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<document id="279ACF33EAFB569A8E5F7114E3117358" ID-CLB-Dataset="162544" ID-DOI="10.11646/zootaxa.5195.2.9" ID-GBIF-Dataset="34b55a6b-c747-4ef5-b12d-040a4855449c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7184813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665472873254" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Levesque-Beaudin, Valerie" docDate="2022" docId="03AA87B7275E6C24BAFCCBE615E6FA8B" docLanguage="en" docName="zootaxa.5195.2.9.pdf" docOrigin="Zootaxa 5195 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cabamofa alexsmithi Jaschhof &amp; Levesque-Beaudin 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="199" masterDocId="FF93FFCF275E6C27BA6BCE121678FFFC" masterDocTitle="Cabamofa alexsmithi sp. nov., the second species of Cabamofa in Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis)" masterLastPageNumber="200" masterPageNumber="196" pageNumber="196" updateTime="1699407240450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="95F55EEF27FD024748FFA43DB0DCD073">Cabamofa alexsmithi sp. nov., the second species of Cabamofa in Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis)</mods:title>
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<mods:name id="BC36DD4D2C06E29B7DC80AD72F6829D6" type="personal">
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<mods:namePart id="97EB236006E2FF1356696833AB56B0CD">Jaschhof, Mathias</mods:namePart>
<mods:affiliation id="7272B239575883D9DE669FC0D890403B">Station Linné, Ölands Skogsby 161, 38693 Färjestaden, Sweden</mods:affiliation>
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<mods:namePart id="7FE682E55B4959C7AC94F7D629FC9B58">Levesque-Beaudin, Valerie</mods:namePart>
<mods:affiliation id="6E538F087A9C0DE83C158244C97C07A3">University of Guelph, Centre for Biodiversity Genomics, Guelph, Ontario, Canada</mods:affiliation>
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<mods:title id="3EFF78664DE51025DE8825F530E933E9">Zootaxa</mods:title>
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<mods:part id="F25341E8734F8AEA2489E8FBD22095AB">
<mods:date id="98ECA0977DDA4370CBBBB862E24B5563">2022</mods:date>
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<mods:number id="A4905C7BD8EFA88CC54FC748767092A5">2022-10-11</mods:number>
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<mods:number id="6EA5EFE6978F1118BBA87FD7670FEAEA">5195</mods:number>
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<mods:number id="DBF319254D19EA419AEC92A1CBF01450">2</mods:number>
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<mods:identifier id="B8757DD19A44D7AD6ACC8EDF42D9CC00" type="DOI">10.11646/zootaxa.5195.2.9</mods:identifier>
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<treatment id="03AA87B7275E6C24BAFCCBE615E6FA8B" ID-DOI="http://doi.org/10.5281/zenodo.7186482" ID-GBIF-Taxon="202593490" ID-Zenodo-Dep="7186482" LSID="urn:lsid:plazi:treatment:03AA87B7275E6C24BAFCCBE615E6FA8B" httpUri="http://treatment.plazi.org/id/03AA87B7275E6C24BAFCCBE615E6FA8B" lastPageId="3" lastPageNumber="199" pageId="0" pageNumber="196">
<subSubSection id="C319652A275E6C27BAFCCBE615FBF9F3" box="[151,899,1524,1551]" pageId="0" pageNumber="196" type="nomenclature">
<paragraph id="8BBC36A1275E6C27BAFCCBE615FBF9F3" blockId="0.[151,1437,1524,2018]" box="[151,899,1524,1551]" pageId="0" pageNumber="196">
<heading id="D0F481CD275E6C27BAFCCBE615FBF9F3" bold="true" box="[151,899,1524,1551]" fontSize="11" level="1" pageId="0" pageNumber="196" reason="1">
<emphasis id="B977EAB3275E6C27BAFCCBE615FBF9F3" bold="true" box="[151,899,1524,1551]" pageId="0" pageNumber="196">
<taxonomicName id="4C034D22275E6C27BAFCCBE61566F9F2" authority="Jaschhof &amp; Levesque-Beaudin" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[151,798,1524,1551]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi" status="sp. nov.">
<emphasis id="B977EAB3275E6C27BAFCCBE617E2F9F3" bold="true" box="[151,410,1524,1551]" italics="true" pageId="0" pageNumber="196">Cabamofa alexsmithi</emphasis>
Jaschhof &amp; Levesque-Beaudin
</taxonomicName>
<taxonomicNameLabel id="A24457C8275E6C27B94DCBE715FBF9F3" box="[806,899,1525,1551]" pageId="0" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C319652A275E6C27BAFCC808177EF9CF" box="[151,262,1562,1587]" pageId="0" pageNumber="196" type="description">
<paragraph id="8BBC36A1275E6C27BAFCC808177EF9CF" blockId="0.[151,1437,1524,2018]" box="[151,262,1562,1587]" pageId="0" pageNumber="196">
(
<figureCitation id="13382A24275E6C27BAF4C8081684F9CF" box="[159,252,1562,1587]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–8" captionStartId-0="1.[152,270,1862,1886]" captionStartId-1="2.[152,270,1495,1519]" captionTargetBox-0="[164,1436,596,1826]" captionTargetBox-1="[162,1425,190,1469]" captionTargetId-0="figure-249@1.[151,1436,596,1839]" captionTargetId-1="figure-17@2.[151,1436,181,1471]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–2. Cabamofa alexsmithi sp. nov., male paratype. 1: Habitus, lateral (specimen in ethanol after DNA extraction). 1A: Posterior abdomen including genitalia, enlargement from 1. 2: Ditto, dorsolateral. Scale for 1, 1.0 mm, for 1A and 2, 0.2 mm. Abbreviations: gen = genitalia, sg6 (‒sg8) = sixth segment (‒eighth segment), st7 (‒st8) = seventh sternite (‒eighth sternite), tg7 (‒tg8) = seventh tergite (‒eighth tergite)." captionText-1="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi-0="http://doi.org/10.5281/zenodo.7184815" figureDoi-1="http://doi.org/10.5281/zenodo.7184817" httpUri-0="https://zenodo.org/record/7184815/files/figure.png" httpUri-1="https://zenodo.org/record/7184817/files/figure.png" pageId="0" pageNumber="196">Figs 1–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C319652A275E6C26BAFCC82C1270FF28" lastPageId="1" lastPageNumber="197" pageId="0" pageNumber="196" type="diagnosis">
<paragraph id="8BBC36A1275E6C26BAFCC82C1270FF28" blockId="0.[151,1437,1524,2018]" lastBlockId="1.[151,1437,151,572]" lastPageId="1" lastPageNumber="197" pageId="0" pageNumber="196">
<emphasis id="B977EAB3275E6C27BAFCC82C17EBF9AA" bold="true" box="[151,403,1598,1622]" inLineHeading="true" pageId="0" pageNumber="196" reason="1">Differential diagnosis.</emphasis>
Adults of the two Neotropical
<taxonomicName id="4C034D22275E6C27B883C82C152FF9AA" authorityName="Jaschhof" authorityYear="2005" box="[744,855,1598,1622]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAB3275E6C27B883C82C152FF9AA" box="[744,855,1598,1622]" italics="true" pageId="0" pageNumber="196">Cabamofa</emphasis>
</taxonomicName>
differ in vein M 1+2, which in
<taxonomicName id="4C034D22275E6C27BEF0C82D1352F9AA" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[1179,1322,1598,1622]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27BEF0C82D1352F9AA" box="[1179,1322,1598,1622]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
resembles the shape of an U (
<figureCitation id="13382A24275E6C27BB0DC87017DEF987" box="[358,422,1634,1659]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="0" pageNumber="196">Fig. 4</figureCitation>
) whereas in
<taxonomicName id="4C034D22275E6C27B859C87114FCF986" authorityName="Jaschhof" authorityYear="2005" box="[562,644,1635,1658]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27B859C87114FCF986" box="[562,644,1635,1658]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
the U has a short stem (
<bibRefCitation id="EF924B50275E6C27B9E7C8701254F986" author="Jaschhof, M." box="[908,1068,1634,1658]" pageId="0" pageNumber="196" pagination="323 - 332" refId="ref2769" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof 2005</bibRefCitation>
: fig. 24). This distinction is stable among all the specimens known of both species (
<taxonomicName id="4C034D22275E6C27B8F6C8951551F962" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[669,809,1670,1694]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27B8F6C8951551F962" box="[669,809,1670,1694]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
, n = 5;
<taxonomicName id="4C034D22275E6C27B913C89515B0F962" authorityName="Jaschhof" authorityYear="2005" box="[888,968,1671,1694]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27B913C89515B0F962" box="[888,968,1671,1694]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
, n = 3). Also, in some, but not all specimens known of
<taxonomicName id="4C034D22275E6C27BB6FC8B917E9F93E" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[260,401,1706,1730]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27BB6FC8B917E9F93E" box="[260,401,1706,1730]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
vein CuA is slightly sinuous (
<figureCitation id="13382A24275E6C27B8BFC8B8156EF93F" box="[724,790,1706,1731]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="0" pageNumber="196">Fig. 4</figureCitation>
), not straight apically as in
<taxonomicName id="4C034D22275E6C27BE55C8B912F7F93E" authorityName="Jaschhof" authorityYear="2005" box="[1086,1167,1707,1730]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27BE55C8B912F7F93E" box="[1086,1167,1707,1730]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF924B50275E6C27BEF6C8B81344F93E" author="Jaschhof, M." box="[1181,1340,1706,1730]" pageId="0" pageNumber="196" pagination="323 - 332" refId="ref2769" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof 2005</bibRefCitation>
: fig. 24). The two Oriental species of
<taxonomicName id="4C034D22275E6C27BBACC8DC144EF91A" authorityName="Jaschhof" authorityYear="2005" box="[455,566,1742,1766]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAB3275E6C27BBACC8DC144EF91A" box="[455,566,1742,1766]" italics="true" pageId="0" pageNumber="196">Cabamofa</emphasis>
</taxonomicName>
have a stemless M
<subScript id="178734E4275E6C27B968C8CD1563F911" attach="left" box="[771,795,1759,1773]" fontSize="6" pageId="0" pageNumber="196">1+2</subScript>
and straight CuA (
<bibRefCitation id="EF924B50275E6C27B988C8DC1296F91B" author="Jaschhof, M. &amp; Sevcik, J." box="[995,1262,1742,1769]" pageId="0" pageNumber="196" pagination="392 - 394" refId="ref2897" refString="Jaschhof, M. &amp; Sevcik, J. (2019) Cabamofa orientalis sp. nov. from Thailand, congeneric with C. mira Jaschhof from Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis). Zootaxa, 4576 (2), 392 - 394. https: // doi. org / 10.11646 / zootaxa. 4576.2.12" type="journal article" year="2019">Jaschhof &amp; Ševčík 2019</bibRefCitation>
: fig. 2,
<bibRefCitation id="EF924B50275E6C27BF54C8DC1776F8F7" author="Jaschhof, M. &amp; Levesque-Beaudin, V. &amp; Broadley, A. &amp; Heller, K. &amp; Lien, V. V. &amp; Schmidt, S." pageId="0" pageNumber="196" pagination="297 - 300" refId="ref2812" refString="Jaschhof, M., Levesque-Beaudin, V., Broadley, A., Heller, K., Lien, V. V. &amp; Schmidt, S. (2022) Description of Cabamofa vietnamensis sp. nov., the second species of Cabamofa in mainland southeast Asia (Diptera: Bibionomorpha: Sciaroidea incertae sedis). Zootaxa, 5182 (3), 297 - 300. https: // doi. org / 10.11646 / zootaxa. 5182.3.7" type="journal article" year="2022">
Jaschhof
<emphasis id="B977EAB3275E6C27BAFCC8E116B6F8F6" box="[151,206,1778,1802]" italics="true" pageId="0" pageNumber="196">et al.</emphasis>
2022
</bibRefCitation>
: fig. 2). Otherwise the females of
<taxonomicName id="4C034D22275E6C27B8EAC8E11568F8F6" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[641,784,1778,1802]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27B8EAC8E11568F8F6" box="[641,784,1778,1802]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C034D22275E6C27B92EC8E115E0F8F6" authorityName="Jaschhof" authorityYear="2005" box="[837,920,1779,1802]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27B92EC8E115E0F8F6" box="[837,920,1779,1802]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
are indistinguishable from each other, although the spermathecae of
<taxonomicName id="4C034D22275E6C27BB10C9051473F8D2" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[379,523,1814,1838]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27BB10C9051473F8D2" box="[379,523,1814,1838]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
are partially covered with dense, tiny spikes, a structure not described for
<taxonomicName id="4C034D22275E6C27BF23C9051730F8AE" authority="(Jaschhof 2005)" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27BF23C90513E4F8D2" box="[1352,1436,1815,1838]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
(
<bibRefCitation id="EF924B50275E6C27BAF5C9281738F8AE" author="Jaschhof, M." box="[158,320,1850,1874]" pageId="0" pageNumber="196" pagination="323 - 332" refId="ref2769" refString="Jaschhof, M. (2005) Rogambara and Cabamofa, two new genera of enigmatic sciaroids from Costa Rica (Insecta: Diptera: Sciaroidea). Bonner zoologische Beitrage, 53 (3 / 4), 323 - 332." type="journal article" year="2005">Jaschhof 2005</bibRefCitation>
)
</taxonomicName>
. Females of the Oriental species are unknown. Males of
<taxonomicName id="4C034D22275E6C27B9D0C9291232F8AE" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[955,1098,1850,1874]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27B9D0C9291232F8AE" box="[955,1098,1850,1874]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C034D22275E6C27BEE9C92912AEF8AE" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[1154,1238,1851,1874]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27BEE9C92912AEF8AE" box="[1154,1238,1851,1874]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
differ in genitalic structures, notably the aedeagal apodeme, whose apex is pointed in
<taxonomicName id="4C034D22275E6C27B908C94D1588F88A" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[867,1008,1886,1910]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27B908C94D1588F88A" box="[867,1008,1886,1910]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
(
<figureCitation id="13382A24275E6C27B996C94C1245F88B" box="[1021,1085,1886,1911]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="0" pageNumber="196">Fig. 7</figureCitation>
) and furcate in
<taxonomicName id="4C034D22275E6C27BE89C94D134BF88A" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[1250,1331,1887,1910]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27BE89C94D134BF88A" box="[1250,1331,1887,1910]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF924B50275E6C27BF2BC94C1746F867" author="Amorim, D. S. &amp; Rindal, E." pageId="0" pageNumber="196" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 45), and the tegmen, whose posterior edge is markedly incised in
<taxonomicName id="4C034D22275E6C27BE20C99112A2F866" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[1099,1242,1922,1946]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275E6C27BE20C99112A2F866" box="[1099,1242,1922,1946]" italics="true" pageId="0" pageNumber="196">C. alexsmithi</emphasis>
</taxonomicName>
(
<figureCitation id="13382A24275E6C27BE81C9901354F867" box="[1258,1324,1922,1947]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="0" pageNumber="196">Fig. 7</figureCitation>
) and only slightly concave in
<taxonomicName id="4C034D22275E6C27BB0FC9B517CCF842" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[356,436,1959,1982]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275E6C27BB0FC9B517CCF842" box="[356,436,1959,1982]" italics="true" pageId="0" pageNumber="196">C. mira</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF924B50275E6C27BBA9C9B414BCF843" author="Amorim, D. S. &amp; Rindal, E." box="[450,708,1958,1983]" pageId="0" pageNumber="196" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 45). Further possible distinctions in male genitalic structures are discussed below. Males of the two Neotropical
<taxonomicName id="4C034D22275E6C27B8FBC9D81487F81E" authorityName="Jaschhof" authorityYear="2005" box="[656,767,1994,2018]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAB3275E6C27B8FBC9D81487F81E" box="[656,767,1994,2018]" italics="true" pageId="0" pageNumber="196">Cabamofa</emphasis>
</taxonomicName>
are abundantly different from males in the Oriental region, for instance in having shorter flagellomeres (
<figureCitation id="13382A24275F6C26B827CE8514F2FF4C" box="[588,650,151,176]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 3</figureCitation>
<emphasis id="B977EAB3275F6C26B8FBCE8A14ACFF53" box="[656,724,152,175]" italics="true" pageId="1" pageNumber="197">versus</emphasis>
<bibRefCitation id="EF924B50275F6C26B8B1CE8515A7FF4C" author="Jaschhof, M. &amp; Sevcik, J." box="[730,991,151,178]" pageId="1" pageNumber="197" pagination="392 - 394" refId="ref2897" refString="Jaschhof, M. &amp; Sevcik, J. (2019) Cabamofa orientalis sp. nov. from Thailand, congeneric with C. mira Jaschhof from Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis). Zootaxa, 4576 (2), 392 - 394. https: // doi. org / 10.11646 / zootaxa. 4576.2.12" type="journal article" year="2019">Jaschhof &amp; Ševčík 2019</bibRefCitation>
: fig. 1) and markedly shorter apical palpal segments (
<bibRefCitation id="EF924B50275F6C26BB63CEA91475FF28" author="Amorim, D. S. &amp; Rindal, E." box="[264,525,187,212]" pageId="1" pageNumber="197" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 39
<emphasis id="B977EAB3275F6C26B80FCEAE14D0FF2F" box="[612,680,188,211]" italics="true" pageId="1" pageNumber="197">versus</emphasis>
<bibRefCitation id="EF924B50275F6C26B8C4CEA915CEFF28" author="Jaschhof, M. &amp; Sevcik, J." box="[687,950,187,214]" pageId="1" pageNumber="197" pagination="392 - 394" refId="ref2897" refString="Jaschhof, M. &amp; Sevcik, J. (2019) Cabamofa orientalis sp. nov. from Thailand, congeneric with C. mira Jaschhof from Costa Rica (Diptera: Bibionomorpha: Sciaroidea incertae sedis). Zootaxa, 4576 (2), 392 - 394. https: // doi. org / 10.11646 / zootaxa. 4576.2.12" type="journal article" year="2019">Jaschhof &amp; Ševčík 2019</bibRefCitation>
: fig. 3).
</paragraph>
</subSubSection>
<subSubSection id="C319652A275F6C24BAACCECD15A1FBF3" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197" type="discussion">
<paragraph id="8BBC36A1275F6C26BAACCECD12AEFDC7" blockId="1.[151,1437,151,572]" pageId="1" pageNumber="197">
<emphasis id="B977EAB3275F6C26BAACCECD173EFF0B" bold="true" box="[199,326,223,247]" inLineHeading="true" pageId="1" pageNumber="197" reason="1">Discussion.</emphasis>
There are likely further differences in male genitalic structures that could be used to distinguish
<taxonomicName id="4C034D22275F6C26BFEFCEF2177DFEE7" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275F6C26BFEFCEF2177DFEE7" italics="true" pageId="1" pageNumber="197">C. alexsmithi</emphasis>
</taxonomicName>
from
<emphasis id="B977EAB3275F6C26BB22CF1617DCFEE7" box="[329,420,260,283]" italics="true" pageId="1" pageNumber="197">
<taxonomicName id="4C034D22275F6C26BB22CF1617D8FEE7" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[329,416,260,283]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="mira">C. mira</taxonomicName>
.
</emphasis>
For instance, the tegmen of
<taxonomicName id="4C034D22275F6C26B8B5CF161515FEE7" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[734,877,259,283]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275F6C26B8B5CF161515FEE7" box="[734,877,259,283]" italics="true" pageId="1" pageNumber="197">C. alexsmithi</emphasis>
</taxonomicName>
has two pairs of hooks ventrally, which are largely transparent and thus not as obvious as
<figureCitation id="13382A24275F6C26B85BCF351416FEC3" box="[560,622,295,319]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 7</figureCitation>
might suggest, whereas the tegmen of
<taxonomicName id="4C034D22275F6C26BE67CF3A1225FEC3" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[1036,1117,296,319]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275F6C26BE67CF3A1225FEC3" box="[1036,1117,296,319]" italics="true" pageId="1" pageNumber="197">C. mira</emphasis>
</taxonomicName>
was illustrated as lacking any substructures (
<bibRefCitation id="EF924B50275F6C26BB5ACF59144BFE98" author="Amorim, D. S. &amp; Rindal, E." box="[305,563,331,356]" pageId="1" pageNumber="197" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 45). Also, the ninth tergite of
<taxonomicName id="4C034D22275F6C26B9F6CF5E1251FE9F" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[925,1065,331,355]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275F6C26B9F6CF5E1251FE9F" box="[925,1065,331,355]" italics="true" pageId="1" pageNumber="197">C. alexsmithi</emphasis>
</taxonomicName>
is not as deeply incised posteriorly (
<figureCitation id="13382A24275F6C26BAF4CF7D1698FE7B" box="[159,224,367,391]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 6</figureCitation>
) as that drawn for
<taxonomicName id="4C034D22275F6C26BBC4CF62147AFE7B" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[431,514,368,391]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275F6C26BBC4CF62147AFE7B" box="[431,514,368,391]" italics="true" pageId="1" pageNumber="197">C. mira</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF924B50275F6C26B87ACF7D1561FE74" author="Amorim, D. S. &amp; Rindal, E." box="[529,793,367,392]" pageId="1" pageNumber="197" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 47), and the two species differ markedly regarding the outline of the hypoproct (
<figureCitation id="13382A24275F6C26BBCFCF81179BFE57" box="[420,483,403,427]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 6</figureCitation>
<emphasis id="B977EAB3275F6C26BB82CF861455FE57" box="[489,557,404,427]" italics="true" pageId="1" pageNumber="197">versus</emphasis>
<bibRefCitation id="EF924B50275F6C26B858CF81154DFE50" author="Amorim, D. S. &amp; Rindal, E." box="[563,821,403,428]" pageId="1" pageNumber="197" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
: fig. 47). Finally, one may expect interspecific differences in the construction and setosity of the gonostylus, although those might be hard to recognize from illustrations; in
<taxonomicName id="4C034D22275F6C26BFEFCFAA177DFE0F" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="alexsmithi">
<emphasis id="B977EAB3275F6C26BFEFCFAA177DFE0F" italics="true" pageId="1" pageNumber="197">C. alexsmithi</emphasis>
</taxonomicName>
, for instance, the three-lobed structure is so complex that different perspectives seem to show completely different gonostyli (
<figureCitation id="13382A24275F6C26BB00CFED17D1FDEB" box="[363,425,511,535]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 5</figureCitation>
<emphasis id="B977EAB3275F6C26BBDBCC12178CFDEB" box="[432,500,512,535]" italics="true" pageId="1" pageNumber="197">versus</emphasis>
<figureCitation id="13382A24275F6C26BB90CFED1442FDE4" box="[507,570,511,536]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="1" pageNumber="197">Fig. 8</figureCitation>
). Also, one cannot be sure whether the gonostylar setae in
<taxonomicName id="4C034D22275F6C26BEC0CC121284FDEB" baseAuthorityName="Jaschhof" baseAuthorityYear="2005" box="[1195,1276,512,535]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275F6C26BEC0CC121284FDEB" box="[1195,1276,512,535]" italics="true" pageId="1" pageNumber="197">C. mira</emphasis>
</taxonomicName>
are as uniform as illustrated by
<bibRefCitation id="EF924B50275F6C26BB2ECC311429FDC0" author="Amorim, D. S. &amp; Rindal, E." box="[325,593,547,572]" pageId="1" pageNumber="197" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal (2007</bibRefCitation>
: fig. 46) or whether this is a result of the drawing technique.
</paragraph>
<caption id="DF7C6629275F6C26BAF3C95414C3F836" ID-DOI="http://doi.org/10.5281/zenodo.7184815" ID-Zenodo-Dep="7184815" httpUri="https://zenodo.org/record/7184815/files/figure.png" pageId="1" pageNumber="197" startId="1.[152,270,1862,1886]" targetBox="[164,1436,596,1826]" targetPageId="1">
<paragraph id="8BBC36A1275F6C26BAF3C95414C3F836" blockId="1.[151,1437,1862,1995]" pageId="1" pageNumber="197">
<emphasis id="B977EAB3275F6C26BAF3C9541739F8A2" bold="true" box="[152,321,1862,1886]" pageId="1" pageNumber="197">FIGURES 1–2.</emphasis>
<taxonomicName id="4C034D22275F6C26BB2CC9541458F8A2" authority="Jaschhof &amp; Levesque-Beaudin, 2022" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[327,544,1862,1886]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="alexsmithi" status="sp. nov.">
<emphasis id="B977EAB3275F6C26BB2CC9541458F8A2" box="[327,544,1862,1886]" italics="true" pageId="1" pageNumber="197">Cabamofa alexsmithi</emphasis>
</taxonomicName>
<emphasis id="B977EAB3275F6C26B84DC954140FF8A2" bold="true" box="[550,631,1862,1886]" pageId="1" pageNumber="197">
<taxonomicNameLabel id="A24457C8275F6C26B84DC954140FF8A2" box="[550,631,1862,1886]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype.
<emphasis id="B977EAB3275F6C26B94BC954154EF8A2" bold="true" box="[800,822,1862,1886]" pageId="1" pageNumber="197">1:</emphasis>
Habitus, lateral (specimen in ethanol after DNA extraction).
<emphasis id="B977EAB3275F6C26BAFCC97816C6F87E" bold="true" box="[151,190,1898,1922]" pageId="1" pageNumber="197">1A:</emphasis>
Posterior abdomen including genitalia, enlargement from 1.
<emphasis id="B977EAB3275F6C26B92BC978152EF87E" bold="true" box="[832,854,1898,1922]" pageId="1" pageNumber="197">2:</emphasis>
Ditto, dorsolateral. Scale for 1, 1.0 mm, for 1A and 2, 0.2 mm. Abbreviations: gen = genitalia, sg6 (‒sg8) = sixth segment (‒eighth segment), st7 (‒st8) = seventh sternite (‒eighth sternite), tg7 (‒tg8) = seventh tergite (‒eighth tergite).
</paragraph>
</caption>
<caption id="DF7C6629275C6C25BAF3CBC51567F9A7" ID-DOI="http://doi.org/10.5281/zenodo.7184817" ID-Zenodo-Dep="7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="2" pageNumber="198" startId="2.[152,270,1495,1519]" targetBox="[162,1425,190,1469]" targetPageId="2">
<paragraph id="8BBC36A1275C6C25BAF3CBC51567F9A7" blockId="2.[151,1437,1495,1628]" pageId="2" pageNumber="198">
<emphasis id="B977EAB3275C6C25BAF3CBC5173DFA13" bold="true" box="[152,325,1495,1519]" pageId="2" pageNumber="198">FIGURES 3–8.</emphasis>
<taxonomicName id="4C034D22275C6C25BB24CBC51453FA13" authority="Jaschhof &amp; Levesque-Beaudin, 2022" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[335,555,1495,1519]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="alexsmithi" status="sp. nov.">
<emphasis id="B977EAB3275C6C25BB24CBC51453FA13" box="[335,555,1495,1519]" italics="true" pageId="2" pageNumber="198">Cabamofa alexsmithi</emphasis>
</taxonomicName>
<emphasis id="B977EAB3275C6C25B85ECBCA14F1FA0C" bold="true" box="[565,649,1496,1520]" pageId="2" pageNumber="198">
<taxonomicNameLabel id="A24457C8275C6C25B85ECBCA14F1FA0C" box="[565,649,1496,1520]" pageId="2" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, males.
<emphasis id="B977EAB3275C6C25B88DCBC51483FA13" bold="true" box="[742,763,1495,1519]" pageId="2" pageNumber="198">3:</emphasis>
Third to fifth flagellomeres, lateral (paratype).
<emphasis id="B977EAB3275C6C25BE93CBC51375FA13" bold="true" box="[1272,1293,1495,1519]" pageId="2" pageNumber="198">4:</emphasis>
Wing, dorsal (holotype).
<emphasis id="B977EAB3275C6C25BB66CBEE175AF9E8" bold="true" box="[269,290,1532,1556]" pageId="2" pageNumber="198">5:</emphasis>
Genitalia, ventral (holotype).
<emphasis id="B977EAB3275C6C25B838CBE91410F9EF" bold="true" box="[595,616,1531,1555]" pageId="2" pageNumber="198">6:</emphasis>
Ninth tergite and adjacent structures, ventral (paratype).
<emphasis id="B977EAB3275C6C25BEC3CBEE12C5F9E8" bold="true" box="[1192,1213,1532,1556]" pageId="2" pageNumber="198">7:</emphasis>
Tegmen and aedeagal apodeme, ventral (holotype).
<emphasis id="B977EAB3275C6C25BBABC80D17ADF9CB" bold="true" box="[448,469,1567,1591]" pageId="2" pageNumber="198">8:</emphasis>
Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm. Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite.
</paragraph>
</caption>
<paragraph id="8BBC36A1275C6C24BAACC8991471FDA3" blockId="2.[151,1437,1675,2024]" lastBlockId="3.[151,1437,151,1400]" lastPageId="3" lastPageNumber="199" pageId="2" pageNumber="198">
<emphasis id="B977EAB3275C6C25BAACC89917EAF958" bold="true" box="[199,402,1675,1700]" inLineHeading="true" pageId="2" pageNumber="198" reason="1">Other characters.</emphasis>
Male. Body length
<quantity id="4CFB9B44275C6C25B800C89E1488F95F" box="[619,752,1676,1700]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="2" pageNumber="198" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7‒2.8 mm</quantity>
.
<emphasis id="B977EAB3275C6C25B890C89E1547F958" bold="true" box="[763,831,1676,1700]" pageId="2" pageNumber="198">Head.</emphasis>
Clypeus setose. Scape slightly larger than pedicel, both setose, pedicel somewhat lighter in color than scape and flagellum. Fourth flagellomere short-cylindrical, with short neck; node 1.1 times as long as wide, with dense, irregular cover of fine seta-like sensilla, interspersed with very few larger setae arising from sockets, surface with irregular network of tenuous ridges (
<figureCitation id="13382A24275C6C25BE47C8E51217F8EC" box="[1068,1135,1783,1808]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="2" pageNumber="198">Fig. 3</figureCitation>
). Compound eyes touching at vertex, eye bridge 5‒6 ommatidia long. Palpus short, with 5 setae-bearing segments, third segment conspicuously swollen, with sensory pit, fifth segment short, only slightly longer than fourth (similar to
<taxonomicName id="4C034D22275C6C25BE55C95213E0F8A4" authority=", Amorim &amp; Rindal 2007" authorityName=", Amorim &amp; Rindal" authorityYear="2007" box="[1086,1432,1855,1880]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275C6C25BE55C95212F6F8AB" box="[1086,1166,1856,1879]" italics="true" pageId="2" pageNumber="198">C. mira</emphasis>
,
<bibRefCitation id="EF924B50275C6C25BEF2C92D13E0F8A4" author="Amorim, D. S. &amp; Rindal, E." box="[1177,1432,1855,1880]" pageId="2" pageNumber="198" pagination="1 - 92" refId="ref2513" refString="Amorim, D. S. &amp; Rindal, E. (2007) Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535 (1), 1 - 92." type="journal article" year="2007">Amorim &amp; Rindal 2007</bibRefCitation>
</taxonomicName>
: fig. 39).
<emphasis id="B977EAB3275C6C25BA9CC976174BF880" bold="true" box="[247,307,1892,1916]" pageId="2" pageNumber="198">Legs.</emphasis>
Coxal lengths relative to thoracal height: forecoxa, 0.7, midcoxa, 0.6, hindcoxa, 0.5. Edge of foretibial anteroapical depression with comb of 12 straight, stiff setae; apices of mid- and hindtibia with combs of 4 and 15 setae respectively similar to those on foretibia but those on midtibia more widely spaced. Claws small, strong, crescent-shaped, toothless. Empodia barely claw-long.
<emphasis id="B977EAB3275C6C25B84DC9DD141CF81B" bold="true" box="[550,612,1999,2023]" pageId="2" pageNumber="198">Wing</emphasis>
(
<figureCitation id="13382A24275C6C25B81AC9DD14C9F814" box="[625,689,1999,2024]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="2" pageNumber="198">Fig. 4</figureCitation>
). Slightly shorter than body, 2.7 times as long as wide. All veins clearly contoured, although some (M
<subScript id="178734E4275D6C24BBB2CEBA1789FF4A" attach="right" box="[473,497,168,182]" fontSize="6" pageId="3" pageNumber="199">1+2</subScript>
, Rs) are weaker than others.
<emphasis id="B977EAB3275D6C24B958CE8515DAFF53" bold="true" box="[819,930,151,175]" pageId="3" pageNumber="199">Abdomen</emphasis>
(
<figureCitation id="13382A24275D6C24B9D8CE85126EFF4C" box="[947,1046,151,176]" captionStart="FIGURES 1–2" captionStartId="1.[152,270,1862,1886]" captionTargetBox="[164,1436,596,1826]" captionTargetId="figure-249@1.[151,1436,596,1839]" captionTargetPageId="1" captionText="FIGURES 1–2. Cabamofa alexsmithi sp. nov., male paratype. 1: Habitus, lateral (specimen in ethanol after DNA extraction). 1A: Posterior abdomen including genitalia, enlargement from 1. 2: Ditto, dorsolateral. Scale for 1, 1.0 mm, for 1A and 2, 0.2 mm. Abbreviations: gen = genitalia, sg6 (‒sg8) = sixth segment (‒eighth segment), st7 (‒st8) = seventh sternite (‒eighth sternite), tg7 (‒tg8) = seventh tergite (‒eighth tergite)." figureDoi="http://doi.org/10.5281/zenodo.7184815" httpUri="https://zenodo.org/record/7184815/files/figure.png" pageId="3" pageNumber="199">Figs 1‒2</figureCitation>
). Segments 1‒6 normal size, tergite and sternite of a particular segment ending on same level; segment 7 considerably shorter than anterior segments, sternite twice as long as tergite; segment 8 similar to 7 but tergite still shorter and almost non-setose.
<emphasis id="B977EAB3275D6C24BEF0CECD1373FF0B" bold="true" box="[1179,1291,223,247]" pageId="3" pageNumber="199">Genitalia.</emphasis>
Ninth tergite (
<figureCitation id="13382A24275D6C24BAF4CF1116A7FEE7" box="[159,223,259,283]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Fig. 6</figureCitation>
) large, subquadratic, outside with large setae, posterior edge with lateral tufts of short megasetae pointing inwards (ventrad). Gonocoxal synsclerite (
<figureCitation id="13382A24275D6C24B86DCF35143FFEC3" box="[518,583,295,319]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Fig. 5</figureCitation>
) with deep, U-shaped emargination ventroposteriorly, around the emargination setae of various sizes, a large non-setose portion ventroanteriorly; dorsal apodemes small, subtriangular; medial bridges densely setose (not illustrated). Gonostylus (
<figureCitation id="13382A24275D6C24B815CF7D1498FE74" box="[638,736,367,392]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Figs 5, 8</figureCitation>
) with 3 lobes of different size, outline and setosity; apical lobe finger-shaped, with long setae; middle lobe roundish, its outside almost non-setose, inside densely setose, setae mostly strongly bent and relatively short; mediobasal lobe elongate, very densely covered in setae of various sizes, setae either bent or straight, all directed inwards (mediad). Tegmen (
<figureCitation id="13382A24275D6C24B96BCFC9153AFE08" box="[768,834,475,500]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Fig. 7</figureCitation>
) 1.8 times as long as broad, broadest at the midlength; parameral apodemes long and thin. Ejaculatory apodeme (
<figureCitation id="13382A24275D6C24B94BCFED151CFDE4" box="[800,868,511,536]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Fig. 7</figureCitation>
) nearly as long as tegmen, moderately sclerotized, markedly swollen subbasally. Cerci (
<figureCitation id="13382A24275D6C24B877CC311423FDC7" box="[540,603,547,571]" captionStart="FIGURES 3–8" captionStartId="2.[152,270,1495,1519]" captionTargetBox="[162,1425,190,1469]" captionTargetId="figure-17@2.[151,1436,181,1471]" captionTargetPageId="2" captionText="FIGURES 3–8. Cabamofa alexsmithi sp. nov., males. 3: Third to fifth flagellomeres, lateral (paratype). 4: Wing, dorsal (holotype). 5: Genitalia, ventral (holotype). 6: Ninth tergite and adjacent structures, ventral (paratype). 7: Tegmen and aedeagal apodeme, ventral (holotype). 8: Gonostylus, dorsal (paratype). Scale for 3 and 5‒8, 0.05 mm, for 4, 1.0 mm.Abbreviations: cerc = cercus, hyp = hypoproct, sens = sensillum, tg9 = ninth tergite." figureDoi="http://doi.org/10.5281/zenodo.7184817" httpUri="https://zenodo.org/record/7184817/files/figure.png" pageId="3" pageNumber="199">Fig. 6</figureCitation>
) small, halfmoon-shaped, with large setae. Hypoproct large, elongate-oviform, non-setose, densely microtrichose.
</paragraph>
<paragraph id="8BBC36A1275D6C24BAACCC7916A9FD5B" blockId="3.[151,1437,151,1400]" pageId="3" pageNumber="199">
Female. Body length
<quantity id="4CFB9B44275D6C24BBDACC7E144EFD7F" box="[433,566,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="3" pageNumber="199" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2‒3.8 mm</quantity>
. Other characters are as in the corresponding males or indistinguishable from
<taxonomicName id="4C034D22275D6C24BFEFCC7E16B1FD5B" authorityName=", Amorim &amp; Rindal" authorityYear="2007" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B977EAB3275D6C24BFEFCC7E16B1FD5B" italics="true" pageId="3" pageNumber="199">C. mira</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBC36A1275D6C24BAACCCA614AAFC3B" blockId="3.[151,1437,151,1400]" pageId="3" pageNumber="199">
<emphasis id="B977EAB3275D6C24BAACCCA6179BFD37" bold="true" box="[199,483,691,716]" inLineHeading="true" pageId="3" pageNumber="199" reason="1">Molecular identification.</emphasis>
COI sequences were obtained for all
<specimenCount id="9D05FD28275D6C24B9E0CCA11255FD37" box="[907,1069,691,715]" pageId="3" pageNumber="199" type="generic">five specimens</specimenCount>
through the Canadian Centre for DNA Barcoding (CCDB: https://ccdb.ca/). Sequence lengths varied between 652‒654bp and collectively are publicly available in BIN: BOLD:AEL9580 (
<bibRefCitation id="EF924B50275D6C24B875CCE91519FCE8" author="Ratnasingham, S. &amp; Hebert, P. D. N." box="[542,865,763,788]" pageId="3" pageNumber="199" pagination="16" refId="ref2960" refString="Ratnasingham, S. &amp; Hebert, P. D. N. (2013) A DNA-based registry for all animal species: the Barcode Index Number (BIN) system. PLoS ONE, 8, 16. https: // doi. org / 10.1371 / journal. pone. 0066213" type="journal article" year="2013">Ratnasingham &amp; Hebert 2013</bibRefCitation>
) for the Barcode of Life Data System (BOLD: http:// boldsystems.org/). This BIN and its barcodes are widely divergent from everything else on BOLD with the nearest neighbor being about 14% distant and matching the family
<taxonomicName id="4C034D22275D6C24B975CD5115BDFCA7" box="[798,965,835,859]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
(BIN: BOLD:ADV0335). This emphasizes the uniqueness of
<taxonomicName id="4C034D22275D6C24BB3CCD7517BEFC83" authorityName="Jaschhof" authorityYear="2005" box="[343,454,871,895]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAB3275D6C24BB3CCD7517BEFC83" box="[343,454,871,895]" italics="true" pageId="3" pageNumber="199">Cabamofa</emphasis>
</taxonomicName>
among Sciaroidea. Even within
<taxonomicName id="4C034D22275D6C24B974CD7515F6FC83" authorityName="Jaschhof" authorityYear="2005" box="[799,910,871,895]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAB3275D6C24B974CD7515F6FC83" box="[799,910,871,895]" italics="true" pageId="3" pageNumber="199">Cabamofa</emphasis>
</taxonomicName>
the divergence is wide, with a 19% distance to
<taxonomicName id="4C034D22275D6C24BFEFCD7A1759FC5F" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="B977EAB3275D6C24BFEFCD7A1759FC5F" italics="true" pageId="3" pageNumber="199">C. vietnamensis</emphasis>
</taxonomicName>
, the only other DNA barcode (BOLD) available for that genus. This suggests that the Neotropical species might be a different lineage than the Oriental species.
</paragraph>
<paragraph id="8BBC36A1275D6C24BAACCDC115A1FBF3" blockId="3.[151,1437,151,1400]" pageId="3" pageNumber="199">
All data is publicly available on BOLD, its public dataset (dx.doi.org/10.5883/DS-CABAMOCR), and GenBank (https://www.ncbi.nlm.nih.gov/genbank/; accession:
<accessionNumber id="9450AB42275D6C24B8A1CDE51531FBF3" box="[714,841,1015,1039]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM782492" pageId="3" pageNumber="199">OM782492</accessionNumber>
-
<accessionNumber id="9450AB42275D6C24B924CDE515B6FBF3" box="[847,974,1015,1039]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM782496" pageId="3" pageNumber="199">OM782496</accessionNumber>
).
</paragraph>
</subSubSection>
<subSubSection id="C319652A275D6C24BAACCA0E122EFB63" pageId="3" pageNumber="199" type="etymology">
<paragraph id="8BBC36A1275D6C24BAACCA0E122EFB63" blockId="3.[151,1437,151,1400]" pageId="3" pageNumber="199">
<emphasis id="B977EAB3275D6C24BAACCA0E1730FBC8" bold="true" box="[199,328,1052,1076]" inLineHeading="true" pageId="3" pageNumber="199" reason="1">Etymology.</emphasis>
<emphasis id="B977EAB3275D6C24BB38CA091443FBCF" box="[339,571,1051,1075]" italics="true" pageId="3" pageNumber="199">
Cabomofa
<taxonomicName id="4C034D22275D6C24BBA6CA091443FBCF" authorityName="Jaschhof &amp; Levesque-Beaudin" authorityYear="2022" box="[461,571,1051,1075]" class="Insecta" family="Rangomaramidae" genus="Cabamofa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="alexsmithi">alexsmithi</taxonomicName>
</emphasis>
was collected by a Malaise trap at
<quantity id="4CFB9B44275D6C24B9B3CA0E1256FBCF" box="[984,1070,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="199" unit="m" value="1200.0">1200 m</quantity>
elevation in cloud forest on the Pacific slope of Volcan Cacao of Área de Conservación Guanacaste. It is named in honor of Dr. M. Alex Smith of the Department of Integrative Biology of the University of Guelph, Guelph,
<collectingCountry id="F3147631275D6C24B9D3CA711273FB87" box="[952,1035,1123,1147]" name="Canada" pageId="3" pageNumber="199">Canada</collectingCountry>
, in recognition of his many years of intense documentation of the upper elevation ecology of the entomofauna of this volcano.
</paragraph>
</subSubSection>
<subSubSection id="C319652A275D6C24BAACCABE15E6FA8B" pageId="3" pageNumber="199" type="materials_examined">
<paragraph id="8BBC36A1275D6C24BAACCABE15E6FA8B" blockId="3.[151,1437,151,1400]" pageId="3" pageNumber="199">
<emphasis id="B977EAB3275D6C24BAACCABE1715FB3F" bold="true" box="[199,365,1195,1220]" inLineHeading="true" pageId="3" pageNumber="199" reason="1">Type material.</emphasis>
<materialsCitation id="3B6B3CFC275D6C24BB19CAB9151CFAF7" ID-GBIF-Occurrence="3923192302" collectingDate="2015-12-03" collectionCode="BIOUG, CNC" collectorName="D. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="1220" latitude="10.929" location="Derrumbe" longLatPrecision="77" longitude="-85.464" municipality="Sector Cacao" pageId="3" pageNumber="199" specimenCode="BIOUG71158, CNC1156197" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="B977EAB3275D6C24BB19CAB917ACFB3F" box="[370,468,1195,1219]" italics="true" pageId="3" pageNumber="199">
<typeStatus id="54B88803275D6C24BB19CAB917ACFB3F" box="[370,468,1195,1219]" pageId="3" pageNumber="199" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<collectingCountry id="F3147631275D6C24B849CABE14E1FB3F" box="[546,665,1195,1220]" name="Costa Rica" pageId="3" pageNumber="199">Costa Rica</collectingCountry>
,
<collectingCounty id="62DD4E2D275D6C24B8C8CABE15E8FB3F" box="[675,912,1195,1222]" pageId="3" pageNumber="199">
Área
<collectorName id="26F65377275D6C24B8B6CAB915E8FB3F" box="[733,912,1195,1219]" pageId="3" pageNumber="199">de Conservación</collectorName>
</collectingCounty>
<collectingRegion id="49C7F843275D6C24B9FECABE126BFB38" box="[917,1043,1196,1220]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="199">Guanacaste</collectingRegion>
,
<collectingRegion id="49C7F843275D6C24BE76CABE1286FB3F" box="[1053,1278,1195,1220]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="199">Guanacaste province</collectingRegion>
,
<collectingMunicipality id="6BD8ACDB275D6C24BF62CABE13E0FB38" box="[1289,1432,1196,1220]" pageId="3" pageNumber="199">Sector Cacao</collectingMunicipality>
,
<location id="8EDC607A275D6C24BAFCCADD177EFB1B" LSID="urn:lsid:plazi:treatment:03AA87B7275E6C24BAFCCBE615E6FA8B:8EDC607A275D6C24BAFCCADD177EFB1B" box="[151,262,1231,1255]" country="Costa Rica" county="Area de Conservacion" latitude="10.929" longLatPrecision="77" longitude="-85.464" municipality="Sector Cacao" name="Derrumbe" pageId="3" pageNumber="199" stateProvince="Guanacaste">Derrumbe</location>
,
<quantity id="4CFB9B44275D6C24BB79CAC2171DFB1B" box="[274,357,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="199" unit="m" value="1220.0">
<elevation id="002ED192275D6C24BB79CAC2171DFB1B" box="[274,357,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="199" unit="m" value="1220.0">1220 m</elevation>
</quantity>
elevation,
<geoCoordinate id="EE375066275D6C24BBB4CADD143FFB1B" box="[479,583,1231,1255]" degrees="10.929" direction="north" orientation="latitude" pageId="3" pageNumber="199" precision="55" value="10.929">10.929°N</geoCoordinate>
:
<geoCoordinate id="EE375066275D6C24B821CADD14C3FB1B" box="[586,699,1231,1255]" degrees="85.464" direction="west" orientation="longitude" pageId="3" pageNumber="199" precision="55" value="-85.464">85.464°W</geoCoordinate>
, cloud forest,
<date id="FFBD1061275D6C24B93CCADD15AAFB1B" box="[855,978,1231,1255]" pageId="3" pageNumber="199" value="2015-12-03">
<collectingDate id="EFF9E989275D6C24B93CCADD15AAFB1B" box="[855,978,1231,1255]" pageId="3" pageNumber="199" value="2015-12-03">03.xii.2015</collectingDate>
</date>
,
<collectingMethod id="52424EB6275D6C24B98BCADD1211FB1B" box="[992,1129,1231,1255]" pageId="3" pageNumber="199">Malaise trap</collectingMethod>
,
<collectorName id="26F65377275D6C24BE1ECAC21299FB1B" box="[1141,1249,1232,1255]" pageId="3" pageNumber="199">D. Janzen</collectorName>
&amp;
<collectorName id="26F65377275D6C24BF6DCAC213E5FB1B" box="[1286,1437,1231,1255]" pageId="3" pageNumber="199">W. Hallwachs</collectorName>
leg. (sample ID:
<specimenCode id="DBA59EDA275D6C24BB3CCAE1178CFAF7" box="[343,500,1267,1291]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="3" pageNumber="199" type="University or college">BIOUG71158</specimenCode>
-F10, museum ID:
<specimenCode id="DBA59EDA275D6C24B8AECAE11525FAF7" box="[709,861,1267,1291]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="199">CNC1156197</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B6B3CFC275D6C24B918CAE615ABFAD3" ID-GBIF-Occurrence="3923192301" collectingDate="2016-06-02" collectionCode="BIOUG, CNC" collectorName="D. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="1220" latitude="10.929" location="Derrumbe" longLatPrecision="77" longitude="-85.464" municipality="Sector Cacao" pageId="3" pageNumber="199" specimenCode="BIOUG70056, CNC1156198" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B977EAB3275D6C24B918CAE615A7FAF7" box="[883,991,1268,1291]" italics="true" pageId="3" pageNumber="199">
<typeStatus id="54B88803275D6C24B918CAE615A7FAF7" box="[883,991,1268,1291]" pageId="3" pageNumber="199" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D05FD28275D6C24B99ACAE61238FAF7" box="[1009,1088,1267,1292]" pageId="3" pageNumber="199" type="male">1 male</specimenCount>
, same data as for the holotype but
<date id="FFBD1061275D6C24BAA9CB05174DFAD3" box="[194,309,1303,1327]" pageId="3" pageNumber="199" value="2016-06-02">
<collectingDate id="EFF9E989275D6C24BAA9CB05174DFAD3" box="[194,309,1303,1327]" pageId="3" pageNumber="199" value="2016-06-02">02.vi.2016</collectingDate>
</date>
(sample ID:
<specimenCode id="DBA59EDA275D6C24BBACCB05141CFAD3" box="[455,612,1303,1327]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="3" pageNumber="199" type="University or college">BIOUG70056</specimenCode>
-B09, museum ID:
<specimenCode id="DBA59EDA275D6C24B958CB0515B3FAD3" box="[819,971,1303,1327]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="199">CNC1156198</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B6B3CFC275D6C24B98BCB0A15E2FA8B" ID-GBIF-Occurrence="3923192303" collectingDate="2015-12-17" collectionCode="BIOUG, CNC" collectorName="D. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="1220" latitude="10.929" location="Derrumbe" longLatPrecision="77" longitude="-85.464" municipality="Sector Cacao" pageId="3" pageNumber="199" specimenCode="BIOUG68329, CNC1156199, BIOUG71132, CNC1156200, BIOUG71841, CNC1156201" specimenCount="3" specimenCount-female="3" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D05FD28275D6C24B98BCB0A1234FAD3" box="[992,1100,1303,1328]" pageId="3" pageNumber="199" type="female">3 females</specimenCount>
, same data as for the holotype but
<collectingDate id="EFF9E989275D6C24BAA8CB291423FAAF" box="[195,603,1339,1363]" isEnumeration="true" pageId="3" pageNumber="199">
<date id="FFBD1061275D6C24BAA8CB291746FAAF" box="[195,318,1339,1363]" pageId="3" pageNumber="199" value="2015-12-17">17.xii.2015</date>
,
<date id="FFBD1061275D6C24BB26CB2917CEFAAF" box="[333,438,1339,1363]" pageId="3" pageNumber="199" value="2016-05-05">05.v.2016</date>
and
<date id="FFBD1061275D6C24BB9BCB291423FAAF" box="[496,603,1339,1363]" pageId="3" pageNumber="199" value="2016-05-19">19.v.2016</date>
</collectingDate>
, respectively (sample ID / museum ID:
<specimenCode id="DBA59EDA275D6C24BE70CB2912CFFAAF" box="[1051,1207,1339,1363]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="3" pageNumber="199" type="University or college">BIOUG68329</specimenCode>
-G11 /
<specimenCode id="DBA59EDA275D6C24BF69CB2913E0FAAF" box="[1282,1432,1339,1363]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="199">CNC1156199</specimenCode>
,
<specimenCode id="DBA59EDA275D6C24BAFCCB721749FA84" box="[151,305,1376,1400]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="3" pageNumber="199" type="University or college">BIOUG71132</specimenCode>
-G03 /
<specimenCode id="DBA59EDA275D6C24BB12CB4D1477FA8B" box="[377,527,1375,1399]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="199">CNC1156200</specimenCode>
,
<specimenCode id="DBA59EDA275D6C24B872CB7214CCFA84" box="[537,692,1376,1400]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="3" pageNumber="199" type="University or college">BIOUG71841</specimenCode>
-C09 /
<specimenCode id="DBA59EDA275D6C24B890CB4D15EBFA8B" box="[763,915,1375,1399]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="199">CNC1156201</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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