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<document id="FFB71FF662D2D70ADD2264DCE7ED6F4B" ID="10.11646/zootaxa.4175.6.5" ID-DOI="10.11646/zootaxa.4175.6.5" ID-GBIF-Dataset="e79fd18a-fa17-4b4b-8490-1e50489bb71b" ID-ISSN="1175-5326" ID-Zenodo-Dep="262702" ID-ZooBank="DD402425-F055-4CC9-B7BF-4710F4CA96CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476866464250" checkinUser="plazi" docAuthor="Iwan, Dariusz" docDate="2016" docId="03A72351FFBFBB5EB5E3507E0EAA971F" docLanguage="en" docName="zootaxa.4175.6.5.pdf" docOrigin="Zootaxa 4175 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrocrates matthewsi Iwan, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="584" masterDocId="FF9E5B29FFB6BB53B57456670D169309" masterDocTitle="Species groups in the genus Atrocrates Koch, 1956 (Coleoptera: Tenebrionidae: Pedinini)" masterLastPageNumber="587" masterPageNumber="571" pageNumber="580" updateTime="1698390191471" updateUser="plazi">
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<mods:title id="6AFAF180350D592A0CA99B0C8CF206D6">Species groups in the genus Atrocrates Koch, 1956 (Coleoptera: Tenebrionidae: Pedinini)</mods:title>
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<mods:name id="66E299D7FE059BB5DC48A86146453AFB" type="personal">
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<mods:namePart id="47BC2EC4239C5F0262DE4281F4E5717C">Iwan, Dariusz</mods:namePart>
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<mods:title id="C7EDF193B5E0221FB6A88CB3A86AD0D2">Zootaxa</mods:title>
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<mods:part id="837233576383BB9EE32E3E8F3A84B3BA">
<mods:date id="0878FBA13B951A41DA5F8575BA7E845D">2016</mods:date>
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<mods:number id="51B409C2DB36BB4C1A3A08CF0550BD44">4175</mods:number>
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<treatment id="03A72351FFBFBB5EB5E3507E0EAA971F" ID-DOI="http://doi.org/10.5281/zenodo.6087556" ID-GBIF-Taxon="124996585" ID-Zenodo-Dep="6087556" LSID="urn:lsid:plazi:treatment:03A72351FFBFBB5EB5E3507E0EAA971F" httpUri="http://treatment.plazi.org/id/03A72351FFBFBB5EB5E3507E0EAA971F" lastPageId="13" lastPageNumber="584" pageId="9" pageNumber="580">
<subSubSection id="C314C1CCFFBFBB5AB5E3507E0CB2955F" pageId="9" pageNumber="580" type="nomenclature">
<paragraph id="8BB19247FFBFBB5AB5E3507E0CFD953B" blockId="9.[151,491,1561,1622]" box="[151,491,1561,1586]" pageId="9" pageNumber="580">
<heading id="D0F9252BFFBFBB5AB5E3507E0CFD953B" bold="true" box="[151,491,1561,1586]" fontSize="10" level="2" pageId="9" pageNumber="580" reason="2">
<emphasis id="B97A4E55FFBFBB5AB5E3507E0CFD953B" bold="true" box="[151,491,1561,1586]" pageId="9" pageNumber="580">
<taxonomicName id="4C0EE9C4FFBFBB5AB5E3507E0C9C953B" box="[151,394,1561,1586]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
<emphasis id="B97A4E55FFBFBB5AB5E3507E0C9C953B" bold="true" box="[151,394,1561,1586]" italics="true" pageId="9" pageNumber="580">Atrocrates matthewsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249F32EFFBFBB5AB4E5507E0CFD953B" box="[401,491,1561,1586]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB19247FFBFBB5AB5E3505A0CB2955F" blockId="9.[151,491,1561,1622]" box="[151,420,1597,1622]" pageId="9" pageNumber="580">
(
<figureCitation id="13358EC2FFBFBB5AB5EB505A0C36955F" box="[159,288,1597,1622]" captionStart-0="FIGURES 47 54" captionStart-1="FIGURES 55 62" captionStart-2="FIGURES 63 70" captionStartId-0="10.[151,264,1962,1985]" captionStartId-1="11.[151,264,1934,1957]" captionStartId-2="12.[151,264,1962,1985]" captionTargetBox-0="[209,1377,193,1940]" captionTargetBox-1="[217,1354,193,1894]" captionTargetBox-2="[209,1377,193,1940]" captionTargetId-0="figure@10.[209,1378,193,1941]" captionTargetId-1="figure@11.[217,1355,193,1895]" captionTargetId-2="figure@12.[209,1378,193,1941]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." captionText-1="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." captionText-2="FIGURES 63 70. Atrocrates matthewsi sp. nov. male. Fore tibia, inside (63). Fore tibia, ventral (64). Mid tibia, ventral (65). Hind tibia, ventral (66). Fore tibia, dorsal (67). Mid tibia, dorsal (68). Hind tibia, dorsal (69). Fore leg (70)." httpUri-0="https://zenodo.org/record/262709/files/figure.png" httpUri-1="https://zenodo.org/record/262710/files/figure.png" httpUri-2="https://zenodo.org/record/262711/files/figure.png" pageId="9" pageNumber="580">Figs 4770</figureCitation>
,
<figureCitation id="13358EC2FFBFBB5AB45F50590C65955F" box="[299,371,1598,1622]" captionStart="FIGURES 71 73" captionStartId="14.[151,264,1926,1949]" captionTargetBox="[246,1345,203,1894]" captionTargetId="figure@14.[225,1362,193,1905]" captionTargetPageId="14" captionText="FIGURES 71 73. Habitus of Atrocrates galbasi sp. nov., male (71). A. matthewsi sp. nov., male (72). Map of distribution (73)." httpUri="https://zenodo.org/record/262712/files/figure.png" pageId="9" pageNumber="580">72, 73</figureCitation>
,
<figureCitation id="13358EC2FFBFBB5AB40B50590C8A955F" box="[383,412,1598,1622]" captionStart="FIGURES 74 81" captionStartId="15.[151,264,1990,2013]" captionTargetBox="[224,1373,209,1949]" captionTargetId="figure@15.[207,1390,193,1963]" captionTargetPageId="15" captionText="FIGURES 74 81. Distribution of the Atrocrates species groups." httpUri="https://zenodo.org/record/262713/files/figure.png" pageId="9" pageNumber="580">80</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C314C1CCFFBFBB5AB5E350E2095B9597" box="[151,1101,1669,1694]" pageId="9" pageNumber="580" type="description">
<paragraph id="8BB19247FFBFBB5AB5E350E2095B9597" blockId="9.[151,1436,1669,1982]" box="[151,1101,1669,1694]" pageId="9" pageNumber="580">
<materialsCitation id="3B66981AFFBFBB5AB5E350E2095B9597" ID-GBIF-Occurrence="1323872872" box="[151,1101,1669,1694]" country="South Africa" location="Cape" pageId="9" pageNumber="580" specimenCount="1" stateProvince="Cape">
<emphasis id="B97A4E55FFBFBB5AB5E350E20C549597" bold="true" box="[151,322,1669,1694]" pageId="9" pageNumber="580">Locus typicus.</emphasis>
Graaff Reinet (South East
<collectingRegion id="49CA5CA5FFBFBB5AB70550E10FBB9597" box="[625,685,1670,1694]" country="South Africa" name="Western Cape" pageId="9" pageNumber="580">Cape</collectingRegion>
,
<collectingCountry id="F319D2D7FFBFBB5AB7CD50E20EC39597" box="[697,981,1669,1694]" name="South Africa" pageId="9" pageNumber="580">Republic of South Africa</collectingCountry>
) (
<figureCitation id="13358EC2FFBFBB5AB69850E209299597" box="[1004,1087,1669,1694]" captionStart="FIGURES 71 73" captionStartId="14.[151,264,1926,1949]" captionTargetBox="[246,1345,203,1894]" captionTargetId="figure@14.[225,1362,193,1905]" captionTargetPageId="14" captionText="FIGURES 71 73. Habitus of Atrocrates galbasi sp. nov., male (71). A. matthewsi sp. nov., male (72). Map of distribution (73)." httpUri="https://zenodo.org/record/262712/files/figure.png" pageId="9" pageNumber="580">Fig. 73</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C314C1CCFFBFBB5AB5B350CE0E8795EF" pageId="9" pageNumber="580" type="etymology">
<paragraph id="8BB19247FFBFBB5AB5B350CE0E8795EF" blockId="9.[151,1436,1669,1982]" pageId="9" pageNumber="580">
<emphasis id="B97A4E55FFBFBB5AB5B350CE0C5895CB" bold="true" box="[199,334,1705,1730]" pageId="9" pageNumber="580">Etymology.</emphasis>
Im pleased to dedicate this species to the distinguished coleopterist Dr. Eric G. Matthews (The South Australian Museum, North Terrace, Adelaide, SA,
<collectingCountry id="F319D2D7FFBFBB5AB66D50AA0E9395EF" box="[793,901,1741,1766]" name="Australia" pageId="9" pageNumber="580">Australia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C314C1CCFFBFBB5AB5B350960E649403" box="[199,882,1777,1802]" pageId="9" pageNumber="580" type="diagnosis">
<paragraph id="8BB19247FFBFBB5AB5B350960E649403" blockId="9.[151,1436,1669,1982]" box="[199,882,1777,1802]" pageId="9" pageNumber="580">
<emphasis id="B97A4E55FFBFBB5AB5B350960C579403" bold="true" box="[199,321,1777,1802]" pageId="9" pageNumber="580">Diagnosis.</emphasis>
<taxonomicName id="4C0EE9C4FFBFBB5AB43C50950CCE9403" box="[328,472,1777,1802]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="matthewsi">
<emphasis id="B97A4E55FFBFBB5AB43C50950CCE9403" box="[328,472,1777,1802]" italics="true" pageId="9" pageNumber="580">A. matthewsi</emphasis>
</taxonomicName>
is close to
<taxonomicName id="4C0EE9C4FFBFBB5AB72250950FDE9403" box="[598,712,1777,1802]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="galbasi">
<emphasis id="B97A4E55FFBFBB5AB72250950FDE9403" box="[598,712,1777,1802]" italics="true" pageId="9" pageNumber="580">A. galbasi</emphasis>
</taxonomicName>
(see dignosis).
</paragraph>
</subSubSection>
<subSubSection id="C314C1CCFFBFBB5EB5B351720EAA971F" lastPageId="13" lastPageNumber="584" pageId="9" pageNumber="580" type="description">
<paragraph id="8BB19247FFBFBB5AB5B351720F8B945B" blockId="9.[151,1436,1669,1982]" pageId="9" pageNumber="580">
<emphasis id="B97A4E55FFBFBB5AB5B351720C479427" bold="true" box="[199,337,1813,1838]" pageId="9" pageNumber="580">Description</emphasis>
. Measurements. Body length
<quantity id="4CF63FA2FFBFBB5AB7DE51720E579424" box="[682,833,1813,1838]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.06" metricValueMin="0.99" pageId="9" pageNumber="580" unit="mm" value="10.25" valueMax="10.6" valueMin="9.9">9.910.6 mm</quantity>
(
<typeStatus id="54B52CE5FFBFBB5AB62551720EA19427" box="[849,951,1813,1838]" pageId="9" pageNumber="580" type="holotype">holotype</typeStatus>
, male—
<quantity id="4CF63FA2FFBFBB5AB16F517209949427" box="[1051,1154,1813,1838]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="580" unit="mm" value="10.1">10.1 mm</quantity>
), pl/pb—0.7, el/eb—1.3, el/pl—1.9, eb/pb—0.95 (male), 1.00 (female).
</paragraph>
<paragraph id="8BB19247FFBFBB5AB5B351390DE994B7" blockId="9.[151,1436,1669,1982]" pageId="9" pageNumber="580">
Upper side of body mat, with a greasy sheen; head, pronotum and elytral intervals sparsely and delicately punctate (
<figureCitation id="13358EC2FFBFBB5AB47C51E60C769493" box="[264,352,1921,1946]" captionStart="FIGURES 71 73" captionStartId="14.[151,264,1926,1949]" captionTargetBox="[246,1345,203,1894]" captionTargetId="figure@14.[225,1362,193,1905]" captionTargetPageId="14" captionText="FIGURES 71 73. Habitus of Atrocrates galbasi sp. nov., male (71). A. matthewsi sp. nov., male (72). Map of distribution (73)." httpUri="https://zenodo.org/record/262712/files/figure.png" pageId="9" pageNumber="580">Fig. 72</figureCitation>
). Underside of body slightly shiny; prosternum smooth, episternum with shallow longitudinal wrinkles.
</paragraph>
<caption id="DF71C2CFFFBCBB59B5E351CD0EF394D6" httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="10" pageNumber="581" targetBox="[209,1377,193,1940]" targetPageId="10">
<paragraph id="8BB19247FFBCBB59B5E351CD0EF394D6" blockId="10.[151,1436,1962,2015]" pageId="10" pageNumber="581">
<emphasis id="B97A4E55FFBCBB59B5E351CD0C4294C9" bold="true" box="[151,340,1962,1985]" pageId="10" pageNumber="581">FIGURES 4754.</emphasis>
<taxonomicName id="4C0EE9C4FFBCBB59B42F51CC0F2594C8" box="[347,563,1963,1985]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
<emphasis id="B97A4E55FFBCBB59B42F51CC0F2594C8" box="[347,563,1963,1985]" italics="true" pageId="10" pageNumber="581">Atrocrates matthewsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249F32EFFBCBB59B74D51CC0F9C94C9" box="[569,650,1963,1984]" pageId="10" pageNumber="581" rank="species">
<emphasis id="B97A4E55FFBCBB59B74D51CC0F9C94C9" bold="true" box="[569,650,1963,1984]" pageId="10" pageNumber="581">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54).
</paragraph>
</caption>
<caption id="DF71C2CFFFBDBB58B5E351E90E1494ED" httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="11" pageNumber="582" targetBox="[217,1354,193,1894]" targetPageId="11">
<paragraph id="8BB19247FFBDBB58B5E351E90E1494ED" blockId="11.[151,1436,1934,2020]" pageId="11" pageNumber="582">
<emphasis id="B97A4E55FFBDBB58B5E351E90C4394AD" bold="true" box="[151,341,1934,1957]" pageId="11" pageNumber="582">FIGURES 5562.</emphasis>
<taxonomicName id="4C0EE9C4FFBDBB58B42851F70F2394AC" box="[348,565,1935,1957]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
<emphasis id="B97A4E55FFBDBB58B42851F70F2394AC" box="[348,565,1935,1957]" italics="true" pageId="11" pageNumber="582">Atrocrates matthewsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249F32EFFBDBB58B74851E80F9894AD" box="[572,654,1935,1956]" pageId="11" pageNumber="582" rank="species">
<emphasis id="B97A4E55FFBDBB58B74851E80F9894AD" bold="true" box="[572,654,1935,1956]" pageId="11" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3rd, 4th and 5th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62).
</paragraph>
</caption>
<caption id="DF71C2CFFFBABB5FB5E351CD09C294D6" httpUri="https://zenodo.org/record/262711/files/figure.png" pageId="12" pageNumber="583" targetBox="[209,1377,193,1940]" targetPageId="12">
<paragraph id="8BB19247FFBABB5FB5E351CD09C294D6" blockId="12.[151,1436,1962,2015]" pageId="12" pageNumber="583">
<emphasis id="B97A4E55FFBABB5FB5E351CD0C4394C9" bold="true" box="[151,341,1962,1985]" pageId="12" pageNumber="583">FIGURES 6370.</emphasis>
<taxonomicName id="4C0EE9C4FFBABB5FB42F51CC0F2594C8" box="[347,563,1963,1985]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
<emphasis id="B97A4E55FFBABB5FB42F51CC0F2594C8" box="[347,563,1963,1985]" italics="true" pageId="12" pageNumber="583">Atrocrates matthewsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249F32EFFBABB5FB74E51CC0F9D94C9" box="[570,651,1963,1984]" pageId="12" pageNumber="583" rank="species">
<emphasis id="B97A4E55FFBABB5FB74E51CC0F9D94C9" bold="true" box="[570,651,1963,1984]" pageId="12" pageNumber="583">sp. nov.</emphasis>
</taxonomicNameLabel>
male. Fore tibia, inside (63). Fore tibia, ventral (64). Mid tibia, ventral (65). Hind tibia, ventral (66). Fore tibia, dorsal (67). Mid tibia, dorsal (68). Hind tibia, dorsal (69). Fore leg (70).
</paragraph>
</caption>
<paragraph id="8BB19247FFBBBB5EB5B356F00836924A" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Head as in
<figureCitation id="13358EC2FFBBBB5EB42156F00CBA93B9" box="[341,428,151,176]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 47</figureCitation>
. Clypeus with shallow emargination (ew/ed—1.0). Genal canthus equal to eyes. Eye between tempus and genal canthus narrow (23 ommatidia visible), circumocular depression distinct, narrow (
<figureCitation id="13358EC2FFBBBB5EB01B56DB0DA193F2" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 48</figureCitation>
). Antenna moderately long (al/apl—0.8) and wide (al/was3—12), 3rd antennomere short (as3/as2—2.3), distal segments (antennomere 711) evenly widened (
<figureCitation id="13358EC2FFBBBB5EB7C357610E1A9216" box="[695,780,262,287]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 51</figureCitation>
). Mentum and submentum as in
<figureCitation id="13358EC2FFBBBB5EB1F1576109CF9216" box="[1157,1241,262,287]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 50</figureCitation>
. Last segment of maxillary palp moderately wide (mp/as3—1.4). Hipostoma near maxillary articulation simple (
<figureCitation id="13358EC2FFBBBB5EB1CB574D0802924A" box="[1215,1300,298,323]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 49</figureCitation>
).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B357370F8392DA" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Pronotum as in
<figureCitation id="13358EC2FFBBBB5EB40857290CD9926E" box="[380,463,334,359]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Fig. 52</figureCitation>
, sides rounded, widest at 1/3 from base; pronotal disc evenly convex, depressed near lateral border, with extremely fine puncturation, punctures 35 diameters apart; anterior angles obtuse and moderately protruding anteriorly (pl/lapa—0.8); posterior angles right, not protruding beyond the level of the middle of base; pronotal base almost straight.
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B357B90F7892FE" blockId="13.[151,1437,151,1046]" box="[199,622,478,503]" pageId="13" pageNumber="584">Scutellum wide at base (pkp/st—3.0).</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B354650C38916A" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Elytron composed of 9 rows, striae punctato-sulcate, shallow with fine punctures; intervals flat with fine punctures, 56 diameters apart; basal margin almost straight; elytral humeri rectangular, not protruding outwards (
<figureCitation id="13358EC2FFBBBB5EB5EB542D0C37916A" box="[159,289,586,611]" captionStart="FIGURES 47 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">Figs 53, 54</figureCitation>
).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5BA54090EA6918E" blockId="13.[151,1437,151,1046]" box="[206,944,622,647]" pageId="13" pageNumber="584">
Prosternal process protruting posteriad, rounded at apex (
<figureCitation id="13358EC2FFBBBB5EB63B54090EB5918E" box="[847,931,622,647]" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B354F50F0A91C6" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Metaventrite (
<figureCitation id="13358EC2FFBBBB5EB41154F50CF691A2" box="[357,480,658,683]" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Fig. 56, 57</figureCitation>
) between the insertions of mid and hind coxae short (metl/cavl—4.5); metepisternum rectangle and short (ml/mw—3.2).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B354BC0C449032" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Legs, all tarsi in both sexes narrow; female legs as in
<figureCitation id="13358EC2FFBBBB5EB65054BD0EB391FA" box="[804,933,730,755]" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Figs 5962</figureCitation>
; male fore tibia with longitudinal depression on inner side (
<figureCitation id="13358EC2FFBBBB5EB44A54990CE5901E" box="[318,499,766,791]" captionStart="FIGURES 63 70" captionStartId="12.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@12.[209,1378,193,1941]" captionTargetPageId="12" captionText="FIGURES 63 70. Atrocrates matthewsi sp. nov. male. Fore tibia, inside (63). Fore tibia, ventral (64). Mid tibia, ventral (65). Hind tibia, ventral (66). Fore tibia, dorsal (67). Mid tibia, dorsal (68). Hind tibia, dorsal (69). Fore leg (70)." httpUri="https://zenodo.org/record/262711/files/figure.png" pageId="13" pageNumber="584">Figs 63, 64, 70</figureCitation>
); male mid tibia widened, with small apical denticle (
<figureCitation id="13358EC2FFBBBB5EB11754990801901E" box="[1123,1303,766,791]" captionStart="FIGURES 63 70" captionStartId="12.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@12.[209,1378,193,1941]" captionTargetPageId="12" captionText="FIGURES 63 70. Atrocrates matthewsi sp. nov. male. Fore tibia, inside (63). Fore tibia, ventral (64). Mid tibia, ventral (65). Hind tibia, ventral (66). Fore tibia, dorsal (67). Mid tibia, dorsal (68). Hind tibia, dorsal (69). Fore leg (70)." httpUri="https://zenodo.org/record/262711/files/figure.png" pageId="13" pageNumber="584">Figs 65, 66, 68</figureCitation>
); hind tibia simple (
<figureCitation id="13358EC2FFBBBB5EB58555450C539032" box="[241,325,802,827]" captionStart="FIGURES 63 70" captionStartId="12.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@12.[209,1378,193,1941]" captionTargetPageId="12" captionText="FIGURES 63 70. Atrocrates matthewsi sp. nov. male. Fore tibia, inside (63). Fore tibia, ventral (64). Mid tibia, ventral (65). Hind tibia, ventral (66). Fore tibia, dorsal (67). Mid tibia, dorsal (68). Hind tibia, dorsal (69). Fore leg (70)." httpUri="https://zenodo.org/record/262711/files/figure.png" pageId="13" pageNumber="584">Fig. 69</figureCitation>
).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B355210EBF908F" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
Abdominal ventrites with delicate puncturation (
<figureCitation id="13358EC2FFBBBB5EB79155210E2F9056" box="[741,825,838,863]" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Fig. 57</figureCitation>
); bordering of the last abdominal ventrite distinct (
<figureCitation id="13358EC2FFBBBB5EB01B55210DA0908F" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Fig. 58</figureCitation>
); process of 1st abdominal ventrite narrow (pav/pm—1.9) (
<figureCitation id="13358EC2FFBBBB5EB63C550A0E8B908F" box="[840,925,877,902]" captionStart="FIGURES 55 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">Fig. 56</figureCitation>
).
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B355F60E7590A0" blockId="13.[151,1437,151,1046]" box="[199,867,913,938]" pageId="13" pageNumber="584">
Female and male genitalia as in other species of
<taxonomicName id="4C0EE9C4FFBBBB5EB79255F50E4D90A0" box="[742,859,914,937]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="B97A4E55FFBBBB5EB79255F50E4D90A0" box="[742,859,914,937]" italics="true" pageId="13" pageNumber="584">Atrocrates</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5B355D20EAA971F" blockId="13.[151,1437,151,1046]" pageId="13" pageNumber="584">
<materialsCitation id="3B66981AFFBBBB5EB5B355D20ED790C7" ID-GBIF-Occurrence="1323872870" box="[199,961,949,974]" collectionCode="DNMNH" location="Cape" pageId="13" pageNumber="584" specimenCount="2" stateProvince="Cape" typeStatus="holotype">
<emphasis id="B97A4E55FFBBBB5EB5B355D20C1A90C7" bold="true" box="[199,268,949,974]" pageId="13" pageNumber="584">
<typeStatus id="54B52CE5FFBBBB5EB5B355D20C1A90C7" box="[199,268,949,974]" pageId="13" pageNumber="584">Types</typeStatus>
</emphasis>
.
<typeStatus id="54B52CE5FFBBBB5EB46A55D20C9D90C7" box="[286,395,949,974]" pageId="13" pageNumber="584" type="holotype">Holotype</typeStatus>
: male, deposited in
<collectionCode id="ED1F0A82FFBBBB5EB70C55D10FF090C4" box="[632,742,950,973]" country="South Africa" httpUri="http://grbio.org/cool/6t48-bahh" name="Ditsong National Museum of Natural History" pageId="13" pageNumber="584">DNMNH</collectionCode>
; “S. A. East
<collectingRegion id="49CA5CA5FFBBBB5EB6F055D10ED790C7" box="[900,961,950,974]" country="South Africa" name="Western Cape" pageId="13" pageNumber="584">Cape</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B66981AFFBBBB5EB6A455D20CB890FB" ID-GBIF-Occurrence="1323872868" collectingDate="1996-01-12" collectionCode="ARNDT" location="Graaff Reinet" pageId="13" pageNumber="584" specimenCount="1" stateProvince="Cape" typeStatus="holotype">
<location id="8ED1C49CFFBBBB5EB6A455D2097990C7" LSID="urn:lsid:plazi:treatment:03A72351FFBFBB5EB5E3507E0EAA971F:8ED1C49CFFBBBB5EB6A455D2097990C7" box="[976,1135,949,974]" name="Graaff Reinet" pageId="13" pageNumber="584" stateProvince="Cape">Graaff Reinet</location>
, 32”13S/24”30E; 29.11-
<collectingDate id="EFF44D6FFFBBBB5EB5E355BE0C1D90FB" box="[151,267,985,1010]" pageId="13" pageNumber="584" value="1996-01-12">
<date id="FFB0B487FFBBBB5EB5E355BE0C1D90FB" box="[151,267,985,1010]" pageId="13" pageNumber="584" value="1996-01-12">1.12.1996</date>
</collectingDate>
; leg
<collectionCode id="ED1F0A82FFBBBB5EB44955BD0CB590FB" box="[317,419,986,1010]" pageId="13" pageNumber="584">ARNDT</collectionCode>
</materialsCitation>
. Paratypes: 2 males and 3 females (DNMNH),
<materialsCitation id="3B66981AFFBBBB5EB6CD55BD0EAA971F" ID-GBIF-Occurrence="1323872871" collectingDate="1996-01-12" collectionCode="ARNDT" location="Graaff Reinet" pageId="13" pageNumber="584" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cape" typeStatus="holotype">
<specimenCount id="9D0859CEFFBBBB5EB6CD55BD091590FB" box="[953,1027,985,1010]" pageId="13" pageNumber="584" type="male">1 male</specimenCount>
and
<specimenCount id="9D0859CEFFBBBB5EB14D55BD098F90FB" box="[1081,1177,985,1010]" pageId="13" pageNumber="584" type="female">1 female</specimenCount>
(MIZPAN) “S. A. East
<collectingRegion id="49CA5CA5FFBBBB5EB5E355990DC2971F" box="[151,212,1022,1046]" country="South Africa" name="Western Cape" pageId="13" pageNumber="584">Cape</collectingRegion>
;
<location id="8ED1C49CFFBBBB5EB5AB559A0C6D971F" LSID="urn:lsid:plazi:treatment:03A72351FFBFBB5EB5E3507E0EAA971F:8ED1C49CFFBBBB5EB5AB559A0C6D971F" box="[223,379,1021,1046]" name="Graaff Reinet" pageId="13" pageNumber="584" stateProvince="Cape">Graaff Reinet</location>
, 32”13S/24”30E; 29.11-
<collectingDate id="EFF44D6FFFBBBB5EB7EB559A0E05971F" box="[671,787,1021,1046]" pageId="13" pageNumber="584" value="1996-01-12">
<date id="FFB0B487FFBBBB5EB7EB559A0E05971F" box="[671,787,1021,1046]" pageId="13" pageNumber="584" value="1996-01-12">1.12.1996</date>
</collectingDate>
; leg
<collectionCode id="ED1F0A82FFBBBB5EB63C55990EB8971F" box="[840,942,1022,1046]" pageId="13" pageNumber="584">ARNDT</collectionCode>
”.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>