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<document id="788CC86323DDF0714BDA5BD6BFF58852" ID-CLB-Dataset="113070" ID-DOI="10.1093/zoolinnean/zlab043" ID-GBIF-Dataset="e070d52f-d008-4f08-988a-a8b17c27418f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6959516" ID-ZooBank="1C89556A-12CB-43B7-9B49-E02EFF1543D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1653991014928" checkinUser="plazi" docAuthor="Medrano, Miguel, Kury, Adriano Brilhante &amp; Mendes, Amanda Cruz" docDate="2022" docId="03CA878CFF9DFFF3FC1DF99BFDA527DE" docLanguage="en" docName="zlab043.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Protus Simon 1879" docType="treatment" docVersion="5" lastPageNumber="626" masterDocId="FFF3FFF4FFB5FFDAFFFEFFB7FFA72171" masterDocTitle="Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)" masterLastPageNumber="672" masterPageNumber="585" pageNumber="625" updateTime="1699338404186" updateUser="ExternalLinkService">
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<mods:title id="D79648EBD8615460D2104A63CDD2E9D9">Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)</mods:title>
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<mods:namePart id="97562758FA278860E8368675639EB1DC">Medrano, Miguel</mods:namePart>
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<mods:namePart id="7BBC3E131A01B5BD91C477C52C772C28">Kury, Adriano Brilhante</mods:namePart>
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<mods:namePart id="1959CE2CBC16B92F42CBAA9CA406B101">Mendes, Amanda Cruz</mods:namePart>
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<mods:title id="95C59321B4E94EEE5E0AC945545764DD">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="C2EA5A525DE11B49312B1D5733BB3ED0">2022</mods:date>
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<mods:number id="67A34FB3FE265C19B30ADC84E5CE6E73">2021-08-30</mods:number>
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<mods:number id="AB48E02F46914E1575442CACA4829596">195</mods:number>
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<mods:identifier id="9A80DF265652613DEC42420FEC228828" type="DOI">10.1093/zoolinnean/zlab043</mods:identifier>
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<mods:identifier id="10A9669F442CF1279B2756DC73C896E1" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03CA878CFF9DFFF3FC1DF99BFDA527DE" ID-DOI="http://doi.org/10.5281/zenodo.6597980" ID-GBIF-Taxon="198579189" ID-Zenodo-Dep="6597980" LSID="urn:lsid:plazi:treatment:03CA878CFF9DFFF3FC1DF99BFDA527DE" httpUri="http://treatment.plazi.org/id/03CA878CFF9DFFF3FC1DF99BFDA527DE" lastPageId="41" lastPageNumber="626" pageId="40" pageNumber="625">
<subSubSection id="C3796511FF9DFFF2FC1DF99BFB712735" box="[995,1238,1579,1604]" pageId="40" pageNumber="625" type="nomenclature">
<paragraph id="8BDC369AFF9DFFF2FC1DF99BFB712735" blockId="40.[995,1238,1579,1604]" box="[995,1238,1579,1604]" pageId="40" pageNumber="625">
<heading id="D09481F6FF9DFFF2FC1DF99BFB712735" box="[995,1238,1579,1604]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="625" reason="2">
<taxonomicName id="4C634D19FF9DFFF2FC1DF99BFB712735" ID-CoL="7PVZ3" authority="SIMON, 1879" authorityName="Simon" authorityYear="1879" box="[995,1238,1579,1604]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="625" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9DFFF2FC1DF99BFB982732" box="[995,1087,1580,1603]" italics="true" pageId="40" pageNumber="625">PROTUS</emphasis>
