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<document id="3E0F2E1EE2903D0048D1A015717086A6" ID-DOI="10.37828/em.2023.64.2" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246869" ID-ZooBank="urn:lsid:zoobank.org:pub:C6640452-261E-4893-9A1C-D68A6F0D8A4D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972205822" checkinUser="felipe" docAuthor="Glöer, Peter &amp; Pešić, Vladimir" docDate="2023" docId="03CA6424FFA40E789893496AFE419FC8" docLanguage="en" docName="EcolMontenegrina.64.185-190.pdf" docOrigin="Ecologica Montenegrina 64" docSource="http://dx.doi.org/10.37828/em.2023.64.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Ancylus naumiensis Glöer &amp; Pešić, 2023, n. sp." docType="treatment" docUuid="2CCBEB51-BA20-4BC4-85F8-E7254717E9A5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="187" masterDocId="FFF31C5CFFA50E7A98044F63FFDC9860" masterDocTitle="A new Ancylus and a new genus of Hydrobiidae from the Ohrid Lake basin (Mollusca: Gastropoda)" masterLastPageNumber="190" masterPageNumber="185" pageNumber="186" updateTime="1722980748538" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C1599DDEAC7EFD097A9F71C3035173A8">A new Ancylus and a new genus of Hydrobiidae from the Ohrid Lake basin (Mollusca: Gastropoda)</mods:title>
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<mods:name id="484FC8DE2E4E9853BB4539AE4E17C77B" type="personal">
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<mods:namePart id="011669A5960A94C0E8612949C6738E2C">Glöer, Peter</mods:namePart>
<mods:affiliation id="56F9635DAEDC281321C0C51AEF318073">Schulstrasse 3, D- 25491 Hetlingen, Germany. E-mail: gloeer @ malaco. de</mods:affiliation>
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<mods:namePart id="75398F87412334384909A750552ADB18">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="178CA77502BB0C6A4BD9EF9042211131">Department of Biology, Faculty of Sciences, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro. E-mail: vladopesic @ gmail. com, https: // orcid. org / 0000 - 0002 - 9724 - 345 X</mods:affiliation>
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<mods:title id="DDD276B98EE50C377355E2E483541342">Ecologica Montenegrina</mods:title>
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<mods:date id="726E797E55991A9F4426DEFC4250900E">2023</mods:date>
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<mods:number id="E22CF4C6A009D24A7F4FDD380CDBEDFE">2023-07-11</mods:number>
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<mods:number id="1FE643F9A195C92B10995165B54DC45E">64</mods:number>
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<mods:classification id="05CB8195E4896769AD8C4F89F9791EAF">journal article</mods:classification>
<mods:identifier id="F449AA2F8C39F0B50D93644BEF384EA1" type="DOI">10.37828/em.2023.64.2</mods:identifier>
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<treatment id="03CA6424FFA40E789893496AFE419FC8" ID-DOI="http://doi.org/10.5281/zenodo.13247472" ID-Zenodo-Dep="13247472" LSID="urn:lsid:zoobank.org:act:2CCBEB51-BA20-4BC4-85F8-E7254717E9A5" httpUri="http://treatment.plazi.org/id/03CA6424FFA40E789893496AFE419FC8" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
<subSubSection id="C37986B9FFA40E7B9893496AFBA09E20" pageId="1" pageNumber="186" type="nomenclature">
<paragraph id="8BDCD532FFA40E7B9893496AFE099E43" blockId="1.[151,1148,1545,1635]" box="[151,469,1545,1571]" pageId="1" pageNumber="186">
<heading id="D094625EFFA40E7B9893496AFE099E43" box="[151,469,1545,1571]" fontSize="11" level="2" pageId="1" pageNumber="186" reason="2">
<emphasis id="B9170920FFA40E7B9893496AFE099E43" bold="true" box="[151,469,1545,1571]" pageId="1" pageNumber="186">
<taxonomicName id="4C63AEB1FFA40E7B9893496AFE509E43" ID-CoL="BM3DM" authorityName="Glöer &amp; Pešić" authorityYear="2023" box="[151,396,1545,1571]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="1" pageNumber="186" phylum="Mollusca" rank="species" species="naumiensis" status="sp. nov.">
<emphasis id="B9170920FFA40E7B9893496AFE509E43" bold="true" box="[151,396,1545,1571]" italics="true" pageId="1" pageNumber="186">Ancylus naumiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224B45BFFA40E7B9997496AFE099E43" box="[403,469,1545,1571]" pageId="1" pageNumber="186" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDCD532FFA40E7B9893494AFBA09E20" blockId="1.[151,1148,1545,1635]" box="[151,1148,1577,1600]" pageId="1" pageNumber="186">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C37986B9FFA40E7B9893492AFD959EC6" pageId="1" pageNumber="186" type="description">
<paragraph id="8BDCD532FFA40E7B9893492AFEC79E03" blockId="1.[151,1148,1545,1635]" box="[151,283,1609,1635]" pageId="1" pageNumber="186">
<figureCitation id="1358C9B7FFA40E7B9893492AFF3A9E03" box="[151,230,1609,1635]" captionStart="Figure 2" captionStartId="2.[151,227,1010,1033]" captionTargetBox="[158,1430,189,991]" captionTargetId="figure-10@2.[157,1430,189,991]" captionTargetPageId="2" captionText="Figure 2. Ancylus naumiensis n. sp., holotype (A-B): A = shell, lateral view; B = shell, apical view; C-D = apex." figureDoi="http://doi.org/10.5281/zenodo.13246874" httpUri="https://zenodo.org/record/13246874/files/figure.png" pageId="1" pageNumber="186">Figs. 2</figureCitation>
A-D
</paragraph>
<paragraph id="8BDCD532FFA40E7B989349EFFD959EC6" blockId="1.[151,585,1676,1702]" box="[151,585,1676,1702]" pageId="1" pageNumber="186">
<taxonomicName id="4C63AEB1FFA40E7B989349EFFEE59EC6" box="[151,313,1676,1702]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="1" pageNumber="186" phylum="Mollusca" rank="species" species="undefined-D">
<emphasis id="B9170920FFA40E7B989349EFFF289EC6" box="[151,244,1676,1702]" italics="true" pageId="1" pageNumber="186">Ancylus</emphasis>
sp. D
</taxonomicName>
–
<bibRefCitation id="EFF2A8C3FFA40E7B995349EFFD999EC6" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[343,581,1676,1702]" pageId="1" pageNumber="186" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA40E7B99C149EFFE249EC6" box="[453,504,1676,1702]" italics="true" pageId="1" pageNumber="186">et al</emphasis>
. 2006
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37986B9FFA40E7B989349ACFD829FD4" pageId="1" pageNumber="186" type="materials_examined">
<paragraph id="8BDCD532FFA40E7B989349ACFEDA9F2F" blockId="1.[151,1437,1743,1871]" pageId="1" pageNumber="186">
<emphasis id="B9170920FFA40E7B989349ACFEEC9E89" bold="true" box="[151,304,1743,1769]" pageId="1" pageNumber="186">Type series</emphasis>
—
<materialsCitation id="3B0BDF6FFFA40E7B997749ACFD809F4D" collectingDate="2019-09-13" collectionCode="NORTH" collectorName="Pesic." country="Macedonia" county="Shell" latitude="40.91001" location="Black Drim" longLatPrecision="1" longitude="20.74754" municipality="Lake area" pageId="1" pageNumber="186" specimenCode="ZMH 141503" specimenCount="1" stateProvince="Ohrid" typeStatus="holotype">
<typeStatus id="54D86B90FFA40E7B997749ACFE3C9E89" box="[371,480,1743,1769]" pageId="1" pageNumber="186" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBC57D49FFA40E7B99FF49ACFD739E89" box="[507,687,1743,1769]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="1" pageNumber="186" type="Museum">ZMH 141503</specimenCode>
):
<collectingCounty id="62BDADBEFFA40E7B9ACA49ACFCD69E89" box="[718,778,1743,1769]" pageId="1" pageNumber="186">Shell</collectingCounty>
height
<quantity id="4C9B78D7FFA40E7B9B7E49ACFC2C9E89" box="[890,1008,1743,1769]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" pageId="1" pageNumber="186" unit="mm" value="2.66">2.66 mm</quantity>
, shell width
<quantity id="4C9B78D7FFA40E7B9CAC49ACFAFC9E89" box="[1192,1312,1743,1769]" metricMagnitude="-3" metricUnit="m" metricValue="6.07" pageId="1" pageNumber="186" unit="mm" value="6.07">6.07 mm</quantity>
;
<collectionCode id="ED724DF7FFA40E7B9D3349ACFA409E89" box="[1335,1436,1743,1769]" pageId="1" pageNumber="186">NORTH</collectionCode>
<collectingCountry id="F37495A2FFA40E7B98934992FE919F6B" box="[151,333,1777,1803]" name="Macedonia" pageId="1" pageNumber="186">MACEDONIA</collectingCountry>
,
<collectingRegion id="49A71BD0FFA40E7B995D4992FE429F6B" box="[345,414,1777,1803]" country="Macedonia" name="Ohrid" pageId="1" pageNumber="186">Ohrid</collectingRegion>
<collectingMunicipality id="6BB84F48FFA40E7B99A14992FDC79F6B" box="[421,539,1777,1803]" pageId="1" pageNumber="186">Lake area</collectingMunicipality>
, the springs of the
<location id="8EBC83E9FFA40E7B9AFD4992FC5F9F6B" LSID="urn:lsid:plazi:treatment:03CA6424FFA40E789893496AFE419FC8:8EBC83E9FFA40E7B9AFD4992FC5F9F6B" box="[761,899,1777,1803]" country="Macedonia" county="Shell" latitude="40.91001" longLatPrecision="1" longitude="20.74754" municipality="Lake area" name="Black Drim" pageId="1" pageNumber="186" stateProvince="Ohrid">Black Drim</location>
near
<location id="8EBC83E9FFA40E7B9BC04992FB079F6B" LSID="urn:lsid:plazi:treatment:03CA6424FFA40E789893496AFE419FC8:8EBC83E9FFA40E7B9BC04992FB079F6B" box="[964,1243,1777,1803]" country="Macedonia" county="Shell" latitude="40.91001" longLatPrecision="1" longitude="20.74754" municipality="Lake area" name="Monastery of St. Naum" pageId="1" pageNumber="186" stateProvince="Ohrid">Monastery of St. Naum</location>
,
<geoCoordinate id="EE57B3F5FFA40E7B9CE24992FA449F6B" box="[1254,1432,1777,1803]" degrees="40" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="186" precision="1" seconds="36.04" value="40.91001">40°54'36.04&quot;N</geoCoordinate>
,
<geoCoordinate id="EE57B3F5FFA40E7B98934870FE999F4D" box="[151,325,1811,1837]" degrees="20" direction="east" minutes="44" orientation="longitude" pageId="1" pageNumber="186" precision="1" seconds="51.14" value="20.74754">20°44'51.14&quot;E</geoCoordinate>
,
<date id="FFDDF3F2FFA40E7B99504870FE0B9F4D" box="[340,471,1811,1837]" pageId="1" pageNumber="186" value="2019-09-13">
<collectingDate id="EF990A1AFFA40E7B99504870FE0B9F4D" box="[340,471,1811,1837]" pageId="1" pageNumber="186" value="2019-09-13">13.09.2019</collectingDate>
</date>
leg.
<collectorName id="2696B0E4FFA40E7B9A124870FD809F4D" box="[534,604,1811,1837]" pageId="1" pageNumber="186">Pešić.</collectorName>
</materialsCitation>
<materialsCitation id="3B0BDF6FFFA40E7B9A614870FC0E9F4D" box="[613,978,1811,1837]" collectionCode="ZMH" country="Macedonia" pageId="1" pageNumber="186" specimenCode="ZMH 141504" specimenCount="6" stateProvince="Ohrid" typeStatus="paratype">
<typeStatus id="54D86B90FFA40E7B9A614870FD079F4D" box="[613,731,1811,1837]" pageId="1" pageNumber="186" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D651EBBFFA40E7B9AEC4870FCFF9F4D" box="[744,803,1811,1837]" count="6" pageId="1" pageNumber="186" type="generic">6 ex.</specimenCount>
<specimenCode id="DBC57D49FFA40E7B9B294870FC0E9F4D" box="[813,978,1811,1837]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="1" pageNumber="186" type="Museum">ZMH 141504</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B0BDF6FFFA40E7B9BE54870FEDE9F2F" collectingDate="2019-09-13" collectionCode="NORTH" collectorName="Gloer" country="Macedonia" county="Shell" latitude="40.91001" location="Black Drim" longLatPrecision="1" longitude="20.74754" municipality="Lake area" pageId="1" pageNumber="186" specimenCount="3" stateProvince="Ohrid" typeStatus="paratype">
<specimenCount id="9D651EBBFFA40E7B9BE54870FBC09F4D" box="[993,1052,1811,1837]" count="3" pageId="1" pageNumber="186" type="generic">3 ex.</specimenCount>
in coll.
<collectorName id="2696B0E4FFA40E7B9C874870FB149F4D" box="[1155,1224,1811,1837]" pageId="1" pageNumber="186">Glöer</collectorName>
, same data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BDCD532FFA40E7B9893481BFD829FD4" blockId="1.[151,1436,1912,1972]" pageId="1" pageNumber="186">
<materialsCitation id="3B0BDF6FFFA40E7B9893481BFD869FD4" country="Macedonia" county="North" latitude="40.91001" location="Black Drim" longLatPrecision="1" longitude="20.74754" municipality="Lake region" pageId="1" pageNumber="186" specimenCount="1" stateProvince="Ohrid" typeStatus="holotype">
<emphasis id="B9170920FFA40E7B9893481BFEE69FF2" bold="true" box="[151,314,1912,1938]" pageId="1" pageNumber="186">
<typeStatus id="54D86B90FFA40E7B9893481BFF0B9FF2" box="[151,215,1912,1938]" pageId="1" pageNumber="186">Type</typeStatus>
locality
</emphasis>
—
<collectingCounty id="62BDADBEFFA40E7B996D481BFE729FF2" box="[361,430,1912,1938]" pageId="1" pageNumber="186">North</collectingCounty>
<collectingCountry id="F37495A2FFA40E7B99B1481BFDE79FF2" box="[437,571,1912,1938]" name="Macedonia" pageId="1" pageNumber="186">Macedonia</collectingCountry>
,
<collectingRegion id="49A71BD0FFA40E7B9A43481BFD509FF2" box="[583,652,1912,1938]" country="Macedonia" name="Ohrid" pageId="1" pageNumber="186">Ohrid</collectingRegion>
<collectingMunicipality id="6BB84F48FFA40E7B9A90481BFCF99FF2" box="[660,805,1912,1938]" pageId="1" pageNumber="186">Lake region</collectingMunicipality>
, the springs of the
<location id="8EBC83E9FFA40E7B9C0C481BFB4F9FF2" LSID="urn:lsid:plazi:treatment:03CA6424FFA40E789893496AFE419FC8:8EBC83E9FFA40E7B9C0C481BFB4F9FF2" box="[1032,1171,1912,1938]" country="Macedonia" county="North" latitude="40.91001" longLatPrecision="1" longitude="20.74754" municipality="Lake region" name="Black Drim" pageId="1" pageNumber="186" stateProvince="Ohrid">Black Drim</location>
near
<location id="8EBC83E9FFA40E7B9CD2481BFF3D9FD4" LSID="urn:lsid:plazi:treatment:03CA6424FFA40E789893496AFE419FC8:8EBC83E9FFA40E7B9CD2481BFF3D9FD4" country="Macedonia" county="North" latitude="40.91001" longLatPrecision="1" longitude="20.74754" municipality="Lake region" name="Monastery of St. Naum" pageId="1" pageNumber="186" stateProvince="Ohrid">Monastery of St. Naum</location>
,
<geoCoordinate id="EE57B3F5FFA40E7B98E948F9FE439FD4" box="[237,415,1946,1972]" degrees="40" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="186" precision="1" seconds="36.04" value="40.91001">40°54'36.04&quot;N</geoCoordinate>
,
<geoCoordinate id="EE57B3F5FFA40E7B99AF48F9FD869FD4" box="[427,602,1946,1972]" degrees="20" direction="east" minutes="44" orientation="longitude" pageId="1" pageNumber="186" precision="1" seconds="51.14" value="20.74754">20°44'51.14&quot;E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37986B9FFA40E78989348BEFF3C9C84" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186" type="etymology">
<paragraph id="8BDCD532FFA40E78989348BEFF3C9C84" blockId="1.[151,1405,2013,2039]" lastBlockId="2.[151,1436,1158,1252]" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
<emphasis id="B9170920FFA40E7B989348BEFEC39F97" bold="true" box="[151,287,2013,2039]" pageId="1" pageNumber="186">Etymology</emphasis>
— Named after the Monastery of Saint Naum in whose vicinity the new species was collected.
<emphasis id="B9170920FFA70E7898934BE5FEF49CC0" bold="true" box="[151,296,1158,1184]" pageId="2" pageNumber="187">Description</emphasis>
— The shell is broad oval. The shell is
<quantity id="4C9B78D7FFA70E789B004BE5FC829CC0" box="[772,862,1158,1184]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="187" unit="mm" value="2.7">2.7 mm</quantity>
high and 6.0 mm long. The apex does not reach the border of the shell from apical view. Upper side of apex straight and not convex and the tip not bent down.
</paragraph>
</subSubSection>
<caption id="DF1C85BAFFA70E7898934C91FA889C69" ID-DOI="http://doi.org/10.5281/zenodo.13246874" ID-Zenodo-Dep="13246874" box="[151,1364,1010,1033]" httpUri="https://zenodo.org/record/13246874/files/figure.png" pageId="2" pageNumber="187" startId="2.[151,227,1010,1033]" targetBox="[158,1430,189,991]" targetPageId="2" targetType="figure">
<paragraph id="8BDCD532FFA70E7898934C91FA889C69" blockId="2.[151,1364,1010,1033]" box="[151,1364,1010,1033]" pageId="2" pageNumber="187">
<emphasis id="B9170920FFA70E7898934C91FF2A9C69" bold="true" box="[151,246,1010,1033]" pageId="2" pageNumber="187">Figure 2</emphasis>
.
<taxonomicName id="4C63AEB1FFA70E78990F4C91FE029C69" authorityName="Glöer &amp; Pešić" authorityYear="2023" box="[267,478,1010,1033]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="naumiensis" status="sp. nov.">
<emphasis id="B9170920FFA70E78990F4C91FE029C69" box="[267,478,1010,1033]" italics="true" pageId="2" pageNumber="187">Ancylus naumiensis</emphasis>
</taxonomicName>
<emphasis id="B9170920FFA70E7899E04C91FDFB9C69" bold="true" box="[484,551,1010,1033]" pageId="2" pageNumber="187">
<taxonomicNameLabel id="A224B45BFFA70E7899E04C91FDFD9C69" box="[484,545,1010,1033]" pageId="2" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype (A-B): A = shell, lateral view; B = shell, apical view; C-D = apex.
</paragraph>
</caption>
<subSubSection id="C37986B9FFA70E7898934A6EFDA29F23" pageId="2" pageNumber="187" type="diagnosis">
<paragraph id="8BDCD532FFA70E7898934A6EFDA69E18" blockId="2.[151,1437,1293,1758]" pageId="2" pageNumber="187">
<emphasis id="B9170920FFA70E7898934A6EFE779D47" bold="true" box="[151,427,1293,1319]" pageId="2" pageNumber="187">Differential diagnosis</emphasis>
— Four species of the genus
<taxonomicName id="4C63AEB1FFA70E789B4F4A6EFC749D47" authorityName="O. F. Muller" authorityYear="1773" box="[843,936,1293,1319]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="genus">
<emphasis id="B9170920FFA70E789B4F4A6EFC749D47" box="[843,936,1293,1319]" italics="true" pageId="2" pageNumber="187">Ancylus</emphasis>
</taxonomicName>
are known from the
<collectingRegion id="49A71BD0FFA70E789CC64A6EFADB9D47" box="[1218,1287,1293,1319]" country="Macedonia" name="Ohrid" pageId="2" pageNumber="187">Ohrid</collectingRegion>
Lake basin (
<bibRefCitation id="EFF2A8C3FFA70E78989B4A4CFE719D29" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[159,429,1327,1353]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E78991C4A4CFE899D29" box="[280,341,1327,1353]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. 2006
</bibRefCitation>
).
<taxonomicName id="4C63AEB1FFA70E7899CE4A4CFCB99D29" authority="Polinski, 1929" authorityName="Polinski" authorityYear="1929" box="[458,869,1327,1353]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="tapirulus">
<emphasis id="B9170920FFA70E7899CE4A4CFD7D9D29" box="[458,673,1327,1353]" italics="true" pageId="2" pageNumber="187">Ancylus tapirulus</emphasis>
Poliński, 1929
</taxonomicName>
is characterized by a sharp apex, while
<taxonomicName id="4C63AEB1FFA70E789D804A4CFD7E9D0A" authority="Stankovic &amp; Radoman, 1953" authorityName="Stankovic &amp; Radoman" authorityYear="1953" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="scalariformis">
<emphasis id="B9170920FFA70E789D804A4CFEE89D0A" italics="true" pageId="2" pageNumber="187">A. scalariformis</emphasis>
Stanković &amp; Radoman, 1953
</taxonomicName>
is differentiated by a characteristic shape of the shell, with a number of concentric eaves-like projections. The both species live in the “shell zone” (
<quantity id="4C9B78D7FFA70E789CA24A11FAD29DEC" box="[1190,1294,1394,1420]" metricMagnitude="1" metricUnit="m" metricValue="2.65" metricValueMax="3.5" metricValueMin="1.8" pageId="2" pageNumber="187" unit="m" value="26.5" valueMax="35.0" valueMin="18.0">18-35 m</quantity>
), mainly on the zebra mussel
<taxonomicName id="4C63AEB1FFA70E78996C4AF7FD839DCE" authorityName="Lvova &amp; Starobogatov" authorityYear="1982" box="[360,607,1428,1454]" class="Bivalvia" family="Dreissenidae" genus="Dreissena" kingdom="Animalia" order="Myida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="stankovici">
<emphasis id="B9170920FFA70E78996C4AF7FD839DCE" box="[360,607,1428,1454]" italics="true" pageId="2" pageNumber="187">Dreissena stankovici</emphasis>
</taxonomicName>
. The third species,
<taxonomicName id="4C63AEB1FFA70E789B494AF7FB2B9DCE" authority="Hubendick, 1960" authorityName="Hubendick" authorityYear="1960" box="[845,1271,1428,1454]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="lapicidus">
<emphasis id="B9170920FFA70E789B494AF7FBC39DCE" box="[845,1055,1428,1454]" italics="true" pageId="2" pageNumber="187">Ancylus lapicidus</emphasis>
<bibRefCitation id="EFF2A8C3FFA70E789C2C4AF7FB2B9DCE" author="Hubendick, B." box="[1064,1271,1428,1454]" pageId="2" pageNumber="187" pagination="497 - 529" refId="ref2288" refString="Hubendick, B. (1960) The Ancylidae of lake Ochrid and their bearing on intralacustrine speciation. Proceedings of the Zoological Society of London, 133, 497 - 529. https: // doi. org / 10.1111 / j. 1469 - 7998.1960. tb 05577. x" type="journal article" year="1960">Hubendick, 1960</bibRefCitation>
</taxonomicName>
, resembles
<taxonomicName id="4C63AEB1FFA70E789D874AF7FEE89DB0" authorityName="Stankovic &amp; Radoman" authorityYear="1953" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="scalariformis">
<emphasis id="B9170920FFA70E789D874AF7FEE89DB0" italics="true" pageId="2" pageNumber="187">A. scalariformis</emphasis>
</taxonomicName>
in possessing comarginal ribss (
<bibRefCitation id="EFF2A8C3FFA70E789ADA4AD5FC719DB0" author="Hubendick, B." box="[734,941,1462,1488]" pageId="2" pageNumber="187" pagination="497 - 529" refId="ref2288" refString="Hubendick, B. (1960) The Ancylidae of lake Ochrid and their bearing on intralacustrine speciation. Proceedings of the Zoological Society of London, 133, 497 - 529. https: // doi. org / 10.1111 / j. 1469 - 7998.1960. tb 05577. x" type="journal article" year="1960">Hubendick 1960</bibRefCitation>
) and is restricted to the littoral zone, inhabiting the rocky substrate on the eastern side of Lake
<collectingRegion id="49A71BD0FFA70E789B444AB4FC559D91" box="[832,905,1495,1521]" country="Macedonia" name="Ohrid" pageId="2" pageNumber="187">Ohrid</collectingRegion>
. Fourth species, putatively named as
<taxonomicName id="4C63AEB1FFA70E789D444AB4FF0A9E73" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="undefined-D">
<emphasis id="B9170920FFA70E789D444AB4FA419D91" box="[1344,1437,1495,1521]" italics="true" pageId="2" pageNumber="187">Ancylus</emphasis>
sp. D
</taxonomicName>
was separated by
<bibRefCitation id="EFF2A8C3FFA70E7899B54A9AFD6E9E73" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[433,690,1529,1555]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E789A244A9AFD889E73" box="[544,596,1529,1555]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. (2006)
</bibRefCitation>
based on DNA data from two mitochondrial genes (COI, LSU rDNA), and includes specimens collected from springs near the southeastern edge of the lake (St. Naum in North
<collectingCountry id="F37495A2FFA70E7898E1495EFEBB9E37" box="[229,359,1597,1623]" name="Macedonia" pageId="2" pageNumber="187">Macedonia</collectingCountry>
and near Tušemišta in
<collectingCountry id="F37495A2FFA70E789A7B495EFD3D9E37" box="[639,737,1597,1623]" name="Albania" pageId="2" pageNumber="187">Albania</collectingCountry>
). The latter species, here described as
<taxonomicName id="4C63AEB1FFA70E789CAF495EFA8C9E37" authorityName="Glöer &amp; Pešić" authorityYear="2023" box="[1195,1360,1597,1623]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Siphonariida" pageId="2" pageNumber="187" phylum="Mollusca" rank="species" species="naumiensis" status="n. sp.">
<emphasis id="B9170920FFA70E789CAF495EFA8C9E37" box="[1195,1360,1597,1623]" italics="true" pageId="2" pageNumber="187">A. naumiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224B45BFFA70E789D5D495EFA409E37" box="[1369,1436,1597,1623]" pageId="2" pageNumber="187" rank="species">
<emphasis id="B9170920FFA70E789D5D495EFA499E37" bold="true" box="[1369,1429,1597,1623]" pageId="2" pageNumber="187">n. sp</emphasis>
.
</taxonomicNameLabel>
is diiferentiated by the shape of the apex.
</paragraph>
<paragraph id="8BDCD532FFA70E7898F149E3FA449EBE" blockId="2.[151,1437,1293,1758]" pageId="2" pageNumber="187">
All four species, as shown by
<bibRefCitation id="EFF2A8C3FFA70E789A7F49E3FC509EFA" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[635,908,1664,1690]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E789AEB49E3FCF59EFA" box="[751,809,1664,1690]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. (2006)
</bibRefCitation>
form a monophyletic group that probably evolved in the
<collectingRegion id="49A71BD0FFA70E78994C49C1FE509EDC" box="[328,396,1698,1724]" country="Macedonia" name="Ohrid" pageId="2" pageNumber="187">Ohrid</collectingRegion>
Lake Basin, also supported by the morphological similarity of the species of this group sharing a very characteristic appearance of the radula with solid teeth without cusplets (
<bibRefCitation id="EFF2A8C3FFA70E789C9E49A7FA579EBE" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[1178,1419,1732,1758]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E789D0E49A7FAE19EBE" box="[1290,1341,1732,1758]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
<paragraph id="8BDCD532FFA70E7898934864FDA29F23" blockId="2.[151,1437,1799,1859]" pageId="2" pageNumber="187">
<emphasis id="B9170920FFA70E7898934864FF2B9F41" bold="true" box="[151,247,1799,1825]" pageId="2" pageNumber="187">Habitat</emphasis>
— Springs; crenobiontic.
<bibRefCitation id="EFF2A8C3FFA70E789A384864FC999F41" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[572,837,1799,1825]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E789AAA4864FD399F41" box="[686,741,1799,1825]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. (2006)
</bibRefCitation>
mentioned that despite an extensive search, they were unable to find this taxon in the lake.
</paragraph>
</subSubSection>
<subSubSection id="C37986B9FFA70E789893480FFE419FC8" pageId="2" pageNumber="187" type="distribution">
<paragraph id="8BDCD532FFA70E789893480FFE419FC8" blockId="2.[151,1436,1900,1960]" pageId="2" pageNumber="187">
<emphasis id="B9170920FFA70E789893480FFEEC9FE6" bold="true" box="[151,304,1900,1926]" pageId="2" pageNumber="187">Distribution</emphasis>
— Known from the St. Naum springs in North
<collectingCountry id="F37495A2FFA70E789B80480FFBDA9FE6" box="[900,1030,1900,1926]" name="Macedonia" pageId="2" pageNumber="187">Macedonia</collectingCountry>
and Tusemista spring in
<collectingCountry id="F37495A2FFA70E789D3B480FFA409FE6" box="[1343,1436,1900,1926]" name="Albania" pageId="2" pageNumber="187">Albania</collectingCountry>
(
<bibRefCitation id="EFF2A8C3FFA70E78989B48EDFE4D9FC8" author="Albrecht, C. &amp; Trajanovski, S. &amp; Kuhn, K. &amp; Streit, B. &amp; Wilke, T." box="[159,401,1934,1960]" pageId="2" pageNumber="187" pagination="294 - 307" refId="ref2081" refString="Albrecht, C., Trajanovski, S., Kuhn, K., Streit, B. &amp; Wilke, T. (2006) Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid. Organisms, Diversity &amp; Evolution, 6, 294 - 307. https: // doi. org / 10.1016 / j. ode. 2005.12.003" type="journal article" year="2006">
Albrecht
<emphasis id="B9170920FFA70E78990B48EDFE9F9FC8" box="[271,323,1934,1960]" italics="true" pageId="2" pageNumber="187">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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