332 lines
51 KiB
XML
332 lines
51 KiB
XML
<document id="8FBC2D3A554F1763AA841B46D15DCF0B" ID-CLB-Dataset="113070" ID-DOI="10.1093/zoolinnean/zlab043" ID-GBIF-Dataset="e070d52f-d008-4f08-988a-a8b17c27418f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6959516" ID-ZooBank="1C89556A-12CB-43B7-9B49-E02EFF1543D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1653991014928" checkinUser="plazi" docAuthor="Medrano, Miguel, Kury, Adriano Brilhante & Mendes, Amanda Cruz" docDate="2022" docId="03CA878CFF8AFF9BFC05FEF6FA2E23CC" docLanguage="en" docName="zlab043.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Gryne Simon 1879" docType="treatment" docVersion="5" lastPageNumber="650" masterDocId="FFF3FFF4FFB5FFDAFFFEFFB7FFA72171" masterDocTitle="Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)" masterLastPageNumber="672" masterPageNumber="585" pageNumber="648" updateTime="1699338404186" updateUser="ExternalLinkService">
|
||
<mods:mods id="FF1999861E296C642626CEBF33A2AE2F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D379153E3849CF7755F337BFB7294F58">
|
||
<mods:title id="BFCB16B67EE8E0B04EE17C85109EC83A">Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7E21865DC01A7A5A1192D921C7D1C185" type="personal">
|
||
<mods:role id="3C672EB5B159ED2125FFC61DEC454701">
|
||
<mods:roleTerm id="E62C20E620B04DC10C87C7995CD66424">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C8B4AEFB568DEBF653CE3DA085AA356B">Medrano, Miguel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B4E8E6254C95957EED022BA3EBEC15F8" type="personal">
|
||
<mods:role id="F0CFA4A1D600FB6F2A71F2609AE33858">
|
||
<mods:roleTerm id="00A2801A165D6B3CAE9164A58A804855">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8DBFB2A1ABC6101E73D074AD6371AB44">Kury, Adriano Brilhante</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="77E92D8EC1D51CB8FFE0F5179940933C" type="personal">
|
||
<mods:role id="6BB1C2F3DAFA903CC75A8DA6AF2B6113">
|
||
<mods:roleTerm id="2EFA0F6F7DF1C562C0CE1149A0A3C024">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="32D66F791D769CE628BBD828D2AB7E1D">Mendes, Amanda Cruz</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E054D48C8088E07B81F49DAEA77E607D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="83E4BFBCA9C86E53F06DA7C7CA3E2063" type="host">
|
||
<mods:titleInfo id="B16B009DBCCBD8A82684A9CD8C9144E3">
|
||
<mods:title id="793B1888A87067FD3F2CEAE43C5DCB03">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CC7C4BA56D85D9ED0999B4B72C256A09">
|
||
<mods:date id="FDB37A4DB160C9C07D57F4D144A3C9D5">2022</mods:date>
|
||
<mods:detail id="608DD69EEC1F6B2C47E048185AA33050" type="pubDate">
|
||
<mods:number id="AA066B60EAC322EAA7285D0B304A0245">2021-08-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D88F328A8D62940D0EA9B6DBFD36F31C" type="volume">
|
||
<mods:number id="19E42B0FA203EC4166DF60849532E65B">195</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5A9A76C47D06945E115C0C64C96C54D6" unit="page">
|
||
<mods:start id="E9A80A7D7815EDFA3DC2720E6C607EFB">585</mods:start>
|
||
<mods:end id="E060E3B4D10071E48656795774E2D4A5">672</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CDB408637A0D56AB3D0AE6C8190CC257">journal article</mods:classification>
|
||
<mods:identifier id="79AECED34D92E1A74C1FCAFD5DEB82DB" type="CLB-Dataset">113070</mods:identifier>
|
||
<mods:identifier id="746D0C99BEEDA8260FB7EBAB852EAB94" type="DOI">10.1093/zoolinnean/zlab043</mods:identifier>
|
||
<mods:identifier id="8A25D4FC932451A81812BA35F642E913" type="GBIF-Dataset">e070d52f-d008-4f08-988a-a8b17c27418f</mods:identifier>
|
||
<mods:identifier id="23DF607763C5DF55563B3128BC8DC8A7" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="1308901DAEDD1816CF5E1EFB78BD7B09" type="Zenodo-Dep">6959516</mods:identifier>
|
||
<mods:identifier id="4401410BFF087BFF9517F9DBAF57335E" type="ZooBank">1C89556A-12CB-43B7-9B49-E02EFF1543D3</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CA878CFF8AFF9BFC05FEF6FA2E23CC" ID-DOI="http://doi.org/10.5281/zenodo.6960502" ID-GBIF-Taxon="198579165" ID-Zenodo-Dep="6960502" LSID="urn:lsid:plazi:treatment:03CA878CFF8AFF9BFC05FEF6FA2E23CC" httpUri="http://treatment.plazi.org/id/03CA878CFF8AFF9BFC05FEF6FA2E23CC" lastPageId="65" lastPageNumber="650" pageId="63" pageNumber="648">
|
||
<subSubSection id="C3796511FF8AFFE5FC05FEF6FB45202B" box="[1019,1250,321,346]" pageId="63" pageNumber="648" type="nomenclature">
|
||
<paragraph id="8BDC369AFF8AFFE5FC05FEF6FB45202B" blockId="63.[1019,1250,321,346]" box="[1019,1250,321,346]" pageId="63" pageNumber="648">
|
||
<heading id="D09481F6FF8AFFE5FC05FEF6FB45202B" box="[1019,1250,321,346]" centered="true" fontSize="9" level="2" pageId="63" pageNumber="648" reason="2">
|
||
<taxonomicName id="4C634D19FF8AFFE5FC05FEF6FB45202B" ID-CoL="7PHNH" authority="SIMON, 1879" authorityName="Simon" authorityYear="1879" box="[1019,1250,321,346]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FC05FEF6FBEC2029" box="[1019,1099,321,344]" italics="true" pageId="63" pageNumber="648">GRYNE</emphasis>
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FBAFFEF6FB45202B" author="Simon E" box="[1105,1250,321,346]" pageId="63" pageNumber="648" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">SIMON, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<treatmentCitationGroup id="AB7311B4FF8AFFE5FCC5FEDDFB2920EC" pageId="63" pageNumber="648">
|
||
<subSubSection id="C3796511FF8AFFE5FCC5FEDDFBFA200F" box="[827,1117,361,383]" pageId="63" pageNumber="648" type="reference_group">
|
||
<paragraph id="8BDC369AFF8AFFE5FCC5FEDDFBFA200F" blockId="63.[827,1443,361,414]" box="[827,1117,361,383]" pageId="63" pageNumber="648">
|
||
<treatmentCitation id="0AC2108BFF8AFFE5FCC5FEDDFBFE200F" author="Simon E" box="[827,1113,361,383]" page="194" pageId="63" pageNumber="648" year="1879">
|
||
<taxonomicName id="4C634D19FF8AFFE5FCC5FEDDFBFE200F" ID-CoL="7PHNH" authority="Simon, 1879: 194" authorityName="Simon" authorityPageNumber="194" authorityYear="1879" box="[827,1113,361,383]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FEDDFC25200E" box="[827,898,362,383]" italics="true" pageId="63" pageNumber="648">Gryne</emphasis>
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FC74FEDEFBFE200F" author="Simon E" box="[906,1113,361,383]" pageId="63" pageNumber="648" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879: 194</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FF8AFFE5FB9BFEDEFB2920EC" pageId="63" pageNumber="648" type="type_taxon">
|
||
<paragraph id="8BDC369AFF8AFFE5FB9BFEDEFB2920EC" blockId="63.[827,1443,361,414]" pageId="63" pageNumber="648">
|
||
Type-species by monotypy:
|
||
<taxonomicName id="4C634D19FF8AFFE5FCC5FE3FFB2020EC" ID-CoL="9X624" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[827,1159,392,414]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="paraensis">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FE3FFC5320EC" box="[827,1012,392,413]" italics="true" pageId="63" pageNumber="648">Gryne paraensis</emphasis>
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FC04FE3FFB2020EC" author="Simon E" box="[1018,1159,392,414]" pageId="63" pageNumber="648" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatmentCitationGroup>
|
||
<subSubSection id="C3796511FF8AFFE5FCC5FE71FC0C208A" pageId="63" pageNumber="648" type="etymology">
|
||
<paragraph id="8BDC369AFF8AFFE5FCC5FE71FC0C208A" blockId="63.[827,1442,454,507]" pageId="63" pageNumber="648">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FE71FBB520AD" box="[827,1042,454,476]" italics="true" pageId="63" pageNumber="648">
|
||
Etymology:
|
||
<taxonomicName id="4C634D19FF8AFFE5FC33FE70FBB520AD" authorityName="Simon" authorityYear="1879" box="[973,1042,455,476]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">Gryne</taxonomicName>
|
||
</emphasis>
|
||
was the name of an Amazon loved by Apollo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FF8AFFE5FCC5FD9BFBBF255D" pageId="63" pageNumber="648" type="diagnosis">
|
||
<paragraph id="8BDC369AFF8AFFE5FCC5FD9BFBBF255D" blockId="63.[827,1444,556,1068]" pageId="63" pageNumber="648">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FD9BFC1F2330" box="[827,952,556,577]" italics="true" pageId="63" pageNumber="648">Diagnosis:</emphasis>
|
||
Median-sized cosmetids with long and unarmed straight legs (Fe IV three times DS length in major forms of males,
|
||
<figureCitation id="13582A1FFF8AFFE5FBD7FDDEFB22230E" box="[1065,1157,617,639]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="63" pageNumber="648">Fig. 54C</figureCitation>
|
||
), dorsal scutum beta-type with short coda and narrow cephalothorax (
|
||
<figureCitation id="13582A1FFF8AFFE5FACAFD3FFCE923CD" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="63" pageNumber="648">Fig. 54D– G</figureCitation>
|
||
). Reticular pattern in dorsal scutum more marked in laterals of cephalothorax and abdomen (
|
||
<figureCitation id="13582A1FFF8AFFE5FACAFD72FA3B23AA" box="[1332,1436,709,731]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="63" pageNumber="648">Figs 50A</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF8AFFE5FCC5FD53FCCE238B" box="[827,873,740,762]" captionStart="Figure 52" captionStartId="69.[163,243,1281,1303]" captionTargetBox="[167,775,195,1242]" captionTargetId="figure-591@69.[167,775,195,1242]" captionTargetPageId="69" captionText="Figure 52. Gryne leprosa. A–C, female holotype (ZMUC): A, dorsal view; B, ventral view; C, lateral view. D–G, male lectotype of G. perlata (MNRJ 41773): D, dorsal view; E, dorsal panoramic view; F, lateral view; G, frontal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959642" httpUri="https://zenodo.org/record/6959642/files/figure.png" pageId="63" pageNumber="648">52A</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF8AFFE5FC86FD53FC01238B" box="[888,934,740,762]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="63" pageNumber="648">54A</figureCitation>
|
||
). Area I with a pair of small tubercles and area III with a pair of high acuminate spines (spines lower in
|
||
<taxonomicName id="4C634D19FF8AFFE5FC5FFC96FB942247" authorityName="Mello-Leitao" authorityYear="1928" box="[929,1075,801,822]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="dimorpha">
|
||
<emphasis id="B917EA88FF8AFFE5FC5FFC96FB942247" box="[929,1075,801,822]" italics="true" pageId="63" pageNumber="648">G. dimorpha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13582A1FFF8AFFE5FBBCFC96FB052246" box="[1090,1186,801,823]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="63" pageNumber="648">Fig. 50B</figureCitation>
|
||
). Pedipalpus sexually monomorphic, tibia without expansion opposed to the slender tarsus forming a chela in males. Major forms of males with long tibia IV (as long as femur), and metatarsus short and triquetrous (
|
||
<figureCitation id="13582A1FFF8AFFE5FAF0FC2BFA3422C3" box="[1294,1427,924,946]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="63" pageNumber="648">Fig. 50E–H</figureCitation>
|
||
). Ventral plate of the penis sub-rectangular, narrower at the tip (except for
|
||
<taxonomicName id="4C634D19FF8AFFE5FBCBFC6EFB72229F" baseAuthorityName="Perty" baseAuthorityYear="1833" box="[1077,1237,985,1006]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="marginalis">
|
||
<emphasis id="B917EA88FF8AFFE5FBCBFC6EFB72229F" box="[1077,1237,985,1006]" italics="true" pageId="63" pageNumber="648">G. marginalis</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="13582A1FFF8AFFE5FB11FC6EFAF0229E" box="[1263,1367,985,1007]" captionStart="Figure 55" captionStartId="73.[163,243,1727,1749]" captionTargetBox="[163,1443,195,1687]" captionTargetId="figure-24@73.[163,1443,195,1687]" captionTargetPageId="73" captionText="Figure 55. SEM images of some structures of Gryne marginalis. A–D, penis (MNRJ 17579): A, dorsal view; B, lateral view; C, ventral view; D, detail of microsetae, MS-D and MS-E, ventral view. E–I, male (MNRJ 2156): E, claws of left leg IV showing two rows of pectination; F, left cheliceral hand, dorsal view; G, left basichelicerite, dorsal view; H, left pedipalpus, femur, mesal view; I, same, tibia and tarsus, ventral view. Scale bars 200 μm (F, H, I), 100 μm (A, B, G), 50 μm (C, E), 10 μm (D). Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959650" httpUri="https://zenodo.org/record/6959650/files/figure.png" pageId="63" pageNumber="648">Figs 55A</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF8AFFE5FA98FC6EFA34229E" box="[1382,1427,985,1007]" captionStart="Figure 51" captionStartId="67.[163,241,1513,1535]" captionTargetBox="[163,1443,196,1472]" captionTargetId="figure-163@67.[163,1443,195,1473]" captionTargetPageId="67" captionText="Figure 51. SEM images of some structures of Gryne dimorpha. A–D, penis (MNRJ 6297): A, dorsal view; B, lateral view; C, ventral view; D, detail of MS-C and stylus, oblique view. E–H, male (MNRJ 2156): E, left cheliceral hand, dorsal view; F, left basichelicerite, dorsal view; G, left pedipalpus, femur, mesal view; H, same, tibia and tarsus, ventral view. Scale bars: 50 μm (A–C), 10 μm (D), 200 μm (E, G, H), 100 μm (F)." figureDoi="http://doi.org/10.5281/zenodo.6959640" httpUri="https://zenodo.org/record/6959640/files/figure.png" pageId="63" pageNumber="648">51A</figureCitation>
|
||
). Three distal curved MS-C and two basal straight MS-A (
|
||
<figureCitation id="13582A1FFF8AFFE5FC75FBA1FBAE255D" box="[907,1033,1046,1068]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="63" pageNumber="648">Fig. 54A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FF8AFFE5FCC5FBD2FB812584" pageId="63" pageNumber="648" type="synonymic_list">
|
||
<paragraph id="8BDC369AFF8AFFE5FCC5FBD2FB812584" blockId="63.[827,1444,1125,1362]" pageId="63" pageNumber="648">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FBD2FBAF250A" box="[827,1032,1125,1147]" italics="true" pageId="63" pageNumber="648">Included species:</emphasis>
|
||
<taxonomicName id="4C634D19FF8AFFE5FBDEFBD2FA39250A" authority="(Perty, 1833)" baseAuthorityName="Perty" baseAuthorityYear="1833" box="[1056,1438,1125,1147]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="marginalis">
|
||
<emphasis id="B917EA88FF8AFFE5FBDEFBD2FB50250B" box="[1056,1271,1125,1146]" italics="true" pageId="63" pageNumber="648">Gryne marginalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FAF4FBD1FA34250B" author="Perty JAM" box="[1290,1427,1125,1147]" pageId="63" pageNumber="648" pagination="201 - 209" refId="ref53820" refString="Perty JAM. 1833. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX [1817 - 1820] jussu et auspiciis Maximiliani Josephi I Bavariae Regis augustissimi peracto, collegerunt Dr. In: de Spix JB, de Martius CF-Ph, eds. Vol. 3. [Opiliones, pp 201 - 209]. [' 1830 - 1834 ']. Munich: Fleischer." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D19FF8AFFE5FCC5FB33FB2725E8" authority="(Sorensen, 1879)" baseAuthorityName="Sorensen" baseAuthorityYear="1879" box="[827,1152,1156,1177]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="orensis">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FB33FC0825E8" box="[827,943,1156,1177]" italics="true" pageId="63" pageNumber="648">G. orensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FC3FFB33FBD125E8" author="Sorensen WE" box="[961,1142,1156,1177]" pageId="63" pageNumber="648" pagination="97 - 222" refId="ref55474" refString="Sorensen WE. 1879. Om Bygnigen af Gonyleptiderne, en Type af Arachnidernes Classe. Naturhistorisk Tidsskrift, KjObenhavn, ser. 3 12: 97 - 222, pls 1 - 2." type="journal article" year="1879">Sørensen, 1879</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D19FF8AFFE5FB70FB33FCDA25C9" authority="(Roewer, 1915)" authorityName="Medrano & Kury & Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1915" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="marmorata" status="comb. nov.">
|
||
<emphasis id="B917EA88FF8AFFE5FB70FB33FA9225E8" box="[1166,1333,1156,1177]" italics="true" pageId="63" pageNumber="648">G. marmorata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FAB9FB33FCD225C9" author="Roewer CF" pageId="63" pageNumber="648" pagination="106 - 132" refId="ref54383" refString="Roewer CF. 1915 [' 1914 ']. Funfzehn neue Opilioniden. Archiv fur Naturgeschichte, Abt. A, Original-Arbeiten 80: 106 - 132." type="journal article" year="1915">Roewer, 1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457F3FF8AFFE5FC79FB15FBA025C9" box="[903,1031,1186,1208]" pageId="63" pageNumber="648" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C634D19FF8AFFE5FBEBFB14FA3B25C9" authority="Mello-Leitao, 1928" authorityName="Mello-Leitao" authorityYear="1928" box="[1045,1436,1186,1208]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="dimorpha">
|
||
<emphasis id="B917EA88FF8AFFE5FBEBFB14FB0925C6" box="[1045,1198,1186,1208]" italics="true" pageId="63" pageNumber="648">G. dimorpha</emphasis>
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FB46FB15FA3B25C9" author="de Mello-Leitao CF" box="[1208,1436,1186,1208]" pageId="63" pageNumber="648" pagination="9 - 13" refId="ref53193" refString="de Mello-Leitao CF. 1928. Opilioes de Matto Grosso e Pernambuco. Boletim do Museu Nacional 4: 9 - 13." type="journal article" year="1928">Mello-Leitão, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D19FF8AFFE5FCC5FB76FBD525A7" authority="Sorensen, 1932" authorityName="Sorensen" authorityYear="1932" box="[827,1138,1217,1238]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="leprosa">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FB76FC1525A7" box="[827,946,1217,1238]" italics="true" pageId="63" pageNumber="648">G. leprosa</emphasis>
|
||
Sørensen, 1932
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D19FF8AFFE5FB4EFB76FB862584" authority="(Mello-Leitao, 1935)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1935" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="coccinelloides">
|
||
<emphasis id="B917EA88FF8AFFE5FB4EFB76FA0425A7" box="[1200,1443,1217,1238]" italics="true" pageId="63" pageNumber="648">Gryne coccinelloides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FCBDFB57FBB12587" author="de Mello-Leitao CF" box="[835,1046,1248,1270]" pageId="63" pageNumber="648" pagination="369 - 411" refId="ref53287" refString="de Mello-Leitao CF. 1935. A proposito de alguns opilioes novos. Memorias do Instituto Butantan 9: 369 - 411." type="journal article" year="1935">Mello-Leitão, 1935</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FF8AFFE5FCADFB49FB062423" pageId="63" pageNumber="648" type="distribution">
|
||
<paragraph id="8BDC369AFF8AFFE5FCADFB49FB062423" blockId="63.[827,1444,1125,1362]" pageId="63" pageNumber="648">
|
||
Combined distribution (
|
||
<figureCitation id="13582A1FFF8AFFE5FB9BFB49FB152465" box="[1125,1202,1278,1300]" captionStart="Figure 48" captionStartId="64.[145,226,1758,1780]" captionTargetBox="[147,1428,447,1718]" captionTargetId="figure-112@64.[145,1428,445,1719]" captionTargetPageId="64" captionText="Figure 48. Map of known distribution of Gryne. Numbers correspond to the following WWF ecoregions: 1 = Uatumã- Trombetas moist forest, 2 = Madeira-Tapajós moist forest, 3 = Xingú-Tocantins-Araguaia moist forest, 4 = Marajó várzea, 5 = Tocantins/Pindaré moist forest, 6 = Mato grosso seasonal forest, 7 = Caatinga, 8 = Pernambuco Coastal forest, 9 = Cerrado, 10 = Bahia interior forest, 11 = Bahia coastal forest, 12 = Alto Paraná Atlantic forest, 13 = Pantanal, 14 = Dry Chaco, 15 = Humid chaco, 16 = Parana flooded savanna, 17 = Humid pampas." figureDoi="http://doi.org/10.5281/zenodo.6959634" httpUri="https://zenodo.org/record/6959634/files/figure.png" pageId="63" pageNumber="648">Fig. 48</figureCitation>
|
||
). Semi-arid diagonal of South America, northern half of Brazilian Atlantic forest and lower Amazon basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FF8AFF9AFCC5FA3FFC632005" lastPageId="64" lastPageNumber="649" pageId="63" pageNumber="648" type="discussion">
|
||
<paragraph id="8BDC369AFF8AFFE5FCC5FA3FFBF927A1" blockId="63.[827,1444,1416,1898]" pageId="63" pageNumber="648">
|
||
<emphasis id="B917EA88FF8AFFE5FCC5FA3FFC1024EC" box="[827,951,1416,1437]" italics="true" pageId="63" pageNumber="648">Remarks:</emphasis>
|
||
The remarkable non-homoplastic synapomorphy of
|
||
<taxonomicName id="4C634D19FF8AFFE5FBE4FA10FBC424CD" authorityName="Simon" authorityYear="1879" box="[1050,1123,1447,1468]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FBE4FA10FBC424CD" box="[1050,1123,1447,1468]" italics="true" pageId="63" pageNumber="648">Gryne</emphasis>
|
||
</taxonomicName>
|
||
refers to the morphology of leg IV, having a sexual dimorphic metatarsus, in the males being thicker at basal portion (
|
||
<figureCitation id="13582A1FFF8AFFE5FABCFA53FCE92766" captionStart="Figure 2" captionStartId="2.[145,226,1414,1436]" captionTargetBox="[225,1345,195,1374]" captionTargetId="figure-248@2.[225,1345,195,1374]" captionTargetPageId="2" captionText="Figure 2. Habitus in vivo of some Discosomaticinae from Brazil. A, Roquettea singularis from Pará. B, Gryne leprosa from Pará. C, Gryne marginalis from Pará. D, Gryne dimorpha from Goiás. E–G, Gryne orensis from Mato Grosso do Sul. Photographs of Arthur Anker & Pedro H. Martins (A, C–D), Bruno Garcia (B) and Adriano Kury (E–G)." figureDoi="http://doi.org/10.5281/zenodo.6959524" httpUri="https://zenodo.org/record/6959524/files/figure.png" pageId="63" pageNumber="648">Figs 2F, G</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF8AFFE5FCA2F9B5FC2E2769" box="[860,905,1538,1560]" captionStart="Figure 57" captionStartId="76.[145,226,1846,1868]" captionTargetBox="[305,1265,923,1807]" captionTargetId="figure-48@76.[305,1265,923,1807]" captionTargetPageId="76" captionText="Figure 57. Gryne orensis (Sørensen, 1879), male (USNMENT 01538077). A, dorsal view. B, dorsal panoramic view. C, lateral view. D, left metatarsus IV and tarsomeres. Scale bars: 1 mm (A, C, D), 2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.6959654" httpUri="https://zenodo.org/record/6959654/files/figure.png" pageId="63" pageNumber="648">57C</figureCitation>
|
||
). Modifications in this segment of leg IV also occur in an unrelated member of
|
||
<taxonomicName id="4C634D19FF8AFFE5FB26F996FAC72747" authorityName="C. L. Koch" authorityYear="1839" box="[1240,1376,1569,1590]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FB26F996FAC72747" box="[1240,1376,1569,1590]" italics="true" pageId="63" pageNumber="648">Paecilaema</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F374760AFF8AFFE5FCC5F9F7FC1C2727" box="[827,955,1600,1622]" name="Venezuela" pageId="63" pageNumber="648">Venezuela</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C634D19FF8AFFE5FC37F9F7FAB82724" authorityName="Gonzalez-Sponga" authorityYear="1992" box="[969,1311,1600,1621]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="species" species="trisegmentatum">
|
||
<emphasis id="B917EA88FF8AFFE5FC37F9F7FAB82724" box="[969,1311,1600,1621]" italics="true" pageId="63" pageNumber="648">Paecilaema trisegmentatum</emphasis>
|
||
</taxonomicName>
|
||
González- Sponga, 1992. The condition of this species, however, is different, having a unique ‘fusiform subdistal area’ (
|
||
<bibRefCitation id="EFF24B6BFF8AFFE5FCBDF92BFB3327C3" author="Gonzalez-Sponga MA" box="[835,1172,1692,1714]" pageId="63" pageNumber="648" refId="ref51725" refString="Gonzalez-Sponga MA. 1992. Aracnidos de Venezuela. Opiliones Laniatores II. Familia Cosmetidae. Caracas: Academia de Ciencias Fisicas, Matematicas y Naturales." type="book" year="1992">González-Sponga, 1992: 396</bibRefCitation>
|
||
) indicating a putative autapomorphic condition.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF8AFF9AFCADF96EFE472046" blockId="63.[827,1444,1416,1898]" lastBlockId="64.[145,761,197,372]" lastPageId="64" lastPageNumber="649" pageId="63" pageNumber="648">
|
||
A large quantity of material of
|
||
<taxonomicName id="4C634D19FF8AFFE5FB3CF96EFAAF279F" authorityName="Simon" authorityYear="1879" box="[1218,1288,1753,1774]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FB3CF96EFAAF279F" box="[1218,1288,1753,1774]" italics="true" pageId="63" pageNumber="648">Gryne</emphasis>
|
||
</taxonomicName>
|
||
from regions of
|
||
<collectingCountry id="F374760AFF8AFFE5FCA7F94FFC05267F" box="[857,930,1784,1806]" name="Brazil" pageId="63" pageNumber="648">Brazil</collectingCountry>
|
||
was deposited in arachnological collection of Museu Nacional before the fire that destroyed it (Kury
|
||
<emphasis id="B917EA88FF8AFFE5FC7AF881FC1A263B" box="[900,957,1845,1867]" italics="true" pageId="63" pageNumber="648">et al.</emphasis>
|
||
, 2018). A great percentage of the lots were examined and identified by us as belonging to
|
||
<taxonomicName id="4C634D19FF8AFFE5FAA9F8E3FA3B2618" authorityName="Simon" authorityYear="1879" box="[1367,1436,1876,1897]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="648" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF8AFFE5FAA9F8E3FA3B2618" box="[1367,1436,1876,1897]" italics="true" pageId="63" pageNumber="648">Gryne</emphasis>
|
||
</taxonomicName>
|
||
. Nonetheless, inter and intraspecific variations were not known or properly illustrated to make possible the identification to species level. Here, we present their distributional data in
|
||
<figureCitation id="13582A1FFFF5FF9AFE72FE96FE7F2047" box="[396,472,289,311]" captionStart="Figure 48" captionStartId="64.[145,226,1758,1780]" captionTargetBox="[147,1428,447,1718]" captionTargetId="figure-112@64.[145,1428,445,1719]" captionTargetPageId="64" captionText="Figure 48. Map of known distribution of Gryne. Numbers correspond to the following WWF ecoregions: 1 = Uatumã- Trombetas moist forest, 2 = Madeira-Tapajós moist forest, 3 = Xingú-Tocantins-Araguaia moist forest, 4 = Marajó várzea, 5 = Tocantins/Pindaré moist forest, 6 = Mato grosso seasonal forest, 7 = Caatinga, 8 = Pernambuco Coastal forest, 9 = Cerrado, 10 = Bahia interior forest, 11 = Bahia coastal forest, 12 = Alto Paraná Atlantic forest, 13 = Pantanal, 14 = Dry Chaco, 15 = Humid chaco, 16 = Parana flooded savanna, 17 = Humid pampas." figureDoi="http://doi.org/10.5281/zenodo.6959634" httpUri="https://zenodo.org/record/6959634/files/figure.png" pageId="64" pageNumber="649">Fig. 48</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFFF5FF9AFF57FEF7FC632005" blockId="64.[145,761,197,372]" lastBlockId="64.[809,1426,197,372]" pageId="64" pageNumber="649">
|
||
Species
|
||
<taxonomicName id="4C634D19FFF5FF9AFEF0FEF7FE7E2024" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1935" box="[270,473,320,341]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="64" pageNumber="649" phylum="Arthropoda" rank="species" species="coccinelloides">
|
||
<emphasis id="B917EA88FFF5FF9AFEF0FEF7FE7E2024" box="[270,473,320,341]" italics="true" pageId="64" pageNumber="649">G. coccinelloides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D19FFF5FF9AFDE5FEF7FD602024" authorityName="Medrano & Kury & Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1915" box="[539,711,320,341]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="64" pageNumber="649" phylum="Arthropoda" rank="species" species="marmorata">
|
||
<emphasis id="B917EA88FFF5FF9AFDE5FEF7FD602024" box="[539,711,320,341]" italics="true" pageId="64" pageNumber="649">G. marmorata</emphasis>
|
||
</taxonomicName>
|
||
are not included in the taxonomic key because detailed examination of specimens was not possible, preventing the confection of diagnoses as well. Future research with fresh material from
|
||
<typeStatus id="54D88838FFF5FF9AFBACFEB4FB232069" box="[1106,1156,259,280]" pageId="64" pageNumber="649">type</typeStatus>
|
||
localities is needed for unveiling the identity and validity of both species. For the same reason, a diagnosis section is lacking for those species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1C6612FFF5FF9AFF6FF969FC11261B" ID-DOI="http://doi.org/10.5281/zenodo.6959634" ID-Zenodo-Dep="6959634" httpUri="https://zenodo.org/record/6959634/files/figure.png" pageId="64" pageNumber="649" startId="64.[145,226,1758,1780]" targetBox="[147,1428,447,1718]" targetPageId="64">
|
||
<paragraph id="8BDC369AFFF5FF9AFF6FF969FC11261B" blockId="64.[145,1425,1758,1898]" pageId="64" pageNumber="649">
|
||
<emphasis id="B917EA88FFF5FF9AFF6FF969FEAC2782" bold="true" box="[145,267,1758,1780]" pageId="64" pageNumber="649">Figure 48.</emphasis>
|
||
Map of known distribution of
|
||
<taxonomicName id="4C634D19FFF5FF9AFDA3F968FD392784" authorityName="Simon" authorityYear="1879" box="[605,670,1759,1781]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="64" pageNumber="649" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FFF5FF9AFDA3F968FD392784" box="[605,670,1759,1781]" italics="true" pageId="64" pageNumber="649">Gryne</emphasis>
|
||
</taxonomicName>
|
||
. Numbers correspond to the following WWF ecoregions: 1 = Uatumã- Trombetas moist forest, 2 = Madeira-Tapajós moist forest, 3 = Xingú-Tocantins-Araguaia moist forest, 4 = Marajó várzea, 5 = Tocantins/Pindaré moist forest, 6 = Mato grosso seasonal forest, 7 = Caatinga, 8 = Pernambuco Coastal forest, 9 = Cerrado, 10 = Bahia interior forest, 11 = Bahia coastal forest, 12 = Alto Paraná Atlantic forest, 13 = Pantanal, 14 = Dry Chaco, 15 = Humid chaco, 16 = Parana flooded savanna, 17 = Humid pampas.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3796511FFF4FF9BFD8EFF6AFA85204D" pageId="65" pageNumber="650" type="multiple">
|
||
<paragraph id="8BDC369AFFF4FF9BFD8EFF6AFC712184" blockId="65.[188,1418,221,701]" box="[624,982,221,246]" pageId="65" pageNumber="650">
|
||
<heading id="D09481F6FFF4FF9BFD8EFF6AFC712184" box="[624,982,221,246]" centered="true" fontSize="9" level="2" pageId="65" pageNumber="650" reason="2">
|
||
KEY TO THE SPECIES OF
|
||
<taxonomicName id="4C634D19FFF4FF9BFC78FF69FC712184" authorityName="Simon" authorityYear="1879" box="[902,982,222,245]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FFF4FF9BFC78FF69FC712184" box="[902,982,222,245]" italics="true" pageId="65" pageNumber="650">GRYNE</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFFF4FF9BFF42FE91FA85204D" blockId="65.[188,1418,221,701]" box="[188,1314,294,316]" pageId="65" pageNumber="650">
|
||
<emphasis id="B917EA88FFF4FF9BFF42FE91FE88204A" box="[188,303,294,315]" italics="true" pageId="65" pageNumber="650">Comment:</emphasis>
|
||
Excluding
|
||
<taxonomicName id="4C634D19FFF4FF9BFE42FE91FDD4204A" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1935" box="[444,627,294,315]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="species" species="coccinelloides">
|
||
<emphasis id="B917EA88FFF4FF9BFE42FE91FDD4204A" box="[444,627,294,315]" italics="true" pageId="65" pageNumber="650">G. coccinelloides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D19FFF4FF9BFD52FE91FCEB204A" authorityName="Medrano & Kury & Mendes" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1915" box="[684,844,294,315]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="species" species="marmorata">
|
||
<emphasis id="B917EA88FFF4FF9BFD52FE91FCEB204A" box="[684,844,294,315]" italics="true" pageId="65" pageNumber="650">G. marmorata</emphasis>
|
||
</taxonomicName>
|
||
(see remarks on the genus
|
||
<taxonomicName id="4C634D19FFF4FF9BFB7BFE91FB6E204A" authorityName="Simon" authorityYear="1879" box="[1157,1225,294,315]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FFF4FF9BFB7BFE91FB6E204A" box="[1157,1225,294,315]" italics="true" pageId="65" pageNumber="650">Gryne</emphasis>
|
||
</taxonomicName>
|
||
above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FFF4FF9BFF42FEFDFA2E23CC" pageId="65" pageNumber="650" type="key">
|
||
<key id="FFF261ABFFF4FF9BFF42FEFDFA2E23CC" pageId="65" pageNumber="650">
|
||
<paragraph id="8BDC369AFFF4FF9BFF42FEFDFA2D20FC" blockId="65.[188,1418,221,701]" pageId="65" pageNumber="650">
|
||
<keyStep id="30972E3FFFF4FF9BFF42FEFDFA2D20FC" pageId="65" pageNumber="650">
|
||
<keyLead id="309295AFFFF4FF9BFF42FEFDFA2D20FC" pageId="65" pageNumber="650">
|
||
1. Reticulate pattern of DS spots on lateral margins of carapace and abdomen (
|
||
<figureCitation id="13582A1FFFF4FF9BFBBAFEFDFB37202E" box="[1092,1168,329,352]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="65" pageNumber="650">Fig. 54</figureCitation>
|
||
) ................................ 2 1’. Reticulate pattern of DS spots invading mesotergum (
|
||
<figureCitation id="13582A1FFFF4FF9BFC9AFECFFC6420FF" box="[868,963,376,398]" captionStart="Figure 52" captionStartId="69.[163,243,1281,1303]" captionTargetBox="[167,775,195,1242]" captionTargetId="figure-591@69.[167,775,195,1242]" captionTargetPageId="69" captionText="Figure 52. Gryne leprosa. A–C, female holotype (ZMUC): A, dorsal view; B, ventral view; C, lateral view. D–G, male lectotype of G. perlata (MNRJ 41773): D, dorsal view; E, dorsal panoramic view; F, lateral view; G, frontal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959642" httpUri="https://zenodo.org/record/6959642/files/figure.png" pageId="65" pageNumber="650">Fig. 52A</figureCitation>
|
||
) ..............................................................3
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFFF4FF9BFF42FE28FA2E2322" blockId="65.[188,1418,221,701]" pageId="65" pageNumber="650">
|
||
<keyStep id="30972E3FFFF4FF9BFF42FE28FA2E2322" pageId="65" pageNumber="650">
|
||
<keyLead id="309295AFFFF4FF9BFF42FE28FA2E2322" pageId="65" pageNumber="650">
|
||
2. Reticulate pattern restricted to laterals of chevron, invading scutal sulcus and going gently to the lateral margin of carapace (
|
||
<figureCitation id="13582A1FFFF4FF9BFE3AFE09FD8520A5" box="[452,546,446,468]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="65" pageNumber="650">Fig. 50A</figureCitation>
|
||
); spines of area III low (
|
||
<figureCitation id="13582A1FFFF4FF9BFCC8FE09FC3320A5" box="[822,916,446,468]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="65" pageNumber="650">Fig. 50B</figureCitation>
|
||
); VP of penis not elongate and apical border narrower than base (
|
||
<figureCitation id="13582A1FFFF4FF9BFE35FE6BFDEE2083" box="[459,585,476,498]" captionStart="Figure 50" captionStartId="66.[145,225,1776,1798]" captionTargetBox="[145,1425,195,1736]" captionTargetId="figure-24@66.[145,1425,195,1736]" captionTargetPageId="66" captionText="Figure 50. Gryne dimorpha. A–C, male (MNRJ 2144): A, dorsal view; B, lateral view; C, panoramic dorsal view. D, two adult males of different sizes and colours; from left to right: minor form (MNRJ 17591) and major form (MNRJ 2144). E, male, major form (MNRJ 2144), leg IV, tibia, metatarsus and tarsomeres. F, same, detail of tibia, triquetrous metatarsus and tarsomeres. G, male, minor form (MNRJ 17591), leg IV, tibia and tarsomeres. H, same, detail of tibia and tarsomeres. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959638" httpUri="https://zenodo.org/record/6959638/files/figure.png" pageId="65" pageNumber="650">Fig. 50A–C</figureCitation>
|
||
) ..................................................................................................
|
||
<taxonomicName id="4C634D19FFF4FF9BFB07FE6BFA2E2080" ID-CoL="7WDWV" authorityName="Mello-Leitao" authorityYear="1928" box="[1273,1417,476,498]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="663" phylum="Arthropoda" rank="species" species="dimorpha">
|
||
G.
