170 lines
32 KiB
XML
170 lines
32 KiB
XML
<document id="9354685A1D55E5A26EC1413FF5103D77" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. & Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFAB07725653FA327D3FFA0A" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Alcomaja latens Ng & Forges 2015, n. sp." docType="treatment" docUuid="BD7762D7-E054-4F37-B9D7-EBD4C3CD5475" docUuidSource="ZooBank" docVersion="5" lastPageNumber="152" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="152" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8382CB1F637BD848216DDA3A0196A006">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="03AB9FFBB3E5BA992A88718D8CB1159B">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="998CA50EF6F4988F756AD46F96063BF5">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="084AC61727B45914BF015F4B00EE8A8F">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="B049001541B7B5EC9B3C9E067B9B899A">2015</mods:date>
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<mods:number id="31C140EA8A3485FFF23D7AC3A6871BB4">2015-05-29</mods:number>
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<mods:number id="9F5CBBC39B0F2336A9FD5A4F921A54E5">63</mods:number>
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<treatment id="03A9654BFFAB07725653FA327D3FFA0A" ID-DOI="http://doi.org/10.5281/zenodo.5466799" ID-Zenodo-Dep="5466799" LSID="urn:lsid:zoobank.org:act:BD7762D7-E054-4F37-B9D7-EBD4C3CD5475" httpUri="http://treatment.plazi.org/id/03A9654BFFAB07725653FA327D3FFA0A" lastPageNumber="152" pageId="42" pageNumber="152">
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<subSubSection id="C31A87D6FFAB07725653FA327B4CFA6A" box="[308,558,1457,1481]" pageId="42" pageNumber="152" type="nomenclature">
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<paragraph id="8BBFD45DFFAB07725653FA327B4CFA6A" blockId="42.[234,631,1457,1513]" box="[308,558,1457,1481]" pageId="42" pageNumber="152">
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<heading id="D0F76331FFAB07725653FA327B4CFA6A" bold="true" box="[308,558,1457,1481]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="152" reason="2">
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<taxonomicName id="4C00AFDEFFAB07725653FA327885FA6A" ID-CoL="BKVW" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[308,487,1457,1481]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="latens" status="sp. nov.">
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<emphasis id="B974084FFFAB07725653FA327885FA6A" bold="true" box="[308,487,1457,1481]" italics="true" pageId="42" pageNumber="152">Alcomaja latens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B534FFAB07725697FA327B4CFA6A" box="[496,558,1457,1481]" pageId="42" pageNumber="152" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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<subSubSection id="C31A87D6FFAB0772578DFA527B15FA4A" box="[234,631,1489,1513]" pageId="42" pageNumber="152" type="description">
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<paragraph id="8BBFD45DFFAB0772578DFA527B15FA4A" blockId="42.[234,631,1457,1513]" box="[234,631,1489,1513]" pageId="42" pageNumber="152">
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(
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<figureCitation id="133BC8D8FFAB07725795FA527802FA4A" box="[242,352,1489,1513]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="42" pageNumber="152">Figs. 17D</figureCitation>
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,
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<figureCitation id="133BC8D8FFAB0772560AFA5278DCFA4A" box="[365,446,1489,1513]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="42" pageNumber="152">20K–N</figureCitation>
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,
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<figureCitation id="133BC8D8FFAB077256ACFA52789EFA4A" box="[459,508,1489,1513]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="42" pageNumber="152">36N</figureCitation>
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,
