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<document id="612112A5FCDC2290F1487F2B0D16F0AA" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFB5076D5494FA1E7ABAFBF6" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Holthuija pauli Ng &amp; Forges 2015, n. sp." docType="treatment" docUuid="99F27323-4605-403A-9EBB-B6CCEB5C3011" docUuidSource="ZooBank" docVersion="4" lastPageNumber="163" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="162" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3069AC2E7D8355B1B69F4BEC90420855">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="249BBD9F88F52C3D336BB395E261BCBD">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="36A2B18E8161E93C6A91823E6B68EFF7">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="EC33487D5623891F01F3A2FA6CE269E0">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="E951016C87BDAD0E92D5D2CC0D3CB211">2015</mods:date>
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<mods:number id="DA10151128562914017F4A07721C37F7">2015-05-29</mods:number>
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<treatment id="03A9654BFFB5076D5494FA1E7ABAFBF6" LSID="urn:lsid:zoobank.org:act:99F27323-4605-403A-9EBB-B6CCEB5C3011" httpUri="http://treatment.plazi.org/id/03A9654BFFB5076D5494FA1E7ABAFBF6" lastPageId="53" lastPageNumber="163" pageId="52" pageNumber="162">
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<paragraph id="8BBFD45DFFB5076C5494FA1E7D87FA16" blockId="52.[831,1432,1437,1525]" box="[1011,1253,1437,1461]" pageId="52" pageNumber="162">
<heading id="D0F76331FFB5076C5494FA1E7D87FA16" bold="true" box="[1011,1253,1437,1461]" centered="true" fontSize="10" level="2" pageId="52" pageNumber="162" reason="2">
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<taxonomicName id="4C00AFDEFFB5076C5494FA1E7DFCFA16" ID-CoL="3MBWS" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1011,1182,1437,1461]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="162" phylum="Arthropoda" rank="species" species="pauli" status="sp. nov.">
<emphasis id="B974084FFFB5076C5494FA1E7DFCFA16" bold="true" box="[1011,1182,1437,1461]" italics="true" pageId="52" pageNumber="162">Holthuija pauli</emphasis>
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<taxonomicNameLabel id="A247B534FFB5076C53C0FA1E7D87FA16" box="[1191,1253,1437,1461]" pageId="52" pageNumber="162" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C31A87D6FFB5076C5458FA3E7DF5FA56" pageId="52" pageNumber="162" type="description">
<paragraph id="8BBFD45DFFB5076C5458FA3E7DF5FA56" blockId="52.[831,1432,1437,1525]" pageId="52" pageNumber="162">
(
<figureCitation id="133BC8D8FFB5076C5420FA3E7AD0FA76" box="[839,946,1469,1493]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="52" pageNumber="162">Figs. 24F</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C54D9FA3E7D6CFA76" box="[958,1038,1469,1493]" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="52" pageNumber="162">26B, C</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C537CFA3E7D0AFA76" box="[1051,1128,1469,1493]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. AC, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; FH, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; IK, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; LO, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [AF, I, L]; 0.5 mm [D, G, H, J, K, MO]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="52" pageNumber="162">27FH</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C5312FA3E7DC7FA76" box="[1141,1189,1469,1493]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="52" pageNumber="162">37G</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C53D5FA3E7D81FA76" box="[1202,1251,1469,1493]" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="52" pageNumber="162">40G</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C5388FA3E7C7CFA76" box="[1263,1310,1469,1493]" captionStart="Fig" captionStartId="82.[113,148,1833,1855]" captionTargetBox="[170,1394,189,1818]" captionTargetId="figure-234@82.[170,1394,189,1819]" captionTargetPageId="82" captionText="Fig. 46. Third maxillipeds. A, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; B, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; C, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; D, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; E, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; F, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; G, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; H, Sakaija sakaii, male (9.0 × 6.6 mm) (NSMT-Cr 8094), Japan; I, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; J, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; K, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; L, Sakaija longispinosa n. sp., paratype ovigerous female (14.2 × 10.7 mm) (NMV J63197), Australia; M, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; N, Planaja plana n. gen. n. sp., paratype male (38.3 × 30.7 mm) (ZRC 2013.1370), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384690" httpUri="https://zenodo.org/record/5384690/files/figure.png" pageId="52" pageNumber="162">46C</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C524CFA3E7C39FA76" box="[1323,1371,1469,1493]" captionStart="Fig" captionStartId="86.[113,148,1750,1772]" captionTargetBox="[113,1452,388,1735]" captionTargetId="figure-138@86.[113,1452,388,1735]" captionTargetPageId="86" captionText="Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea." figureDoi="http://doi.org/10.5281/zenodo.5384702" httpUri="https://zenodo.org/record/5384702/files/figure.png" pageId="52" pageNumber="162">50G</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C520FFA3E7CF6FA76" box="[1384,1428,1469,1493]" captionStart="Fig" captionStartId="90.[113,148,1833,1855]" captionTargetBox="[151,1414,189,1818]" captionTargetId="figure-229@90.[151,1414,189,1821]" captionTargetPageId="90" captionText="Fig. 54. Chelae. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., paratype male (76.5 × 61.3 mm) (ZRC 2001.0577), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; G, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; L, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; M, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, holotype female (51.2 × 40.5 mm) (NHM 1860.15), China; P, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384710" httpUri="https://zenodo.org/record/5384710/files/figure.png" pageId="52" pageNumber="162">54F</figureCitation>
,
<figureCitation id="133BC8D8FFB5076C5322FA5E7DEDFA56" box="[1093,1167,1501,1525]" captionStart="Fig" captionStartId="108.[113,148,1935,1957]" captionTargetBox="[113,1451,260,1919]" captionTargetId="figure-15@108.[113,1451,260,1920]" captionTargetPageId="108" captionText="Fig. 69. Colours in life. A, Alcomaja nagashimaensis, male (specimen not examined), Philippines; B, Alcomaja miriky n. sp., paratype male (17.3 × 12.7 mm) (MNHN-IU-2010-60), Madagascar; C, D, Holthuija miersii, ovigerous female (50.2 × 40.4 mm) (ZRC 2013.1272), Singapore; E, Holthuija pauli n. sp., paratype male (31.5 × 23.4 mm) (ZRC 2001.0590), Philippines; F, Holthuija pauli n. sp., female (38.8 × 28.9 mm) (ZRC 2013.1379), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384744" httpUri="https://zenodo.org/record/5384744/files/figure.png" pageId="52" pageNumber="162">69E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFB5076D544AF99E78F9FA76" lastPageId="53" lastPageNumber="163" pageId="52" pageNumber="162" type="materials_examined">
<paragraph id="8BBFD45DFFB5076D544AF99E78F9FA76" blockId="52.[811,1453,1565,2101]" lastBlockId="53.[135,778,189,1493]" lastPageId="53" lastPageNumber="163" pageId="52" pageNumber="162">
<emphasis id="B974084FFFB5076C544AF99E7D7AF996" bold="true" box="[813,1048,1565,1589]" pageId="52" pageNumber="162">Material examined.</emphasis>
<materialsCitation id="3B68DE00FFB5076C5344F99E7C50F9D6" collectingDate="2003-07" collectionCode="NMCR" county="Philippines" elevation="250" location="Balicasag Island" municipality="Bohol" pageId="52" pageNumber="162" specimenCount="1" typeStatus="holotype">
<typeStatus id="54BB6AFFFFB5076C5344F99E7DEDF996" box="[1059,1167,1565,1589]" pageId="52" pageNumber="162" type="holotype">Holotype</typeStatus>
: male (37.3 × 28.0 mm) (
<collectionCode id="ED114C98FFB5076C5453F9BE7AE5F9F6" box="[820,903,1597,1621]" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="52" pageNumber="162" type="Herbarium">NMCR</collectionCode>
),
<location id="8EDF8286FFB5076C54F0F9BE7D21F9F6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB5076C54F0F9BE7D21F9F6" box="[919,1091,1597,1621]" county="Philippines" municipality="Bohol" name="Balicasag Island" pageId="52" pageNumber="162">Balicasag Island</location>
,
<location id="8EDF8286FFB5076C532BF9BE7DC1F9F6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB5076C532BF9BE7DC1F9F6" box="[1100,1187,1597,1621]" county="Philippines" municipality="Bohol" name="Panglao" pageId="52" pageNumber="162">Panglao</location>
,
<collectingMunicipality id="6BDB4E27FFB5076C53CCF9BE7D8CF9F6" box="[1195,1262,1597,1621]" pageId="52" pageNumber="162">Bohol</collectingMunicipality>
,
<collectingCounty id="62DEACD1FFB5076C5390F9BE7C0DF9F6" box="[1271,1391,1597,1621]" pageId="52" pageNumber="162">Philippines</collectingCounty>
,
<quantity id="4CF879B8FFB5076C5210F9BE7A11F9D6" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="52" pageNumber="162" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB5076C5210F9BE7A11F9D6" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="52" pageNumber="162" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200 300 m</elevation>
</quantity>
, coll. fishermen in tangle nets,
<date id="FFBEF29DFFB5076C53A2F9DE7C50F9D6" box="[1221,1330,1629,1653]" pageId="52" pageNumber="162" value="2003-07">
<collectingDate id="EFFA0B75FFB5076C53A2F9DE7C50F9D6" box="[1221,1330,1629,1653]" pageId="52" pageNumber="162" value="2003-07">July 2003</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B68DE00FFB5076C525BF9DD7D46F956" country="Philippines" location="Philippines" pageId="52" pageNumber="162" specimenCount="10" specimenCount-male="10" typeStatus="paratype">
<typeStatus id="54BB6AFFFFB5076C525BF9DD7CCAF9D5" box="[1340,1448,1630,1654]" pageId="52" pageNumber="162" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B974084FFFB5076C544AF9FE7ACEF936" bold="true" box="[813,940,1661,1685]" pageId="52" pageNumber="162">
<collectingCountry id="F31794CDFFB5076C544AF9FE7ACEF936" box="[813,940,1661,1685]" name="Philippines" pageId="52" pageNumber="162">Philippines</collectingCountry>
</emphasis>
<specimenCount id="9D061FD4FFB5076C54AEF9FE7D48F936" box="[969,1066,1661,1685]" pageId="52" pageNumber="162" type="male">10 males</specimenCount>
(27.0 ×
<quantity id="4CF879B8FFB5076C53EFF9FE7D89F936" box="[1160,1259,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="52" pageNumber="162" unit="mm" value="19.7">19.7 mm</quantity>
, 27.6 ×
<quantity id="4CF879B8FFB5076C5222F9FE7CC5F936" box="[1349,1447,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="52" pageNumber="162" unit="mm" value="20.2">20.2 mm</quantity>
, 27.9 ×
<quantity id="4CF879B8FFB5076C541BF91E7ABDF916" box="[892,991,1693,1717]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="52" pageNumber="162" unit="mm" value="20.3">20.3 mm</quantity>
, 30.6 ×
<quantity id="4CF879B8FFB5076C535BF91E7DFEF916" box="[1084,1180,1693,1717]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="52" pageNumber="162" unit="mm" value="22.8">22.8 mm</quantity>
32.4 × 24.0 mm, 33.2 ×
<quantity id="4CF879B8FFB5076C544AF93E7AF0F976" box="[813,914,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" pageId="52" pageNumber="162" unit="mm" value="24.1">24.1 mm</quantity>
, 34.2 ×
<quantity id="4CF879B8FFB5076C5495F93E7D35F976" box="[1010,1111,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="52" pageNumber="162" unit="mm" value="26.1">26.1 mm</quantity>
, 34.4 ×
<quantity id="4CF879B8FFB5076C53D0F93E7C7EF976" box="[1207,1308,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="52" pageNumber="162" unit="mm" value="25.3">25.3 mm</quantity>
, 34.5 ×
<quantity id="4CF879B8FFB5076C521AF93E7A3BF956" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="52" pageNumber="162" unit="mm" value="25.9">25.9 mm</quantity>
, 41.0 ×
<quantity id="4CF879B8FFB5076C54D5F95E7D7AF956" box="[946,1048,1757,1781]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" pageId="52" pageNumber="162" unit="mm" value="30.8">30.8 mm</quantity>
,)
</materialsCitation>
,
<materialsCitation id="3B68DE00FFB5076C5356F95E7D78F8F6" collectingDate="2003-07" collectionCode="ZRC" country="Philippines" county="Philippines" elevation="250" location="Balicasag Island" municipality="Bohol" pageId="52" pageNumber="162" specimenCode="ZRC 2013.1371" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="9D061FD4FFB5076C5356F95E7DFBF956" box="[1073,1177,1757,1781]" pageId="52" pageNumber="162" type="female">3 females</specimenCount>
(41.9 ×
<quantity id="4CF879B8FFB5076C5390F95E7C38F956" box="[1271,1370,1757,1781]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="52" pageNumber="162" unit="mm" value="31.4">31.4 mm</quantity>
, 41.9 ×
<quantity id="4CF879B8FFB5076C544AF97E7AECF8B6" box="[813,910,1789,1813]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="52" pageNumber="162" unit="mm" value="32.4">32.4 mm</quantity>
, 34.8 ×
<quantity id="4CF879B8FFB5076C5484F97E7D27F8B6" box="[995,1093,1789,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="52" pageNumber="162" unit="mm" value="26.2">26.2 mm</quantity>
), 1 ovigerous female (34.7 ×
<quantity id="4CF879B8FFB5076C521AF97E7A3BF896" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="52" pageNumber="162" unit="mm" value="26.4">26.4 mm</quantity>
), 1 damaged female (28.0 ×
<quantity id="4CF879B8FFB5076C5318F89E7DBCF896" box="[1151,1246,1821,1845]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="52" pageNumber="162" unit="mm" value="20.8">20.8 mm</quantity>
) (
<specimenCode id="DBA67C26FFB5076C5396F89E7CC3F896" box="[1265,1441,1821,1845]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="52" pageNumber="162">ZRC 2013.1371</specimenCode>
), same data as holotype
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB5076C5321F8BE7DE1F836" collectingDate="2000-12" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="52" pageNumber="162" specimenCode="ZRC 2001.411" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB5076C5321F8BE7DC1F8F6" box="[1094,1187,1853,1877]" pageId="52" pageNumber="162" type="female">1 female</specimenCount>
(27.8 ×
<quantity id="4CF879B8FFB5076C5267F8BE7C02F8F6" box="[1280,1376,1853,1877]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="52" pageNumber="162" unit="mm" value="21.5">21.5 mm</quantity>
) (
<specimenCode id="DBA67C26FFB5076C5211F8BE7AF7F8D6" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="52" pageNumber="162">ZRC 2001.411</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB5076C54C1F8DE7D35F8D6" box="[934,1111,1885,1909]" pageId="52" pageNumber="162">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB5076C5307F8DE7DDBF8D6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB5076C5307F8DE7DDBF8D6" box="[1120,1209,1885,1909]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="52" pageNumber="162" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB5076C53A5F8DE7C64F8D6" box="[1218,1286,1885,1909]" country="Philippines" name="Bohol" pageId="52" pageNumber="162">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB5076C54BCF8FE7DE1F836" box="[987,1155,1917,1941]" pageId="52" pageNumber="162" value="2000-12">
<collectingDate id="EFFA0B75FFB5076C54BCF8FE7DE1F836" box="[987,1155,1917,1941]" pageId="52" pageNumber="162" value="2000-12">December 2000</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB5076C53CCF8FE7C0CF876" collectingDate="2001-11-28" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="52" pageNumber="162" specimenCode="ZRC 2013.1266, ZRC 2001.0577" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB5076C53CCF8FE7D92F836" box="[1195,1264,1917,1941]" pageId="52" pageNumber="162" type="male">1 male</specimenCount>
(
<specimenCode id="DBA67C26FFB5076C539BF8FE7CCAF836" box="[1276,1448,1917,1941]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="52" pageNumber="162">ZRC 2013.1266</specimenCode>
, ex part
<specimenCode id="DBA67C26FFB5076C541BF81E7D4CF816" box="[892,1070,1949,1973]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="52" pageNumber="162">ZRC 2001.0577</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB5076C5327F81E7D91F816" box="[1088,1267,1949,1973]" pageId="52" pageNumber="162">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB5076C5399F81E7C35F816" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB5076C5399F81E7C35F816" box="[1278,1367,1949,1973]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="52" pageNumber="162" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB5076C5205F81E7CC4F816" box="[1378,1446,1949,1973]" country="Philippines" name="Bohol" pageId="52" pageNumber="162">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB5076C53F0F83E7C0CF876" box="[1175,1390,1981,2005]" pageId="52" pageNumber="162" value="2001-11-28">
<collectingDate id="EFFA0B75FFB5076C53F0F83E7C0CF876" box="[1175,1390,1981,2005]" pageId="52" pageNumber="162" value="2001-11-28">28 November 2001</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB5076D52F8F83E7BABFF76" collectingDate="2001-11-28" collectionCode="ZRC" collectorName="Panglao" country="Philippines" lastPageId="53" lastPageNumber="163" location="Balicasag Island" pageId="52" pageNumber="162" specimenCode="ZRC 2001.0590" specimenCount="4" specimenCount-male="4" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB5076C52F8F83E7A0CF856" pageId="52" pageNumber="162" type="male">4 males</specimenCount>
(27.5 ×
<quantity id="4CF879B8FFB5076C54BDF85E7D26F856" box="[986,1092,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="52" pageNumber="162" unit="mm" value="19.9">19.9 mm</quantity>
, 28.2 ×
<quantity id="4CF879B8FFB5076C53CDF85E7C77F856" box="[1194,1301,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="52" pageNumber="162" unit="mm" value="20.7">20.7 mm</quantity>
, 31.5 ×
<quantity id="4CF879B8FFB5076C521CF85E7A38F7B6" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="52" pageNumber="162" unit="mm" value="23.4">23.4 mm</quantity>
, 31.9 ×
<quantity id="4CF879B8FFB5076C54DBF87E7D46F7B6" box="[956,1060,2045,2069]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="52" pageNumber="162" unit="mm" value="23.8">23.8 mm</quantity>
), 1 ovigerous female (35.0 × 26.0 mm) (
<specimenCode id="DBA67C26FFB5076C5417F79E7D47F796" box="[880,1061,2077,2101]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="52" pageNumber="162">ZRC 2001.0590</specimenCode>
),
<location id="8EDF8286FFB5076C535DF79E7D92F796" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB5076C535DF79E7D92F796" box="[1082,1264,2077,2101]" country="Philippines" name="Balicasag Island" pageId="52" pageNumber="162" stateProvince="Bohol">Balicasag Island</location>
,
<collectorName id="26F5B18BFFB5076C539AF79E7C34F796" box="[1277,1366,2077,2101]" pageId="52" pageNumber="162">Panglao</collectorName>
,
<collectingRegion id="49C41ABFFFB5076C5203F79E7CCAF796" box="[1380,1448,2077,2101]" country="Philippines" name="Bohol" pageId="52" pageNumber="162">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D5695FF3E7BABFF76" box="[498,713,189,213]" pageId="53" pageNumber="163" value="2001-11-28">
<collectingDate id="EFFA0B75FFB4076D5695FF3E7BABFF76" box="[498,713,189,213]" pageId="53" pageNumber="163" value="2001-11-28">28 November 2001</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D559EFF3E79A3FE96" collectingDate="2002-06" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1376" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D559EFF3E79B6FF56" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
(23.8 ×
<quantity id="4CF879B8FFB4076D5627FF5E78CAFF56" box="[320,424,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="53" pageNumber="163" unit="mm" value="17.6">17.6 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D56A4FF5E7BE0FF56" box="[451,642,221,245]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1376</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D55FEFF5E79AEFEB6" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D57BFFF7E7853FEB6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D57BFFF7E7853FEB6" box="[216,305,253,277]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D565AFF7E78E3FEB6" box="[317,385,253,277]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen in tangle nets,
<date id="FFBEF29DFFB4076D55B1FF7D79A3FE96" pageId="53" pageNumber="163" value="2002-06">
<collectingDate id="EFFA0B75FFB4076D55B1FF7D79A3FE96" pageId="53" pageNumber="163" value="2002-06">June 2002</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5789FE9E78D9FED6" collectingDate="2002-06" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1314" specimenCount="1" specimenCount-juvenile="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5789FE9E783BFE96" box="[238,345,285,309]" pageId="53" pageNumber="163" type="juvenile">1 juvenile</specimenCount>
female with bopyrid (
<specimenCode id="DBA67C26FFB4076D552EFE9E7B9EFE96" box="[585,764,285,309]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1314</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D57EEFEBE7855FEF6" box="[137,311,317,341]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5627FEBE78F5FEF6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5627FEBE78F5FEF6" box="[320,407,317,341]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D56C7FEBE7881FEF6" box="[416,483,317,341]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFB4076D568BFEBE7B06FEF6" box="[492,612,317,341]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB4076D568BFEBE7B06FEF6" box="[492,612,317,341]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D562FFEDD78D9FED6" box="[328,443,349,373]" pageId="53" pageNumber="163" value="2002-06">
<collectingDate id="EFFA0B75FFB4076D562FFEDD78D9FED6" box="[328,443,349,373]" pageId="53" pageNumber="163" value="2002-06">June 2002</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D568AFEDE7B70FE16" collectingDate="2002-06" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1378" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D568AFEDE7B5AFED6" box="[493,568,349,373]" pageId="53" pageNumber="163" type="male">1 male</specimenCount>
(35.0 × 27.0 mm) (
<specimenCode id="DBA67C26FFB4076D57E8FEFE7827FE36" box="[143,325,381,405]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1378</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D563CFEFE7B73FE36" box="[347,529,381,405]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5578FEFE7B1AFE36" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5578FEFE7B1AFE36" box="[543,632,381,405]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D55E1FEFE7BA8FE36" box="[646,714,381,405]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen in tangle nets,
<date id="FFBEF29DFFB4076D56F9FE1D7B70FE16" box="[414,530,413,437]" pageId="53" pageNumber="163" value="2002-06">
<collectingDate id="EFFA0B75FFB4076D56F9FE1D7B70FE16" box="[414,530,413,437]" pageId="53" pageNumber="163" value="2002-06">June 2002</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5525FE1E784BFDB6" collectingDate="2003-02" collectorName="M. Takeda &amp; H. Komatsu" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5525FE1E7BEDFE16" box="[578,655,413,437]" pageId="53" pageNumber="163" type="male">1 male</specimenCount>
,
<specimenCount id="9D061FD4FFB4076D55FAFE1E7A65FE16" box="[669,775,413,437]" pageId="53" pageNumber="163" type="female">2 females</specimenCount>
(1 ovigerous) (NSMT-Cr 15392),
<collectingMunicipality id="6BDB4E27FFB4076D5688FE3E7BC3FE76" box="[495,673,445,469]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D55CDFE3E7A61FE76" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D55CDFE3E7A61FE76" box="[682,771,445,469]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D57E0FE5E79A9FE56" box="[135,203,477,501]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. local fisherman via
<collectorName id="26F5B18BFFB4076D5684FE5D7B3AFE56" box="[483,600,477,501]" pageId="53" pageNumber="163">M. Takeda</collectorName>
&amp;
<collectorName id="26F5B18BFFB4076D551AFE5D7A60FE56" box="[637,770,478,501]" pageId="53" pageNumber="163">H. Komatsu</collectorName>
,
<date id="FFBEF29DFFB4076D57EFFE7E784BFDB6" box="[136,297,509,533]" pageId="53" pageNumber="163" value="2003-02">
<collectingDate id="EFFA0B75FFB4076D57EFFE7E784BFDB6" box="[136,297,509,533]" pageId="53" pageNumber="163" value="2003-02">February 2003</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5631FE7E7999FDD6" collectingDate="2003-07" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1375" specimenCount="2" specimenCount-female="2" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5631FE7E78DCFDB6" box="[342,446,509,533]" pageId="53" pageNumber="163" type="female">2 females</specimenCount>
(33.8 ×
<quantity id="4CF879B8FFB4076D557BFE7E7B1DFDB6" box="[540,639,509,533]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="53" pageNumber="163" unit="mm" value="25.3">25.3 mm</quantity>
, 40.7 × 30.7 [carapace badly damaged]) (
<specimenCode id="DBA67C26FFB4076D56ABFD9E7BE5FD96" box="[460,647,541,565]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1375</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D55FAFD9E79ACFDF6" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D57BAFDBE785AFDF6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D57BAFDBE785AFDF6" box="[221,312,573,597]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D5620FDBE78EEFDF6" box="[327,396,573,597]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D57EFFDDE7999FDD6" box="[136,251,605,629]" pageId="53" pageNumber="163" value="2003-07">
<collectingDate id="EFFA0B75FFB4076D57EFFDDE7999FDD6" box="[136,251,605,629]" pageId="53" pageNumber="163" value="2003-07">July 2003</collectingDate>
</date>
</materialsCitation>
.
<emphasis id="B974084FFFB4076D566EFDDE7846FDD6" bold="true" box="[265,292,605,629]" pageId="53" pageNumber="163"></emphasis>
<materialsCitation id="3B68DE00FFB4076D5649FDDE78D3FD16" collectingDate="2003-07" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1377" specimenCount="1" stateProvince="Bohol" typeStatus="paratype">
1 damaged male (19.5 ×
<quantity id="4CF879B8FFB4076D5536FDDE7BD5FDD6" box="[593,695,605,629]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="53" pageNumber="163" unit="mm" value="15.6">15.6 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D55B7FDDE799EFD36" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1377</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D566CFDFE78DAFD36" box="[267,440,637,661]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D56A6FDFE7B7AFD36" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D56A6FDFE7B7AFD36" box="[449,536,637,661]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D5546FDFE7B06FD36" box="[545,612,637,661]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D5624FD1E78D3FD16" box="[323,433,669,693]" pageId="53" pageNumber="163" value="2003-07">
<collectingDate id="EFFA0B75FFB4076D5624FD1E78D3FD16" box="[323,433,669,693]" pageId="53" pageNumber="163" value="2003-07">July 2003</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D56B8FD1E7B14FD56" collectingDate="2003-07-25" collectingDateMax="2003-07-30" collectingDateMin="2003-07-25" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1379" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D56B8FD1E7B5CFD16" box="[479,574,669,693]" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
(38.8 ×
<quantity id="4CF879B8FFB4076D55FAFD1E7B9DFD16" box="[669,767,669,693]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="53" pageNumber="163" unit="mm" value="28.9">28.9 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D57F6FD3E7825FD76" box="[145,327,701,725]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1379</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D563BFD3E7B70FD76" box="[348,530,701,725]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5546FD3E7B18FD76" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5546FD3E7B18FD76" box="[545,634,701,725]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D55EFFD3E7BAEFD76" box="[648,716,701,725]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D56DDFD5E7B14FD56" box="[442,630,733,757]" pageId="53" pageNumber="163" value="2003-07-25" valueMax="2003-07-30" valueMin="2003-07-25">
<collectingDate id="EFFA0B75FFB4076D56DDFD5E7B14FD56" box="[442,630,733,757]" pageId="53" pageNumber="163" value="2003-07-25" valueMax="2003-07-30" valueMin="2003-07-25">2530 July 2003</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D55C0FD5E79A3FCF6" collectingDate="2003-11" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1380" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D55C0FD5E7A65FD56" box="[679,775,733,757]" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
(35.5 ×
<quantity id="4CF879B8FFB4076D578FFD7E7833FCB6" box="[232,337,765,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="53" pageNumber="163" unit="mm" value="27.2">27.2 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D560CFD7E7B48FCB6" box="[363,554,765,789]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1380</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D5525FD7E7A60FCB6" box="[578,770,765,789]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D57EFFC9E7983FC96" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D57EFFC9E7983FC96" box="[136,225,797,821]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D578CFC9E784DFC96" box="[235,303,797,821]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D55F1FC9E79A3FCF6" pageId="53" pageNumber="163" value="2003-11">
<collectingDate id="EFFA0B75FFB4076D55F1FC9E79A3FCF6" pageId="53" pageNumber="163" value="2003-11">November 2003</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5797FCBE7A64FCD6" collectingDate="2004-01" collectionCode="AM" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCount="2" specimenCount-male="2" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5797FCBE7826FCF6" box="[240,324,829,853]" pageId="53" pageNumber="163" type="male">2 males</specimenCount>
(
<collectionCode id="ED114C98FFB4076D5632FCBE78E7FCF6" box="[341,389,829,853]" country="Australia" name="Australian Museum" pageId="53" pageNumber="163" type="Museum">AM</collectionCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D56FEFCBE7B2FFCF6" box="[409,589,829,853]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D553DFCBE7BD1FCF6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D553DFCBE7BD1FCF6" box="[602,691,829,853]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D55D8FCBE7A61FCF6" box="[703,771,829,853]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFB4076D57EDFCDE7864FCD6" box="[138,262,861,885]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB4076D57EDFCDE7864FCD6" box="[138,262,861,885]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D5515FCDD7A64FCD6" box="[626,774,861,886]" pageId="53" pageNumber="163" value="2004-01">
<collectingDate id="EFFA0B75FFB4076D5515FCDD7A64FCD6" box="[626,774,861,886]" pageId="53" pageNumber="163" value="2004-01">January 2004</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D57CAFCFE79A0FC76" collectingDate="2004-02" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1324" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D57CAFCFE7998FC36" box="[173,250,893,917]" pageId="53" pageNumber="163" type="male">1 male</specimenCount>
(
<specimenCode id="DBA67C26FFB4076D566AFCFE78A5FC36" box="[269,455,893,917]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1324</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D56BAFCFE7BFBFC36" box="[477,665,893,917]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D55C0FCFE7A60FC36" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D55C0FCFE7A60FC36" box="[679,770,893,917]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D57EFFC1E79A9FC16" box="[136,203,925,949]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFB4076D57B3FC1E782FFC16" box="[212,333,925,949]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB4076D57B3FC1E782FFC16" box="[212,333,925,949]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D55CFFC1E79A0FC76" pageId="53" pageNumber="163" value="2004-02">
<collectingDate id="EFFA0B75FFB4076D55CFFC1E79A0FC76" pageId="53" pageNumber="163" value="2004-02">February 2004</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5791FC3E782BFBB6" collectingDate="2004-03" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1384" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5791FC3E783AFC76" box="[246,344,957,981]" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
(19.9 ×
<quantity id="4CF879B8FFB4076D56A6FC3E7B45FC76" box="[449,551,957,981]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="53" pageNumber="163" unit="mm" value="14.7">14.7 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D5526FC3E7B99FC76" box="[577,763,957,981]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1384</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D57EFFC5E785AFC56" box="[136,312,989,1013]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5626FC5E78FBFC56" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5626FC5E78FBFC56" box="[321,409,989,1013]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D56C4FC5E7885FC56" box="[419,487,989,1013]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D57A5FC7E782BFBB6" box="[194,329,1021,1045]" pageId="53" pageNumber="163" value="2004-03">
<collectingDate id="EFFA0B75FFB4076D57A5FC7E782BFBB6" box="[194,329,1021,1045]" pageId="53" pageNumber="163" value="2004-03">March 2004</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5610FC7E78A2FBF6" collectingDate="2004-04" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1222" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5610FC7E78A1FBB6" box="[375,451,1021,1045]" pageId="53" pageNumber="163" type="male">1 male</specimenCount>
,
<specimenCount id="9D061FD4FFB4076D56B7FC7E7B5AFBB6" box="[464,568,1021,1045]" pageId="53" pageNumber="163" type="female">2 females</specimenCount>
(
<specimenCode id="DBA67C26FFB4076D552FFC7E7B9EFBB6" box="[584,764,1021,1045]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1222</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D57EDFB9E785AFB96" box="[138,312,1053,1077]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5627FB9E78F5FB96" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5627FB9E78F5FB96" box="[320,407,1053,1077]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D56C6FB9E7886FB96" box="[417,484,1053,1077]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFB4076D568AFB9E7B07FB96" box="[493,613,1053,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB4076D568AFB9E7B07FB96" box="[493,613,1053,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D5621FBBE78A2FBF6" box="[326,448,1085,1109]" pageId="53" pageNumber="163" value="2004-04">
<collectingDate id="EFFA0B75FFB4076D5621FBBE78A2FBF6" box="[326,448,1085,1109]" pageId="53" pageNumber="163" value="2004-04">April 2004</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D5697FBBE7B43FB36" collectingDate="2004-05" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1373" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5697FBBE7B59FBF6" box="[496,571,1085,1109]" pageId="53" pageNumber="163" type="male">1 male</specimenCount>
(29.0 ×
<quantity id="4CF879B8FFB4076D55FAFBBE7B9DFBF6" box="[669,767,1085,1109]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="53" pageNumber="163" unit="mm" value="21.9">21.9 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D57E8FBDE7827FBD6" box="[143,325,1117,1141]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1373</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D563CFBDE7B73FBD6" box="[347,529,1117,1141]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5578FBDE7B1AFBD6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5578FBDE7B1AFBD6" box="[543,632,1117,1141]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D55E1FBDE7BA8FBD6" box="[646,714,1117,1141]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D56D7FBFD7B43FB36" box="[432,545,1149,1174]" pageId="53" pageNumber="163" value="2004-05">
<collectingDate id="EFFA0B75FFB4076D56D7FBFD7B43FB36" box="[432,545,1149,1174]" pageId="53" pageNumber="163" value="2004-05">May 2004</collectingDate>
</date>
</materialsCitation>
. —
<specimenCount id="9D061FD4FFB4076D552AFBFE7BFDFB36" box="[589,671,1149,1173]" pageId="53" pageNumber="163" type="male">4 males</specimenCount>
(smallest 23.4 ×
<quantity id="4CF879B8FFB4076D57BDFB1E785DFB16" box="[218,319,1181,1205]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="53" pageNumber="163" unit="mm" value="17.4">17.4 mm</quantity>
, largest 42.0 ×
<quantity id="4CF879B8FFB4076D5696FB1E7B3AFB16" box="[497,600,1181,1205]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="53" pageNumber="163" unit="mm" value="31.8">31.8 mm</quantity>
),
<specimenCount id="9D061FD4FFB4076D550AFB1E7BAFFB16" box="[621,717,1181,1205]" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
with rhizocephalan (28.2 ×
<quantity id="4CF879B8FFB4076D56E0FB3E788DFB76" box="[391,495,1213,1237]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="53" pageNumber="163" unit="mm" value="21.4">21.4 mm</quantity>
),
<materialsCitation id="3B68DE00FFB4076D5562FB3E79A1FA96" collectingDate="2004-05-29" collectionCode="ZRC" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1372" specimenCount="2" specimenCount-female="2" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D5562FB3E7B12FB76" box="[517,624,1213,1237]" pageId="53" pageNumber="163" type="female">2 females</specimenCount>
(33.4 ×
<quantity id="4CF879B8FFB4076D55B0FB3E79D6FB56" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="53" pageNumber="163" unit="mm" value="25.7">25.7 mm</quantity>
, 45.9 × 35.0 mm) (
<specimenCode id="DBA67C26FFB4076D56EFFB5E7B5EFB56" box="[392,572,1245,1269]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1372</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D5528FB5E7A61FB56" box="[591,771,1245,1269]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D57EDFB7E7981FAB6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D57EDFB7E7981FAB6" box="[138,227,1277,1301]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D5796FB7E7857FAB6" box="[241,309,1277,1301]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D55D5FB7E79A1FA96" pageId="53" pageNumber="163" value="2004-05-29">
<collectingDate id="EFFA0B75FFB4076D55D5FB7E79A1FA96" pageId="53" pageNumber="163" value="2004-05-29">29 May 2004</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFB4076D579FFA9E78FAFAD6" collectingDate="2005-02-29" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1374" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFB4076D579FFA9E783EFA96" box="[248,348,1309,1333]" pageId="53" pageNumber="163" type="female">1 female</specimenCount>
(
<specimenCode id="DBA67C26FFB4076D5609FA9E7B4EFA96" box="[366,556,1309,1333]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1374</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFB4076D5524FA9E7A60FA96" box="[579,770,1309,1333]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D57EFFABE7981FAF6" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D57EFFABE7981FAF6" box="[136,227,1341,1365]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D5795FABE785AFAF6" box="[242,312,1341,1365]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFB4076D5621FABE78AAFAF6" box="[326,456,1341,1365]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFB4076D5621FABE78AAFAF6" box="[326,456,1341,1365]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="53" pageNumber="163" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D57A0FADE78FAFAD6" box="[199,408,1373,1397]" pageId="53" pageNumber="163" value="2005-02-29">
<collectingDate id="EFFA0B75FFB4076D57A0FADE78FAFAD6" box="[199,408,1373,1397]" pageId="53" pageNumber="163" value="2005-02-29">29 February 2005</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B68DE00FFB4076D56C1FADE78FAFA76" collectingDate="2001-11-28" collectionCode="ZRC" country="Philippines" county="Others" location="Panglao" municipality="Balicasag Island" pageId="53" pageNumber="163" specimenCode="ZRC 2013.1323, ZRC 2000.585" specimenCount="1" stateProvince="Bohol" typeStatus="paratype">
<collectingCounty id="62DEACD1FFB4076D56C1FADE7896FAD6" box="[422,500,1373,1397]" pageId="53" pageNumber="163">Others</collectingCounty>
: 1 badly damaged male (21.5 ×
<quantity id="4CF879B8FFB4076D57BAFAFE785FFA36" box="[221,317,1405,1429]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="53" pageNumber="163" unit="mm" value="17.8">17.8 mm</quantity>
) (
<specimenCode id="DBA67C26FFB4076D5634FAFE7B61FA36" box="[339,515,1405,1429]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2013.1323</specimenCode>
, ex part
<specimenCode id="DBA67C26FFB4076D5538FAFE7A62FA36" box="[607,768,1405,1429]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="53" pageNumber="163">ZRC 2000.585</specimenCode>
)
<collectingMunicipality id="6BDB4E27FFB4076D57EFFA1E785BFA16" box="[136,313,1437,1461]" pageId="53" pageNumber="163">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFB4076D5625FA1E78F8FA16" LSID="urn:lsid:plazi:treatment:03A9654BFFB5076D5494FA1E7ABAFBF6:8EDF8286FFB4076D5625FA1E78F8FA16" box="[322,410,1437,1461]" country="Philippines" county="Others" municipality="Balicasag Island" name="Panglao" pageId="53" pageNumber="163" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFB4076D56C4FA1E7885FA16" box="[419,487,1437,1461]" country="Philippines" name="Bohol" pageId="53" pageNumber="163">Bohol</collectingRegion>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFB4076D57A5FA3E78FAFA76" box="[194,408,1469,1493]" pageId="53" pageNumber="163" value="2001-11-28">
<collectingDate id="EFFA0B75FFB4076D57A5FA3E78FAFA76" box="[194,408,1469,1493]" pageId="53" pageNumber="163" value="2001-11-28">28 November 2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFB4076D57EDFA7E787CF836" pageId="53" pageNumber="163" type="diagnosis">
<paragraph id="8BBFD45DFFB4076D57EDFA7E787CF836" blockId="53.[135,776,1533,1941]" pageId="53" pageNumber="163">
<emphasis id="B974084FFFB4076D57EDFA7E7866F9B6" bold="true" box="[138,260,1533,1557]" pageId="53" pageNumber="163">Diagnosis.</emphasis>
Carapace covered with small granules (
<figureCitation id="133BC8D8FFB4076D55BEFA7E79D5F996" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="53" pageNumber="163">Fig. 24F</figureCitation>
). Pseudorostral spines long, strongly diverging and usually curved upwards (
<figureCitation id="133BC8D8FFB4076D56EAF9BE789AF9F6" box="[397,504,1597,1621]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="53" pageNumber="163">Figs. 37G</figureCitation>
). Antorbital spine curved; intercalated spine triangular with large gaps; postorbital spine very long; hepatic spine relatively small (
<figureCitation id="133BC8D8FFB4076D55F0F9FE7A61F936" box="[663,771,1661,1685]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="53" pageNumber="163">Figs. 24F</figureCitation>
,
<figureCitation id="133BC8D8FFB4076D57EFF91E79D8F916" box="[136,186,1693,1717]" captionStart="Fig" captionStartId="73.[135,170,1833,1855]" captionTargetBox="[138,1471,189,1817]" captionTargetId="figure-8@73.[138,1471,188,1818]" captionTargetPageId="73" captionText="Fig. 37. Frontal regions of carapaces. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; J, Holthuija poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea; K, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; L, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; M, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; N, Sakaija longispinosa n. sp., holotype ovigerous female (11.4 × 8.6 mm) (NMV J63792), Australia; O, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384672" httpUri="https://zenodo.org/record/5384672/files/figure.png" pageId="53" pageNumber="163">37G</figureCitation>
). Lateral margin with 3 long spines, posterior longest; 1 branchial long spine; median row with 2 spines: 1 gastric, 1 cardiac; 2 short spines on posterior carapace margin (
<figureCitation id="133BC8D8FFB4076D57F7F97E7997F8B6" box="[144,245,1789,1813]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="53" pageNumber="163">Fig. 24F</figureCitation>
). Basal antennal article broad, granulated, with longitudinal furrow, 2 strong distal spines (
<figureCitation id="133BC8D8FFB4076D55EBF89E7B98F896" box="[652,762,1821,1845]" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="53" pageNumber="163">Fig. 40G</figureCitation>
). Epistome with 2 low tubercles on anterior margin (
<figureCitation id="133BC8D8FFB4076D55BDF8BE79D9F8D6" captionStart="Fig" captionStartId="76.[113,148,1692,1714]" captionTargetBox="[113,1451,331,1676]" captionTargetId="figure-15@76.[113,1451,331,1677]" captionTargetPageId="76" captionText="Fig. 40. Antennae, antennules and epistomes. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Sakaija africana, male (32.2 × 25.4 mm) (MNHN-IU-2010-928), Madagascar; L, Sakaija serenei n. sp., holotype male (17.4 × 14.7 mm) (NMCR), Philippines; M, Sakaija santo n. sp., holotype male (9.4 × 6.6 mm) (MNHN), Vanuatu; N, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; O, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384678" httpUri="https://zenodo.org/record/5384678/files/figure.png" pageId="53" pageNumber="163">Fig. 40G</figureCitation>
). G1 almost straight with distal part slightly dilated (
<figureCitation id="133BC8D8FFB4076D57F7F8FE7870F836" box="[144,274,1917,1941]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. AC, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; FH, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; IK, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; LO, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [AF, I, L]; 0.5 mm [D, G, H, J, K, MO]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="53" pageNumber="163">Fig. 27FH</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFB4076D57EFF83E78B3F7B6" pageId="53" pageNumber="163" type="etymology">
<paragraph id="8BBFD45DFFB4076D57EFF83E78B3F7B6" blockId="53.[136,774,1981,2070]" pageId="53" pageNumber="163">
<emphasis id="B974084FFFB4076D57EFF83E787BF876" bold="true" box="[136,281,1981,2005]" pageId="53" pageNumber="163">Etymology.</emphasis>
The name honors the memory of Paul Beaudemoulin, father-in-law of the second author, who passed away in
<date id="FFBEF29DFFB4076D5651F87E78ACF7B6" box="[310,462,2045,2069]" pageId="53" pageNumber="163" value="2012-10">October 2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFB4076D5422FF3E7ABAFBF6" pageId="53" pageNumber="163" type="discussion">
<paragraph id="8BBFD45DFFB4076D5422FF3E7AAEFE76" blockId="53.[835,1475,189,469]" pageId="53" pageNumber="163">
<emphasis id="B974084FFFB4076D5422FF3E7AD8FF76" bold="true" box="[837,954,189,213]" pageId="53" pageNumber="163">Remarks.</emphasis>
Of the
<taxonomicName id="4C00AFDEFFB4076D537DFF3E7D85FF76" box="[1050,1255,189,213]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B974084FFFB4076D537DFF3E7DE5FF76" box="[1050,1159,189,213]" italics="true" pageId="53" pageNumber="163">Holthuija</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C00AFDEFFB4076D5390FF3D7C3AFF76" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1271,1368,189,213]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli" status="sp. nov.">
<emphasis id="B974084FFFB4076D5390FF3D7C3AFF76" box="[1271,1368,189,213]" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFB4076D5204FF3E7CC6FF77" box="[1379,1444,189,212]" pageId="53" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
is distinct in possessing very long carapace spines which are prominently curved obliquely upwards. The large number of specimens we have examined show that the length and orientation of these spines are not variable, even for small specimens and females. It also has proportionately, the longest ambulatory legs (
<figureCitation id="133BC8D8FFB4076D5335FEFE7DD0FE36" box="[1106,1202,381,405]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="53" pageNumber="163">Fig. 24F</figureCitation>
). In the other species, the ambulatory legs, particularly the merus, are shorter (
<figureCitation id="133BC8D8FFB4076D52EAFE1E7AF6FE76" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="53" pageNumber="163">Figs. 24AD</figureCitation>
,
<figureCitation id="133BC8D8FFB4076D54C6FE3E7ADDFE76" box="[929,959,445,469]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="53" pageNumber="163">25</figureCitation>
).
</paragraph>
<paragraph id="8BBFD45DFFB4076D5424FE7E7C66FD76" blockId="53.[835,1476,509,725]" pageId="53" pageNumber="163">
The G1 structure of
<taxonomicName id="4C00AFDEFFB4076D5379FE7D7D14FDB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1054,1142,509,533]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D5379FE7D7D14FDB6" box="[1054,1142,509,533]" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
most closely resembles that of
<taxonomicName id="4C00AFDEFFB4076D5423FD9D7ADBFD96" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[836,953,542,565]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="cognata" status="sp. nov.">
<emphasis id="B974084FFFB4076D5423FD9D7ADBFD96" box="[836,953,542,565]" italics="true" pageId="53" pageNumber="163">H. cognata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFB4076D54D9FD9E7A9AFD97" box="[958,1016,541,564]" pageId="53" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
, with the distal part laterally flattened, even though their carapace and legs are quite different (carapace relatively broader with longer and more curved spines, and with proportionately longer legs in
<taxonomicName id="4C00AFDEFFB4076D53A5FDFD7C78FD36" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1218,1306,637,661]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D53A5FDFD7C78FD36" box="[1218,1306,637,661]" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
). The G1 of
<taxonomicName id="4C00AFDEFFB4076D52CEFDFD7A18FD16" authorityName="Ng &amp; Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D52CEFDFD7A18FD16" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
, however, is relatively more slender than that of
<taxonomicName id="4C00AFDEFFB4076D52CFFD1D7AFBFD76" authorityName="Ng &amp; Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="cognata">
<emphasis id="B974084FFFB4076D52CFFD1D7AFBFD76" italics="true" pageId="53" pageNumber="163">H. cognata</emphasis>
</taxonomicName>
(
<figureCitation id="133BC8D8FFB4076D54CDFD3E7D4AFD76" box="[938,1064,701,725]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. AC, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; FH, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; IK, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; LO, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [AF, I, L]; 0.5 mm [D, G, H, J, K, MO]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="53" pageNumber="163">Fig. 27FH</figureCitation>
versus
<figureCitation id="133BC8D8FFB4076D531AFD3E7D9BFD76" box="[1149,1273,701,725]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. AC, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; FH, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; IK, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; LO, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [AF, I, L]; 0.5 mm [D, G, H, J, K, MO]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="53" pageNumber="163">Fig. 27IK</figureCitation>
).
</paragraph>
<paragraph id="8BBFD45DFFB4076D5424FD7E7CA0FC76" blockId="53.[834,1474,765,981]" pageId="53" pageNumber="163">
<taxonomicName id="4C00AFDEFFB4076D5424FD7E7A8AFCB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[835,1000,765,789]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D5424FD7E7A8AFCB6" box="[835,1000,765,789]" italics="true" pageId="53" pageNumber="163">Holthuija pauli</emphasis>
</taxonomicName>
might be confused with small specimens of
<taxonomicName id="4C00AFDEFFB4076D5423FC9D7D46FC96" baseAuthorityName="De Haan" baseAuthorityYear="1837" box="[836,1060,798,821]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="spinigera">
<emphasis id="B974084FFFB4076D5423FC9D7D46FC96" box="[836,1060,798,821]" italics="true" pageId="53" pageNumber="163">Paramaya spinigera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C00AFDEFFB4076D5339FC9D7C6EFC96" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1118,1292,797,821]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="ouch">
<emphasis id="B974084FFFB4076D5339FC9D7C6EFC96" box="[1118,1292,797,821]" italics="true" pageId="53" pageNumber="163">Paramaya ouch</emphasis>
</taxonomicName>
, especially with regard to the relatively long carapace spines which are curved and the very divergent pseudorostral spines.
<taxonomicName id="4C00AFDEFFB4076D5228FCDD7AF6FC36" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B974084FFFB4076D5228FCDD7CA0FCD6" box="[1359,1474,862,885]" italics="true" pageId="53" pageNumber="163">Paramaya</emphasis>
species
</taxonomicName>
, however, have long distal spines on the meri of ambulatory legs (absent in
<taxonomicName id="4C00AFDEFFB4076D530BFC1D7DA1FC16" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1132,1219,925,949]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D530BFC1D7DA1FC16" box="[1132,1219,925,949]" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
) as well as many other characters at the generic level (see discussion for
<taxonomicName id="4C00AFDEFFB4076D5224FC3D7CD3FC76" authorityName="De Haan" authorityYear="1837" box="[1347,1457,958,981]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFB4076D5224FC3D7CD3FC76" box="[1347,1457,958,981]" italics="true" pageId="53" pageNumber="163">Paramaya</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBFD45DFFB4076D5424FC7E7ABAFBF6" blockId="53.[835,1474,1021,1109]" pageId="53" pageNumber="163">
In life,
<taxonomicName id="4C00AFDEFFB4076D54FFFC7D7A96FBB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[920,1012,1021,1045]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="163" phylum="Arthropoda" rank="species" species="pauli">
<emphasis id="B974084FFFB4076D54FFFC7D7A96FBB6" box="[920,1012,1021,1045]" italics="true" pageId="53" pageNumber="163">H. pauli</emphasis>
</taxonomicName>
is beige and light reddish-brown overall, with the third maxillipeds and abdomen brighter in colour (
<figureCitation id="133BC8D8FFB4076D542CFBBE7AA9FBF6" box="[843,971,1085,1109]" captionStart="Fig" captionStartId="108.[113,148,1935,1957]" captionTargetBox="[113,1451,260,1919]" captionTargetId="figure-15@108.[113,1451,260,1920]" captionTargetPageId="108" captionText="Fig. 69. Colours in life. A, Alcomaja nagashimaensis, male (specimen not examined), Philippines; B, Alcomaja miriky n. sp., paratype male (17.3 × 12.7 mm) (MNHN-IU-2010-60), Madagascar; C, D, Holthuija miersii, ovigerous female (50.2 × 40.4 mm) (ZRC 2013.1272), Singapore; E, Holthuija pauli n. sp., paratype male (31.5 × 23.4 mm) (ZRC 2001.0590), Philippines; F, Holthuija pauli n. sp., female (38.8 × 28.9 mm) (ZRC 2013.1379), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384744" httpUri="https://zenodo.org/record/5384744/files/figure.png" pageId="53" pageNumber="163">Fig. 69E, F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>