479 lines
63 KiB
XML
479 lines
63 KiB
XML
<document id="4B2D6E2E9084BCC63561EC164131BCE3" ID-DOI="10.5281/zenodo.181720" ID-GBIF-Dataset="c52d4a56-6118-4d27-9730-ea00111f2572" ID-ISSN="1175-5326" ID-Zenodo-Dep="181720" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838565835" checkinUser="plazi" docAuthor="Witaliński, Wojciech" docDate="2008" docId="03A7073BFFCFFFF9FF6FF95C398CFDF7" docLanguage="en" docName="zt01750p058.pdf" docOrigin="Zootaxa 1750" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Heteroparasitus mariae Witaliński, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FF9E7F43FFCFFFFDFFF8FFBB3877FFED" masterDocTitle="A new species of Heteroparasitus Juvara-Bals 1975 from Karavanke, Austria (Acari: Parasitidae)" masterLastPageNumber="58" masterPageNumber="53" pageNumber="53" updateTime="1698224032747" updateUser="plazi">
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<mods:mods id="5380780C29C399D559B5664DB0CF4509" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="72FA6F0510C735ADB5C28951B58ABA7D">
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<mods:title id="3C71F09FD34070296D05A761D3F9118D">A new species of Heteroparasitus Juvara-Bals 1975 from Karavanke, Austria (Acari: Parasitidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="73249B4DD840D99F6D8853C9D0F32059" type="personal">
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<mods:role id="5CB73B35FA774696DA0A538A796A56BD">
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<mods:roleTerm id="F4C1B67DFAAD8BCF0C9A1D6CEB51ABDD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6E1D5FBBC0068B55A09C16EBDAAC77A">Witaliński, Wojciech</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B5640F5C3E7788A993F65BD05CFB33AE">text</mods:typeOfResource>
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<mods:titleInfo id="0E3410747E04099E43814515658BA317">
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<mods:title id="0E4A6F65F10EC46F53AF6DCFA703D58A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B2D80DBF36F8B1D0E8F33A56ABE3E89C">
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<mods:date id="04550E9DE5D1C0EE9953541C759EEF4F">2008</mods:date>
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<mods:detail id="453C28A9AC69C4A7FCD8E89C0C6BE8AB" type="volume">
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<mods:number id="049F4D5D25114C516F2AF7E58FBD2727">1750</mods:number>
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</mods:detail>
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<mods:extent id="9B216346410667483A9DC94FAA7BC579" unit="page">
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<mods:start id="DBACAA9F66759815F19E00BEF979D59C">53</mods:start>
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<mods:end id="D2873BD4EE3B6DEEA0BAEB347A010CF9">58</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="6694AFB469DA75C1D0CBFD9745754F0F">journal article</mods:classification>
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<mods:identifier id="91EB6981FD3B762C05E82CE9B6549DFF" type="DOI">10.5281/zenodo.181720</mods:identifier>
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<mods:identifier id="B00B2B06233933F98529BB59922DA6D3" type="GBIF-Dataset">c52d4a56-6118-4d27-9730-ea00111f2572</mods:identifier>
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<mods:identifier id="11599372670B4F10B07151E7FB425581" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9CB7E9A5F80B21F134B9C289BBFA9ADB" type="Zenodo-Dep">181720</mods:identifier>
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</mods:mods>
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<treatment id="03A7073BFFCFFFF9FF6FF95C398CFDF7" ID-DOI="http://doi.org/10.5281/zenodo.6232783" ID-GBIF-Taxon="119359637" ID-Zenodo-Dep="6232783" LSID="urn:lsid:plazi:treatment:03A7073BFFCFFFF9FF6FF95C398CFDF7" httpUri="http://treatment.plazi.org/id/03A7073BFFCFFFF9FF6FF95C398CFDF7" lastPageId="4" lastPageNumber="57" pageId="0" pageNumber="53">
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<subSubSection id="C314E5A6FFCFFFFDFF6FF95C3969F8C9" pageId="0" pageNumber="53" type="nomenclature">
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<paragraph id="8BB1B62DFFCFFFFDFF6FF95C3A6EF8EC" blockId="0.[151,537,1767,1828]" box="[151,537,1767,1793]" pageId="0" pageNumber="53">
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<heading id="D0F90141FFCFFFFDFF6FF95C3A6EF8EC" bold="true" box="[151,537,1767,1793]" fontSize="11" level="1" pageId="0" pageNumber="53" reason="1">
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||
<emphasis id="B97A6A3FFFCFFFFDFF6FF95C3A6EF8EC" bold="true" box="[151,537,1767,1793]" pageId="0" pageNumber="53">
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<taxonomicName id="4C0ECDAEFFCFFFFDFF6FF95C39C2F8EC" box="[151,437,1767,1793]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
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<emphasis id="B97A6A3FFFCFFFFDFF6FF95C39C2F8EC" bold="true" box="[151,437,1767,1793]" italics="true" pageId="0" pageNumber="53">Heteroparasitus mariae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A249D744FFCFFFFDFE44F95C3A6EF8EC" box="[444,537,1767,1793]" pageId="0" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB1B62DFFCFFFFDFF6FF8B13969F8C9" blockId="0.[151,537,1767,1828]" box="[151,286,1802,1828]" pageId="0" pageNumber="53">
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(
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<figureCitation id="1335AAA8FFCFFFFDFF67F8B13961F8C9" box="[159,278,1802,1828]" captionStart-0="FIGURES 1 – 11" captionStart-1="FIGURES 12 – 20" captionStartId-0="2.[151,269,1803,1827]" captionStartId-1="3.[152,270,1317,1341]" captionTargetBox-0="[196,1387,525,1729]" captionTargetBox-1="[194,1395,400,1269]" captionTargetId-0="figure@2.[188,1398,517,1777]" captionTargetId-1="figure@3.[188,1398,394,1313]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." captionText-1="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri-0="https://zenodo.org/record/181722/files/figure.png" httpUri-1="https://zenodo.org/record/181723/files/figure.png" pageId="0" pageNumber="53">Figs 1–19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C314E5A6FFCFFFFDFF6FF8E13C85F802" pageId="0" pageNumber="53" type="materials_examined">
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<paragraph id="8BB1B62DFFCFFFFDFF6FF8E13C85F802" blockId="0.[151,1436,1882,2031]" pageId="0" pageNumber="53">
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<emphasis id="B97A6A3FFFCFFFFDFF6FF8E139FAF899" bold="true" box="[151,397,1882,1908]" pageId="0" pageNumber="53">Material examined.</emphasis>
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<emphasis id="B97A6A3FFFCFFFFDFE6EF8E03A75F899" box="[406,514,1883,1908]" italics="true" pageId="0" pageNumber="53">
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<typeStatus id="54B5088FFFCFFFFDFE6EF8E03A75F899" box="[406,514,1883,1908]" pageId="0" pageNumber="53" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Female (slide no. 2140),
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<date id="FFB090EDFFCFFFFDFCC7F8E13BA9F899" box="[831,990,1882,1908]" pageId="0" pageNumber="53" value="2007-07-23">23 July 2007</date>
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, close to St. Leonhard Church, Karavanke mountains,
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<collectingCountry id="F319F6BDFFCFFFFDFE8DF83E39B9F872" box="[373,462,1925,1951]" name="Austria" pageId="0" pageNumber="53">Austria</collectingCountry>
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,
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<geoCoordinate id="EE3AD0EAFFCFFFFDFE26F83E3AE9F872" box="[478,670,1922,1951]" direction="north" orientation="latitude" pageId="0" pageNumber="53" precision="1" value="46.44592">46o 26' 45.3" N</geoCoordinate>
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,
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<geoCoordinate id="EE3AD0EAFFCFFFFDFD56F83E3B07F872" box="[686,880,1925,1951]" direction="east" orientation="longitude" pageId="0" pageNumber="53" precision="1" value="14.649834">14° 38' 59.4" E</geoCoordinate>
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, alt.
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<emphasis id="B97A6A3FFFCFFFFDFC48F83D3BBCF872" box="[944,971,1926,1951]" italics="true" pageId="0" pageNumber="53">ca</emphasis>
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.
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<quantity id="4CF61BC8FFCFFFFDFC26F83E3C4CF872" box="[990,1083,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="0" pageNumber="53" unit="m" value="1410.0">1410 m</quantity>
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a.s.l., wet moss (
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<taxonomicName id="4C0ECDAEFFCFFFFDFAEEF83D3DE5F872" box="[1302,1426,1926,1951]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="0" pageNumber="53" phylum="Bryophyta" rank="genus">
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<emphasis id="B97A6A3FFFCFFFFDFAEEF83D3DE5F872" box="[1302,1426,1926,1951]" italics="true" pageId="0" pageNumber="53">Sphagnum</emphasis>
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</taxonomicName>
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) around water outflow.
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<emphasis id="B97A6A3FFFCFFFFDFE67F8153A61F82A" box="[415,534,1966,1991]" italics="true" pageId="0" pageNumber="53">
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<typeStatus id="54B5088FFFCFFFFDFE67F8153A61F82A" box="[415,534,1966,1991]" pageId="0" pageNumber="53" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.
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<specimenCount id="9D087DA4FFCFFFFDFDDCF8163AEFF82A" box="[548,664,1965,1991]" pageId="0" pageNumber="53" type="female">7 females</specimenCount>
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,
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<specimenCount id="9D087DA4FFCFFFFDFD5BF8163A8AF82A" box="[675,765,1965,1991]" pageId="0" pageNumber="53" type="male">3 males</specimenCount>
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(slides no.
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<date id="FFB090EDFFCFFFFDFC7BF8163C70F82A" box="[899,1031,1965,1991]" pageId="0" pageNumber="53" value="2141" valueMax="2146">2141-2146</date>
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), same data as
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<typeStatus id="54B5088FFFCFFFFDFB4DF8163D57F82A" box="[1205,1312,1965,1991]" pageId="0" pageNumber="53" type="holotype">holotype</typeStatus>
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. The
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<typeStatus id="54B5088FFFCFFFFDFAA6F8163DEBF82A" box="[1374,1436,1965,1991]" pageId="0" pageNumber="53">types</typeStatus>
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have been deposited in the Zoological Museum of the Jagiellonian University, Cracow,
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<collectingCountry id="F319F6BDFFCFFFFDFB61F86E3C99F802" box="[1177,1262,2005,2031]" name="Poland" pageId="0" pageNumber="53">Poland</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C314E5A6FFCEFFFCFF3EFF2C3A3CFDF8" pageId="1" pageNumber="54" type="diagnosis">
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<paragraph id="8BB1B62DFFCEFFFCFF3EFF2C3CBBFF5F" blockId="1.[151,1437,151,2036]" box="[198,1228,151,178]" pageId="1" pageNumber="54">
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<emphasis id="B97A6A3FFFCEFFFCFF3EFF2C3931FF5C" bold="true" box="[198,326,151,177]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
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Gnathotectum of both sexes trispinate; trochanters III and IV with tubercles.
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</paragraph>
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<paragraph id="8BB1B62DFFCEFFFCFF3EFF0438B8FE9A" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">Female: Sternum with thickened anterior margin; paragynia interdigitate along zigzag axial margins; epigynial subapical structure rectangular in outline, with lamellar lateral protrusions extending far from margin of epigynium; endogynium cup-shaped and elliptical in ventral perspective, with several minute denticles at its entrance; movable digit of chelicera with 3 or 4 teeth; trochanter IV with posteroventral spur-like protrusion.</paragraph>
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<paragraph id="8BB1B62DFFCEFFFCFF3DFE3E3A3CFDF8" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">Male: Corniculi slender, without protrusion on adaxial surface; fixed digit of chelicera edentate, movable digit with several small uniform teeth followed by a larger proximal tooth; femur of leg II with conical and somewhat anterolaterally curved spur; spur on genu roundish and close to distal margin of segment; spur on tibia wedge-like and distally pointed.</paragraph>
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</subSubSection>
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<subSubSection id="C314E5A6FFCEFFFEFF3EFD983899FEBF" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="description">
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<paragraph id="8BB1B62DFFCEFFFCFF3EFD9839C9FDD1" blockId="1.[151,1437,151,2036]" box="[198,446,547,573]" pageId="1" pageNumber="54">
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<emphasis id="B97A6A3FFFCEFFFCFF3EFD983929FDD0" bold="true" box="[198,350,547,573]" pageId="1" pageNumber="54">Description.</emphasis>
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<emphasis id="B97A6A3FFFCEFFFCFE9EFD9839C9FDD1" box="[358,446,547,572]" italics="true" pageId="1" pageNumber="54">Female</emphasis>
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</paragraph>
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<paragraph id="8BB1B62DFFCEFFFCFF3DFDF03D06FD61" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
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<taxonomicName id="4C0ECDAEFFCEFFFCFF3DFDF03945FD89" box="[197,306,587,612]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="B97A6A3FFFCEFFFCFF3DFDF03945FD89" box="[197,306,587,612]" italics="true" pageId="1" pageNumber="54">Idiosoma</emphasis>
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</taxonomicName>
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brownish, 465–505 x 690–735 µm, length/width 1.39–1.57 (n=6). Length of podonotal and opisthonotal setae: 48–52 μm (
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<emphasis id="B97A6A3FFFCEFFFCFDF9FDC93A7EFD66" box="[513,521,626,651]" italics="true" pageId="1" pageNumber="54">j</emphasis>
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1), remaining setae 33–44 μm; length of peritrematal groove 216–225 μm.
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</paragraph>
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<paragraph id="8BB1B62DFFCEFFFCFF3DFD213C72FCBF" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
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<emphasis id="B97A6A3FFFCEFFFCFF3DFD21397AFD5E" box="[197,269,666,691]" italics="true" pageId="1" pageNumber="54">Ve n t e r</emphasis>
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. Lateral presternal platelets small and triangular. Sternum (
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<figureCitation id="1335AAA8FFCEFFFCFC32FD213C79FD59" box="[970,1038,666,692]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
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) regularly reticulated, its anterior margin more pigmented and thickened due to incorporation of a presternal plate. Antero-lateral reticulation forming a row of cells distinctly inclined toward the opening of coxae II; line of reticulation running through the second pair of pores (
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<date id="FFB090EDFFCEFFFCFE45FCAA3996FCC7" box="[445,481,784,810]" pageId="1" pageNumber="54">
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<emphasis id="B97A6A3FFFCEFFFCFE45FCAA39A5FCC7" box="[445,466,785,810]" italics="true" pageId="1" pageNumber="54">iv</emphasis>
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2
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</date>
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) more pronounced and indented anteriorly. Gland pores
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<emphasis id="B97A6A3FFFCEFFFCFB8DFCAA3CE7FCC7" box="[1141,1168,785,810]" italics="true" pageId="1" pageNumber="54">gv</emphasis>
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||
1-
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<emphasis id="B97A6A3FFFCEFFFCFB51FCAA3CB2FCC7" box="[1193,1221,785,810]" italics="true" pageId="1" pageNumber="54">gv</emphasis>
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3 present. Setae of sternogenital region: 42–47 µm (
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<emphasis id="B97A6A3FFFCEFFFCFDE1FC833A5AFCBC" box="[537,557,824,849]" italics="true" pageId="1" pageNumber="54">st</emphasis>
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1), 42–44 µm (
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<emphasis id="B97A6A3FFFCEFFFCFD25FC833A87FCBC" box="[733,752,824,849]" italics="true" pageId="1" pageNumber="54">st</emphasis>
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2), 41–42 µm (
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<emphasis id="B97A6A3FFFCEFFFCFC59FC833BC2FCBC" box="[929,949,824,849]" italics="true" pageId="1" pageNumber="54">st</emphasis>
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3,
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<emphasis id="B97A6A3FFFCEFFFCFC2AFC833B92FCBC" box="[978,997,824,849]" italics="true" pageId="1" pageNumber="54">st</emphasis>
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4).
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</paragraph>
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<paragraph id="8BB1B62DFFCEFFFCFF3DFCDB3B28FB5B" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
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<emphasis id="B97A6A3FFFCEFFFCFF3DFCDB3907FC94" box="[197,368,864,889]" italics="true" pageId="1" pageNumber="54">Genital region</emphasis>
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. Paragynial shields (
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<figureCitation id="1335AAA8FFCEFFFCFD90FCDB3AD8FC97" box="[616,687,864,890]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 2</figureCitation>
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) showing zigzag connection along midline and relatively small posterior paragynial lobes. Central prong of epigynial shield (
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<figureCitation id="1335AAA8FFCEFFFCFC95FC3C3BC4FC4C" box="[877,947,903,929]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 3</figureCitation>
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) triangular. Subapical structure rectangular in outline, lateral protrusions moderately sclerotised, ending far beyond margin of epigynium. Endogynium (
|
||
<figureCitation id="1335AAA8FFCEFFFCFF1DFC6D3927FC1D" box="[229,336,982,1008]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Figs 4, 5</figureCitation>
|
||
) cup-like; in ventral perspective its entrance is oval and extended laterally, showing several minute denticles. Endogynial lamella absent. Gland pores
|
||
<emphasis id="B97A6A3FFFCEFFFCFCB0FC453B13FBFA" box="[840,868,1022,1047]" italics="true" pageId="1" pageNumber="54">gv</emphasis>
|
||
<quantity id="4CF61BC8FFCEFFFCFC9CFC453BE5FBF5" box="[868,914,1022,1048]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="54" unit="in" value="2.0">2 in</quantity>
|
||
unmodified cuticle. Opisthogaster normally with seven pairs of ventral setae, 33–35 µm (
|
||
<date id="FFB090EDFFCEFFFCFD4DFB9D3AA3FBD2" box="[693,724,1061,1087]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFD4DFB9D3AB0FBD2" box="[693,711,1062,1087]" italics="true" pageId="1" pageNumber="54">V</emphasis>
|
||
1
|
||
</date>
|
||
), 34–43 µm (others). In three of six females the number of opisthogastric setae varied due to the absence of
|
||
<emphasis id="B97A6A3FFFCEFFFCFD37FBF63A86FB8B" box="[719,753,1101,1126]" italics="true" pageId="1" pageNumber="54">ZV</emphasis>
|
||
1 on one side or the presence of an additional seta between
|
||
<emphasis id="B97A6A3FFFCEFFFCFF6FFBCE38CEFB63" box="[151,185,1141,1166]" italics="true" pageId="1" pageNumber="54">ZV</emphasis>
|
||
2 and
|
||
<emphasis id="B97A6A3FFFCEFFFCFEFCFBCE3951FB63" box="[260,294,1141,1166]" italics="true" pageId="1" pageNumber="54">ZV</emphasis>
|
||
3). Numbers of left/right setae were 7/6 (
|
||
<emphasis id="B97A6A3FFFCEFFFCFCEBFBCE3B42FB63" box="[787,821,1141,1166]" italics="true" pageId="1" pageNumber="54">ZV</emphasis>
|
||
1 lacking), 7/8 (additional seta present) and 7/8 (on both sides additional setae present, on left side
|
||
<emphasis id="B97A6A3FFFCEFFFCFD47FB273A96FB58" box="[703,737,1180,1205]" italics="true" pageId="1" pageNumber="54">ZV</emphasis>
|
||
1 lacking).
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCEFFFCFF3DFB7F3899FA4E" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
|
||
<taxonomicName id="4C0ECDAEFFCEFFFCFF3DFB7F392CFB30" box="[197,347,1220,1245]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="1" pageNumber="54" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCEFFFCFF3DFB7F392CFB30" box="[197,347,1220,1245]" italics="true" pageId="1" pageNumber="54">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Gnathotectum (
|
||
<figureCitation id="1335AAA8FFCEFFFCFDD8FB783A11FB30" box="[544,614,1219,1245]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 8</figureCitation>
|
||
) trispinate with long acute central prong, lateral prongs much smaller, corniculi conical (
|
||
<figureCitation id="1335AAA8FFCEFFFCFE94FB5039C2FAE8" box="[364,437,1259,1285]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 9</figureCitation>
|
||
). Hypognathal groove (
|
||
<figureCitation id="1335AAA8FFCEFFFCFD35FB503B63FAE8" box="[717,788,1259,1285]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 9</figureCitation>
|
||
) with 11–13 rows of denticles; anterior rows well pronounced, posterior rows barely visible in some specimens. Palpcoxal setae barbed, hypostomatic setae simple. Fixed digit of chelicera with two teeth in front and two behind the
|
||
<emphasis id="B97A6A3FFFCEFFFCFC43FA813C24FABE" box="[955,1107,1338,1363]" italics="true" pageId="1" pageNumber="54">pilus dentilis</emphasis>
|
||
, followed by lamellar edge (
|
||
<figureCitation id="1335AAA8FFCEFFFCFF67FAD9397BFA91" box="[159,268,1378,1404]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Figs 6, 7</figureCitation>
|
||
); movable digit with three or four teeth (
|
||
<figureCitation id="1335AAA8FFCEFFFCFD0DFAD93B49FA91" box="[757,830,1378,1404]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Fig. 7</figureCitation>
|
||
). Palp trochanter with seta
|
||
<date id="FFB090EDFFCEFFFCFB7DFAD93CD6FA91" box="[1157,1185,1378,1404]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFB7DFAD93CE5FA96" box="[1157,1170,1378,1403]" italics="true" pageId="1" pageNumber="54">v</emphasis>
|
||
1
|
||
</date>
|
||
simple and
|
||
<date id="FFB090EDFFCEFFFCFACBFAD93D38FA91" box="[1331,1359,1378,1404]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFACBFAD93D37FA96" box="[1331,1344,1378,1403]" italics="true" pageId="1" pageNumber="54">v</emphasis>
|
||
2
|
||
</date>
|
||
finely barbed.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCEFFFCFF3DFA0A39EEF9F7" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFF3DFA0A388BFA27" box="[197,252,1457,1482]" italics="true" pageId="1" pageNumber="54">Legs</emphasis>
|
||
. Leg structure and setation normal for genus, except trochanter III with a posteroventral tubercle and trochanter IV with an anterodorsal tubercle and posteroventral spur-like protrusion; coxa IV with anterolateral tubercle (
|
||
<figureCitation id="1335AAA8FFCEFFFCFEFDF9BB39FCF9F7" box="[261,395,1536,1562]" captionStart="FIGURES 1 – 11" captionStartId="2.[151,269,1803,1827]" captionTargetBox="[196,1387,525,1729]" captionTargetId="figure@2.[188,1398,517,1777]" captionTargetPageId="2" captionText="FIGURES 1 – 11. Heteroparasitus (H.) mariae sp. nov., female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9, gnathosoma, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9 – 11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype )). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="1" pageNumber="54">Figs 10, 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCEFFFCFF3DF9933976F9AC" blockId="1.[151,1437,151,2036]" box="[197,257,1576,1601]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFF3DF9933976F9AC" box="[197,257,1576,1601]" italics="true" pageId="1" pageNumber="54">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCEFFFCFF3DF9F43A57F955" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
|
||
<taxonomicName id="4C0ECDAEFFCEFFFCFF3DF9F43945F985" box="[197,306,1615,1640]" class="Arachnida" family="Idiopidae" genus="Idiosoma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCEFFFCFF3DF9F43945F985" box="[197,306,1615,1640]" italics="true" pageId="1" pageNumber="54">Idiosoma</emphasis>
|
||
</taxonomicName>
|
||
brownish, 385 x 625–645 µm, length/width 1.61–1.66 (n=2). Length of podonotal setae: 42–55 μm (
|
||
<emphasis id="B97A6A3FFFCEFFFCFF28F9CC38AFF97D" box="[208,216,1655,1680]" italics="true" pageId="1" pageNumber="54">j</emphasis>
|
||
1), 25–33 μm (
|
||
<emphasis id="B97A6A3FFFCEFFFCFE74F9CC39E3F97D" box="[396,404,1655,1680]" italics="true" pageId="1" pageNumber="54">j</emphasis>
|
||
2), remaining setae 27–44 μm; opisthonotal setae slightly shorter, 24–39 μm; length of peritrematal groove 190–200 μm.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCEFFFCFF3DF97D3960F819" blockId="1.[151,1437,151,2036]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFF3DF97D397AF932" box="[197,269,1734,1759]" italics="true" pageId="1" pageNumber="54">Ve n t e r</emphasis>
|
||
. Presternal plates moderately sclerotised (
|
||
<figureCitation id="1335AAA8FFCEFFFCFD03F97D3B25F90D" box="[763,850,1734,1760]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="1" pageNumber="54">Fig. 12</figureCitation>
|
||
). Genital lamina (
|
||
<figureCitation id="1335AAA8FFCEFFFCFBDAF97D3CD1F90D" box="[1058,1190,1734,1760]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="1" pageNumber="54">Figs 12, 13</figureCitation>
|
||
) with a hyaline anterior edge and apically rounded lateral wing-like projections, sclerotisation of lamina paired and distinct; subgenital sclerite ribbon-shaped. Anterior sternal margin normally sclerotised, slightly more so along the concavity containing the genital lamina, this concavity flanked by tubercular protrusions (
|
||
<figureCitation id="1335AAA8FFCEFFFCFB2FF8873D46F8BB" box="[1239,1329,1852,1878]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="1" pageNumber="54">Fig. 12</figureCitation>
|
||
). Sternal reticulation inconspicuous, lines running through pores
|
||
<date id="FFB090EDFFCEFFFCFCCCF8DF3B2FF893" box="[820,856,1892,1918]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFCCCF8DF3B3EF890" box="[820,841,1892,1917]" italics="true" pageId="1" pageNumber="54">iv</emphasis>
|
||
2
|
||
</date>
|
||
more pronounced. Gland pores
|
||
<emphasis id="B97A6A3FFFCEFFFCFB27F8DF3C8CF890" box="[1247,1275,1892,1917]" italics="true" pageId="1" pageNumber="54">gv</emphasis>
|
||
1 lie between setae
|
||
<emphasis id="B97A6A3FFFCEFFFCFF22F837388BF848" box="[218,252,1932,1957]" italics="true" pageId="1" pageNumber="54">st3</emphasis>
|
||
. Two pairs of small thickenings located between pores
|
||
<date id="FFB090EDFFCEFFFCFC6EF8373BB6F848" box="[918,961,1931,1957]" pageId="1" pageNumber="54">
|
||
<emphasis id="B97A6A3FFFCEFFFCFC6EF8373BDCF848" box="[918,939,1932,1957]" italics="true" pageId="1" pageNumber="54">iv</emphasis>
|
||
3.
|
||
</date>
|
||
Gland pores
|
||
<emphasis id="B97A6A3FFFCEFFFCFB9BF8373C08F848" box="[1123,1151,1932,1957]" italics="true" pageId="1" pageNumber="54">gv</emphasis>
|
||
<quantity id="4CF61BC8FFCEFFFCFB87F8303CDAF848" box="[1151,1197,1931,1957]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="54" unit="in" value="2.0">2 in</quantity>
|
||
unmodified cuticle. Approximate length of sternal setae: 42 µm (
|
||
<emphasis id="B97A6A3FFFCEFFFCFD5FF8083ACCF821" box="[679,699,1971,1996]" italics="true" pageId="1" pageNumber="54">st</emphasis>
|
||
1), 33 µm (
|
||
<emphasis id="B97A6A3FFFCEFFFCFCB8F8083B24F821" box="[832,851,1971,1996]" italics="true" pageId="1" pageNumber="54">st</emphasis>
|
||
2), 31 µm (
|
||
<emphasis id="B97A6A3FFFCEFFFCFC20F8083B9BF821" box="[984,1004,1971,1996]" italics="true" pageId="1" pageNumber="54">st</emphasis>
|
||
3,
|
||
<emphasis id="B97A6A3FFFCEFFFCFBF1F8083C6BF821" box="[1033,1052,1971,1996]" italics="true" pageId="1" pageNumber="54">st</emphasis>
|
||
4); length of opisthogastric setae 34–39 µm.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCDFFFFFF3DFF233DE6FE27" blockId="2.[151,1437,152,458]" pageId="2" pageNumber="55">
|
||
<taxonomicName id="4C0ECDAEFFCDFFFFFF3DFF23392CFF5C" box="[197,347,152,177]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="55" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCDFFFFFF3DFF23392CFF5C" box="[197,347,152,177]" italics="true" pageId="2" pageNumber="55">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Gnathotectum trispinate as in female. Hypognathal groove (
|
||
<figureCitation id="1335AAA8FFCDFFFFFBCAFF233CFFFF5F" box="[1074,1160,152,178]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="2" pageNumber="55">Fig. 14</figureCitation>
|
||
) with 11 or 12 rows of denticles; palpcoxal setae finely barbed, hypostomatic setae simple, located on area of cuticle isolated by deep incisions; hypostomatic setae 1 on tubercles. Corniculi conical and slim, without any adaxial protrusions. Chelicera with fixed digit straight and relatively narrow, edentate (
|
||
<figureCitation id="1335AAA8FFCDFFFFFC4DFEAB3C79FEC7" box="[949,1038,272,298]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="2" pageNumber="55">Fig. 15</figureCitation>
|
||
). Movable digit with five minute denticles followed proximally by larger tooth. Arthrodial membrane with a prominent brush-like process, synarthrodial membrane acute, ending closely behind the proximal tooth of the movable digit. Palp trochanter (
|
||
<figureCitation id="1335AAA8FFCDFFFFFF67FE333882FE4F" box="[159,245,392,418]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="2" pageNumber="55">Fig. 16</figureCitation>
|
||
) with ventral tubercle placed proximally, ventral setae next to each other: seta
|
||
<date id="FFB090EDFFCDFFFFFB63FE333CC0FE4F" box="[1179,1207,392,418]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97A6A3FFFCDFFFFFB63FE333CDFFE4C" box="[1179,1192,392,417]" italics="true" pageId="2" pageNumber="55">v</emphasis>
|
||
1
|
||
</date>
|
||
simple and located on tubercle, seta
|
||
<date id="FFB090EDFFCDFFFFFEA5FE0B390EFE27" box="[349,377,432,458]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97A6A3FFFCDFFFFFEA5FE0B391DFE24" box="[349,362,432,457]" italics="true" pageId="2" pageNumber="55">v</emphasis>
|
||
2
|
||
</date>
|
||
barbed. Palp femur with small tubercle located ventrally at the level of anterolateral seta.
|
||
</paragraph>
|
||
<caption id="DF71E6A5FFCDFFFFFF6FF8B03AA7F840" httpUri="https://zenodo.org/record/181722/files/figure.png" pageId="2" pageNumber="55" targetBox="[196,1387,525,1729]" targetPageId="2">
|
||
<paragraph id="8BB1B62DFFCDFFFFFF6FF8B03AA7F840" blockId="2.[151,1436,1803,1965]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97A6A3FFFCDFFFFFF6FF8B0393AF8CE" bold="true" box="[151,333,1803,1827]" pageId="2" pageNumber="55">FIGURES 1–11.</emphasis>
|
||
<taxonomicName id="4C0ECDAEFFCDFFFFFEADF8B73AF1F8CE" box="[341,646,1804,1827]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov." subGenus="Heteroparasitus">
|
||
<emphasis id="B97A6A3FFFCDFFFFFEADF8B73A76F8CE" box="[341,513,1804,1827]" italics="true" pageId="2" pageNumber="55">Heteroparasitus</emphasis>
|
||
(
|
||
<emphasis id="B97A6A3FFFCDFFFFFDE9F8B73A53F8CE" box="[529,548,1804,1827]" italics="true" pageId="2" pageNumber="55">H</emphasis>
|
||
.)
|
||
<emphasis id="B97A6A3FFFCDFFFFFDC3F8B73AF1F8CE" box="[571,646,1804,1827]" italics="true" pageId="2" pageNumber="55">mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A6A3FFFCDFFFFFD75F8B03A95F8CE" bold="true" box="[653,738,1803,1827]" pageId="2" pageNumber="55">
|
||
<taxonomicNameLabel id="A249D744FFCDFFFFFD75F8B03A95F8CE" box="[653,738,1803,1827]" pageId="2" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: 1, sternal shield; 2, paragynia; 3, epigynium; 4, endogynium, ventral perspective (intact specimen); 5, endogynium, posterior perspective (squashed specimen); 6, chelicera, adaxial view; 7, movable digit of chelicera; 8, gnathotectum; 9,
|
||
<taxonomicName id="4C0ECDAEFFCDFFFFFCB8F8EA3BB4F885" box="[832,963,1873,1896]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="55" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
, ventral view; 10, trochanter III, ventral; 11, coxa IV and trochanter IV, ventral. (1, 2, 9–11, slide number 2140 (holotype); 3, 4, 7: slide number 2141 (paratype); 5, 6, 8: slide number 2142 (paratype)). Scale bars 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB1B62DFFCCFFFEFF3DFF233899FEBF" blockId="3.[151,1436,152,338]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97A6A3FFFCCFFFEFF3DFF23388BFF5C" box="[197,252,152,177]" italics="true" pageId="3" pageNumber="56">Legs</emphasis>
|
||
. Leg II spurred as follows (
|
||
<figureCitation id="1335AAA8FFCCFFFEFDB9FF233AEFFF5F" box="[577,664,152,178]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="3" pageNumber="56">Fig. 19</figureCitation>
|
||
): main spur on femur conical and slightly curved, axillary process setiform. Spur on genu roundish, spur on tibia wedge-like and pointed distally, ending close to distal margin of segment. On tibia II, anterolateral seta
|
||
<emphasis id="B97A6A3FFFCCFFFEFD7CFF533AECFEEC" box="[644,667,232,257]" italics="true" pageId="3" pageNumber="56">al</emphasis>
|
||
1 not on elevation and posteroventral seta
|
||
<emphasis id="B97A6A3FFFCCFFFEFB69FF533CDBFEEC" box="[1169,1196,232,257]" italics="true" pageId="3" pageNumber="56">pv</emphasis>
|
||
1 larger than that on genu II. Trochanters III and IV with posteroventral tubercles (
|
||
<figureCitation id="1335AAA8FFCCFFFEFC70FEAB3C6AFEC7" box="[904,1053,272,298]" captionStart="FIGURES 12 – 20" captionStartId="3.[152,270,1317,1341]" captionTargetBox="[194,1395,400,1269]" captionTargetId="figure@3.[188,1398,394,1313]" captionTargetPageId="3" captionText="FIGURES 12 – 20. Heteroparasitus (H.) mariae sp. nov., male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral. Heteroparasitus (H.) quadratus (Witaliński), male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15 – 19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="3" pageNumber="56">Figs. 17, 18</figureCitation>
|
||
). Other segments of legs as for female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF71E6A5FFCCFFFEFF60FA9E3A37FA2A" httpUri="https://zenodo.org/record/181723/files/figure.png" pageId="3" pageNumber="56" targetBox="[194,1395,400,1269]" targetPageId="3">
|
||
<paragraph id="8BB1B62DFFCCFFFEFF60FA9E3A37FA2A" blockId="3.[151,1437,1317,1479]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97A6A3FFFCCFFFEFF60FA9E3929FAD0" bold="true" box="[152,350,1317,1341]" pageId="3" pageNumber="56">FIGURES 12–20.</emphasis>
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFE9DFA9E3AE1FAD1" box="[357,662,1317,1340]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov." subGenus="Heteroparasitus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFE9DFA9E3A66FAD1" box="[357,529,1317,1340]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
(
|
||
<emphasis id="B97A6A3FFFCCFFFEFDD9FA9E3A43FAD1" box="[545,564,1317,1340]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.)
|
||
<emphasis id="B97A6A3FFFCCFFFEFDB3FA9E3AE1FAD1" box="[587,662,1317,1340]" italics="true" pageId="3" pageNumber="56">mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A6A3FFFCCFFFEFD66FA9E3A84FAD0" bold="true" box="[670,755,1317,1341]" pageId="3" pageNumber="56">
|
||
<taxonomicNameLabel id="A249D744FFCCFFFEFD66FA9E3A84FAD0" box="[670,755,1317,1341]" pageId="3" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male: 12, sternal shield and genital lamina; 13, genital lamina; 14, corniculus and hypognathal groove; 15, chelicera, antiaxial; 16, palp trochanter, lateral; 17, trochanter III, ventral; 18, trochanter IV, ventral; 19, leg II, femur, genu and tibia, anterolateral.
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFC83FAD03D21FA6F" authority="Witalinski" authorityName="Witalinski" box="[891,1366,1387,1410]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Heteroparasitus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFC83FAD03C50FA6F" box="[891,1063,1387,1410]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
(
|
||
<emphasis id="B97A6A3FFFCCFFFEFBCFFAD03C3DFA6F" box="[1079,1098,1387,1410]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.)
|
||
<emphasis id="B97A6A3FFFCCFFFEFB98FAD03CBBFA6F" box="[1120,1228,1387,1410]" italics="true" pageId="3" pageNumber="56">quadratus</emphasis>
|
||
(Witaliński)
|
||
</taxonomicName>
|
||
, male: 20, leg II, genu and tibia, posteroventral. (12, 13, 15–19: slide no. 2145 (paratype); 14: slide number 2143 (paratype); 20: slide number 2067). Scale bars 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C314E5A6FFCCFFFEFF3EFA413888F9D0" pageId="3" pageNumber="56" type="etymology">
|
||
<paragraph id="8BB1B62DFFCCFFFEFF3EFA413888F9D0" blockId="3.[151,1437,1530,2037]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97A6A3FFFCCFFFEFF3EFA413924F9F9" bold="true" box="[198,339,1530,1556]" pageId="3" pageNumber="56">Etymology.</emphasis>
|
||
This species is named in honour of Maria Florek, who collected and kindly provided the
|
||
<typeStatus id="54B5088FFFCCFFFEFA91FA403DECF9F8" box="[1385,1435,1531,1557]" pageId="3" pageNumber="56">type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314E5A6FFCCFFF9FF3EF9F1398CFDF7" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="discussion">
|
||
<paragraph id="8BB1B62DFFCCFFFEFF3EF9F13CAAF960" blockId="3.[151,1437,1530,2037]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97A6A3FFFCCFFFEFF3EF9F13925F989" bold="true" box="[198,338,1610,1636]" pageId="3" pageNumber="56">Discussion.</emphasis>
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFEA1F9F03A61F989" box="[345,534,1611,1636]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFEA1F9F03A61F989" box="[345,534,1611,1636]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
is a relatively rare European genus of local distribution, inhabiting wet litter or moss in relatively undisturbed deciduous or mixed forests, frequently on limestone terrain.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCCFFFEFF3DF9203BD2F848" blockId="3.[151,1437,1530,2037]" pageId="3" pageNumber="56">
|
||
The revision of
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFE79F9203A48F959" box="[385,575,1691,1716]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFE79F9203A48F959" box="[385,575,1691,1716]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF9FCBDCFFCCFFFEFD94F9203B3BF958" author="Juvara-Bals" box="[620,844,1691,1717]" pageId="3" pageNumber="58" refString="Juvara-Bals, I. (2002) A revision of the genus Heteroparasitus new status, with the description of Heteroparasitus (Medioparasitus) athiasae subgen. n., sp. n. from Spain and with a key to the genera of Pergamasinae (Acari, Gamasida, Parasitidae). Revue Suisse de Zoologie, 109, 23 - 46." type="journal article" year="2002">Juvara-Bals (2002)</bibRefCitation>
|
||
lists characters of
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFBD2F9203C90F959" box="[1066,1255,1691,1716]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B97A6A3FFFCCFFFEFBD2F9203C90F959" box="[1066,1255,1691,1716]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A249D744FFCCFFFEFB16F9203D5BF958" box="[1262,1324,1691,1717]" pageId="3" pageNumber="56" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
based on the three species that were known at that time. The description of
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFC5CF9783C6BF931" box="[932,1052,1731,1757]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B97A6A3FFFCCFFFEFC5CF9783BCEF931" box="[932,953,1731,1756]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFC31F9783C6BF931" box="[969,1052,1731,1756]" italics="true" pageId="3" pageNumber="56">mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A6A3FFFCCFFFEFBDCF9793CF5F931" bold="true" box="[1060,1154,1730,1756]" pageId="3" pageNumber="56">
|
||
<taxonomicNameLabel id="A249D744FFCCFFFEFBDCF9793CF5F931" box="[1060,1154,1730,1756]" pageId="3" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
permits a review of the genus and subgenus diagnoses. At the generic level, the number of teeth on the movable digit of the chelicera in females, and the number of opisthogastric setae, are variable. In
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFC4BF8A83C34F8C1" box="[947,1091,1811,1837]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="tirolensis">
|
||
<emphasis id="B97A6A3FFFCCFFFEFC4BF8A83BBFF8C1" box="[947,968,1811,1836]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFC2FF8A83C34F8C1" box="[983,1091,1811,1836]" italics="true" pageId="3" pageNumber="56">tirolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFB86F8A83D55F8C1" box="[1150,1314,1811,1837]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="coronarius">
|
||
<emphasis id="B97A6A3FFFCCFFFEFB86F8A83CE4F8C1" box="[1150,1171,1811,1836]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFB5BF8A83D55F8C1" box="[1187,1314,1811,1836]" italics="true" pageId="3" pageNumber="56">coronarius</emphasis>
|
||
</taxonomicName>
|
||
, the movable digit has four teeth, whereas in
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFDB1F8803A92F8B9" box="[585,741,1851,1877]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFDB1F8803A29F8B9" box="[585,606,1851,1876]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFD96F8803A92F8B9" box="[622,741,1851,1876]" italics="true" pageId="3" pageNumber="56">quadratus</emphasis>
|
||
</taxonomicName>
|
||
the number of teeth varies - three or more frequently four teeth occur. In
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFEB2F8D839B4F891" box="[330,451,1891,1917]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B97A6A3FFFCCFFFEFEB2F8D83928F891" box="[330,351,1891,1916]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFE88F8D839B4F891" box="[368,451,1891,1916]" italics="true" pageId="3" pageNumber="56">mariae</emphasis>
|
||
</taxonomicName>
|
||
, the number of teeth can also vary even on both chelicerae of the same specimen. This diagnostic character must therefore be considered as variable.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCCFFFEFF3DF8083DECF818" blockId="3.[151,1437,1530,2037]" pageId="3" pageNumber="56">
|
||
The typical number of opisthogastric setae in
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFD25F8083BEDF821" box="[733,922,1971,1996]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFD25F8083BEDF821" box="[733,922,1971,1996]" italics="true" pageId="3" pageNumber="56">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
is seven pairs. This character is stable in
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFA87F8083974F819" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="tirolensis">
|
||
<emphasis id="B97A6A3FFFCCFFFEFA87F8083DE3F821" box="[1407,1428,1971,1996]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFF6FF8603974F819" box="[151,259,2011,2036]" italics="true" pageId="3" pageNumber="56">tirolensis</emphasis>
|
||
</taxonomicName>
|
||
but variable in
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFE41F8603A47F819" box="[441,560,2011,2037]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B97A6A3FFFCCFFFEFE41F86039B9F819" box="[441,462,2011,2036]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFE25F8603A47F819" box="[477,560,2011,2036]" italics="true" pageId="3" pageNumber="56">mariae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFD90F8603B74F819" box="[616,771,2011,2037]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="B97A6A3FFFCCFFFEFD90F8603A0AF819" box="[616,637,2011,2036]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFD74F8603B74F819" box="[652,771,2011,2036]" italics="true" pageId="3" pageNumber="56">quadratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C0ECDAEFFCCFFFEFCE9F8603BC3F819" box="[785,948,2011,2037]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="coronarius">
|
||
<emphasis id="B97A6A3FFFCCFFFEFCE9F8603B51F819" box="[785,806,2011,2036]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCCFFFEFCCCF8603BC3F819" box="[820,948,2011,2036]" italics="true" pageId="3" pageNumber="56">coronarius</emphasis>
|
||
</taxonomicName>
|
||
has not been studied in this respect. In
|
||
<emphasis id="B97A6A3FFFCCFFFEFA87F8603DE3F819" box="[1407,1428,2011,2036]" italics="true" pageId="3" pageNumber="56">H</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCBFFF9FF6FFF23394FFF37" blockId="4.[151,1436,150,538]" pageId="4" pageNumber="57">
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FF6FFF233979FF5C" box="[151,270,152,177]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="B97A6A3FFFCBFFF9FF6FFF233979FF5C" box="[151,270,152,177]" italics="true" pageId="4" pageNumber="57">quadratus</emphasis>
|
||
</taxonomicName>
|
||
, females showing a complete set of eight pairs of opisthogastric setae can also be found (Witaliński unpublished).
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCBFFF9FF3DFF533CC2FE4F" blockId="4.[151,1436,150,538]" pageId="4" pageNumber="57">
|
||
The diagnostic definition of the male corniculi should also be modified. The corniculi of males may possess adaxial protrusions as in
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FE05FEAB3AF8FEC4" box="[509,655,272,298]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="tirolensis">
|
||
<emphasis id="B97A6A3FFFCBFFF9FE05FEAB3A65FEC4" box="[509,530,272,297]" italics="true" pageId="4" pageNumber="57">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCBFFF9FDDBFEAB3AF8FEC4" box="[547,655,272,297]" italics="true" pageId="4" pageNumber="57">tirolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FD34FEAB3B05FEC4" box="[716,882,272,298]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="coronarius">
|
||
<emphasis id="B97A6A3FFFCBFFF9FD34FEAB3A96FEC4" box="[716,737,272,297]" italics="true" pageId="4" pageNumber="57">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCBFFF9FD0BFEAB3B05FEC4" box="[755,882,272,297]" italics="true" pageId="4" pageNumber="57">coronarius</emphasis>
|
||
</taxonomicName>
|
||
, or the corniculi are conical with either indiscernible or weakly visible protrusions, as in
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FD54FE833B3FFEBC" box="[684,840,312,338]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="B97A6A3FFFCBFFF9FD54FE833AB6FEBC" box="[684,705,312,337]" italics="true" pageId="4" pageNumber="57">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCBFFF9FD29FE833B3FFEBC" box="[721,840,312,337]" italics="true" pageId="4" pageNumber="57">quadratus</emphasis>
|
||
</taxonomicName>
|
||
. The corniculi of
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FBDBFE833CECFEBC" box="[1059,1179,312,338]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B97A6A3FFFCBFFF9FBDBFE833C4FFEBC" box="[1059,1080,312,337]" italics="true" pageId="4" pageNumber="57">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCBFFF9FBB0FE833CECFEBC" box="[1096,1179,312,337]" italics="true" pageId="4" pageNumber="57">mariae</emphasis>
|
||
</taxonomicName>
|
||
males are even more slender and devoid of any extensions or protuberances on their adaxial sides. Consequently, the corniculi in the subgenus
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FECEFE333984FE4C" box="[310,499,392,417]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCBFFF9FECEFE333984FE4C" box="[310,499,392,417]" italics="true" pageId="4" pageNumber="57">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
can be defined as either conical or with adaxial protrusions.
|
||
</paragraph>
|
||
<paragraph id="8BB1B62DFFCBFFF9FF3DFE0B398CFDF7" blockId="4.[151,1436,150,538]" pageId="4" pageNumber="57">
|
||
Other taxonomic characters for
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FDC2FE0B3A80FE24" box="[570,759,432,457]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCBFFF9FDC2FE0B3A80FE24" box="[570,759,432,457]" italics="true" pageId="4" pageNumber="57">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
at the generic and subgeneric levels remain as defined by
|
||
<bibRefCitation id="EF9FCBDCFFCBFFF9FF6FFE63390BFE1F" author="Juvara-Bals" box="[151,380,472,498]" pageId="4" pageNumber="58" refString="Juvara-Bals, I. (2002) A revision of the genus Heteroparasitus new status, with the description of Heteroparasitus (Medioparasitus) athiasae subgen. n., sp. n. from Spain and with a key to the genera of Pergamasinae (Acari, Gamasida, Parasitidae). Revue Suisse de Zoologie, 109, 23 - 46." type="journal article" year="2002">Juvara-Bals (2002)</bibRefCitation>
|
||
. Chaetotaxy, poroidotaxy and adenotaxy in
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FC7FFE633B88FE1C" box="[903,1023,472,498]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B97A6A3FFFCBFFF9FC7FFE633BEBFE1C" box="[903,924,472,497]" italics="true" pageId="4" pageNumber="57">H</emphasis>
|
||
.
|
||
<emphasis id="B97A6A3FFFCBFFF9FC54FE633B88FE1C" box="[940,1023,472,497]" italics="true" pageId="4" pageNumber="57">mariae</emphasis>
|
||
</taxonomicName>
|
||
are in accord with those given for the
|
||
<taxonomicName id="4C0ECDAEFFCBFFF9FF3AFDBB3908FDF4" box="[194,383,512,537]" class="Arachnida" family="Parasitidae" genus="Heteroparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A6A3FFFCBFFF9FF3AFDBB3908FDF4" box="[194,383,512,537]" italics="true" pageId="4" pageNumber="57">Heteroparasitus</emphasis>
|
||
</taxonomicName>
|
||
subgenus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |