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<document id="961FAFAF10AEB2E93C36391DECD7E5EB" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFF94074E5361FF3E7B41F8D6" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Paramaja Kubo 1936" docType="treatment" docVersion="4" lastPageNumber="132" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="131" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="28B2798AA7B46CA56415F3ECDB0F9E5D">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="9F7858A8D9B2F6DC7B10DE0DD944C6A7">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="DE312FAAB50A606093DC395F1F722775">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="6DC8DB4D1C7EA26FDDDDA87C285236C4">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="F218EC8B1BC8E9DAD7F1C78F1FD0ABBC">2015</mods:date>
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<treatment id="03A9654BFF94074E5361FF3E7B41F8D6" ID-DOI="http://doi.org/10.5281/zenodo.5466791" ID-Zenodo-Dep="5466791" LSID="urn:lsid:plazi:treatment:03A9654BFF94074E5361FF3E7B41F8D6" httpUri="http://treatment.plazi.org/id/03A9654BFF94074E5361FF3E7B41F8D6" lastPageId="22" lastPageNumber="132" pageId="21" pageNumber="131">
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<paragraph id="8BBFD45DFF94074D5361FF3E7D9DFF76" blockId="21.[1030,1279,189,213]" box="[1030,1279,189,213]" pageId="21" pageNumber="131">
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<taxonomicName id="4C00AFDEFF94074D5361FF3E7D9DFF76" ID-CoL="7PRYM" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[1030,1279,189,213]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D5361FF3E7D9DFF76" bold="true" box="[1030,1279,189,213]" pageId="21" pageNumber="131">
<emphasis id="B974084FFF94074D5361FF3E7D10FF76" bold="true" box="[1030,1138,189,213]" italics="true" pageId="21" pageNumber="131">Paramaja</emphasis>
<bibRefCitation id="EF91A9ACFF94074D531CFF3E7D9DFF76" author="Kubo I" box="[1147,1279,189,213]" pageId="21" pageNumber="131" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
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<paragraph id="8BBFD45DFF94074D5424FF7F7D3AFEB2" blockId="21.[835,1475,251,389]" box="[835,1112,251,273]" pageId="21" pageNumber="131">
<treatmentCitationGroup id="AB10F373FF94074D5424FF7F7D3AFEB2" box="[835,1112,251,273]" pageId="21" pageNumber="131">
<treatmentCitation id="0AA1F24CFF94074D5424FF7F7D36FEB2" author="Kubo I" box="[835,1108,251,273]" page="361" pageId="21" pageNumber="131" year="1936">
<taxonomicName id="4C00AFDEFF94074D5424FF7F7D36FEB2" ID-CoL="7PRYM" authority="Kubo, 1936: 361" authorityName="Kubo" authorityPageNumber="361" authorityYear="1936" box="[835,1108,251,273]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D5424FF7F7AC1FEB2" box="[835,931,252,273]" italics="true" pageId="21" pageNumber="131">Paramaja</emphasis>
<bibRefCitation id="EF91A9ACFF94074D54CCFF787D36FEB2" author="Kubo I" box="[939,1108,251,273]" pageId="21" pageNumber="131" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936: 361</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BBFD45DFF94074D5424FE9B7D15FEE9" blockId="21.[835,1475,251,389]" pageId="21" pageNumber="131">
<treatmentCitationGroup id="AB10F373FF94074D5424FE9B7D15FEE9" pageId="21" pageNumber="131">
<treatmentCitation id="0AA1F24CFF94074D5424FE9B7D1EFE8D" author="Balss H" box="[835,1148,280,302]" page="1628" pageId="21" pageNumber="131" year="1957">
<taxonomicName id="4C00AFDEFF94074D5424FE9B7D1EFE8D" ID-CoL="d95d1411-1736-4864-a903-965a7d6ff2ab" authority="Balss, 1957: 1628" authorityName="Balss, 1957:" authorityPageNumber="1628" authorityYear="1628" box="[835,1148,280,302]" class="Malacostraca" family="Majidae" genus="Paramajella" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D5424FE9B7ADFFE8E" box="[835,957,280,301]" italics="true" pageId="21" pageNumber="131">Paramajella</emphasis>
<bibRefCitation id="EF91A9ACFF94074D54A1FE9B7D1EFE8D" author="Balss H" box="[966,1148,280,302]" pageId="21" pageNumber="131" pagination="1505 - 1672" refId="ref83135" refString="Balss H (1957) Decapoda. VIII. Systematik. In: Dr. H. G. Bronns, Klassen und Ordnungen des Tierreichs. Leipzig and Heidelberg, Winter. Funfter Band, I. Abteilung, 7. Buch, 12. Lief.: 1505 - 1672, figs. 1131 - 1199." type="book chapter" year="1957">Balss, 1957: 1628</bibRefCitation>
</taxonomicName>
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[unnecessary replacement name for
<taxonomicName id="4C00AFDEFF94074D54EAFEB57D0FFEE9" ID-CoL="7PRYM" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[909,1133,308,331]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D54EAFEB57A8FFEE8" box="[909,1005,310,331]" italics="true" pageId="21" pageNumber="131">Paramaja</emphasis>
<bibRefCitation id="EF91A9ACFF94074D5492FEB67D0FFEE9" author="Kubo I" box="[1013,1133,308,331]" pageId="21" pageNumber="131" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
</taxonomicName>
].
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<paragraph id="8BBFD45DFF94074D5424FED07D16FE26" blockId="21.[835,1475,251,389]" pageId="21" pageNumber="131">
<treatmentCitationGroup id="AB10F373FF94074D5424FED07D16FE26" pageId="21" pageNumber="131">
<taxonomicName id="4C00AFDEFF94074D5424FED07A14FECB" ID-CoL="7PDPL" authorityName="Lamarck" authorityYear="1801" box="[835,886,339,360]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D5424FED07A14FECB" box="[835,886,339,360]" italics="true" pageId="21" pageNumber="131">Maja</emphasis>
</taxonomicName>
– T.
<treatmentCitation id="0AA1F24CFF94074D54D4FED17D3FFECB" author="Sakai T" box="[947,1117,338,360]" page="296" pageId="21" pageNumber="131" year="1938">
<bibRefCitation id="EF91A9ACFF94074D54D4FED17D3FFECB" author="Sakai T" box="[947,1117,338,360]" pageId="21" pageNumber="131" pagination="193 - 364" refId="ref88279" refString="Sakai T (1938) Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, table I." type="book chapter" year="1938">Sakai, 1938: 296</bibRefCitation>
</treatmentCitation>
(part). – T.
<treatmentCitation id="0AA1F24CFF94074D53B8FED17C1FFEC4" author="Sakai T" box="[1247,1405,338,360]" page="83" pageId="21" pageNumber="131" year="1965">
<bibRefCitation id="EF91A9ACFF94074D53B8FED17C1FFEC4" author="Sakai T" box="[1247,1405,338,360]" pageId="21" pageNumber="131" pagination="1 - 92" refId="ref88322" refString="Sakai T (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo. i - xvi + 1 - 206 pp. [English section], figs. 1 - 27, pls. 1 - 100, 1 - 92 [in Japanese] + 1 - 26 [Bibliography and Index in English] + 27 - 32 [Index in Japanese] pp., 1 map." type="book chapter" year="1965">Sakai, 1965: 83</bibRefCitation>
</treatmentCitation>
(part). (not
<taxonomicName id="4C00AFDEFF94074D54FFFEEC7D0AFE26" ID-CoL="7PDPL" authority="Lamarck, 1801" authorityName="Lamarck" authorityYear="1801" box="[920,1128,367,389]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF94074D54FFFEEC7AA9FE27" box="[920,971,367,388]" italics="true" pageId="21" pageNumber="131">Maja</emphasis>
<bibRefCitation id="EF91A9ACFF94074D54B5FEEC7D0AFE26" author="Lamarck JBPA de" box="[978,1128,367,389]" pageId="21" pageNumber="131" pagination="1 - 432" refId="ref85551" refString="Lamarck JBPA de (1801) Systeme des animaux vertebres, ou Tableau general des classes, des ordres et des genres de ces animaux; Presentant leurs caracteres essentiels et leur distribution, d'apres la consideration de leurs rapports naturels et leur organisation, et suivant l'arrangement etablis dans les galeries du Museum d'Histoire Naturelle, parmi leur depouilles conservees; Precede du discours d'ouverture du cours de zoologie, donne dans le Museum National d'Histoire Naturelle l'an 8 de la Republique. Deterville, Paris. Vol. 6: i - viii + 1 - 432." type="journal article" year="1801">Lamarck, 1801</bibRefCitation>
</taxonomicName>
).
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</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FF94074E5424FE2E7BFCFED6" lastPageId="22" lastPageNumber="132" pageId="21" pageNumber="131" type="diagnosis">
<paragraph id="8BBFD45DFF94074E5424FE2E7BFCFED6" blockId="21.[834,1477,429,2117]" lastBlockId="22.[113,754,189,373]" lastPageId="22" lastPageNumber="132" pageId="21" pageNumber="131">
<emphasis id="B974084FFF94074D5424FE2E7ADDFE66" bold="true" box="[835,959,429,453]" pageId="21" pageNumber="131">Diagnosis.</emphasis>
<materialsCitation id="3B68DE00FF94074E54ADFE2E7BF8FED6" collectionCode="V" county="Pseudorostral" lastPageId="22" lastPageNumber="132" location="Carapace" longitude="12.55" municipality="Intestinal region" pageId="21" pageNumber="131" specimenCount="1">
<location id="8EDF8286FF94074D54ADFE2E7D51FE66" LSID="urn:lsid:plazi:treatment:03A9654BFF94074E5361FF3E7B41F8D6:8EDF8286FF94074D54ADFE2E7D51FE66" box="[970,1075,429,453]" county="Pseudorostral" longitude="12.55" municipality="Intestinal region" name="Carapace" pageId="21" pageNumber="131">Carapace</location>
rounded in adults; dorsal surface evenly inflated, covered by granules or tubercles; gastric and branchial regions clearly delimited by grooves (
<figureCitation id="133BC8D8FF94074D52EBFE6E7A1EFD86" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" pageId="21" pageNumber="131">Figs. 8–12</figureCitation>
).
<collectingMunicipality id="6BDB4E27FF94074D54E8FD8E7D5FFD86" box="[911,1085,525,549]" pageId="21" pageNumber="131">Intestinal region</collectingMunicipality>
without distinct median spine (
<figureCitation id="133BC8D8FF94074D52EAFD8E7A1DFDE6" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionStartId-5="31.[135,170,1940,1962]" captionStartId-6="32.[113,148,1835,1857]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetBox-5="[135,1473,254,1923]" captionTargetBox-6="[113,1451,330,1819]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetId-5="figure-7@31.[135,1473,254,1923]" captionTargetId-6="figure-15@32.[113,1451,330,1820]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionTargetPageId-5="31" captionTargetPageId-6="32" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." captionText-5="Fig. 13. Frontal views of carapaces, Paramaja species. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; C, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; D, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." captionText-6="Fig. 14. Lateral views of carapaces, Paramaja species. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, female (62.0 × 58.0 mm) (SMF 47810), Japan; C, P. kominatoensis, female (63.5 × 58.5 mm) (SMF 47728), Japan; D, P. kominatoensis, male (50.5 × 44.6 mm) (SMF 47738), Japan; E, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan [laterally transposed to match other figures]; F, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; G, P. gibba, female (67.7 × 63.2 mm) (SIO), Indian Ocean; H, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" figureDoi-5="http://doi.org/10.5281/zenodo.5384622" figureDoi-6="http://doi.org/10.5281/zenodo.5384624" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" httpUri-5="https://zenodo.org/record/5384622/files/figure.png" httpUri-6="https://zenodo.org/record/5384624/files/figure.png" pageId="21" pageNumber="131">Figs. 8–14</figureCitation>
).
<collectingCounty id="62DEACD1FF94074D54F0FDAE7D52FDE6" box="[919,1072,557,581]" pageId="21" pageNumber="131">Pseudorostral</collectingCounty>
spines relatively short, diverging, forming a
<collectionCode id="ED114C98FF94074D54DEFDCE7AAEFDC7" box="[953,972,589,612]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="21" pageNumber="131" type="Museum">V</collectionCode>
(
<figureCitation id="133BC8D8FF94074D54B9FDCE7D34FDC6" box="[990,1110,589,613]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionStartId-5="31.[135,170,1940,1962]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetBox-5="[135,1473,254,1923]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetId-5="figure-7@31.[135,1473,254,1923]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionTargetPageId-5="31" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." captionText-5="Fig. 13. Frontal views of carapaces, Paramaja species. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; C, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; D, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" figureDoi-5="http://doi.org/10.5281/zenodo.5384622" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" httpUri-5="https://zenodo.org/record/5384622/files/figure.png" pageId="21" pageNumber="131">Figs. 8–13</figureCitation>
). Supraorbital eave with anterior part longitudinally narrow, rectangular, not prominently expanded; antorbital spine sharp (
<figureCitation id="133BC8D8FF94074D53CAFD0E7C51FD06" box="[1197,1331,653,677]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="21" pageNumber="131">Fig. 36G–K</figureCitation>
). Intercalated spine distinct, separated from supraorbital eave and postorbital spine by distinct gaps; postorbital spine strong, lobiform; hepatic region with 1 strong spine, much shorter than postorbital spine; 1 or more smaller spines below (
<figureCitation id="133BC8D8FF94074D52FEFC8E7AFAFCE6" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="21" pageNumber="131">Fig. 36G–K</figureCitation>
). Lateral carapace margin with 3 or 4 spines and numerous tubercles, branchial region with larger tubercle, not discernible from other surrounding tubercles in large specimens (
<figureCitation id="133BC8D8FF94074D54A2FC0E7D5FFC06" box="[965,1085,909,933]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionStartId-5="31.[135,170,1940,1962]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetBox-5="[135,1473,254,1923]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetId-5="figure-7@31.[135,1473,254,1923]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionTargetPageId-5="31" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." captionText-5="Fig. 13. Frontal views of carapaces, Paramaja species. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; C, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; D, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" figureDoi-5="http://doi.org/10.5281/zenodo.5384622" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" httpUri-5="https://zenodo.org/record/5384622/files/figure.png" pageId="21" pageNumber="131">Figs. 8–13</figureCitation>
). Posterior carapace margin with 2 short median spines, not visible from dorsal view when adult due to swollen carapace (
<figureCitation id="133BC8D8FF94074D5333FC4E7DA8FC46" box="[1108,1226,973,997]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" pageId="21" pageNumber="131">Figs. 8–12</figureCitation>
). Eyes relatively short, slender, with large ovoid cornea (
<figureCitation id="133BC8D8FF94074D53A6FC6E7C29FBA6" box="[1217,1355,1005,1029]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="21" pageNumber="131">Fig. 36G–K</figureCitation>
). Antennal flagellum short, slender. Basal antennal article longer than broad, rectangular; surface with several tubercles, with 2 blunt spines distally; inner and outer lateral margins may have low granules or lobes, or entire; proximal outer angle rounded; antero-external crested rim of antennular fossa touches but does not significantly overlap distal part of basal antennal article (
<figureCitation id="133BC8D8FF94074D54F7FB4E7D72FB46" box="[912,1040,1229,1253]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="21" pageNumber="131">Fig. 39G–K</figureCitation>
). Epistome much wider than long, anterior margin unarmed; posterior margin composed of 4 rectangular plates separated by shallow fissures (
<figureCitation id="133BC8D8FF94074D538DFA8E7C1DFA86" box="[1258,1407,1293,1317]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="21" pageNumber="131">Figs. 39G–K</figureCitation>
,
<figureCitation id="133BC8D8FF94074D52E9FA8E7A34FAE6" captionStart="Fig" captionStartId="77.[136,171,1977,1999]" captionTargetBox="[136,1475,673,1962]" captionTargetId="figure-8@77.[135,1475,673,1962]" captionTargetPageId="77" captionText="Fig. 41. Epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; G, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; H, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384680" httpUri="https://zenodo.org/record/5384680/files/figure.png" pageId="21" pageNumber="131">41E, F</figureCitation>
). Suborbital margin separated from basal antennal article by short fissure, confluent with margin of postorbital tooth (
<figureCitation id="133BC8D8FF94074D542CFAEE7AB5FA26" box="[843,983,1389,1413]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="21" pageNumber="131">Fig. 39G–K</figureCitation>
). Outer surface of third maxilliped covered by short setae in adults; ischium subrectangular, distinctly longer than broad; postero-external angle of merus relatively broad, “inserted” into shallower concavity on outer margin of ischium; antero-internal part of ischium rounded, auriculiform (
<figureCitation id="133BC8D8FF94074D542DF98E7AA2F986" box="[842,960,1549,1573]" captionStart="Fig" captionStartId="79.[136,171,1833,1855]" captionTargetBox="[215,1395,190,1817]" captionTargetId="figure-7@79.[215,1395,190,1817]" captionTargetPageId="79" captionText="Fig. 43. Third maxillipeds. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja squinado, female (144.5 × 129.0 mm) (ZRC 2013.1126), Spain; C, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; D, Maja brachydactyla, male (161.2 × 140.1 mm), (ZRC 2008.0179), Spain; E, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; F, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; G, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; H,Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; I, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; J, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K, Paramaja kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; L, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; M, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; N, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; O, Paramaja turgida n. sp., paratype male (71.4 × 64.4 mm) (NSMT-Cr 22329), Philippines; P, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384684" httpUri="https://zenodo.org/record/5384684/files/figure.png" pageId="21" pageNumber="131">Fig. 43I–P</figureCitation>
). Male chelipeds relatively short in adult males, surfaces of merus and carpus almost smooth or with very small granules; carpus short, with low, uneven longitudinal ridge; propodus of palm elongated, curved, smooth, without lateral cristae in young, short, inflated in adults, palm longer than fingers; fingers long, slender, gently curved, with distinct basal gape when closed (
<figureCitation id="133BC8D8FF94074D533AF94E7DB4F946" box="[1117,1238,1741,1765]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" pageId="21" pageNumber="131">Figs. 8–12</figureCitation>
,
<figureCitation id="133BC8D8FF94074D5383F94E7C49F946" box="[1252,1323,1741,1765]" captionStart="Fig" captionStartId="89.[135,170,1806,1828]" captionTargetBox="[136,1474,216,1791]" captionTargetId="figure-8@89.[136,1474,215,1791]" captionTargetPageId="89" captionText="Fig. 53. Chelae. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm), (ZRC 2008.0179), Spain; D, Maja cornuta, male (88.8 × 86.8 mm) (NHM 1928.12.1.177–) (lectotype of Mamaia queketti Stebbing, 1908), South Africa; E, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; F, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; G, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; H, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; I, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; J, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; L, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; M, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; N, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; O, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; P, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384708" httpUri="https://zenodo.org/record/5384708/files/figure.png" pageId="21" pageNumber="131">53I–L</figureCitation>
). Ambulatory legs relatively long, slender; merus without dorsal subdistal spine; dactylus elongate, curved, covered with long setae when young except for corneous tip, almost completely smooth in adults (
<figureCitation id="133BC8D8FF94074D5365F8CE7D02F8C6" box="[1026,1120,1869,1893]" captionStart="Fig" captionStartId="19.[136,171,1824,1846]" captionTargetBox="[136,1474,338,1808]" captionTargetId="figure-8@19.[136,1474,337,1809]" captionTargetPageId="19" captionText="Fig. 6. General habitus and right first ambulatory legs. A, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; B, Maja crispata, male (48.5 × 39.0 mm) (MNHN-IU-2013-4041), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; E, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; F, Neomaja goltziana, male (74.7 × 66.7 mm) (MNHN-IU-2013-4043), Senegal; G, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; H, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384602" httpUri="https://zenodo.org/record/5384602/files/figure.png" pageId="21" pageNumber="131">Figs. 6H</figureCitation>
,
<figureCitation id="133BC8D8FF94074D530BF8CE7DC1F8C6" box="[1132,1187,1869,1893]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="26.[113,148,1947,1969]" captionStartId-1="27.[135,170,2000,2022]" captionStartId-2="28.[113,148,1916,1938]" captionStartId-3="29.[135,170,1907,1929]" captionStartId-4="30.[113,148,1916,1938]" captionTargetBox-0="[113,1451,274,1931]" captionTargetBox-1="[135,1473,220,1985]" captionTargetBox-2="[113,1451,304,1901]" captionTargetBox-3="[135,1473,285,1891]" captionTargetBox-4="[113,1451,304,1900]" captionTargetId-0="figure-15@26.[113,1451,274,1931]" captionTargetId-1="figure-8@27.[132,1476,220,1985]" captionTargetId-2="figure-15@28.[113,1451,304,1901]" captionTargetId-3="figure-7@29.[135,1473,285,1891]" captionTargetId-4="figure-15@30.[113,1451,304,1900]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionText-0="Fig. 8. General habitus, Paramaja kominatoensis. A, holotype male (63.0 × 57.0 mm) (after Kubo, 1936: fig. 2); B, neotype male (56.6 × 40.4 mm) (SMF 47808), Japan; C, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; D, male (76.9 × 69.6 mm) (SMF 47738), Japan; E, female (52.7 × 47.5 mm) (SMF 47736), Japan; F, female (62.0 × 58.0 mm) (SMF 47810), Japan." captionText-1="Fig. 9. General habitus, Paramaja kominatoensis. A, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; B, dried female (61.0 × 54.6 mm) (KPM NH124171), Japan; C, male (73.0 × 66.0 mm) (PCM), Taiwan; D, male (53.1 × 46.8 mm) (SMF 47728), Japan; E, male (50.5 × 44.6 mm) (SMF 47738), Japan." captionText-2="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." captionText-3="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." captionText-4="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." figureDoi-0="http://doi.org/10.5281/zenodo.5384610" figureDoi-1="http://doi.org/10.5281/zenodo.5384612" figureDoi-2="http://doi.org/10.5281/zenodo.5384614" figureDoi-3="http://doi.org/10.5281/zenodo.5384616" figureDoi-4="http://doi.org/10.5281/zenodo.5384620" httpUri-0="https://zenodo.org/record/5384610/files/figure.png" httpUri-1="https://zenodo.org/record/5384612/files/figure.png" httpUri-2="https://zenodo.org/record/5384614/files/figure.png" httpUri-3="https://zenodo.org/record/5384616/files/figure.png" httpUri-4="https://zenodo.org/record/5384620/files/figure.png" pageId="21" pageNumber="131">
8–
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</figureCitation>
,
<figureCitation id="133BC8D8FF94074D53C9F8CE7D97F8C6" box="[1198,1269,1869,1893]" captionStart="Fig" captionStartId="91.[135,171,1862,1884]" captionTargetBox="[170,1440,189,1847]" captionTargetId="figure-8@91.[170,1440,189,1847]" captionTargetPageId="91" captionText="Fig. 55. First ambulatory dactyli and propodi. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; G, Paramaja kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; H, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; I, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; J, Paramaja turgida n. sp., paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; K, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; L, Alcomaja irrorata, male (43.0 × 37.2 mm) (ZRC 2013.1223), Philippines; M, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; N, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384712" httpUri="https://zenodo.org/record/5384712/files/figure.png" pageId="21" pageNumber="131">55E–J</figureCitation>
). Thoracic sternum wide; surfaces of somites 5–8 with numerous prominent rounded tubercles and granules; sternites 3 and 4 slightly depressed; margin between sternites 2 and 3 demarcated by deep notch; anterior margin of sterno-abdominal cavity not forming complete rim (
<figureCitation id="133BC8D8FF94074D5317F86E7DAEF7A6" box="[1136,1228,2029,2053]" captionStart="Fig" captionStartId="84.[113,148,1743,1765]" captionTargetBox="[112,1451,397,1728]" captionTargetId="figure-140@84.[112,1451,396,1731]" captionTargetPageId="84" captionText="Fig. 48. Male anterior thoracic sternums and abdomens, Paramaja. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, male (76.9 × 69.6 mm) (SMF 47738), Japan; C, P. kominatoensis, male (50.5 × 44.6 mm) (SMF 47738), Japan; D, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; E, P. kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; F, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; G, P. gibba, male male (64.7 × 60.7 mm) (SIO), Indian Ocean; H, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; I, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384698" httpUri="https://zenodo.org/record/5384698/files/figure.png" pageId="21" pageNumber="131">Figs. 48</figureCitation>
,
<figureCitation id="133BC8D8FF94074D53BEF86E7C6EF7A6" box="[1241,1292,2029,2053]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="21" pageNumber="131">52D</figureCitation>
). Male abdomen subrectangular, with 6 free somites and telson; somites 3–6 and telson subequal in width (
<figureCitation id="133BC8D8FF94074D53F2F7AE7D8AF7E6" box="[1173,1256,2093,2117]" captionStart="Fig" captionStartId="84.[113,148,1743,1765]" captionTargetBox="[112,1451,397,1728]" captionTargetId="figure-140@84.[112,1451,396,1731]" captionTargetPageId="84" captionText="Fig. 48. Male anterior thoracic sternums and abdomens, Paramaja. A, P. kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; B, P. kominatoensis, male (76.9 × 69.6 mm) (SMF 47738), Japan; C, P. kominatoensis, male (50.5 × 44.6 mm) (SMF 47738), Japan; D, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; E, P. kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; F, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; G, P. gibba, male male (64.7 × 60.7 mm) (SIO), Indian Ocean; H, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; I, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384698" httpUri="https://zenodo.org/record/5384698/files/figure.png" pageId="21" pageNumber="131">Fig. 48</figureCitation>
). Male press-button abdominal locking mechanism submedian in position on sterno-abdominal cavity (
<figureCitation id="133BC8D8FF97074E56C6FF5E7B73FF56" box="[417,529,221,245]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="22" pageNumber="132">Fig. 52D</figureCitation>
). Female abdomen dome-shaped, covering most of thoracic sternum. G1 very long, slender, almost straight for most of length, distal part gently curved, subconical with rounded tip, with small, low subdistal dorsal flap, with scattered setae (
<figureCitation id="133BC8D8FF97074E5527FEDE7BF3FED6" box="[576,657,349,373]" captionStart="Fig" captionStartId="33.[135,170,1988,2010]" captionTargetBox="[138,1474,205,1972]" captionTargetId="figure-8@33.[133,1476,201,1975]" captionTargetPageId="33" captionText="Fig. 15. Left G1s, Paramaja species. A, P. kominatoensis (after Kubo, 1936: fig. 1B); B-E, P. kominatoensis, male (76.9 × 69.6 mm) (SMF), Japan; F-J, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K-M, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; N-R, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; S-U, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines. Scales: B, C, F, G, K, N, O, S = 5.0 mm; D, E, H-J, L, M, P-R, T, U = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384626" httpUri="https://zenodo.org/record/5384626/files/figure.png" pageId="22" pageNumber="132">Fig. 15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FF97074E5716FE1E7BDBFE76" pageId="22" pageNumber="132" type="type_taxon">
<paragraph id="8BBFD45DFF97074E5716FE1E7BDBFE76" blockId="22.[113,752,413,470]" pageId="22" pageNumber="132">
<emphasis id="B974084FFF97074E5716FE1E7872FE16" bold="true" box="[113,272,413,437]" pageId="22" pageNumber="132">
<typeStatus id="54BB6AFFFF97074E5716FE1E79CEFE16" box="[113,172,413,437]" pageId="22" pageNumber="132">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C00AFDEFF97074E567CFE1D7BA4FE16" ID-CoL="4D294" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[283,710,413,437]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="species" species="kominatoensis">
<emphasis id="B974084FFF97074E567CFE1D7B50FE16" box="[283,562,413,437]" italics="true" pageId="22" pageNumber="132">Paramaja kominatoensis</emphasis>
<bibRefCitation id="EF91A9ACFF97074E555AFE1E7BA4FE16" author="Kubo I" box="[573,710,413,437]" pageId="22" pageNumber="132" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
</taxonomicName>
, by original designation and monotypy. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FF97074E5714FE7E7B41F8D6" pageId="22" pageNumber="132" type="discussion">
<paragraph id="8BBFD45DFF97074E5714FE7E7B55FCD6" blockId="22.[112,752,509,885]" pageId="22" pageNumber="132">
<emphasis id="B974084FFF97074E5714FE7E7980FDB6" bold="true" box="[115,226,509,533]" pageId="22" pageNumber="132">Remarks.</emphasis>
<bibRefCitation id="EF91A9ACFF97074E578DFE7E78D3FDB6" author="Kubo I" box="[234,433,509,533]" pageId="22" pageNumber="132" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo (1936: 361)</bibRefCitation>
established a new genus for a new species from Kominato,
<collectingCountry id="F31794CDFF97074E56B4FD9D7B71FD96" box="[467,531,542,565]" name="Japan" pageId="22" pageNumber="132">Japan</collectingCountry>
, with the following comments: “The present genus stands somewhat intermediate between the two genera
<taxonomicName id="4C00AFDEFF97074E560AFDDD78C7FDD6" authorityName="Lamarck" authorityYear="1801" box="[365,421,606,629]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E560AFDDD78C7FDD6" box="[365,421,606,629]" italics="true" pageId="22" pageNumber="132">Maja</emphasis>
</taxonomicName>
and
<emphasis id="B974084FFF97074E56B1FDDD7B5AFDD6" box="[470,568,606,629]" italics="true" pageId="22" pageNumber="132">Majopsis</emphasis>
(sic for
<taxonomicName id="4C00AFDEFF97074E55EAFDDD799EFD36" authority="Faxon, 1893" authorityName="Faxon" authorityYear="1893" class="Malacostraca" family="Majidae" genus="Maiopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E55EAFDDD7B8DFDD6" box="[653,751,606,629]" italics="true" pageId="22" pageNumber="132">Maiopsis</emphasis>
Faxon, 1893
</taxonomicName>
), and it is easily distinguished from the genus
<taxonomicName id="4C00AFDEFF97074E5714FD1D79CCFD16" authorityName="Lamarck" authorityYear="1801" box="[115,174,670,693]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5714FD1D79CCFD16" box="[115,174,670,693]" italics="true" pageId="22" pageNumber="132">Maja</emphasis>
</taxonomicName>
in lacking any spines on carapace, and from the genus
<emphasis id="B974084FFF97074E57DBFD3D7843FD76" box="[188,289,702,725]" italics="true" pageId="22" pageNumber="132">Majopsis</emphasis>
in having no accessory spine on rostral spines, but two spines on distal margin of basal segment of outer antennae, and fingers of chela not canaliculate within.” Although none of these characters work at present to distinguish
<taxonomicName id="4C00AFDEFF97074E5669FCBD781BFCF6" authorityName="Kubo" authorityYear="1936" box="[270,377,830,853]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5669FCBD781BFCF6" box="[270,377,830,853]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
from
<taxonomicName id="4C00AFDEFF97074E56D9FCBD7894FCF6" authorityName="Lamarck" authorityYear="1801" box="[446,502,830,853]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B974084FFF97074E56D9FCBD7894FCF6" box="[446,502,830,853]" italics="true" pageId="22" pageNumber="132">Maja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FF97074E569BFCBD7B5AFCF6" box="[508,568,830,853]" pageId="22" pageNumber="132" sensu="stricto">s. str.</taxonomicNameLabel>
or allied genera, it can still be separated by other features.
</paragraph>
<paragraph id="8BBFD45DFF97074E5716FC1D784FFBD6" blockId="22.[112,754,925,1141]" pageId="22" pageNumber="132">
<taxonomicName id="4C00AFDEFF97074E5716FC1D7804FC16" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[113,358,925,949]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5716FC1D79BEFC16" box="[113,220,926,949]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
<bibRefCitation id="EF91A9ACFF97074E5783FC1E7804FC16" author="Kubo I" box="[228,358,925,949]" pageId="22" pageNumber="132" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
</taxonomicName>
, is here resurrected to accommodate Indo-Pacific species which have a uniformly inflated carapace which has the dorsal surface covered with granules and tubercles:
<taxonomicName id="4C00AFDEFF97074E5787FC7D7884FBB6" authority="(Alcock, 1895)" baseAuthorityName="Alcock" baseAuthorityYear="1895" box="[224,486,1021,1045]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="B974084FFF97074E5787FC7D7859FBB6" box="[224,315,1021,1045]" italics="true" pageId="22" pageNumber="132">P. gibba</emphasis>
(
<bibRefCitation id="EF91A9ACFF97074E562EFC7E78BCFBB6" author="Alcock A" box="[329,478,1021,1045]" pageId="22" pageNumber="132" pagination="157 - 291" refId="ref82409" refString="Alcock A (1895) Materials for a carcinological fauna of India, 1. The Brachyura Oxyrhyncha. Journal of the Asiatic Society of Bengal, Calcutta, 64: 157 - 291, pls. 3 - 5." type="journal article" year="1895">Alcock, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C00AFDEFF97074E5696FC7D79C8FB96" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="species" species="kominatoensis">
<emphasis id="B974084FFF97074E5696FC7D7BCAFBB6" box="[497,680,1021,1045]" italics="true" pageId="22" pageNumber="132">P. kominatoensis</emphasis>
<bibRefCitation id="EF91A9ACFF97074E55C9FC7E79C8FB96" author="Kubo I" pageId="22" pageNumber="132" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C00AFDEFF97074E5781FB9D7836FB96" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[230,340,1053,1077]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="species" species="turgida" status="sp. nov.">
<emphasis id="B974084FFF97074E5781FB9D7836FB96" box="[230,340,1053,1077]" italics="true" pageId="22" pageNumber="132">P. turgida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FF97074E563CFB9D78F4FB96" box="[347,406,1054,1077]" pageId="22" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
The genus is also characterised by features of its suborbital margin, epistome, male abdomen and G1 structure.
</paragraph>
<paragraph id="8BBFD45DFF97074E5714FB1D7B41F8D6" blockId="22.[112,754,1181,1909]" pageId="22" pageNumber="132">
<taxonomicName id="4C00AFDEFF97074E5714FB1D79BCFB16" authorityName="Kubo" authorityYear="1936" box="[115,222,1182,1205]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5714FB1D79BCFB16" box="[115,222,1182,1205]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="4C00AFDEFF97074E5638FB1E78A7FB16" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[351,453,1181,1205]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B974084FFF97074E5638FB1E78A7FB16" box="[351,453,1181,1205]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FF97074E56ABFB1D7B77FB15" box="[460,533,1182,1206]" pageId="22" pageNumber="132" rank="genus">n. gen.</taxonomicNameLabel>
, with which it shares a suite of carapace, antennal, epistomal, third maxilliped, pereiopodal, thoracic sternal and male abdominal characters. The single most important character separating
<taxonomicName id="4C00AFDEFF97074E55E2FB7D7B92FAB6" authorityName="Kubo" authorityYear="1936" box="[645,752,1278,1301]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E55E2FB7D7B92FAB6" box="[645,752,1278,1301]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
from
<taxonomicName id="4C00AFDEFF97074E57CAFA9E7871FA96" authorityName="Ng &amp; Forges" authorityYear="2015" box="[173,275,1309,1333]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E57CAFA9E7871FA96" box="[173,275,1309,1333]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
</taxonomicName>
is the form of the G1. In
<taxonomicName id="4C00AFDEFF97074E5548FA9D7BFAFA96" authorityName="Kubo" authorityYear="1936" box="[559,664,1310,1333]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5548FA9D7BFAFA96" box="[559,664,1310,1333]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
, the G1 is very long and mostly straight, the distal part being conical (
<figureCitation id="133BC8D8FF97074E571EFADE79A9FAD6" box="[121,203,1373,1397]" captionStart="Fig" captionStartId="33.[135,170,1988,2010]" captionTargetBox="[138,1474,205,1972]" captionTargetId="figure-8@33.[133,1476,201,1975]" captionTargetPageId="33" captionText="Fig. 15. Left G1s, Paramaja species. A, P. kominatoensis (after Kubo, 1936: fig. 1B); B-E, P. kominatoensis, male (76.9 × 69.6 mm) (SMF), Japan; F-J, P. kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K-M, P. gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; N-R, P. turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; S-U, P. turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines. Scales: B, C, F, G, K, N, O, S = 5.0 mm; D, E, H-J, L, M, P-R, T, U = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384626" httpUri="https://zenodo.org/record/5384626/files/figure.png" pageId="22" pageNumber="132">Fig. 15</figureCitation>
). In
<taxonomicName id="4C00AFDEFF97074E5663FADE7809FAD6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[260,363,1373,1397]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5663FADE7809FAD6" box="[260,363,1373,1397]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
</taxonomicName>
, the G1 is proportionately shorter, more curved, and part of the dorsal fold bends downwards to form a distinct ventral projection (
<figureCitation id="133BC8D8FF97074E5561FA1E7B03FA16" box="[518,609,1437,1461]" captionStart="Fig" captionStartId="42.[113,148,794,816]" captionTargetBox="[114,1451,189,779]" captionTargetId="figure-15@42.[104,1473,179,782]" captionTargetPageId="42" captionText="Fig. 19. Left G1s, Alcomaja species. A–C, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; D–F, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; G, A. confragosa (as Maja confragosa, after Griffin &amp; Tranter, 1986: fig. 73a). Scale bars = 1.0 mm [A–C]." figureDoi="http://doi.org/10.5281/zenodo.5384634" httpUri="https://zenodo.org/record/5384634/files/figure.png" pageId="22" pageNumber="132">Figs. 19</figureCitation>
,
<figureCitation id="133BC8D8FF97074E5509FA1E7BEEFA16" box="[622,652,1437,1461]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="22" pageNumber="132">20</figureCitation>
). Species of
<taxonomicName id="4C00AFDEFF97074E57F5FA3E799AFA76" authorityName="Ng &amp; Forges" authorityYear="2015" box="[146,248,1469,1493]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E57F5FA3E799AFA76" box="[146,248,1469,1493]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
</taxonomicName>
also have relatively longer and more slender pseudorostral spines (
<figureCitation id="133BC8D8FF97074E5638FA5E7883FA56" box="[351,481,1501,1525]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="22" pageNumber="132">Fig. 36L–O</figureCitation>
) than in
<taxonomicName id="4C00AFDEFF97074E552EFA5D7BD6FA56" authorityName="Kubo" authorityYear="1936" box="[585,692,1502,1525]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E552EFA5D7BD6FA56" box="[585,692,1502,1525]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
(
<figureCitation id="133BC8D8FF97074E55A2FA5E79A5F9B6" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="22" pageNumber="132">Fig. 36G–K</figureCitation>
). Adults of the two genera are easy to separate, with
<taxonomicName id="4C00AFDEFF97074E57CCF99D7874F996" authorityName="Kubo" authorityYear="1936" box="[171,278,1566,1589]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E57CCF99D7874F996" box="[171,278,1566,1589]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
growing to much larger sizes (over
<quantity id="4CF879B8FF97074E55CAF99E79F9F9F6" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="22" pageNumber="132" unit="mm" value="65.0" valueMax="70.0" valueMin="60.0">60–70 mm</quantity>
carapace length) while
<taxonomicName id="4C00AFDEFF97074E56CFF9BE7B04F9F6" box="[424,614,1597,1621]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B974084FFF97074E56CFF9BE7B6DF9F6" box="[424,527,1597,1621]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
species
</taxonomicName>
are smaller. In
<taxonomicName id="4C00AFDEFF97074E57F3F9DD7866F9D6" authorityName="Kubo" authorityYear="1936" box="[148,260,1630,1653]" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E57F3F9DD7866F9D6" box="[148,260,1630,1653]" italics="true" pageId="22" pageNumber="132">Paramaja</emphasis>
</taxonomicName>
specimens, even when small (
<quantity id="4CF879B8FF97074E5515F9DE7B8DF9D6" box="[626,751,1629,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="22" pageNumber="132" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 mm</quantity>
carapace length), the branchial and cardiac regions are proportionately more inflated and the lateral and branchial spines are relatively shorter (
<figureCitation id="133BC8D8FF97074E56DEF93E7B2FF976" box="[441,589,1725,1749]" captionStart="Fig" captionStartId="28.[113,148,1916,1938]" captionTargetBox="[113,1451,304,1901]" captionTargetId="figure-15@28.[113,1451,304,1901]" captionTargetPageId="28" captionText="Fig. 10. General habitus, Paramaja gibba. A, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; B, as Maja gibba (after Alcock &amp; Anderson, 1898: pl. 21 fig. 5); C, male (65.0 × 59.8 mm) (NSMT-Cr 16618), Andaman Sea; D, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal." figureDoi="http://doi.org/10.5281/zenodo.5384614" httpUri="https://zenodo.org/record/5384614/files/figure.png" pageId="22" pageNumber="132">Figs. 10A, B</figureCitation>
,
<figureCitation id="133BC8D8FF97074E553BF93E7BCFF976" box="[604,685,1725,1749]" captionStart="Fig" captionStartId="29.[135,170,1907,1929]" captionTargetBox="[135,1473,285,1891]" captionTargetId="figure-7@29.[135,1473,285,1891]" captionTargetPageId="29" captionText="Fig. 11. General habitus, Paramaja gibba. A, juvenile male (10.6 × 8.1 mm) (MNHN-IU-2010-54), Madagascar; B, male juvenile (16.4 × 12.5 mm) (MNHN-IU-2010-55), Madagascar; C, male (24.6 × 20.9 mm) (MNHN-IU-2010-61), Madagascar; D, male (58.6 × 52.2 mm) (MNHN-IU-2010-63), Madagascar; E, juvenile female (33.9 × 29.4 mm) (NMV J54316), Australia; F, 1 ovigerous female (45.1 × 40.6 mm) (MNHN-IU-2010-467), Madagascar; G, female (46.7 × 41.6 mm) (NMV J61055), Australia." figureDoi="http://doi.org/10.5281/zenodo.5384616" httpUri="https://zenodo.org/record/5384616/files/figure.png" pageId="22" pageNumber="132">11A–C</figureCitation>
,
<figureCitation id="133BC8D8FF97074E55DCF93E79E7F956" captionStart="Fig" captionStartId="30.[113,148,1916,1938]" captionTargetBox="[113,1451,304,1900]" captionTargetId="figure-15@30.[113,1451,304,1900]" captionTargetPageId="30" captionText="Fig. 12. General habitus, Paramaja turgida n. sp. A, paratype male (26.2 × 21.3 mm) (ZRC 2001.0616b); B, paratype female (33.5 × 29.0 mm) (ZRC 2013.1230); C, paratype male (67.9 × 60.5 mm) (MNHN), Philippines; D, paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; E, holotype male (74.1 × 66.8 mm) (NMCR), Philippines; F, male (25.1 × 19.6 mm) (MNHN-IU-2013-4051), Solomon Islands." figureDoi="http://doi.org/10.5281/zenodo.5384620" httpUri="https://zenodo.org/record/5384620/files/figure.png" pageId="22" pageNumber="132">12A, B</figureCitation>
). Specimens of
<taxonomicName id="4C00AFDEFF97074E5655F95E78FAF956" authorityName="Ng &amp; Forges" authorityYear="2015" box="[306,408,1757,1781]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFF97074E5655F95E78FAF956" box="[306,408,1757,1781]" italics="true" pageId="22" pageNumber="132">Alcomaja</emphasis>
</taxonomicName>
of comparable size on the other hand have less inflated branchial and cardiac regions, and the lateral and branchial spines are stronger and longer (
<figureCitation id="133BC8D8FF97074E571EF8BE79B7F8F6" box="[121,213,1853,1877]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="22" pageNumber="132">Figs. 16</figureCitation>
,
<figureCitation id="133BC8D8FF97074E5785F8BE799DF8F6" box="[226,255,1853,1877]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="22" pageNumber="132">17</figureCitation>
). That said, juvenile female specimens of the two species can be difficult to separate.
</paragraph>
</subSubSection>
</treatment>
</document>
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