13215985" ID-GBIF-Taxon="237657496" ID-Zenodo-Dep="13215985" LSID="urn:lsid:plazi:treatment:03CA8795FFB5FFA3FF3F689BFB2D9584" httpUri="http://treatment.plazi.org/id/03CA8795FFB5FFA3FF3F689BFB2D9584" lastPageId="4" lastPageNumber="91" pageId="1" pageNumber="88">
<subSubSection id="C3796508FFB5FFA6FF3F689BFC8194C7" box="[136,786,1453,1480]" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="8BDC3683FFB5FFA6FF3F689BFC8194C7" blockId="1.[136,1452,1453,1551]" box="[136,786,1453,1480]" pageId="1" pageNumber="88">
<heading id="D09481EFFFB5FFA6FF3F689BFC8194C7" box="[136,786,1453,1480]" fontSize="11" level="2" pageId="1" pageNumber="88" reason="4">
<emphasis id="B917EA91FFB5FFA6FF3F689BFF0D94C7" bold="true" box="[136,158,1453,1479]" pageId="1" pageNumber="88">1.</emphasis>
<taxonomicName id="4C634D00FFB5FFA6FF0A689BFD2194C7" authority="Sunarti &amp; Rugayah" authorityName="Sunarti &amp; Rugayah" box="[189,690,1453,1480]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="lampeapiense" status="sp. nov.">
<emphasis id="B917EA91FFB5FFA6FF0A689BFE4394C8" bold="true" box="[189,464,1453,1480]" italics="true" pageId="1" pageNumber="88">Syzygium lampeapiense</emphasis>
Sunarti &amp; Rugayah
</taxonomicName>
, 
<emphasis id="B917EA91FFB5FFA6FD0A6898FC8194C7" box="[701,786,1454,1479]" italics="true" pageId="1" pageNumber="88">
<taxonomicNameLabel id="A22457EAFFB5FFA6FD0A6898FC8194C7" box="[701,786,1454,1479]" pageId="1" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3796508FFB5FFA6FF3F68E3FD5A970E" pageId="1" pageNumber="88" type="materials_examined">
<paragraph id="8BDC3683FFB5FFA6FF3F68E3FD5A970E" blockId="1.[136,1452,1453,1551]" pageId="1" pageNumber="88">
<collectionCode id="ED72AE46FFB5FFA6FF3F68E3FF5594EA" box="[136,198,1493,1514]" pageId="1" pageNumber="88">TYPE</collectionCode>
:— 
<collectingCountry id="F3747613FFB5FFA6FF5568E3FEFB94EA" box="[226,360,1493,1514]" name="Indonesia" pageId="1" pageNumber="88">INDONESIA</collectingCountry>
, 
<collectingRegion id="49A7F861FFB5FFA6FEC568E3FDBD94EB" box="[370,558,1493,1515]" country="Indonesia" name="Sulawesi Tenggara" pageId="1" pageNumber="88">Southeast Sulawesi</collectingRegion>
, Wawonii Island, Lampeapi, 
<date id="FFDD1043FFB5FFA6FCF068E3FC5C94EA" box="[839,975,1493,1515]" pageId="1" pageNumber="88" value="2004-04-28">28 April 2004</date>
, 
<emphasis id="B917EA91FFB5FFA6FC6E68E3FBF794EA" box="[985,1124,1493,1514]" italics="true" pageId="1" pageNumber="88">Rugayah 1004</emphasis>
(
<typeStatus id="54D88821FFB5FFA6FBC668E3FB5494EB" box="[1137,1223,1493,1515]" pageId="1" pageNumber="88" type="holotype">holotype</typeStatus>
<collectionCode id="ED72AE46FFB5FFA6FB7A68E3FB7D94EA" box="[1229,1262,1493,1514]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="1" pageNumber="88" type="Herbarium">BO</collectionCode>
1993392!, 
<typeStatus id="54D88821FFB5FFA6FAEA68E3FA3F94EB" box="[1373,1452,1493,1515]" pageId="1" pageNumber="88" type="isotype">isotypes</typeStatus>
<collectionCode id="ED72AE46FFB5FFA6FF0A68CFFF4D970E" box="[189,222,1529,1550]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="1" pageNumber="88" type="Herbarium">BO</collectionCode>
1993393!, 1993394!, 
<collectionCode id="ED72AE46FFB5FFA6FE0268CFFE66970E" box="[437,501,1529,1550]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="1" pageNumber="88" type="Herbarium">SING</collectionCode>
!). 
<figureCitation id="13582A06FFB5FFA6FDBE68CFFDF0970E" box="[521,611,1529,1551]" captionStart="FIGURE 1" captionStartId="1.[136,229,1192,1214]" captionTargetBox="[151,1422,469,1150]" captionTargetId="figure-138@1.[151,1436,455,1167]" captionTargetPageId="1" captionText="FIGURE 1. Location of plant exploration and distribution of Syzygium lampeapiense (●) and S. wawoniense (■) P. Sulawesi (sources: Berkas:Sulawesi blank map.png - Wikipedia bahasa Indonesia,ensiklopedia bebas id.m.wikipedia.org); P.wawonii (sources: Bakosurtanal. Edisi 1-1992 &amp; Google Map (02 Maret 2015)." figureDoi="http://doi.org/10.5281/zenodo.13218312" httpUri="https://zenodo.org/record/13218312/files/figure.png" pageId="1" pageNumber="88">Figures 1</figureCitation>
, 
<figureCitation id="13582A06FFB5FFA6FDD868CFFD3E970E" box="[623,685,1529,1550]" captionStart="FIGURE 2" captionStartId="2.[136,229,1952,1974]" captionTargetBox="[171,1416,202,1916]" captionTargetId="figure-23@2.[171,1416,190,1928]" captionTargetPageId="2" captionText="FIGURE 2. Syzygium lampeapiense: flowers (A), fruits (B), lower leaf surface (C), upper leaf surface (D), flower bud (E); S. comosum: lower leaf surface (F), upper leaf surface (G), flower bud (H)." figureDoi="http://doi.org/10.5281/zenodo.13218314" httpUri="https://zenodo.org/record/13218314/files/figure.png" pageId="1" pageNumber="88">2 A–E</figureCitation>
, 
<figureCitation id="13582A06FFB5FFA6FD0068CFFD56970E" box="[695,709,1529,1550]" captionStart="FIGURE 3" captionStartId="3.[136,229,1952,1974]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-17@3.[178,1409,190,1928]" captionTargetPageId="3" captionText="FIGURE 3. Type specimen of Syzygium lampeapiense." figureDoi="http://doi.org/10.5281/zenodo.13218316" httpUri="https://zenodo.org/record/13218316/files/figure.png" pageId="1" pageNumber="88">3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796508FFB5FFA6FF3F6B00FC759925" pageId="1" pageNumber="88" type="diagnosis">
<paragraph id="8BDC3683FFB5FFA6FF3F6B00FC759925" blockId="1.[136,1452,1590,2085]" pageId="1" pageNumber="88">
<emphasis id="B917EA91FFB5FFA6FF3F6B00FEB29750" bold="true" box="[136,289,1590,1616]" pageId="1" pageNumber="88">Diagnosis:—</emphasis>
This species is similar to 
<taxonomicName id="4C634D00FFB5FFA6FDF46B01FCB89751" authority="Widodo" authorityName="Widodo" box="[579,811,1591,1617]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB5FFA6FDF46B01FD589751" box="[579,715,1591,1617]" italics="true" pageId="1" pageNumber="88">S. comosum</emphasis>
Widodo
</taxonomicName>
(in 
<bibRefCitation id="EFF24B72FFB5FFA6FCEC6B01FB509751" author="Widodo, P. &amp; Chikmawati, T." box="[859,1219,1591,1617]" pageId="1" pageNumber="88" refId="ref4904" refString="Widodo, P. &amp; Chikmawati, T. (2016) Six new species of Syzygium (Myrtaceae) from Sumatra. Edinburgh Journal of Botany: 1 &amp; 13. https: // doi. org / 10.1017 / S 0960428616000111" type="journal volume" year="2016">Widodo &amp; Chikmawati 2016: 6</bibRefCitation>
) from Payakumbuh, 
<collectingRegion id="49A7F861FFB5FFA6FF3F6B6DFF7B9775" box="[136,232,1627,1653]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="88">Sumatra</collectingRegion>
, especially in its rheophytic habits and solitary flowers in general, but differs in its young twigs and leaf arrangement, the presence of oil dots on the leaf surfaces, the hypanthium and the pedicel. 
<taxonomicName id="4C634D00FFB5FFA6FBC16B49FA139799" box="[1142,1408,1663,1689]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="lampeapiense">
<emphasis id="B917EA91FFB5FFA6FBC16B49FA139799" box="[1142,1408,1663,1689]" italics="true" pageId="1" pageNumber="88">Syzygium lampeapiense</emphasis>
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has slightly sub-terete young twigs with internodes are 
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long and leaves loosely arranged along the branches, the petioles usually forming angles about 60° with the branches, while 
<taxonomicName id="4C634D00FFB5FFA6FC0D6BF1FBD197E1" box="[954,1090,1735,1761]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB5FFA6FC0D6BF1FC5B97E1" box="[954,968,1735,1761]" italics="true" pageId="1" pageNumber="88">S</emphasis>
. 
<emphasis id="B917EA91FFB5FFA6FC6F6BFEFBD197E1" box="[984,1090,1736,1761]" italics="true" pageId="1" pageNumber="88">comosum</emphasis>
</taxonomicName>
has 4-angled young twigs with internodes 
<quantity id="4C9B9B66FFB5FFA6FEB16BDDFEE09605" box="[262,371,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="88" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long and leaves densely arranged towards the tip of the branches, the petioles forming angles about 30° with the branches. The upper and lower leaf surfaces of 
<taxonomicName id="4C634D00FFB5FFA6FC356A39FBA99629" box="[898,1082,1807,1833]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="lampeapiense">
<emphasis id="B917EA91FFB5FFA6FC356A39FBA99629" box="[898,1082,1807,1833]" italics="true" pageId="1" pageNumber="88">S. lampeapiense</emphasis>
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have numerous oil dots (10–15/ mm² lower and 27–30/mm² upper), while 
<taxonomicName id="4C634D00FFB5FFA6FDDF6A05FD74964D" box="[616,743,1843,1869]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB5FFA6FDDF6A05FD74964D" box="[616,743,1843,1869]" italics="true" pageId="1" pageNumber="88">S.comosum</emphasis>
</taxonomicName>
has few oil dots (0–7/mm² lower, and 
<quantity id="4C9B9B66FFB5FFA6FB136A05FA83964F" box="[1188,1296,1843,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="88" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
² upper). This species has more flexible leaves, while 
<taxonomicName id="4C634D00FFB5FFA6FD896A61FD509671" box="[574,707,1879,1905]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB5FFA6FD896A61FD509671" box="[574,707,1879,1905]" italics="true" pageId="1" pageNumber="88">S. comosum</emphasis>
</taxonomicName>
has more rigid leaves. Flowers of 
<taxonomicName id="4C634D00FFB5FFA6FBF56A61FB649671" box="[1090,1271,1879,1905]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="lampeapiense">
<emphasis id="B917EA91FFB5FFA6FBF56A61FB649671" box="[1090,1271,1879,1905]" italics="true" pageId="1" pageNumber="88">S. lampeapiense</emphasis>
</taxonomicName>
bear pedicels to 
<quantity id="4C9B9B66FFB5FFA6FF3F6A4DFF469695" box="[136,213,1915,1942]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="88" unit="cm" value="5.5">5.5 cm</quantity>
long and present obconical hypanthium, while those of 
<taxonomicName id="4C634D00FFB5FFA6FCE66A4DFC449695" box="[849,983,1915,1941]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB5FFA6FCE66A4DFC449695" box="[849,983,1915,1941]" italics="true" pageId="1" pageNumber="88">S. comosum</emphasis>
</taxonomicName>
have pedicels to 
<quantity id="4C9B9B66FFB5FFA6FB2A6A4DFB479695" box="[1181,1236,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="88" unit="cm" value="1.0">1 cm</quantity>
long and a funnel-shaped hypanthium. It is also similar to the Bornean rheophytic species 
<taxonomicName id="4C634D00FFB5FFA6FC126AA9FE3896DD" authority="(Korthals 1847: 199) Merril &amp; L. M. Perry (1939: 166)" authorityName="Merril &amp; L. M. Perry" authorityPageNumber="166" authorityYear="1939" baseAuthorityName="Korthals" baseAuthorityPageNumber="199" baseAuthorityYear="1847" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="medium">
<emphasis id="B917EA91FFB5FFA6FC126AA9FBFD96B9" box="[933,1134,1951,1977]" italics="true" pageId="1" pageNumber="88">Syzygium medium</emphasis>
(
<bibRefCitation id="EFF24B72FFB5FFA6FBCB6AA9FAC696B9" author="Korthals, P. W." box="[1148,1365,1951,1977]" pageId="1" pageNumber="88" pagination="185 - 206" refId="ref4268" refString="Korthals, P. W. (1847) Myrtaceae. Nederlandsch Kruidkundig Archief 1: 185 - 206. [https: // www. biodiversitylibrary. org / item / 106826 # page / 205 / mode / 1 up]" type="journal article" year="1847">Korthals 1847: 199</bibRefCitation>
) Merril &amp; L.M.Perry (1939: 166)
</taxonomicName>
, but differs by its leaves 4–11.5 × 
<quantity id="4C9B9B66FFB5FFA6FC806AF5FC3296DD" box="[823,929,1987,2014]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="88" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
with one intramarginal vein and flowers with obconical hypanthium and styles 
<quantity id="4C9B9B66FFB5FFA6FDB36AD1FD159901" box="[516,646,2023,2050]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="1" pageNumber="88" unit="cm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 cm</quantity>
, while 
<taxonomicName id="4C634D00FFB5FFA6FD6E6AD1FCDC9901" authorityName="Merril &amp; L. M. Perry" authorityYear="1939" baseAuthorityName="Korthals" baseAuthorityYear="1847" box="[729,847,2023,2049]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="medium">
<emphasis id="B917EA91FFB5FFA6FD6E6AD1FCDC9901" box="[729,847,2023,2049]" italics="true" pageId="1" pageNumber="88">S. medium</emphasis>
</taxonomicName>
has leaves 0.6–11 x 
<quantity id="4C9B9B66FFB5FFA6FB896AD1FB2F9901" box="[1086,1212,2023,2049]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="1" pageNumber="88" unit="cm" value="0.75" valueMax="1.1" valueMin="0.4">0.4–1.1 cm</quantity>
with 2 intramarginal veins and flowers with hypanthium funnel-shaped and styles to 
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long.
</paragraph>
</subSubSection>
<caption id="DF1C660BFFB6FFA5FF3F6A96FD4A96DA" ID-DOI="http://doi.org/10.5281/zenodo.13218314" ID-Zenodo-Dep="13218314" httpUri="https://zenodo.org/record/13218314/files/figure.png" pageId="2" pageNumber="89" startId="2.[136,229,1952,1974]" targetBox="[171,1416,202,1916]" targetPageId="2" targetType="figure">
<paragraph id="8BDC3683FFB6FFA5FF3F6A96FD4A96DA" blockId="2.[136,1452,1952,2010]" pageId="2" pageNumber="89">
<emphasis id="B917EA91FFB6FFA5FF3F6A96FF6D96B5" bold="true" box="[136,254,1952,1974]" pageId="2" pageNumber="89">FIGURE 2.</emphasis>
<taxonomicName id="4C634D00FFB6FFA5FEB36A97FE7A96B5" box="[260,489,1952,1974]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="lampeapiense">
<emphasis id="B917EA91FFB6FFA5FEB36A97FE7A96B5" box="[260,489,1952,1974]" italics="true" pageId="2" pageNumber="89">Syzygium lampeapiense</emphasis>
</taxonomicName>
: flowers (A), fruits (B), lower leaf surface (C), upper leaf surface (D), flower bud (E); 
<taxonomicName id="4C634D00FFB6FFA5FA856A97FA3696B6" box="[1330,1445,1953,1974]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="comosum">
<emphasis id="B917EA91FFB6FFA5FA856A97FA3696B6" box="[1330,1445,1953,1974]" italics="true" pageId="2" pageNumber="89">S. comosum</emphasis>
</taxonomicName>
: lower leaf surface (F), upper leaf surface (G), flower bud (H).
</paragraph>
</caption>
<caption id="DF1C660BFFB7FFA4FF3F6A96FD0C96B5" ID-DOI="http://doi.org/10.5281/zenodo.13218316" ID-Zenodo-Dep="13218316" box="[136,671,1952,1974]" httpUri="https://zenodo.org/record/13218316/files/figure.png" pageId="3" pageNumber="90" startId="3.[136,229,1952,1974]" targetBox="[178,1409,190,1928]" targetPageId="3" targetType="figure">
<paragraph id="8BDC3683FFB7FFA4FF3F6A96FD0C96B5" blockId="3.[136,671,1952,1974]" box="[136,671,1952,1974]" pageId="3" pageNumber="90">
<emphasis id="B917EA91FFB7FFA4FF3F6A96FF6D96B5" bold="true" box="[136,254,1952,1974]" pageId="3" pageNumber="90">FIGURE 3.</emphasis>
Type specimen of 
<taxonomicName id="4C634D00FFB7FFA4FE046A97FD0896B5" box="[435,667,1952,1974]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="90" phylum="Tracheophyta" rank="species" species="lampeapiense">Syzygium lampeapiense</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3796508FFB0FFA3FF0A6DA8FECB9368" pageId="4" pageNumber="91" type="description">
<paragraph id="8BDC3683FFB0FFA3FF0A6DA8FECB9368" blockId="4.[136,1452,158,1157]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6DA8FEFE91B8" bold="true" box="[189,365,158,184]" pageId="4" pageNumber="91">Description:—</emphasis>
Rheophytic tree to 
<quantity id="4C9B9B66FFB0FFA3FDF96DA8FDEE91B9" box="[590,637,158,185]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="91" unit="m" value="5.0">5 m</quantity>
tall, diameter at breast height (dbh) 
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. Twigs slightly flat-terete. Leaves opposite, decussate; blades elliptic to narrowly elliptic, (4–)6–11.5 × (1.5–) 1.7–2.5(–3) cm; apex acuminatecuspidate (tip 
<quantity id="4C9B9B66FFB0FFA3FE9C6DD0FE0F9000" box="[299,412,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="4" pageNumber="91" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
), base attenuate-cuneate; midrib on the upper surface slightly raised along a furrow, raised on the lower surface; secondary veins 9–12 pairs, straight and curved, forming an angle of 40° with midrib, raised on the lower surface; upper surface has 27–30/mm² and lower surface has 10–15/ mm² oil dots; intramarginal vein 1, ca. 
<quantity id="4C9B9B66FFB0FFA3FF3F6C64FF5A906D" box="[136,201,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="91" unit="mm" value="1.0">1 mm</quantity>
from the leaf margin; petioles 
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long, 
<quantity id="4C9B9B66FFB0FFA3FD7D6C64FC98906D" box="[714,779,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="91" unit="mm" value="1.0">1 mm</quantity>
thick, on the upper surface slightly grooved. Inflorescences terminal or axillary, mostly uniflorous; pedicels 
<quantity id="4C9B9B66FFB0FFA3FD1F6C40FC879091" box="[680,788,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.5" metricValueMin="3.0" pageId="4" pageNumber="91" unit="cm" value="4.25" valueMax="5.5" valueMin="3.0">3–5.5 cm</quantity>
. Flower buds obconical (
<quantity id="4C9B9B66FFB0FFA3FB986C40FB399091" box="[1071,1194,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="4" pageNumber="91" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
long, 
<quantity id="4C9B9B66FFB0FFA3FB466C40FACF9091" box="[1265,1372,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="4" pageNumber="91" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
diam.) with oil dots on outer surface. Flowers with hypanthium obconical, 
<quantity id="4C9B9B66FFB0FFA3FC376CACFC6A90B5" box="[896,1017,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="4" pageNumber="91" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">13–17 mm</quantity>
long, with numerous oil dots, greyishred; sepals four, free, to 5 × 
<quantity id="4C9B9B66FFB0FFA3FE736C89FD9B90D9" box="[452,520,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="91" unit="mm" value="7.0">7 mm</quantity>
, obovate, persistent along anthesis, membranous, glandulose in the medial portion; petals four, free, to 12 × 
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, membranous, glandulose in the medial portion, white, soon falling after anthesis. Stamens numerous (
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one flower counted); filaments to 
<quantity id="4C9B9B66FFB0FFA3FC8B6F30FC1A9321" box="[828,905,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="91" unit="cm" value="2.5">2.5 cm</quantity>
long, white; anthers basifixed, white. Style 
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, greenish. Ovary 2–locular. Fruits unripe, globose, 
<quantity id="4C9B9B66FFB0FFA3FC936F1CFC1D9345" box="[804,910,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="4" pageNumber="91" unit="cm" value="1.8" valueMax="2.0" valueMin="1.6">1.6–2 cm</quantity>
diam.; greyish-rose until yellowish green, with numerous oil dots.
</paragraph>
</subSubSection>
<subSubSection id="C3796508FFB0FFA3FF0A6F44FE6F9264" pageId="4" pageNumber="91" type="distribution">
<paragraph id="8BDC3683FFB0FFA3FF0A6F44FE4593F8" blockId="4.[136,1452,158,1157]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6F44FEE6938C" bold="true" box="[189,373,626,652]" pageId="4" pageNumber="91">Distribution:—</emphasis>
Wawonii Island (Lampeapi, Wungkolo, Dompo-dompo Jaya), 
<collectingRegion id="49A7F861FFB0FFA3FBFE6F44FABB938C" box="[1097,1320,626,652]" country="Indonesia" name="Sulawesi Tenggara" pageId="4" pageNumber="91">Southeast Sulawesi</collectingRegion>
, 
<collectingCountry id="F3747613FFB0FFA3FA806F44FA3B938C" box="[1335,1448,626,652]" name="Indonesia" pageId="4" pageNumber="91">Indonesia</collectingCountry>
. Besides the specimens examined, a photograph taken by Wiriadinata along the riverbanks of Roko-roko in the Dompo-dompo Jaya area identified by him as 
<taxonomicName id="4C634D00FFB0FFA3FDF16F8DFD4A93D5" box="[582,729,699,725]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B917EA91FFB0FFA3FDF16F8DFD2393D5" box="[582,688,699,725]" italics="true" pageId="4" pageNumber="91">Syzygium</emphasis>
sp.
</taxonomicName>
(Wiriadinata &amp; Hamzah 2006: 49) represents the species here described as 
<emphasis id="B917EA91FFB0FFA3FEAE6FE9FE4593F8" box="[281,470,734,761]" italics="true" pageId="4" pageNumber="91">
<taxonomicName id="4C634D00FFB0FFA3FEAE6FE9FE4093F8" box="[281,467,734,761]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="lampeapiense">S. lampeapiense</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A6E34FCFD921C" blockId="4.[136,1452,158,1157]" box="[189,878,770,797]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6E34FE41921C" bold="true" box="[189,466,770,796]" pageId="4" pageNumber="91">Habitat and ecology:—</emphasis>
vegetation on riverbanks along river.
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A6E10FDA59240" blockId="4.[136,1452,158,1157]" box="[189,566,806,832]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6E10FE609240" bold="true" box="[189,499,806,832]" pageId="4" pageNumber="91">Flowering and fruiting:—</emphasis>
April.
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A6E7CFE6F9264" blockId="4.[136,1452,158,1157]" box="[189,508,842,869]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6E7CFE229264" bold="true" box="[189,433,842,868]" pageId="4" pageNumber="91">Vernacular name:—</emphasis>
pe api.
</paragraph>
</subSubSection>
<subSubSection id="C3796508FFB0FFA3FF0A6E58FB2D9584" pageId="4" pageNumber="91" type="etymology">
<paragraph id="8BDC3683FFB0FFA3FF0A6E58FF6692AC" blockId="4.[136,1452,158,1157]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6E58FEF79288" bold="true" box="[189,356,878,904]" pageId="4" pageNumber="91">Etymology:—</emphasis>
The specific epithet refers to the name of a village in Wawonii island where the species was first collected.
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A6E80FF0C953C" blockId="4.[136,1452,158,1157]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6E80FD0B92D0" bold="true" box="[189,664,950,976]" pageId="4" pageNumber="91">Preliminary conservation assessment:—</emphasis>
VU B2a (Vulnerable). This species is found in a small island, distributed at only three fragmented locations. Even though the species still has a large population (growing along the Lampeapi river site) and there was no threat observed at that moment, nickel mining may represent a possible future threat for it.
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A6970FA9B9560" blockId="4.[136,1452,158,1157]" box="[189,1288,1094,1120]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A6970FEDC9560" bold="true" box="[189,335,1094,1120]" pageId="4" pageNumber="91">
<typeStatus id="54D88821FFB0FFA3FF0A6970FEBE9560" box="[189,301,1094,1120]" pageId="4" pageNumber="91" type="paratype">Paratype</typeStatus>
:—
</emphasis>
<collectingCountry id="F3747613FFB0FFA3FEE76970FE789560" box="[336,491,1094,1120]" name="Indonesia" pageId="4" pageNumber="91">INDONESIA</collectingCountry>
, Wawonii Island, Wungkolo, 
<date id="FFDD1043FFB0FFA3FC816970FC479560" box="[822,980,1094,1120]" pageId="4" pageNumber="91" value="2006-04-30">30 April 2006</date>
, 
<emphasis id="B917EA91FFB0FFA3FC686970FB2B9560" box="[991,1208,1094,1120]" italics="true" pageId="4" pageNumber="91">Arief Hidayat 1313</emphasis>
(BO!).
</paragraph>
<paragraph id="8BDC3683FFB0FFA3FF0A695CFB2D9584" blockId="4.[136,1452,158,1157]" box="[189,1214,1130,1157]" pageId="4" pageNumber="91">
<emphasis id="B917EA91FFB0FFA3FF0A695CFE889584" bold="true" box="[189,283,1130,1156]" pageId="4" pageNumber="91">Note:—</emphasis>
<taxonomicName id="4C634D00FFB0FFA3FEAC695DFDB69584" box="[283,549,1130,1157]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="lampeapiense">
<emphasis id="B917EA91FFB0FFA3FEAC695DFDB69584" box="[283,549,1130,1157]" italics="true" pageId="4" pageNumber="91">Syzygium lampeapiense</emphasis>
</taxonomicName>
was listed in 
<bibRefCitation id="EFF24B72FFB0FFA3FD08695CFC7E9584" author="Rugayah &amp; Sunarti, S. &amp; Sulistiarini, D. &amp; Hidayat, A. &amp; Rahayu, M." box="[703,1005,1130,1157]" pageId="4" pageNumber="91" refId="ref4526" refString="Rugayah, Sunarti, S., Sulistiarini, D., Hidayat, A. &amp; Rahayu, M. (2015) Daftar Jenis Tumbuhan di Pulau Wawonii, Sulawesi Tenggara. LIPI Press, Jakarta, 363 pp." type="book" year="2015">
Rugayah 
<emphasis id="B917EA91FFB0FFA3FC9D695DFCF19584" box="[810,866,1130,1156]" italics="true" pageId="4" pageNumber="91">et al.</emphasis>
(2015: 206)
</bibRefCitation>
as 
<taxonomicName id="4C634D00FFB0FFA3FBA5695DFB2B9584" box="[1042,1208,1130,1157]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="undefined-1">
<emphasis id="B917EA91FFB0FFA3FBA5695DFBEF9585" box="[1042,1148,1131,1157]" italics="true" pageId="4" pageNumber="91">Syzygium</emphasis>
sp. 1
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>