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<document id="E0D16011CF06B6D12EC7F644AEC6BE73" ID-CLB-Dataset="53641" ID-DOI="10.11646/zootaxa.5124.4.2" ID-GBIF-Dataset="7485af56-f9e8-450c-babe-9d6ae7415729" ID-ISSN="1175-5326" ID-Zenodo-Dep="6417643" ID-ZooBank="55967359-1F1E-4812-89F8-3205DC6DB37A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649142451215" checkinUser="plazi" docAuthor="Marta, Kimberly S., Bustamante, Abel A., Ruiz, Gustavo R. S., Teixeira, Renato A., Hagopián, Damián, Brescovit, Antonio D., Valiati, Victor Hugo &amp; Rodrigues, Everton N. L." docDate="2022" docId="03A088765745FFA0FF085D46FE432384" docLanguage="en" docName="zootaxa.5124.4.2.pdf" docOrigin="Zootaxa 5124 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Huriini Simon 1901" docType="treatment" docVersion="7" lastPageNumber="434" masterDocId="FF99F00E5747FFA3FF9F5A48FFB72149" masterDocTitle="A new huriine genus and notes on morphological characters (Araneae: Salticidae: Salticinae)" masterLastPageNumber="457" masterPageNumber="431" pageNumber="433" updateTime="1699319351946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="11F75E1047A87CC39EF0C8F8D1A9A921">A new huriine genus and notes on morphological characters (Araneae: Salticidae: Salticinae)</mods:title>
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<mods:namePart id="10EF66D37209388EEFA8ABB52A6393C6">Marta, Kimberly S.</mods:namePart>
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<mods:affiliation id="0F851A3A2BEE9B633DE0EACBE6387C79">Programa de Pós-Graduação em Biologia: Diversidade e Manejo de Vida Silvestre, UNISINOS. Laboratório de Genética e Biologia Molecular (LGBM) e Grupo de Pesquisa de Evolução, Ecologia e Genética da Conservação. Universidade do Vale do Rio dos Sinos, Centro de Ciências da Saúde. Av. Unisinos, 950, Ciências da Saúde, 93022750, São Leopoldo, RS, Brazil &amp; Laboratório de Diversidade e Sistemática de Arachnida (LaDiSA). Universidade do Vale do Rio dos Sinos (UNISINOS). Av. Unisinos, 950, Cristo Rei, 93022 - 000, São Leopoldo, RS, Brazil &amp; kimberly. dsm @ hotmail. com; https: // orcid. org / 0000 - 0002 - 5141 - 0056</mods:affiliation>
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<mods:namePart id="42E2DE71B73C91CF89FAF7F73BF8E1C0">Bustamante, Abel A.</mods:namePart>
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<mods:affiliation id="B41474C07D476E96DBA782DDBF4EE829">Instituto de Ciências Biológicas, Universidade Federal do Pará (UFPA), Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil Laboratório de Aracnologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul (MCTP), Porto Alegre, RS, Brazil. &amp; a. bustamanteferrada @ gmail. com; https: // orcid. org / 0000 - 0002 - 0046 - 8995</mods:affiliation>
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<mods:namePart id="1081E625F33689CB6B2924CD9B63B81F">Ruiz, Gustavo R. S.</mods:namePart>
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<mods:affiliation id="801588FDFDA42FFB230A52C473F85B58">Instituto de Ciências Biológicas, Universidade Federal do Pará (UFPA), Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil Laboratório de Aracnologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul (MCTP), Porto Alegre, RS, Brazil. &amp; gustavoruiz 86 @ hotmail. com; https: // orcid. org / 0000 - 0002 - 7326 - 7000</mods:affiliation>
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<mods:namePart id="7F3E101B4CD4E6E597FE6E6A4ED3430D">Teixeira, Renato A.</mods:namePart>
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<mods:affiliation id="BE5367346934ACB09DAE60D76D077C77">renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<mods:namePart id="DDE22D91CEB193500F4E539219E9BEEE">Hagopián, Damián</mods:namePart>
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<mods:affiliation id="01CAE1856BD89F7C9E8F1EA1262AD02E">Sección Entomología, Facultad de Ciencias (FCE), Universidad de la República, Iguá 4225, PC 11400, Montevideo, Uruguay &amp; dhagopian @ fcien. edu. uy; https: // orcid. org / 0000 - 0001 - 9060 - 0306</mods:affiliation>
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<mods:namePart id="8868926AC28C78FBA9836004A356871D">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="1DA6F02124DA8DD8205AB91DAE93CEB8">Instituto Butantan, Laboratório de Coleções Zoológicas, Av. Vital Brasil 1500. Butantã, 05503 - 900, São Paulo, SP, Brazil &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 002 - 1511 - 5324</mods:affiliation>
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<mods:namePart id="CA8F86D926972248A0F30C908633197C">Valiati, Victor Hugo</mods:namePart>
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<mods:affiliation id="07007F89C633485A3A9BEC931910F45F">Programa de Pós-Graduação em Biologia: Diversidade e Manejo de Vida Silvestre, UNISINOS. Laboratório de Genética e Biologia Molecular (LGBM) e Grupo de Pesquisa de Evolução, Ecologia e Genética da Conservação. Universidade do Vale do Rio dos Sinos, Centro de Ciências da Saúde. Av. Unisinos, 950, Ciências da Saúde, 93022750, São Leopoldo, RS, Brazil &amp; valiati @ unisinos. br; https: // orcid. org / 0000 - 0002 - 4467 - 4547</mods:affiliation>
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<mods:affiliation id="53C1B33FD2D1EE717D2A31C7EAC0AEEB">Laboratório de Diversidade e Sistemática de Arachnida (LaDiSA). Universidade do Vale do Rio dos Sinos (UNISINOS). Av. Unisinos, 950, Cristo Rei, 93022 - 000, São Leopoldo, RS, Brazil &amp; enlopesrodrigues @ gmail. com; https: // orcid. org / 0000 - 0002 - 9814 - 0954</mods:affiliation>
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<treatment id="03A088765745FFA0FF085D46FE432384" ID-DOI="http://doi.org/10.5281/zenodo.6417645" ID-GBIF-Taxon="194464420" ID-Zenodo-Dep="6417645" LSID="urn:lsid:plazi:treatment:03A088765745FFA0FF085D46FE432384" httpUri="http://treatment.plazi.org/id/03A088765745FFA0FF085D46FE432384" lastPageId="3" lastPageNumber="434" pageId="2" pageNumber="433">
<subSubSection id="C3136AEB5745FFA1FF085D46FE222661" box="[151,405,1806,1832]" pageId="2" pageNumber="433" type="nomenclature">
<paragraph id="8BB639605745FFA1FF085D46FE222661" blockId="2.[151,405,1806,1832]" box="[151,405,1806,1832]" pageId="2" pageNumber="433">
<heading id="D0FE8E0C5745FFA1FF085D46FE222661" bold="true" box="[151,405,1806,1832]" fontSize="11" level="1" pageId="2" pageNumber="433" reason="1">
<taxonomicName id="4C0942E35745FFA1FF085D46FE222661" authority="Simon, 1901" authorityName="Simon" authorityYear="1901" box="[151,405,1806,1832]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="456" phylum="Arthropoda" rank="tribe" tribe="Huriini">
<emphasis id="B97DE5725745FFA1FF085D46FE222661" bold="true" box="[151,405,1806,1832]" pageId="2" pageNumber="433">Huriini Simon, 1901</emphasis>
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<subSubSection id="C3136AEB5745FFA0FF085D1BFE432384" lastPageId="3" lastPageNumber="434" pageId="2" pageNumber="433" type="type_taxon">
<paragraph id="8BB639605745FFA1FF085D1BFD5E2622" blockId="2.[151,745,1875,1899]" box="[151,745,1875,1899]" pageId="2" pageNumber="433">
<emphasis id="B97DE5725745FFA1FF085D1BFEA12622" bold="true" box="[151,278,1875,1899]" pageId="2" pageNumber="433">
<typeStatus id="54B287C25745FFA1FF085D1BFF7B2622" box="[151,204,1875,1899]" pageId="2" pageNumber="433">Type</typeStatus>
genus.
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<taxonomicName id="4C0942E35745FFA1FE825D1CFE592622" ID-CoL="636KD" authority="Simon, 1901" authorityName="Simon" authorityYear="1901" box="[285,494,1875,1899]" class="Arachnida" family="Salticidae" genus="Hurius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725745FFA1FE825D1CFED42622" box="[285,355,1876,1899]" italics="true" pageId="2" pageNumber="433">Hurius</emphasis>
Simon, 1901
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, by original designation.
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<paragraph id="8BB639605745FFA0FF085DD1FAA62098" blockId="2.[151,1437,1945,2008]" lastBlockId="3.[151,1437,151,717]" lastPageId="3" lastPageNumber="434" pageId="2" pageNumber="433">
<emphasis id="B97DE5725745FFA1FF085DD1FED826FA" bold="true" box="[151,367,1945,1971]" pageId="2" pageNumber="433">Revised diagnosis.</emphasis>
Due to the “standard” salticid body form and leg proportions of huriines (
<bibRefCitation id="EF9844915745FFA1FB155DD1FA2726FD" author="Ruiz, G. R. S. &amp; Maddison, W. P." box="[1162,1424,1945,1972]" pageId="2" pageNumber="433" pagination="251 - 279" refId="ref18154" refString="Ruiz, G. R. S. &amp; Maddison, W. P. (2015) The new Andean jumping spider genus Urupuyu and its placement within a revised classification of the Amycoida (Araneae: Salticidae). Zootaxa, 4040 (3), 251 - 279 https: // doi. org / 10.11646 / zootaxa. 4040.3.1" type="journal article" year="2015">Ruiz &amp; Maddison 2015</bibRefCitation>
), i.e., short to compact body with moderately high carapaces, recognition of its members is possible by comparison with other amycoid tribes. Huriines differ from gophoines and bredines by not having flattened and elongate bodies (as in most gophoines and bredines), and from amycines by having leg IV longer than III. From sitticines, by having one retrolateral tooth on the chelicera (absent in sitticines). Females of huriines and simonellines differ from those of scopocirines, thiodinines and sarindines by not having the spermathecae anteriorly-placed in the epigyne, and copulatory ducts not concentrically coiled; on the other hand, males of huriines and simonellines differ from those of scopocirines, thiodinines and sarindines by having the male palp bulb usually more spherical. Huriines differ from simonellines by having only one retromarginal tooth in the chelicera of both sexes, and by not having females with dorsal and/or ventral abdominal scuta. In addition, huriines have orange, yellow, brownish-red and dark coloration, with dorsal abdominal pattern composed of dark chevrons (see arrows in
<figureCitation id="133225E55744FFA0FC525BFFFBA52098" box="[973,1042,439,465]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1871,1895]" captionTargetBox="[166,1423,462,1848]" captionTargetId="figure-160@5.[151,1435,453,1848]" captionTargetPageId="5" captionText="FIGURES 1–8. Male of Guriurius minuano Marta, Bustamante, Ruiz &amp; Rodrigues sp. nov. Habitus (MCTP 21675): 1 dorsal view, white arrow: chevrons coloration pattern, 2 ventral view; 3 lateral view; 4 face, frontal view; scale bars: 1.00 mm. Left male palp (MCN 32225): 5 ventral view, white arrow: RvTA angular axis; 6 dorsal view; 7 prolateral view; 8 retrolateral view, scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6417650" httpUri="https://zenodo.org/record/6417650/files/figure.png" pageId="3" pageNumber="434">Figs 1</figureCitation>
,
<figureCitation id="133225E55744FFA0FBBF5BFFFB872098" box="[1056,1072,439,465]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1401,1425]" captionTargetBox="[168,1421,195,1365]" captionTargetId="figure-22@6.[158,1430,181,1377]" captionTargetPageId="6" captionText="FIGURES 9–14. Female of Guriurius minuano Marta, Bustamante, Ruiz &amp; Rodrigues sp. nov. Habitus (MCTP 40587): 9 dorsal view, white arrow: chevrons coloration pattern, 10 ventral view; 11 lateral view; 12 face, frontal view; scale bars: 1 mm. Epigyne female (MCTP 40587): 13 ventral view; 14 dorsal view, white arrow: dorsally chitinized plate, scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6417664" httpUri="https://zenodo.org/record/6417664/files/figure.png" pageId="3" pageNumber="434">9</figureCitation>
,
<figureCitation id="133225E55744FFA0FBA35BFFFBED2098" box="[1084,1114,439,465]" captionStart="FIGURES 27–34" captionStartId="9.[151,264,1884,1908]" captionTargetBox="[176,1417,535,1861]" captionTargetId="figure-112@9.[158,1430,525,1861]" captionTargetPageId="9" captionText="FIGURES 27–34. Male of Guriurius nancyae Marta, Bustamante, Ruiz &amp; Rodrigues sp. nov. Habitus (MCTP 30545): 27 dorsal view, white arrow: chevrons coloration pattern; 28 ventral view; 29 lateral view; 30 face, frontal view, scale bars: 0.50 mm. Left male palp: 31 ventral, white arrow: RvTA rounded axis; 32 dorsal; 33 prolateral 34 retrolateral view, arrow: RvTA rounded axis, scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6417678" httpUri="https://zenodo.org/record/6417678/files/figure.png" pageId="3" pageNumber="434">27</figureCitation>
,
<figureCitation id="133225E55744FFA0FBF95BFEFB332098" box="[1126,1156,438,465]" captionStart="FIGURES 35–42" captionStartId="13.[151,264,1664,1688]" captionTargetBox="[167,1416,193,1640]" captionTargetId="figure-18@13.[151,1435,181,1640]" captionTargetPageId="13" captionText="FIGURES 35–42. Male of Atelurius segmentatus. Habitus (MCN 49030): 35 dorsal view, white arrow: chevrons coloration pattern; 36 ventral view; 37 lateral view; 38 face, frontal view, scale bars: 0.50 mm. Left male palp: 39 ventral view; 40 dorsal view; 41 prolateral view 42 retrolateral view, scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6417686" httpUri="https://zenodo.org/record/6417686/files/figure.png" pageId="3" pageNumber="434">35</figureCitation>
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<figureCitation id="133225E55744FFA0FB0F5BFFFB192098" box="[1168,1198,439,465]" captionStart="FIGURES 43–48" captionStartId="14.[151,264,1931,1955]" captionTargetBox="[165,1411,691,1889]" captionTargetId="figure-217@14.[151,1435,669,1907]" captionTargetPageId="14" captionText="FIGURES 43–48. Female of Atelurius segmentatus. Habitus (MCN 36915): 43 dorsal view, white arrow: chevrons coloration pattern; 44 ventral view, white arrow: ventrally chitinized plate; 45 lateral view; 46 face, frontal view. Scale bars: 1 mm. Epigyne female 47 ventral; 48 dorsal. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6417694" httpUri="https://zenodo.org/record/6417694/files/figure.png" pageId="3" pageNumber="434">43</figureCitation>
,
<figureCitation id="133225E55744FFA0FB255BFFFAB42098" box="[1210,1283,438,465]" captionStart="FIGURES 49–58" captionStartId="18.[151,264,1869,1893]" captionTargetBox="[173,1409,421,1834]" captionTargetId="figure-133@18.[151,1435,417,1845]" captionTargetPageId="18" captionText="FIGURES 49–58. Simonurius gladifer. Habitus male (MCTP 31078): 49 dorsal view, white arrow: chevrons coloration pattern; 50 ventral view; 51 lateral view; 52 face, frontal view; Habitus female (MCTP 31069): 53 dorsal view, white arrow: chevrons coloration pattern; 54 ventral view; 55 lateral view; 56 face, frontal view, scale bars: 1 mm. Epigyne female (MCTP 31082): 57 ventral view; 58 dorsal view, white arrow: S2* is broken but illustrated and S1 angular axis in anterior region, scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6417698" httpUri="https://zenodo.org/record/6417698/files/figure.png" pageId="3" pageNumber="434">49, 53</figureCitation>
).
</paragraph>
<paragraph id="8BB639605744FFA0FF585B93FE432384" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="434">
<emphasis id="B97DE5725744FFA0FF585B93FEA620BC" bold="true" box="[199,273,475,501]" pageId="3" pageNumber="434">Notes.</emphasis>
Here we reinterpreted
<taxonomicName id="4C0942E35744FFA0FDBC5B93FD3A20BC" ID-CoL="644X6" authorityName="Simon" authorityYear="1901" box="[547,653,475,501]" class="Arachnida" family="Salticidae" genus="Scoturius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FDBC5B93FD3A20BC" box="[547,653,475,501]" italics="true" pageId="3" pageNumber="434">Scoturius</emphasis>
</taxonomicName>
as having two RTA lobes (see discussion below), as described for
<taxonomicName id="4C0942E35744FFA0FF085BB7FF4C2351" ID-CoL="63VHS" authorityName="Ruiz &amp; Maddison" authorityYear="2015" box="[151,251,511,536]" class="Arachnida" family="Salticidae" genus="Urupuyu" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FF085BB7FF4C2351" box="[151,251,511,536]" italics="true" pageId="3" pageNumber="434">Urupuyu</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9844915744FFA0FEA35BB7FD392350" author="Ruiz, G. R. S. &amp; Maddison, W. P." box="[316,654,510,537]" pageId="3" pageNumber="434" pagination="251 - 279" refId="ref18154" refString="Ruiz, G. R. S. &amp; Maddison, W. P. (2015) The new Andean jumping spider genus Urupuyu and its placement within a revised classification of the Amycoida (Araneae: Salticidae). Zootaxa, 4040 (3), 251 - 279 https: // doi. org / 10.11646 / zootaxa. 4040.3.1" type="journal article" year="2015">Ruiz &amp; Maddison 2015: 253</bibRefCitation>
). This new comprehension of a complex, bilobed RTA in
<taxonomicName id="4C0942E35744FFA0FAAD5BB7FA2B2350" ID-CoL="644X6" authorityName="Simon" authorityYear="1901" box="[1330,1436,511,537]" class="Arachnida" family="Salticidae" genus="Scoturius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FAAD5BB7FA2B2350" box="[1330,1436,511,537]" italics="true" pageId="3" pageNumber="434">Scoturius</emphasis>
</taxonomicName>
accompanied by a well-developed RvTA seems more plausible than the traditional idea of three independent tibial apophyses in that genus (see
<bibRefCitation id="EF9844915744FFA0FE79580FFD372328" author="Simon, E." box="[486,640,583,609]" pageId="3" pageNumber="434" pagination="381 - 668" refId="ref18272" refString="Simon, E. (1901 a) s. n. In: Histoire naturelle des araignees. Tome second. Deuxieme Edition. Roret, Paris, pp. 381 - 668. https: // doi. org / 10.5962 / bhl. title. 51973" type="book chapter" year="1901">Simon 1901a</bibRefCitation>
;
<bibRefCitation id="EF9844915744FFA0FD11580FFC9C2328" author="Galiano, M. E." box="[654,811,583,609]" pageId="3" pageNumber="434" pagination="285 - 301" refId="ref17400" refString="Galiano, M. E. (1988) Revision de los generos del grupo Hurieae (Araneae, Salticidae). Journal of Arachnology, 15, 285 - 301." type="journal article" year="1988">Galiano 1988</bibRefCitation>
). Some bredines, such as
<taxonomicName id="4C0942E35744FFA0FBCB580FFF6523CC" ID-CoL="3CY3" authority="Peckham &amp; Peckham, 1894" authorityName="Peckham &amp; Peckham" authorityYear="1894" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FBCB580FFB202328" box="[1108,1175,583,609]" italics="true" pageId="3" pageNumber="434">Breda</emphasis>
Peckham &amp; Peckham, 1894
</taxonomicName>
, also have well-developed RvTA (
<bibRefCitation id="EF9844915744FFA0FDC95823FCD523CC" author="Ruiz, G. R. S. &amp; Brescovit, A. D." box="[598,866,619,645]" pageId="3" pageNumber="434" pagination="401 - 433" refId="ref18098" refString="Ruiz, G. R. S. &amp; Brescovit, A. D. (2013) Revision of Breda and proposal of a new genus (Araneae: Salticidae). Zootaxa, 3664 (4), 401 - 433. https: // doi. org / 10.11646 / zootaxa. 3664.4.1" type="journal article" year="2013">Ruiz &amp; Brescovit 2013</bibRefCitation>
). Huriines differ from the flattened-bodied species of
<taxonomicName id="4C0942E35744FFA0FF2658C7FF4B23E0" ID-CoL="3CY3" authorityName="Peckham &amp; Peckham" authorityYear="1894" box="[185,252,655,681]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FF2658C7FF4B23E0" box="[185,252,655,681]" italics="true" pageId="3" pageNumber="434">Breda</emphasis>
</taxonomicName>
by the curved RvTA positioned obliquely on the tibia (straight and parallel to tibia in
<taxonomicName id="4C0942E35744FFA0FB6E58C7FA8323E0" ID-CoL="3CY3" authorityName="Peckham &amp; Peckham" authorityYear="1894" box="[1265,1332,655,681]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FB6E58C7FA8323E0" box="[1265,1332,655,681]" italics="true" pageId="3" pageNumber="434">Breda</emphasis>
</taxonomicName>
) and the developed RTA (lost in
<taxonomicName id="4C0942E35744FFA0FE3F58FBFE542384" ID-CoL="3CY3" authorityName="Peckham &amp; Peckham" authorityYear="1894" box="[416,483,691,717]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DE5725744FFA0FE3F58FBFE542384" box="[416,483,691,717]" italics="true" pageId="3" pageNumber="434">Breda</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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