<bibRefCitation id="EFF24B6BFF9DFFF2FBB8F99CFB712735" author="Simon E" box="[1094,1238,1579,1604]" pageId="40" pageNumber="625" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">SIMON, 1879</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3796511FF9DFFF2FCD7F9E3FBE42719" box="[809,1091,1619,1641]" pageId="40" pageNumber="625" type="reference_group">
<paragraph id="8BDC369AFF9DFFF2FCD7F9E3FBE42719" blockId="40.[809,1091,1619,1641]" box="[809,1091,1619,1641]" pageId="40" pageNumber="625">
<heading id="D09481F6FF9DFFF2FCD7F9E3FBE42719" box="[809,1091,1619,1641]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="625" reason="2">
<treatmentCitationGroup id="AB7311B4FF9DFFF2FCD7F9E3FBE42719" box="[809,1091,1619,1641]" pageId="40" pageNumber="625">
<treatmentCitation id="0AC2108BFF9DFFF2FCD7F9E3FB992719" author="Simon E" box="[809,1086,1619,1641]" page="193" pageId="40" pageNumber="625" year="1879">
<taxonomicName id="4C634D19FF9DFFF2FCD7F9E3FB992719" ID-CoL="7PVZ3" authority="Simon, 1879: 193" authorityName="Simon" authorityPageNumber="193" authorityYear="1879" box="[809,1086,1619,1641]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="625" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9DFFF2FCD7F9E3FCD42718" box="[809,883,1620,1641]" italics="true" pageId="40" pageNumber="625">Protus</emphasis>
<bibRefCitation id="EFF24B6BFF9DFFF2FC84F9E4FB992719" author="Simon E" box="[890,1086,1619,1641]" pageId="40" pageNumber="625" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879: 193</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3796511FF9DFFF2FCD7F93FFA3527EC" box="[809,1426,1671,1693]" pageId="40" pageNumber="625" type="type_taxon">
<paragraph id="8BDC369AFF9DFFF2FCD7F93FFA3527EC" blockId="40.[809,1426,1671,1693]" box="[809,1426,1671,1693]" pageId="40" pageNumber="625">
<emphasis id="B917EA88FF9DFFF2FCD7F93FFC1127EC" box="[809,950,1672,1693]" italics="true" pageId="40" pageNumber="625">
<typeStatus id="54D88838FF9DFFF2FCD7F93FFCF927EC" box="[809,862,1672,1693]" pageId="40" pageNumber="625">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C634D19FF9DFFF2FC44F93FFB5327ED" ID-CoL="7YHWG" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[954,1268,1671,1693]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="625" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B917EA88FF9DFFF2FC44F93FFBC227ED" box="[954,1125,1671,1693]" italics="true" pageId="40" pageNumber="625">Protus insolens</emphasis>
<bibRefCitation id="EFF24B6BFF9DFFF2FB94F930FB5327ED" author="Simon E" box="[1130,1268,1671,1693]" pageId="40" pageNumber="625" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FF9DFFF2FCD7F90CFC582781" pageId="40" pageNumber="625" type="etymology">
<paragraph id="8BDC369AFF9DFFF2FCD7F90CFC582781" blockId="40.[809,1426,1723,1776]" pageId="40" pageNumber="625">
<emphasis id="B917EA88FF9DFFF2FCD7F90CFBAF27A0" box="[809,1032,1723,1745]" italics="true" pageId="40" pageNumber="625">
Etymology:
<taxonomicName id="4C634D19FF9DFFF2FC42F90BFBAF27A0" authorityName="Simon" authorityYear="1879" box="[956,1032,1724,1745]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="625" phylum="Arthropoda" rank="genus">Protus</taxonomicName>
</emphasis>
is a character of Plutarch´s Dion. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FF9DFFF3FCD7F8AEFE15240C" lastPageId="41" lastPageNumber="626" pageId="40" pageNumber="625" type="diagnosis">
<paragraph id="8BDC369AFF9DFFF3FCD7F8AEFE15240C" blockId="40.[809,1425,1817,1869]" lastBlockId="41.[163,780,1077,1405]" lastPageId="41" lastPageNumber="626" pageId="40" pageNumber="625">
<emphasis id="B917EA88FF9DFFF2FCD7F8AEFC38265F" box="[809,927,1817,1838]" italics="true" pageId="40" pageNumber="625">Diagnosis:</emphasis>
It differs from the other
<taxonomicName id="4C634D19FF9DFFF2FB34F8AEFA37265E" authorityName="Roewer" authorityYear="1923" box="[1226,1424,1817,1839]" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="596" rank="subFamily" subFamily="Discosomaticinae">Discosomaticinae</taxonomicName>
by having two white tubercles in area III (
<figureCitation id="13582A1FFF9DFFF2FAF2F880FA2E263D" box="[1292,1417,1847,1869]" captionStart="Figure 31" captionStartId="42.[145,225,1671,1693]" captionTargetBox="[149,757,856,1632]" captionTargetId="figure-329@42.[149,757,856,1632]" captionTargetPageId="42" captionText="Figure 31. Protus bolivari, male (USNMENT 01538074). A, dorsal view. B, lateral view. C, frontal view. D, panoramic dorsal view. E, ventral view. Scale bars: 1 mm (A–C, E), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.6959594" httpUri="https://zenodo.org/record/6959594/files/figure.png" pageId="40" pageNumber="625">Fig. 31B, C</figureCitation>
) and by having the dorsal process of glans slender and acuminate (
<figureCitation id="13582A1FFF9CFFF3FED8FBE4FE2E2518" box="[294,393,1107,1129]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="41" pageNumber="626">Figs 35G</figureCitation>
,
<figureCitation id="13582A1FFF9CFFF3FE6CFBE3FE672518" box="[402,448,1108,1129]" captionStart="Figure 41" captionStartId="53.[163,242,1667,1689]" captionTargetBox="[163,1443,196,1626]" captionTargetId="figure-25@53.[163,1443,195,1627]" captionTargetPageId="53" captionText="Figure 41. SEM images of some structures of Protus speciosus, male (ICN-Ao- 467). A–C, penis: A, dorsal view; B, lateral view; C, ventral view. D, left cheliceral hand, frontal view. E, left basichelicerite, dorsal view. F, left pedipalpus, mesal view. G, same, dorsal view. H, same, detail of tarsus, ventral view. Scale bars: 100 μm (A–C, E, H), 200 μm (D), 500 μm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.6959618" httpUri="https://zenodo.org/record/6959618/files/figure.png" pageId="41" pageNumber="626">41A</figureCitation>
). Some species have the facies of
<taxonomicName id="4C634D19FF9CFFF3FF41FBC5FE9625F6" authorityName="Roewer" authorityYear="1917" box="[191,305,1138,1159]" class="Arachnida" family="Cosmetidae" genus="Sibambea" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FF41FBC5FE9625F6" box="[191,305,1138,1159]" italics="true" pageId="41" pageNumber="626">Sibambea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D19FF9CFFF3FE95FBC4FE6B25F9" authorityName="Medrano, Kury &amp; Mendes" authorityYear="2022" box="[363,460,1139,1160]" class="Insecta" family="Flatidae" genus="Kayania" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FE95FBC4FE6B25F9" box="[363,460,1139,1160]" italics="true" pageId="41" pageNumber="626">Kayania</emphasis>
</taxonomicName>
because of the coloration of coxa IV with yellow spots (
<figureCitation id="13582A1FFF9CFFF3FE2FFB26FD9325D7" box="[465,564,1169,1191]" captionStart="Figure 33" captionStartId="44.[145,223,1096,1118]" captionTargetBox="[225,1345,195,1056]" captionTargetId="figure-361@44.[225,1345,195,1056]" captionTargetPageId="44" captionText="Figure 33. Protus distinctus, male (CAS AK 105).A, dorsal view. B, lateral view. C, panoramic dorsal view.Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959598" httpUri="https://zenodo.org/record/6959598/files/figure.png" pageId="41" pageNumber="626">Figs 33B</figureCitation>
,
<figureCitation id="13582A1FFF9CFFF3FDBFFB26FDCA25D7" box="[577,621,1169,1190]" captionStart="Figure 44" captionStartId="59.[163,243,1126,1148]" captionTargetBox="[243,1363,195,1086]" captionTargetId="figure-422@59.[243,1363,195,1086]" captionTargetPageId="59" captionText="Figure 44. Sibambea rotunda, male (CAS AK 003). A, dorsal view. B, lateral view. C, panoramic dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959624" httpUri="https://zenodo.org/record/6959624/files/figure.png" pageId="41" pageNumber="626">44C</figureCitation>
), for having a yellowish border in anterior margin of DS (
<figureCitation id="13582A1FFF9CFFF3FD74FB18FD7B25B4" box="[650,732,1199,1221]" captionStart="Figure 25" captionStartId="37.[163,241,1855,1877]" captionTargetBox="[324,1282,1377,1815]" captionTargetId="figure-529@37.[323,1283,1375,1816]" captionTargetPageId="37" captionText="Figure 25. Kayania compressicoxa, schematic female illustration modified from Avram &amp; Soares (1983). A, dorsal view. B, ovipositor and detail of macrosetae. C, left chelicera, mesal view. D, right pedipalpus, mesal view." figureDoi="http://doi.org/10.5281/zenodo.6959580" httpUri="https://zenodo.org/record/6959580/files/figure.png" pageId="41" pageNumber="626">Figs 25</figureCitation>
,
<figureCitation id="13582A1FFF9CFFF3FD14FB07FCA225B4" box="[746,773,1200,1221]" captionStart="Figure 37" captionStartId="49.[164,244,1658,1680]" captionTargetBox="[243,1363,195,1618]" captionTargetId="figure-97@49.[243,1363,195,1618]" captionTargetPageId="49" captionText="Figure 37. Protus marllusi. A–B, Male holotype (MNRJ 7602): A, dorsal view; B, lateral view. C–E, variation of spots of dorsal scutum, female paratypes, dorsal view: C. MNRJ 4842. D. MNRJ 7602. E. MNRJ 2173. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959608" httpUri="https://zenodo.org/record/6959608/files/figure.png" pageId="41" pageNumber="626">37</figureCitation>
,
<figureCitation id="13582A1FFF9CFFF3FF5DFB79FF772592" box="[163,208,1230,1251]" captionStart="Figure 42" captionStartId="55.[163,244,1159,1181]" captionTargetBox="[163,1443,195,1119]" captionTargetId="figure-342@55.[163,1443,195,1119]" captionTargetPageId="55" captionText="Figure 42. Sibambea cincta, male (MPUJ_ENT 644772). A, dorsal view. B, panoramic dorsal view. C, lateral view. D, posterior view. E, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959620" httpUri="https://zenodo.org/record/6959620/files/figure.png" pageId="41" pageNumber="626">42A</figureCitation>
) and for the absence of constriction I in the outline of DS in dorsal view. It differs from
<taxonomicName id="4C634D19FF9CFFF3FDD6FB5AFD3F2473" authorityName="Roewer" authorityYear="1917" box="[552,664,1261,1282]" class="Arachnida" family="Cosmetidae" genus="Sibambea" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FDD6FB5AFD3F2473" box="[552,664,1261,1282]" italics="true" pageId="41" pageNumber="626">Sibambea</emphasis>
</taxonomicName>
by having a long VP of penis (
<figureCitation id="13582A1FFF9CFFF3FE89FABCFE722450" box="[375,469,1291,1313]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="41" pageNumber="626">Fig. 35G</figureCitation>
, except for
<taxonomicName id="4C634D19FF9CFFF3FDADFABBFD7D2451" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Avram &amp; Soares" baseAuthorityYear="1983" box="[595,730,1291,1313]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B917EA88FF9CFFF3FDADFABBFD7D2451" box="[595,730,1291,1313]" italics="true" pageId="41" pageNumber="626">P. distinctus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D19FF9CFFF3FF5DFA9CFEB2244E" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1952" box="[163,277,1322,1344]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B917EA88FF9CFFF3FF5DFA9CFEB2244E" box="[163,277,1322,1344]" italics="true" pageId="41" pageNumber="626">P. bolivari</emphasis>
</taxonomicName>
).
<taxonomicName id="4C634D19FF9CFFF3FED4FA9CFED32431" authorityName="Simon" authorityYear="1879" box="[298,372,1323,1344]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FED4FA9CFED32431" box="[298,372,1323,1344]" italics="true" pageId="41" pageNumber="626">Protus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C634D19FF9CFFF3FDF8FA9CFDC12431" authorityName="Medrano, Kury &amp; Mendes" authorityYear="2022" box="[518,614,1323,1344]" class="Insecta" family="Flatidae" genus="Kayania" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FDF8FA9CFDC12431" box="[518,614,1323,1344]" italics="true" pageId="41" pageNumber="626">Kayania</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D19FF9CFFF3FD60FA9CFCAD2431" box="[670,778,1323,1344]" class="Arachnida" family="Cosmetidae" genus="Marronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FD60FA9CFCAD2431" box="[670,778,1323,1344]" italics="true" pageId="41" pageNumber="626">Marronia</emphasis>
</taxonomicName>
by not having strong ornamentation on the scutal area III (
<figureCitation id="13582A1FFF9CFFF3FF31FAD0FE8C240C" box="[207,299,1383,1405]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="41" pageNumber="626">Fig. 35F</figureCitation>
vs.
<figureCitation id="13582A1FFF9CFFF3FEA9FAD0FE05240D" box="[343,418,1383,1405]" captionStart="Figure 24" captionStartId="36.[145,223,1821,1843]" captionTargetBox="[305,1265,1052,1782]" captionTargetId="figure-407@36.[305,1265,1052,1782]" captionTargetPageId="36" captionText="Figure 24. Kayania bicolumnata, male holotype (SMF 12782/16). A, dorsal view. B, ventral view. C, distal part of the penis emerging through genital operculum. D, habitus, lateral view. E, postero-lateral view. F, original labels by Weyrauch and Roewer. Scale bars: 1 mm. Photos courtesy Abel Pérez." figureDoi="http://doi.org/10.5281/zenodo.6959576" httpUri="https://zenodo.org/record/6959576/files/figure.png" pageId="41" pageNumber="626">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1C6612FF9CFFF3FF5DFC17FE52257E" ID-DOI="http://doi.org/10.5281/zenodo.6959592" ID-Zenodo-Dep="6959592" httpUri="https://zenodo.org/record/6959592/files/figure.png" pageId="41" pageNumber="626" startId="41.[163,241,928,950]" targetBox="[165,1446,197,888]" targetPageId="41">
<paragraph id="8BDC369AFF9CFFF3FF5DFC17FE52257E" blockId="41.[163,1443,928,1039]" pageId="41" pageNumber="626">
<emphasis id="B917EA88FF9CFFF3FF5DFC17FEBF22C4" bold="true" box="[163,280,928,950]" pageId="41" pageNumber="626">Figure 30.</emphasis>
Map of known distribution of
<taxonomicName id="4C634D19FF9CFFF3FDA4FC16FD3922C7" authorityName="Simon" authorityYear="1879" box="[602,670,929,950]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FF9CFFF3FDA4FC16FD3922C7" box="[602,670,929,950]" italics="true" pageId="41" pageNumber="626">Protus</emphasis>
</taxonomicName>
. Numbers correspond to the following WWF ecoregions: 1 = Ucayali moist forest, 2 = Iquitos varzea, 3 = Southwest Amazon moist forest, 4 = Solimões-Japurá moist forest, 5 = Juruá-Purus moist forest, 6 = Madeira-Tapajos moist forest, 7 = Japurá-Solimões-Negro moist forest, 8 = Uatuma-Trombetas moist forest, 9 = Mato Grosso seasonal forest.
</paragraph>
</caption>
<subSubSection id="C3796511FF9CFFF3FF5DFA11FDD42724" pageId="41" pageNumber="626" type="synonymic_list">
<paragraph id="8BDC369AFF9CFFF3FF5DFA11FDD42724" blockId="41.[163,780,1446,1621]" pageId="41" pageNumber="626">
<emphasis id="B917EA88FF9CFFF3FF5DFA11FEC924CD" box="[163,366,1446,1468]" italics="true" pageId="41" pageNumber="626">Included species:</emphasis>
<taxonomicName id="4C634D19FF9CFFF3FE7AFA11FCAD24CA" authority="(Roewer, 1952)" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1952" box="[388,778,1446,1468]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="bolivari" status="comb. nov .">
<emphasis id="B917EA88FF9CFFF3FE7AFA11FDF724CD" bold="true" box="[388,592,1446,1468]" italics="true" pageId="41" pageNumber="626">Protus bolivari</emphasis>
(
<bibRefCitation id="EFF24B6BFF9CFFF3FD9CFA11FCA524CD" author="Roewer CF" box="[610,770,1446,1468]" pageId="41" pageNumber="626" pagination="37 - 58" refId="ref54681" refString="Roewer CF. 1952. Neotropische Arachnida Arthrogastra, zumeist aus Peru [I]. Senckenbergiana 33: 37 - 58." type="journal article" year="1952">Roewer, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457F3FF9CFFF3FF5DFA72FEE624AB" box="[163,321,1477,1498]" pageId="41" pageNumber="626">
<emphasis id="B917EA88FF9CFFF3FF5DFA72FE9024AB" bold="true" box="[163,311,1477,1498]" pageId="41" pageNumber="626">comb. nov</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C634D19FF9CFFF3FEA9FA72FF4C2488" authority="(Avram &amp; Soares, 1983)" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Avram &amp; Soares" baseAuthorityYear="1983" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="distinctus" status="comb. nov .">
<emphasis id="B917EA88FF9CFFF3FEA9FA72FDA824AB" bold="true" box="[343,527,1476,1498]" italics="true" pageId="41" pageNumber="626">P. distinctus</emphasis>
(
<bibRefCitation id="EFF24B6BFF9CFFF3FDD6FA72FF462489" author="Avram S &amp; Soares HEM" pageId="41" pageNumber="626" pagination="47 - 64" refId="ref50844" refString="Avram S, Soares HEM. 1983. Opilionides du Perou et d'Argentine. Resultats des Expedititions Biospeologiques Cubano-Roumaines a Cuba 4: 47 - 64." type="journal article" year="1983">Avram &amp; Soares, 1983</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457F3FF9CFFF3FF05FA54FE382488" box="[251,415,1507,1529]" pageId="41" pageNumber="626">
<emphasis id="B917EA88FF9CFFF3FF05FA54FE322489" bold="true" box="[251,405,1507,1528]" pageId="41" pageNumber="626">comb. nov</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C634D19FF9CFFF3FE49FA53FCA42488" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[439,771,1507,1529]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B917EA88FF9CFFF3FE49FA53FDEE2488" box="[439,585,1507,1529]" italics="true" pageId="41" pageNumber="626">P. insolens</emphasis>
<bibRefCitation id="EFF24B6BFF9CFFF3FDA7FA54FCA42488" author="Simon E" box="[601,771,1507,1529]" pageId="41" pageNumber="626" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C634D19FF9CFFF3FF5DF9B5FDC02766" authority="(Roewer, 1947)" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1947" box="[163,615,1538,1560]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="quadripunctatus" status="comb. nov.">
<emphasis id="B917EA88FF9CFFF3FF5DF9B5FE0D2769" bold="true" box="[163,426,1538,1560]" italics="true" pageId="41" pageNumber="626">P. quadripunctatus</emphasis>
(
<bibRefCitation id="EFF24B6BFF9CFFF3FE43F9B5FDF82766" author="Roewer CF" box="[445,607,1538,1559]" pageId="41" pageNumber="626" pagination="7 - 57" refId="ref54632" refString="Roewer CF. 1947. Diagnosen neuer Gattungen und Arten der Opiliones Laniatores (Arachn.) aus C. F. Roewer's Sammlung im Senckenberg-Museum. 1. Cosmetidae. [Weitere Weberknechte XII]. Senckenbergiana 28: 7 - 57, pls 1 - 12." type="journal article" year="1947">Roewer, 1947</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457F3FF9CFFF3FD8CF9B5FCA32766" box="[626,772,1538,1559]" pageId="41" pageNumber="626" rank="species">
<emphasis id="B917EA88FF9CFFF3FD8CF9B5FD5B2766" bold="true" box="[626,764,1538,1559]" pageId="41" pageNumber="626">comb. nov</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C634D19FF9CFFF3FF5DF996FDCC2746" authority="Medrano, Kury &amp; Mendes" authorityName="Medrano, Kury &amp; Mendes" authorityYear="2022" box="[163,619,1568,1591]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="marllusi" status="sp. nov.">
<emphasis id="B917EA88FF9CFFF3FF5DF996FE942747" bold="true" box="[163,307,1568,1590]" italics="true" pageId="41" pageNumber="626">P. marllusi</emphasis>
Medrano, Kury &amp; Mendes
</taxonomicName>
<taxonomicNameLabel id="A22457F3FF9CFFF3FD8AF996FD702747" box="[628,727,1569,1590]" pageId="41" pageNumber="626" rank="species">
<emphasis id="B917EA88FF9CFFF3FD8AF996FD772747" bold="true" box="[628,720,1569,1590]" pageId="41" pageNumber="626">sp. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C634D19FF9CFFF3FF5DF9F7FE432724" authority="(Roewer, 1928)" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1928" box="[163,484,1599,1621]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="626" phylum="Arthropoda" rank="species" species="speciosus" status="comb. nov.">
<emphasis id="B917EA88FF9CFFF3FF5DF9F7FE912725" bold="true" box="[163,310,1599,1621]" italics="true" pageId="41" pageNumber="626">P. speciosus</emphasis>
(
<bibRefCitation id="EFF24B6BFF9CFFF3FEBBF9F7FE7B2725" author="Roewer CF" box="[325,476,1599,1621]" pageId="41" pageNumber="626" pagination="527 - 632" refId="ref54532" refString="Roewer CF. 1928 [' 1927 ']. Weitere Weberknechte II. (2. Erganzung der Weberknechte der Erde, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen 26: 527 - 632." type="journal article" year="1928">Roewer, 1928</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457F3FF9CFFF3FE15F988FDD42724" box="[491,627,1599,1621]" pageId="41" pageNumber="626" rank="species">
<emphasis id="B917EA88FF9CFFF3FE15F988FDCB2725" bold="true" box="[491,620,1599,1620]" pageId="41" pageNumber="626">comb. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3796511FF9CFFF3FF5DF9CCFDA527DE" pageId="41" pageNumber="626" type="distribution">
<paragraph id="8BDC369AFF9CFFF3FF5DF9CCFDA527DE" blockId="41.[163,778,1659,1712]" pageId="41" pageNumber="626">
<emphasis id="B917EA88FF9CFFF3FF5DF9CCFE0927E1" box="[163,430,1659,1680]" italics="true" pageId="41" pageNumber="626">Combined distribution:</emphasis>
Upper Amazon basin, eastern Andean slope in
<collectingCountry id="F374760AFF9CFFF3FE9EF92DFE3127DE" box="[352,406,1690,1711]" name="Peru" pageId="41" pageNumber="626">Peru</collectingCountry>
(
<figureCitation id="13582A1FFF9CFFF3FE5BF92DFE5527DE" box="[421,498,1690,1712]" captionStart="Figure 30" captionStartId="41.[163,241,928,950]" captionTargetBox="[165,1446,197,888]" captionTargetId="figure-513@41.[163,1446,195,889]" captionTargetPageId="41" captionText="Figure 30. Map of known distribution of Protus. Numbers correspond to the following WWF ecoregions: 1 = Ucayali moist forest, 2 = Iquitos varzea, 3 = Southwest Amazon moist forest, 4 = Solimões-Japurá moist forest, 5 = Juruá-Purus moist forest, 6 = Madeira-Tapajos moist forest, 7 = Japurá-Solimões-Negro moist forest, 8 = Uatuma-Trombetas moist forest, 9 = Mato Grosso seasonal forest." figureDoi="http://doi.org/10.5281/zenodo.6959592" httpUri="https://zenodo.org/record/6959592/files/figure.png" pageId="41" pageNumber="626">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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