|
||
<emphasis id="B917EA88FFF4FF9BFAE7FE6BFA2E2080" box="[1305,1417,476,497]" italics="true" pageId="65" pageNumber="650">dimorpha</emphasis>
|
||
</taxonomicName>
|
||
2’. Reticulate pattern restricted to lateral margin of carapace and abdomen, not invading scutal sulcus (
|
||
<figureCitation id="13582A1FFFF4FF9BFEF4FDA8FEF22344" box="[266,341,543,565]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="65" pageNumber="650">Fig. 54</figureCitation>
|
||
); spines of area III low (
|
||
<figureCitation id="13582A1FFFF4FF9BFD99FDA8FD632344" box="[615,708,543,565]" captionStart="Figure 54" captionStartId="72.[145,225,1703,1725]" captionTargetBox="[145,1425,196,1662]" captionTargetId="figure-24@72.[145,1425,195,1663]" captionTargetPageId="72" captionText="Figure 54. Gryne marginalis. A, male (UFMG 9388), dorsal view. B, same, lateral view. C, same, panoramic dorsal view. D–G, variations of spots pattern in dorsal view: D, male (MNRJ 60252) from Nova Timboteua (Pará, Brazil); E, male, same data; F, female (MNRJ 60251) from Rondon do Pará (Pará, Brazil); G, female, same data. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959646" httpUri="https://zenodo.org/record/6959646/files/figure.png" pageId="65" pageNumber="650">Fig. 54B</figureCitation>
|
||
); VP of penis elongate and with apical margin wider than base (
|
||
<figureCitation id="13582A1FFFF4FF9BFEF4FD89FECE2322" box="[266,361,573,596]" captionStart="Figure 55" captionStartId="73.[163,243,1727,1749]" captionTargetBox="[163,1443,195,1687]" captionTargetId="figure-24@73.[163,1443,195,1687]" captionTargetPageId="73" captionText="Figure 55. SEM images of some structures of Gryne marginalis. A–D, penis (MNRJ 17579): A, dorsal view; B, lateral view; C, ventral view; D, detail of microsetae, MS-D and MS-E, ventral view. E–I, male (MNRJ 2156): E, claws of left leg IV showing two rows of pectination; F, left cheliceral hand, dorsal view; G, left basichelicerite, dorsal view; H, left pedipalpus, femur, mesal view; I, same, tibia and tarsus, ventral view. Scale bars 200 μm (F, H, I), 100 μm (A, B, G), 50 μm (C, E), 10 μm (D). Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959650" httpUri="https://zenodo.org/record/6959650/files/figure.png" pageId="65" pageNumber="650">Fig. 55A</figureCitation>
|
||
) .................................................................................................................................
|
||
<taxonomicName id="4C634D19FFF4FF9BFB13FD89FA2E2322" ID-CoL="7WDWX" baseAuthorityName="Perty" baseAuthorityYear="1833" box="[1261,1417,574,595]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="species" species="marginalis">
|
||
<emphasis id="B917EA88FFF4FF9BFB13FD89FA2E2322" box="[1261,1417,574,595]" italics="true" pageId="65" pageNumber="650">G. marginalis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFFF4FF9BFF42FDD3FA2E23CC" blockId="65.[188,1418,221,701]" pageId="65" pageNumber="650">
|
||
<keyStep id="30972E3FFFF4FF9BFF42FDD3FA2E23CC" pageId="65" pageNumber="650">
|
||
<keyLead id="309295AFFFF4FF9BFF42FDD3FA2E23CC" pageId="65" pageNumber="650">
|
||
3. Reticulate pattern uniformly occupying abdomen (
|
||
<figureCitation id="13582A1FFFF4FF9BFCE7FDD3FCDF230B" box="[793,888,612,634]" captionStart="Figure 52" captionStartId="69.[163,243,1281,1303]" captionTargetBox="[167,775,195,1242]" captionTargetId="figure-591@69.[167,775,195,1242]" captionTargetPageId="69" captionText="Figure 52. Gryne leprosa. A–C, female holotype (ZMUC): A, dorsal view; B, ventral view; C, lateral view. D–G, male lectotype of G. perlata (MNRJ 41773): D, dorsal view; E, dorsal panoramic view; F, lateral view; G, frontal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6959642" httpUri="https://zenodo.org/record/6959642/files/figure.png" pageId="65" pageNumber="650">Fig. 52A</figureCitation>
|
||
) ..........................................................
|
||
<taxonomicName id="4C634D19FFF4FF9BFAE7FDD2FA2D2308" ID-CoL="7WDWW" authorityName="Sorensen" authorityYear="1932" box="[1305,1418,612,634]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="species" species="leprosa">
|
||
<emphasis id="B917EA88FFF4FF9BFAE7FDD2FA2D2308" box="[1305,1418,612,634]" italics="true" pageId="65" pageNumber="650">G. leprosa</emphasis>
|
||
</taxonomicName>
|
||
3’. Reticulate pattern clearly marked in the region of the omega stripe and scutal sulcus of abdomen, but not occupying areas of mesotergum (
|
||
<figureCitation id="13582A1FFFF4FF9BFD5AFD10FCA523CC" box="[676,770,679,701]" captionStart="Figure 57" captionStartId="76.[145,226,1846,1868]" captionTargetBox="[305,1265,923,1807]" captionTargetId="figure-48@76.[305,1265,923,1807]" captionTargetPageId="76" captionText="Figure 57. Gryne orensis (Sørensen, 1879), male (USNMENT 01538077). A, dorsal view. B, dorsal panoramic view. C, lateral view. D, left metatarsus IV and tarsomeres. Scale bars: 1 mm (A, C, D), 2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.6959654" httpUri="https://zenodo.org/record/6959654/files/figure.png" pageId="65" pageNumber="650">Fig. 57A</figureCitation>
|
||
) ............................................................................
|
||
<taxonomicName id="4C634D19FFF4FF9BFAE5FD10FA2E23CC" ID-CoL="7WDWY" baseAuthorityName="Sorensen" baseAuthorityYear="1879" box="[1307,1417,679,701]" class="Arachnida" family="Cosmetidae" genus="Gryne" kingdom="Animalia" order="Opiliones" pageId="65" pageNumber="650" phylum="Arthropoda" rank="species" species="orensis">
|
||
<emphasis id="B917EA88FFF4FF9BFAE5FD10FA2E23CC" box="[1307,1417,679,701]" italics="true" pageId="65" pageNumber="650">G. orensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |