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<document id="A1C8B6EAB9D646CF65F5266890D8A0C9" ID-CLB-Dataset="30691" ID-DOI="10.11646/zootaxa.4382.2.3" ID-GBIF-Dataset="6dc55986-74fa-4ac0-bca6-b301f3831180" ID-ISSN="1175-5326" ID-Zenodo-Dep="1182167" ID-ZooBank="8E248E6F-FDA5-4829-9DD7-4CC0EFDB08B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519199964901" checkinUser="plazi" docAuthor="Okamoto, Makoto &amp; Gon, Ofer" docDate="2018" docId="03A2BF386E7F407AFF185A7615989BD7" docLanguage="en" docName="zootaxa.4382.2.3.pdf" docOrigin="Zootaxa 4382 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigonus marisrubri Krupp, Zajons &amp; Khalaf 2009" docType="treatment" docVersion="7" lastPageNumber="280" masterDocId="FF9BC7406E6D4069FF8F5C1A170D9F12" masterDocTitle="A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species" masterLastPageNumber="291" masterPageNumber="261" pageNumber="279" updateTime="1698523467852" updateUser="ExternalLinkService">
<mods:mods id="015FE23196FE2EF58D10B31E2C53266A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A373D15B4F5E3C0BC6B28DA71A7A3DC9">A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species</mods:title>
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<mods:name id="B245385C76277B76AB11F4449C5B2652" type="personal">
<mods:role id="31BFA8A85495296F20CB0DEDC5CD0067">
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<mods:namePart id="066741FCD69788644486235F9750E027">Okamoto, Makoto</mods:namePart>
</mods:name>
<mods:name id="0D3E38B64E128328912F734670ED9EEB" type="personal">
<mods:role id="A35E63F6C5A03A36954C7C1D78ECF1BD">
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</mods:role>
<mods:namePart id="E3BF8F7439B4AFF763DA1161A8B740B3">Gon, Ofer</mods:namePart>
</mods:name>
<mods:typeOfResource id="494265A7FE62AB2EFECA9CD3B31CBAF6">text</mods:typeOfResource>
<mods:relatedItem id="640A6AEFEC24D4C0538ACBFB2B8C7239" type="host">
<mods:titleInfo id="FE9C0B8A28D80D036C6821911A5C9691">
<mods:title id="C6620CC770384B78E6D391FD5BF9EFAD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41F3BA09F80521529E201B9C89FD4A3F">
<mods:date id="B7B8A56F17DFE91A67FA4E86528711F2">2018</mods:date>
<mods:detail id="EF3C076BFE46885CC96F1F1764F519F8" type="pubDate">
<mods:number id="DAF14A78B17C9489A2FAC4B4244D0686">2018-02-21</mods:number>
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<mods:detail id="8339DAC4F847E09290FF549B410201D1" type="volume">
<mods:number id="FEF3E3868BC3F4DB259C3EB534DA675A">4382</mods:number>
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<mods:number id="44B77A286CAB9731FAC76A001876BE72">2</mods:number>
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<mods:start id="1A406D90687175121603F1B3C3B46E4A">261</mods:start>
<mods:end id="033B9BD22D9F2E43EEF8886F33483E7A">291</mods:end>
</mods:extent>
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<mods:classification id="2F8F4B38C5B80449E42DA811055AB7F9">journal article</mods:classification>
<mods:identifier id="8E4CC37D1407FB9AC01C81B731369214" type="CLB-Dataset">30691</mods:identifier>
<mods:identifier id="CA8B4D5F5BB03940B66250268799D81D" type="DOI">10.11646/zootaxa.4382.2.3</mods:identifier>
<mods:identifier id="40E9E3A654C1D1BB1CC2A475D8B425BE" type="GBIF-Dataset">6dc55986-74fa-4ac0-bca6-b301f3831180</mods:identifier>
<mods:identifier id="1ED3C072175D24F0FC1F5891D59E738A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BB9EC98487F82FA95DDE725CCDCDB16C" type="Zenodo-Dep">1182167</mods:identifier>
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<treatment id="03A2BF386E7F407AFF185A7615989BD7" ID-DOI="http://doi.org/10.5281/zenodo.5998255" ID-GBIF-Taxon="141118055" ID-Zenodo-Dep="5998255" LSID="urn:lsid:plazi:treatment:03A2BF386E7F407AFF185A7615989BD7" httpUri="http://treatment.plazi.org/id/03A2BF386E7F407AFF185A7615989BD7" lastPageId="19" lastPageNumber="280" pageId="18" pageNumber="279">
<subSubSection id="C3115DA56E7F407BFF185A7616A499D9" pageId="18" pageNumber="279" type="nomenclature">
<paragraph id="8BB40E2E6E7F407BFF185A7614119994" blockId="18.[151,796,1644,1739]" box="[151,796,1644,1670]" pageId="18" pageNumber="279">
<heading id="D0FCB9426E7F407BFF185A7614119994" bold="true" box="[151,796,1644,1670]" fontSize="11" level="1" pageId="18" pageNumber="279" reason="1">
<taxonomicName id="4C0B75AD6E7F407BFF185A7614119994" ID-CoL="6G5LB" authority="Krupp, Zajons &amp; Khalaf, 2009" authorityName="Krupp, Zajons &amp; Khalaf" authorityYear="2009" box="[151,796,1644,1670]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="279" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7F407BFF185A7614119994" bold="true" box="[151,796,1644,1670]" pageId="18" pageNumber="279">
<emphasis id="B97FD23C6E7F407BFF185A7616989994" bold="true" box="[151,405,1644,1670]" italics="true" pageId="18" pageNumber="279">Epigonus marisrubri</emphasis>
Krupp, Zajons &amp;
<bibRefCitation id="EF9A73DF6E7F407BFDF35A7614119994" box="[636,796,1644,1670]" pageId="18" pageNumber="279" refId="ref19794" refString="Krupp, F., Zajonz, U. &amp; Khalaf, M. A. (2009) A new species of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) from the Gulf of Aqaba, Red Sea. Aqua, 15, 223 - 227." type="journal article">Khalaf, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB40E2E6E7F407BFF185A9516A499D9" blockId="18.[151,796,1644,1739]" pageId="18" pageNumber="279">
(New English name: Red Sea Deepwater Cardinalfish) (
<figureCitation id="133012AB6E7F407BFF105AA817E399D9" box="[159,238,1714,1739]" captionStart="FIGURE 2" captionStartId="2.[151,250,915,937]" captionTargetBox="[331,1199,202,881]" captionTargetId="figure@2.[302,1284,193,894]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of three species of the Epigonus pandionis group in the Western Indian Ocean." httpUri="https://zenodo.org/record/1182171/files/figure.png" pageId="18" pageNumber="279">Figs. 2</figureCitation>
,
<figureCitation id="133012AB6E7F407BFF755AA8161499D9" box="[250,281,1714,1739]" captionStart="FIGURE 15" captionStartId="19.[151,250,1661,1683]" captionTargetBox="[190,1398,1276,1640]" captionTargetId="figure@19.[188,1398,1276,1640]" captionTargetPageId="19" captionText="FIGURE 15. Epigonus marisrubri, SMF 24661, holotype, 136 mm SL." httpUri="https://zenodo.org/record/1182199/files/figure.png" pageId="18" pageNumber="279">15</figureCitation>
; Tables 1, 2)
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E7F407BFF185AE6150A985C" pageId="18" pageNumber="279" type="reference_group">
<paragraph id="8BB40E2E6E7F407BFF185AE6150A985C" blockId="18.[151,1436,1787,1870]" pageId="18" pageNumber="279">
<treatmentCitationGroup id="AB1B29006E7F407BFF185AE6150A985C" infered="true" pageId="18" pageNumber="279">
<taxonomicName id="4C0B75AD6E7F407BFF185AE615E89803" ID-CoL="6G5LB" authority="Krupp, Zajonz &amp; Khalaf, 2009: 224" authorityName="Krupp, Zajonz &amp; Khalaf" authorityPageNumber="224" authorityYear="2009" box="[151,741,1787,1809]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="279" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7F407BFF185AE616669803" box="[151,363,1787,1809]" italics="true" pageId="18" pageNumber="279">Epigonus marisrubri</emphasis>
<treatmentCitation id="0AAA283F6E7F407BFEFC5AE615E89803" author="" box="[371,741,1787,1809]" page="224" pageId="18" pageNumber="279">
<bibRefCitation id="EF9A73DF6E7F407BFEFC5AE615BF9803" box="[371,690,1787,1809]" pageId="18" pageNumber="279" refId="ref19794" refString="Krupp, F., Zajonz, U. &amp; Khalaf, M. A. (2009) A new species of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) from the Gulf of Aqaba, Red Sea. Aqua, 15, 223 - 227." type="journal article">Krupp, Zajonz &amp; Khalaf, 2009</bibRefCitation>
: 224
</treatmentCitation>
</taxonomicName>
, fig. 1 (original description; type locality: ca. 29°27.30 N, 34°58.30 E, Northern of Gulf of Aqaba, Red Sea, Jordan,
<quantity id="4CF3A3CB6E7F407BFD325B0315FF983D" box="[701,754,1817,1839]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="18" pageNumber="279" unit="m" value="52.0">52 m</quantity>
depth; holotype: SMF 24661);
<treatmentCitation id="0AAA283F6E7F407BFBB65B031291983D" author="" box="[1081,1436,1817,1839]" page="307" pageId="18" pageNumber="279">
<bibRefCitation id="EF9A73DF6E7F407BFBB65B031267983D" box="[1081,1386,1817,1839]" pageId="18" pageNumber="279" refId="ref21226" refString="Okamoto, M. &amp; Aungtonya, C. (2013) Re-examination of the holotype of Epigonus thai Prokofiev and Bussarawit: redescription and comparison with related species (Teleostei: Perciformes: Epigonidae). Phuket Marine Biological Center Research Bulletin, 72, 49 - 53." type="journal article">Okamoto &amp; Motomura, 2013</bibRefCitation>
: 307
</treatmentCitation>
(record and key, Gulf of Aden).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E7F407AFF185B60165E9FC6" lastPageId="19" lastPageNumber="280" pageId="18" pageNumber="279" type="materials_examined">
<paragraph id="8BB40E2E6E7F407AFF185B60165E9FC6" blockId="18.[151,1437,1914,2011]" lastBlockId="19.[151,1436,151,1221]" lastPageId="19" lastPageNumber="280" pageId="18" pageNumber="279">
<materialsCitation id="3B6304736E7F407BFF185B6015EF98A5" ID-GBIF-Occurrence="1820370030" collectingDate="1990-05-02" collectionCode="SMF" country="Jordan" elevation="52" latitude="29.455" location="Material" longLatPrecision="12" longitude="34.97167" municipality="Red Sea" pageId="18" pageNumber="279" specimenCount="1" stateProvince="Aqaba" typeStatus="holotype">
<emphasis id="B97FD23C6E7F407BFF185B60168C9881" bold="true" box="[151,385,1914,1939]" pageId="18" pageNumber="279">
<location id="8ED458F56E7F407BFF185B6017F29881" LSID="urn:lsid:plazi:treatment:03A2BF386E7F407AFF185A7615989BD7:8ED458F56E7F407BFF185B6017F29881" box="[151,255,1914,1939]" country="Jordan" latitude="29.455" longLatPrecision="12" longitude="34.97167" municipality="Red Sea" name="Material" pageId="18" pageNumber="279" stateProvince="Aqaba">Material</location>
examined.
</emphasis>
<collectionCode id="ED1A96EB6E7F407BFE075B6116CD9881" box="[392,448,1915,1939]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="279">SMF</collectionCode>
24661,
<typeStatus id="54B0B08C6E7F407BFD935B60158C9881" box="[540,641,1914,1939]" pageId="18" pageNumber="279" type="holotype">holotype</typeStatus>
, female,
<quantity id="4CF3A3CB6E7F407BFD685B6014499880" box="[743,836,1914,1939]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="18" pageNumber="279" unit="mm" value="136.0">136 mm</quantity>
SL, ca.
<geoCoordinate id="EE3F68E96E7F407BFC2E5B61132B9880" box="[929,1062,1915,1939]" direction="north" orientation="latitude" pageId="18" pageNumber="279" precision="9" value="29.455">29°27.30 N</geoCoordinate>
,
<geoCoordinate id="EE3F68E96E7F407BFBBE5B6013BE9880" box="[1073,1203,1914,1939]" direction="east" orientation="longitude" pageId="18" pageNumber="279" precision="9" value="34.97167">34°58.30 E</geoCoordinate>
, Gulf of
<collectingRegion id="49CFC0CC6E7F407BFA975B6012699881" box="[1304,1380,1914,1939]" country="Jordan" name="Al 'Aqabah" pageId="18" pageNumber="279">Aqaba</collectingRegion>
,
<collectingMunicipality id="6BD094546E7F407BFAE05B6017CF98A5" pageId="18" pageNumber="279">Red Sea</collectingMunicipality>
,
<collectingCountry id="F31C4EBE6E7F407BFF405B84161098A5" box="[207,285,1950,1975]" name="Jordan" pageId="18" pageNumber="279">Jordan</collectingCountry>
,
<elevation id="0026E91D6E7F407BFEA65B84166E98A5" box="[297,355,1950,1975]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="18" pageNumber="279" unit="m" value="52.0">
<quantity id="4CF3A3CB6E7F407BFEA65B84166E98A5" box="[297,355,1950,1975]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="18" pageNumber="279" unit="m" value="52.0">52 m</quantity>
</elevation>
depth,
<collectingDate id="EFF1D1066E7F407BFE365B85154E98A5" box="[441,579,1951,1976]" pageId="18" pageNumber="279" value="1990-05-02">
<date id="FFB528EE6E7F407BFE365B85154E98A5" box="[441,579,1951,1976]" pageId="18" pageNumber="279" value="1990-05-02">2 May 1990</date>
</collectingDate>
(photograph)
</materialsCitation>
;
<materialsCitation id="3B6304736E7F407BFD615B85129598A5" ID-GBIF-Occurrence="1820370024" box="[750,1432,1950,1976]" collectingDate="1990-05-02" collectionCode="SMF" country="Jordan" elevation="52" latitude="29.455" location="Material" longLatPrecision="12" longitude="34.97167" municipality="Red Sea" pageId="18" pageNumber="279" specimenCount="1" stateProvince="Aqaba" typeStatus="paratype">
<collectionCode id="ED1A96EB6E7F407BFD615B85142B98A5" box="[750,806,1951,1975]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="279">SMF</collectionCode>
31636,
<typeStatus id="54B0B08C6E7F407BFC0B5BBA14E598AA" box="[900,1000,1952,1976]" pageId="18" pageNumber="279" type="paratype">paratype</typeStatus>
,
<quantity id="4CF3A3CB6E7F407BFC7B5B85135E98A5" box="[1012,1107,1951,1975]" metricMagnitude="-1" metricUnit="m" metricValue="1.34" pageId="18" pageNumber="279" unit="mm" value="134.0">134 mm</quantity>
SL, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B6304736E7F407AFF185BD916D59FA2" ID-GBIF-Occurrence="1820370039" collectingDate="1964-12-17" collectionCode="USNM" country="Somalia" elevation="125" lastPageId="19" lastPageNumber="280" location="western South Indian Ocean" pageId="18" pageNumber="279" specimenCode="USNM 269827" specimenCount="2" stateProvince="South" typeStatus="paratype">
<specimenCode id="DBADA6556E7F407BFF185BD9164B98C9" box="[151,326,1986,2011]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="279">USNM 269827</specimenCode>
,
<specimenCount id="9D0DC5A76E7F407BFEDF5BD916D198C9" box="[336,476,1986,2011]" pageId="18" pageNumber="279" type="generic">2 specimens</specimenCount>
,
<quantity id="4CF3A3CB6E7F407BFE675BD9158698C8" box="[488,651,1987,2011]" metricMagnitude="-2" metricUnit="m" metricValue="8.635" metricValueMax="8.84" metricValueMin="8.43" pageId="18" pageNumber="279" unit="mm" value="86.35" valueMax="88.4" valueMin="84.3">84.3–88.4 mm</quantity>
SL, 11°24 N, 51°35 E,
<collectingCountry id="F31C4EBE6E7F407BFC1B5BD814FE98C9" box="[916,1011,1986,2011]" name="Somalia" pageId="18" pageNumber="279">Somalia</collectingCountry>
,
<location id="8ED458F56E7F407BFC715BD9123098C9" LSID="urn:lsid:plazi:treatment:03A2BF386E7F407AFF185A7615989BD7:8ED458F56E7F407BFC715BD9123098C9" box="[1022,1341,1986,2011]" country="Somalia" name="western South Indian Ocean" pageId="18" pageNumber="279" stateProvince="South">
western
<collectingRegion id="49CFC0CC6E7F407BFBD35BD8139098C9" box="[1116,1181,1986,2011]" country="Tanzania" name="Kusini Unguja" pageId="18" pageNumber="279">South</collectingRegion>
Indian Ocean
</location>
,
<elevation id="0026E91D6E7F407AFAC75BD817A09FBD" lastPageId="19" lastPageNumber="280" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.75" metricValueMin="0.75" pageId="18" pageNumber="279" unit="m" value="125.0" valueMax="175.0" valueMin="75.0">
<quantity id="4CF3A3CB6E7F407AFAC75BD817A09FBD" lastPageId="19" lastPageNumber="280" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.75" metricValueMin="0.75" pageId="18" pageNumber="279" unit="m" value="125.0" valueMax="175.0" valueMin="75.0">75–175 m</quantity>
</elevation>
depth,
<collectingDate id="EFF1D1066E7E407AFE8F5C8216D59FA2" box="[256,472,151,176]" pageId="19" pageNumber="280" value="1964-12-17">
<date id="FFB528EE6E7E407AFE8F5C8216D59FA2" box="[256,472,151,176]" pageId="19" pageNumber="280" value="1964-12-17">17 December 1964</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B6304736E7E407AFE6C5C82165E9FC6" ID-GBIF-Occurrence="1820369953" collectingDate="1989-11-07" collectionCode="USNM" country="Somalia" elevation="150" location="Gulf" pageId="19" pageNumber="280" specimenCode="USNM 305950, 139.8" specimenCount="1" stateProvince="Gulf" typeStatus="paratype">
<specimenCode id="DBADA6556E7E407AFE6C5C82159F9FA2" box="[483,658,151,176]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="280">
USNM
<quantity id="4CF3A3CB6E7E407AFDB45C8D159F9FA2" box="[571,658,151,176]" metricMagnitude="5" metricUnit="m" metricValue="3.0595013979999997" pageId="19" pageNumber="280" unit="mm" value="3.059501398E8">305950</quantity>
</specimenCode>
,
<specimenCode id="DBADA6556E7E407AFD125C8215D19FA2" box="[669,732,152,176]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="280">139.8</specimenCode>
mm SL, 12°03 N, 50°54 E,
<collectingRegion id="49CFC0CC6E7E407AFB955C8D13409FA2" box="[1050,1101,151,176]" country="Papua New Guinea" name="Gulf" pageId="19" pageNumber="280">Gulf</collectingRegion>
of Aden,
<elevation id="0026E91D6E7E407AFB335C8212369FBD" box="[1212,1339,152,176]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="19" pageNumber="280" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<quantity id="4CF3A3CB6E7E407AFB335C8212369FBD" box="[1212,1339,152,176]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="19" pageNumber="280" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</quantity>
</elevation>
depth,
<collectingDate id="EFF1D1066E7E407AFA015C8216429FC6" pageId="19" pageNumber="280" value="1989-11-07">
<date id="FFB528EE6E7E407AFA015C8216429FC6" pageId="19" pageNumber="280" value="1989-11-07">7 November 1989</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E7E407AFF485CC515B59E77" pageId="19" pageNumber="280" type="diagnosis">
<paragraph id="8BB40E2E6E7E407AFF485CC515B59E77" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="280">
<emphasis id="B97FD23C6E7E407AFF485CC5164C9FEA" bold="true" box="[199,321,223,248]" pageId="19" pageNumber="280">Diagnosis.</emphasis>
Dorsal-fin rays VII-I, 10; pectoral-fin rays 17–18; total gill rakers 28–30; vertebrae 10 + 15; pyloric caeca 9–10; pored lateral-line scales 47–49 + 3; scales below lateral line 9–10; opercular spine absent; maxillary mustache-like processes absent; ribs on last abdominal vertebra present; pectoral fin reaching level of origin of second dorsal fin; tooth patch present on tongue.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E7E407AFF485D7514299DDF" pageId="19" pageNumber="280" type="description">
<paragraph id="8BB40E2E6E7E407AFF485D7514299DDF" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="280">
<emphasis id="B97FD23C6E7E407AFF485D7516D19E9A" bold="true" box="[199,476,367,392]" pageId="19" pageNumber="280">Measurements (% SL)</emphasis>
; counts are given in Table 1. Head length 31.9–36.8; head width 16.1–18.3; head height 17.4–18.5; body depth 20.3–24.5; body width 13.2–17.6; caudal-peduncle depth 8.3.5–10.7; caudalpeduncle length 25.0–27.7; orbital diameter 11.6–15.1; interorbital width 8.4–8.8; postorbital length 13.6–16.4; upper-jaw length 14.4–15.9; lower-jaw length 17.4–18.5; snout length 7.5–8.7; pre-first dorsal-fin length 38.2– 40.8; pre-second dorsal-fin length 59.6–62.9; pre-pectoral-fin length 32.1–39.3; pre-pelvic-fin length 34.3–41.3; pre-anus length 59.7–63.2; pre-anal-fin length 65.0–68.9; first spine length on first dorsal fin 5.0–7.5; second spine length on first dorsal fin 13.3–15.9; third spine length on first dorsal fin 14.1–17.0; second dorsal-fin spine length 4.6–5.8; first anal-fin spine length 2.1–2.4; second anal-fin spine length 5.3–5.9; pelvic-fin spine length 6.9–9.9; first dorsal-fin base length 11.6–15.4; second dorsal-fin base length 10.7–11.5; anal-fin base length 10.2–10.9; pectoral-fin length 19.0–26.0; pelvic-fin length 11.6–17.1.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E7E407AFF485ECD15989BD7" pageId="19" pageNumber="280" type="distribution">
<paragraph id="8BB40E2E6E7E407AFF485ECD16C89C07" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="280">
<emphasis id="B97FD23C6E7E407AFF485ECD166D9DE2" bold="true" box="[199,352,727,752]" pageId="19" pageNumber="280">Distribution.</emphasis>
Red Sea (
<bibRefCitation id="EF9A73DF6E7E407AFE535EC215BC9DE2" box="[476,689,728,752]" pageId="19" pageNumber="280" refId="ref19794" refString="Krupp, F., Zajonz, U. &amp; Khalaf, M. A. (2009) A new species of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) from the Gulf of Aqaba, Red Sea. Aqua, 15, 223 - 227." type="journal article">
Krupp
<emphasis id="B97FD23C6E7E407AFDA05EC3156E9DE2" box="[559,611,728,752]" italics="true" pageId="19" pageNumber="280">et al</emphasis>
. 2009
</bibRefCitation>
);
<collectingRegion id="49CFC0CC6E7E407AFD475ECD15F69DE2" box="[712,763,727,752]" country="Papua New Guinea" name="Gulf" pageId="19" pageNumber="280">Gulf</collectingRegion>
of Aden (
<bibRefCitation id="EF9A73DF6E7E407AFCFA5ECD13C89DE2" box="[885,1221,727,752]" pageId="19" pageNumber="280" refId="ref21226" refString="Okamoto, M. &amp; Aungtonya, C. (2013) Re-examination of the holotype of Epigonus thai Prokofiev and Bussarawit: redescription and comparison with related species (Teleostei: Perciformes: Epigonidae). Phuket Marine Biological Center Research Bulletin, 72, 49 - 53." type="journal article">Okamoto &amp; Motomura 2013</bibRefCitation>
),
<collectingCountry id="F31C4EBE6E7E407AFB535ECD12359DE2" box="[1244,1336,727,752]" name="Somalia" pageId="19" pageNumber="280">Somalia</collectingCountry>
(present study); at
<quantity id="4CF3A3CB6E7E407AFE885EE616759C06" box="[263,376,764,789]" metricMagnitude="2" metricUnit="m" metricValue="1.26" metricValueMax="2.0" metricValueMin="0.52" pageId="19" pageNumber="280" unit="m" value="126.0" valueMax="200.0" valueMin="52.0">52–200 m</quantity>
depth.
</paragraph>
<paragraph id="8BB40E2E6E7E407AFF485F0514D19B4A" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="280">
<emphasis id="B97FD23C6E7E407AFF485F05151B9C2A" bold="true" box="[199,534,799,824]" pageId="19" pageNumber="280">Comparisons and Remarks.</emphasis>
<taxonomicName id="4C0B75AD6E7E407AFD925F3A14059C2A" authorityName="Krupp, Zajonz &amp; Khalaf" authorityYear="2009" box="[541,776,800,824]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7E407AFD925F3A14059C2A" box="[541,776,800,824]" italics="true" pageId="19" pageNumber="280">Epigonus marisrubri</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C0B75AD6E7E407AFC395F3B13499C2A" baseAuthorityName="Goode &amp; Bean" baseAuthorityYear="1881" box="[950,1092,800,824]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="pandionis">
<emphasis id="B97FD23C6E7E407AFC395F3B13499C2A" box="[950,1092,800,824]" italics="true" pageId="19" pageNumber="280">E. pandionis</emphasis>
</taxonomicName>
group in having more than 43 pored lateral-line scales and lacking an opercular spine and an isolated dorsal fin spine between first and second dorsal fins (
<bibRefCitation id="EF9A73DF6E7E407AFE955F7D156C9C92" box="[282,609,871,896]" pageId="19" pageNumber="280" refId="ref21226" refString="Okamoto, M. &amp; Aungtonya, C. (2013) Re-examination of the holotype of Epigonus thai Prokofiev and Bussarawit: redescription and comparison with related species (Teleostei: Perciformes: Epigonidae). Phuket Marine Biological Center Research Bulletin, 72, 49 - 53." type="journal article">Okamoto &amp; Motomura 2013</bibRefCitation>
). This species is similar to
<taxonomicName id="4C0B75AD6E7E407AFC025F7314F09C92" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[909,1021,872,896]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E7E407AFC025F7314F09C92" box="[909,1021,872,896]" italics="true" pageId="19" pageNumber="280">E. exodon</emphasis>
</taxonomicName>
by having tooth patch on the tongue; however, it differs in having 47–49 pored lateral-line scales and 29–30 total gill rakers (vs. 35–37 pored lateral-line scales and 26–28 total gill rakers in
<taxonomicName id="4C0B75AD6E7E407AFDA95FAB15989CDA" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[550,661,944,968]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E7E407AFDA95FAB15989CDA" box="[550,661,944,968]" italics="true" pageId="19" pageNumber="280">E. exodon</emphasis>
</taxonomicName>
), and in lacking anteriorly projecting teeth on the symphysis of lower jaw (vs. present in
<taxonomicName id="4C0B75AD6E7E407AFEE45FCF16D69CFE" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[363,475,980,1004]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E7E407AFEE45FCF16D69CFE" box="[363,475,980,1004]" italics="true" pageId="19" pageNumber="280">E. exodon</emphasis>
</taxonomicName>
,
<figureCitation id="133012AB6E7E407AFE665FCE15369CFE" box="[489,571,980,1005]" captionStart="FIGURE 10" captionStartId="12.[151,250,1827,1849]" captionTargetBox="[188,1398,926,1805]" captionTargetId="figure@12.[188,1398,901,1805]" captionTargetPageId="12" captionText="FIGURE 10. Epigonus exodon, MNHN 1991-0155, holotype, 97.0 mm SL. Overall body (A); and dorsal (B) and ventral (C) views of tipof the lower jaw, arrows show the anteriorly projecting teeth." httpUri="https://zenodo.org/record/1182187/files/figure.png" pageId="19" pageNumber="280">Fig. 10</figureCitation>
).
<taxonomicName id="4C0B75AD6E7E407AFDDF5FCF14269CFE" authorityName="Mayer" authorityYear="1974" box="[592,811,980,1004]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="pectinifer">
<emphasis id="B97FD23C6E7E407AFDDF5FCF14269CFE" box="[592,811,980,1004]" italics="true" pageId="19" pageNumber="280">Epigonus pectinifer</emphasis>
</taxonomicName>
also has a slender tooth patch on the tongue, but can be distinguished from
<taxonomicName id="4C0B75AD6E7E407AFEFD5FE315069B02" authorityName="Krupp, Zajonz &amp; Khalaf" authorityYear="2009" box="[370,523,1016,1040]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7E407AFEFD5FE315069B02" box="[370,523,1016,1040]" italics="true" pageId="19" pageNumber="280">E. marisrubri</emphasis>
</taxonomicName>
in having a pungent opercular spine and 9 soft rays on the second dorsal fin (vs. opercular spine absent and 10 soft rays on the second dorsal fin in
<taxonomicName id="4C0B75AD6E7E407AFC2E580713319B26" authorityName="Krupp, Zajonz &amp; Khalaf" authorityYear="2009" box="[929,1084,1052,1076]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7E407AFC2E580713319B26" box="[929,1084,1052,1076]" italics="true" pageId="19" pageNumber="280">E. marisrubri</emphasis>
</taxonomicName>
;
<figureCitation id="133012AB6E7E407AFBC2580613AE9B27" box="[1101,1187,1052,1077]" captionStart="FIGURE 15" captionStartId="19.[151,250,1661,1683]" captionTargetBox="[190,1398,1276,1640]" captionTargetId="figure@19.[188,1398,1276,1640]" captionTargetPageId="19" captionText="FIGURE 15. Epigonus marisrubri, SMF 24661, holotype, 136 mm SL." httpUri="https://zenodo.org/record/1182199/files/figure.png" pageId="19" pageNumber="280">Fig. 15</figureCitation>
). In the remaining 11 species of the
<collectingRegion id="49CFC0CC6E7E407AFEB9585A169C9B45" box="[310,401,1088,1111]" country="Papua New Guinea" name="Western" pageId="19" pageNumber="280">Western</collectingRegion>
Indian Ocean, there is no tooth patch on the tongue.
</paragraph>
<paragraph id="8BB40E2E6E7E407AFF48587F15989BD7" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="280">
We found two additional specimens of
<taxonomicName id="4C0B75AD6E7E407AFD00587F14279B6E" authorityName="Krupp, Zajonz &amp; Khalaf" authorityYear="2009" box="[655,810,1124,1148]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="marisrubri">
<emphasis id="B97FD23C6E7E407AFD00587F14279B6E" box="[655,810,1124,1148]" italics="true" pageId="19" pageNumber="280">E. marisrubri</emphasis>
</taxonomicName>
from
<collectingCountry id="F31C4EBE6E7E407AFCFA587E14DC9B6F" box="[885,977,1124,1149]" name="Somalia" pageId="19" pageNumber="280">Somalia</collectingCountry>
(USNM 269827), representing the first record of the species from that area. All specimens were collected from coastal reef areas and the continental shelf (
<bibRefCitation id="EF9A73DF6E7E407AFF1158B615849BD7" box="[158,649,1196,1221]" pageId="19" pageNumber="280" refId="ref21226" refString="Okamoto, M. &amp; Aungtonya, C. (2013) Re-examination of the holotype of Epigonus thai Prokofiev and Bussarawit: redescription and comparison with related species (Teleostei: Perciformes: Epigonidae). Phuket Marine Biological Center Research Bulletin, 72, 49 - 53." type="journal article">Okamoto &amp; Motomura 2013; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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