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<figureCitation id="133BC8D8FFAB0772556FFA527B5AFA4A" box="[520,568,1489,1513]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="42" pageNumber="152">45G</figureCitation>
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,
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<figureCitation id="133BC8D8FFAB07725522FA527B12FA4A" box="[581,624,1489,1513]" captionStart="Fig" captionStartId="85.[135,170,1745,1767]" captionTargetBox="[136,1474,392,1730]" captionTargetId="figure-8@85.[136,1474,391,1732]" captionTargetPageId="85" captionText="Fig. 49. Male anterior thoracic sternums and abdomens, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A.gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; E, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; F, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; G, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; H, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384700" httpUri="https://zenodo.org/record/5384700/files/figure.png" pageId="42" pageNumber="152">49F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A87D6FFAB07725716F992782FF92A" pageId="42" pageNumber="152" type="materials_examined">
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<paragraph id="8BBFD45DFFAB07725716F992782FF92A" blockId="42.[112,752,1553,1673]" pageId="42" pageNumber="152">
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<emphasis id="B974084FFFAB07725716F992783FF98A" bold="true" box="[113,349,1553,1577]" pageId="42" pageNumber="152">Material examined.</emphasis>
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<materialsCitation id="3B68DE00FFAB07725600F992782FF92A" collectingDate="2004-10-24" collectionCode="RV" country="Solomon Islands" elevation="202" latitude="-8.4" location="Alis" longLatPrecision="1301" longitude="159.45" pageId="42" pageNumber="152" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54BB6AFFFFAB07725600F99278B1F98A" box="[359,467,1553,1577]" pageId="42" pageNumber="152" type="holotype">Holotype</typeStatus>
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: male (25.3 ×
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<quantity id="4CF879B8FFAB077255E6F9927B8BF98A" box="[641,745,1553,1577]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="42" pageNumber="152" unit="mm" value="21.4">21.4 mm</quantity>
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) (MNHN-IU-2013-4050), station DW 2190,
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<geoCoordinate id="EE34B29AFFAB07725550F9B27BE3F9EA" box="[567,641,1585,1609]" degrees="8" direction="south" minutes="24" orientation="latitude" pageId="42" pageNumber="152" precision="925" value="-8.4">8°24’S</geoCoordinate>
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<geoCoordinate id="EE34B29AFFAB077255E2F9B27B8EF9EA" box="[645,748,1585,1609]" degrees="159" direction="east" minutes="27" orientation="longitude" pageId="42" pageNumber="152" precision="925" value="159.45">159°27’E</geoCoordinate>
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,
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<quantity id="4CF879B8FFAB07725717F9D2798DF9CA" box="[112,239,1617,1641]" metricMagnitude="2" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.63" metricValueMin="1.4" pageId="42" pageNumber="152" unit="m" value="201.5" valueMax="263.0" valueMin="140.0">
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<elevation id="002D336EFFAB07725717F9D2798DF9CA" box="[112,239,1617,1641]" metricMagnitude="2" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.63" metricValueMin="1.4" pageId="42" pageNumber="152" unit="m" value="201.5" valueMax="263.0" valueMin="140.0">140–263 m</elevation>
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</quantity>
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,
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<collectingCountry id="F31794CDFFAB07725799F9D278DAF9CA" box="[254,440,1617,1641]" name="Solomon Islands" pageId="42" pageNumber="152">Solomon Islands</collectingCountry>
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, coll.
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<collectionCode id="ED114C98FFAB07725698F9D17B46F9CA" box="[511,548,1618,1641]" country="Italy" name="Collection of Leptospira Strains" pageId="42" pageNumber="152">RV</collectionCode>
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<emphasis id="B974084FFFAB07725556F9D27B39F9CA" box="[561,603,1617,1641]" italics="true" pageId="42" pageNumber="152">
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<location id="8EDF8286FFAB07725556F9D27B39F9CA" LSID="urn:lsid:plazi:treatment:03A9654BFFAB07725653FA327D3FFA0A:8EDF8286FFAB07725556F9D27B39F9CA" box="[561,603,1617,1641]" country="Solomon Islands" latitude="-8.4" longLatPrecision="1301" longitude="159.45" name="Alis" pageId="42" pageNumber="152">Alis</location>
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</emphasis>
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, SALOMON 2,
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<date id="FFBEF29DFFAB077257E9F9F2782BF92A" box="[142,329,1649,1673]" pageId="42" pageNumber="152" value="2004-10-24">
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<collectingDate id="EFFA0B75FFAB077257E9F9F2782BF92A" box="[142,329,1649,1673]" pageId="42" pageNumber="152" value="2004-10-24">24 October 2004</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A87D6FFAB07725714F9327D1EFBAA" pageId="42" pageNumber="152" type="diagnosis">
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<paragraph id="8BBFD45DFFAB07725714F9327D1EFBAA" blockId="42.[112,753,1713,2090]" lastBlockId="42.[812,1452,945,1033]" pageId="42" pageNumber="152">
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<emphasis id="B974084FFFAB07725714F932798BF96A" bold="true" box="[115,233,1713,1737]" pageId="42" pageNumber="152">Diagnosis.</emphasis>
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Carapace pyriform; dorsal surface of carapace and ambulatory legs covered with short, hooked setae; with small rounded granules (
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<figureCitation id="133BC8D8FFAB07725614F97278B6F8AA" box="[371,468,1777,1801]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="42" pageNumber="152">Fig. 17D</figureCitation>
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). Pseudorostral spines with rounded cross-section (
|
||
<figureCitation id="133BC8D8FFAB07725617F89278B7F88A" box="[368,469,1809,1833]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="42" pageNumber="152">Fig. 36N</figureCitation>
|
||
). Antorbital spine almost straight; intercalated spine subtruncate basally; postocular spine strong, pointing anteriorly and outwards; 1 distinct hepatic spine with small basal spine (
|
||
<figureCitation id="133BC8D8FFAB07725566F8F27B03F82A" box="[513,609,1905,1929]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="42" pageNumber="152">Fig. 36N</figureCitation>
|
||
). Lateral and branchial spines strong in juveniles and adults (
|
||
<figureCitation id="133BC8D8FFAB07725519F8127B86F80A" box="[638,740,1937,1961]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="42" pageNumber="152">Fig. 17D</figureCitation>
|
||
). Subhepatic region with rugose and weakly granulated surface. Basal antennal article with low granules and 2 tubercles, with 2 distal spines. Ischium of third maxilliped almost smooth to slightly rugose (
|
||
<figureCitation id="133BC8D8FFAB0772567BF792781DF78A" box="[284,383,2065,2089]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="42" pageNumber="152">Fig. 45G</figureCitation>
|
||
). Ambulatory legs relatively short, first leg never twice length of carapace; surfaces distinctly setose (
|
||
<figureCitation id="133BC8D8FFAB0772541AFC527ABDFC4A" box="[893,991,977,1001]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="42" pageNumber="152">Fig. 17D</figureCitation>
|
||
). G1 gently curved, with distal part gently curved upwards (
|
||
<figureCitation id="133BC8D8FFAB0772548DFC727D13FBAA" box="[1002,1137,1009,1033]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="42" pageNumber="152">Fig. 20K–N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A87D6FFAB0772544AFBB27D8CFB2A" pageId="42" pageNumber="152" type="etymology">
|
||
<paragraph id="8BBFD45DFFAB0772544AFBB27D8CFB2A" blockId="42.[811,1451,1073,1161]" pageId="42" pageNumber="152">
|
||
<emphasis id="B974084FFFAB0772544AFBB27ACFFBEA" bold="true" box="[813,941,1073,1097]" pageId="42" pageNumber="152">Etymology.</emphasis>
|
||
The name is derived from the Latin for “hidden”, alluding to a general absence of external characters to separate the new species from
|
||
<taxonomicName id="4C00AFDEFFAB07725378FBF17D85FB2A" authorityName="T. Sakai" authorityYear="1969" box="[1055,1255,1137,1161]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="nagashimaensis">
|
||
<emphasis id="B974084FFFAB07725378FBF17D85FB2A" box="[1055,1255,1137,1161]" italics="true" pageId="42" pageNumber="152">A. nagashimaensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A87D6FFAB0772544AFB327D3FFA0A" pageId="42" pageNumber="152" type="discussion">
|
||
<paragraph id="8BBFD45DFFAB0772544AFB327D3FFA0A" blockId="42.[811,1452,1201,1449]" pageId="42" pageNumber="152">
|
||
<emphasis id="B974084FFFAB0772544AFB327AC1FB6A" bold="true" box="[813,931,1201,1225]" pageId="42" pageNumber="152">Remarks.</emphasis>
|
||
<taxonomicName id="4C00AFDEFFAB077254D7FB327D12FB6A" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[944,1136,1201,1225]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="latens" status="sp. nov.">
|
||
<emphasis id="B974084FFFAB077254D7FB327D12FB6A" box="[944,1136,1201,1225]" italics="true" pageId="42" pageNumber="152">Alcomaja latens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFAB0772531BFB317DDCFB6A" box="[1148,1214,1202,1225]" pageId="42" pageNumber="152" rank="species">n. sp.</taxonomicNameLabel>
|
||
is very close to
|
||
<taxonomicName id="4C00AFDEFFAB077252F4FB317DE2FB4A" authority="(T. Sakai, 1969)" baseAuthorityName="T. Sakai" baseAuthorityYear="1969" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="nagashimaensis">
|
||
<emphasis id="B974084FFFAB077252F4FB317AB7FB4A" italics="true" pageId="42" pageNumber="152">A. nagashimaensis</emphasis>
|
||
(T.
|
||
<bibRefCitation id="EF91A9ACFFAB07725499FB527D1BFB4A" author="Sakai T" box="[1022,1145,1233,1257]" pageId="42" pageNumber="152" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and the two species can only be effectively separated by the form of the G1. In
|
||
<taxonomicName id="4C00AFDEFFAB07725222FB727A0FFA8A" authorityName="Ng & Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="latens">
|
||
<emphasis id="B974084FFFAB07725222FB727A0FFA8A" italics="true" pageId="42" pageNumber="152">Alcomaja latens</emphasis>
|
||
</taxonomicName>
|
||
, the dorsal part is gently bent upwards (
|
||
<figureCitation id="133BC8D8FFAB07725246FA927CC7FA8A" box="[1313,1445,1297,1321]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="42" pageNumber="152">Fig. 20K–N</figureCitation>
|
||
) while in
|
||
<taxonomicName id="4C00AFDEFFAB077254E9FAB17D34FAEA" baseAuthorityName="T. Sakai" baseAuthorityYear="1969" box="[910,1110,1329,1353]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="nagashimaensis">
|
||
<emphasis id="B974084FFFAB077254E9FAB17D34FAEA" box="[910,1110,1329,1353]" italics="true" pageId="42" pageNumber="152">A. nagashimaensis</emphasis>
|
||
</taxonomicName>
|
||
, it is gently curved downwards (
|
||
<figureCitation id="133BC8D8FFAB07725451FAD27AD1FACA" box="[822,947,1361,1385]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="42" pageNumber="152">Fig. 20A–J</figureCitation>
|
||
). At around the same size as the
|
||
<typeStatus id="54BB6AFFFFAB07725273FAD27C13FACA" box="[1300,1393,1361,1385]" pageId="42" pageNumber="152" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C00AFDEFFAB077252F2FAD17A0FFA2A" authorityName="Ng & Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="latens">
|
||
<emphasis id="B974084FFFAB077252F2FAD17A0FFA2A" italics="true" pageId="42" pageNumber="152">A. latens</emphasis>
|
||
</taxonomicName>
|
||
, a male specimen of
|
||
<taxonomicName id="4C00AFDEFFAB0772533BFAF17C44FA2A" baseAuthorityName="T. Sakai" baseAuthorityYear="1969" box="[1116,1318,1393,1417]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="152" phylum="Arthropoda" rank="species" species="nagashimaensis">
|
||
<emphasis id="B974084FFFAB0772533BFAF17C44FA2A" box="[1116,1318,1393,1417]" italics="true" pageId="42" pageNumber="152">A. nagashimaensis</emphasis>
|
||
</taxonomicName>
|
||
already has distinctly inflated chelipeds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |