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<document id="F91B305D13E37BB33DAB9B0C29D1B37D" ID-DOI="10.11646/zootaxa.5437.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10985633" ID-ZooBank="7114B49F-4A44-4165-9FC6-8A52A6E898F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713347432768" checkinUser="plazi" docAuthor="Kasparyan, Dmitry R. &amp; Khalaim, Andrey I." docDate="2024" docId="03A03624FFFCB1054DA4AADCFA3DF897" docLanguage="en" docName="zootaxa.5437.4.9.pdf" docOrigin="Zootaxa 5437 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.18:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="18" docTitle="Hylophasma Townes 1969" docType="key" docVersion="3" lastPageNumber="580" masterDocId="FF994E5CFFF5B10C4D33A81EFFA1FF95" masterDocTitle="Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico" masterLastPageNumber="581" masterPageNumber="571" pageNumber="580" updateTime="1713383471488" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="2877C79F5EF0963A50C81B6864B25003">Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico</mods:title>
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<mods:namePart id="67B758D9294D9E346D54CA4178C16D47">Kasparyan, Dmitry R.</mods:namePart>
<mods:affiliation id="DD09B9F1865E55B26C6128A9B7F86538">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart id="4485928255AE77CF140DB38553EAC4A2">Khalaim, Andrey I.</mods:namePart>
<mods:nameIdentifier id="2301B5EDFA04F19C4EA8383750CDFA12" type="ORCID">0000-0003-1802-2649</mods:nameIdentifier>
<mods:affiliation id="F7FA1E78E72622579D82E0BB319B09D9">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
<mods:nameIdentifier id="99F090FB06BF7824AC27631699236487" type="email">akhalaim@gmail.com</mods:nameIdentifier>
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<mods:date id="337BA31BC3EB02C816976973D9161B92">2024</mods:date>
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<treatment id="03A03624FFFCB1054DA4AADCFA3DF897" LSID="urn:lsid:plazi:treatment:03A03624FFFCB1054DA4AADCFA3DF897" httpUri="http://treatment.plazi.org/id/03A03624FFFCB1054DA4AADCFA3DF897" lastPageNumber="580" pageId="9" pageNumber="580">
<subSubSection id="C313D4B9FFFCB1054DA4AADCFDF9FD49" box="[151,600,705,732]" pageId="9" pageNumber="580" type="nomenclature">
<paragraph id="8BB68732FFFCB1054DA4AADCFDF9FD49" blockId="9.[151,600,705,732]" box="[151,600,705,732]" pageId="9" pageNumber="580">
<heading id="D0FE305EFFFCB1054DA4AADCFDF9FD49" bold="true" box="[151,600,705,732]" fontSize="11" level="1" pageId="9" pageNumber="580" reason="1">
<emphasis id="B97D5B20FFFCB1054DA4AADCFDF9FD49" bold="true" box="[151,600,705,732]" pageId="9" pageNumber="580">
Key to world species of
<taxonomicName id="4C09FCB1FFFCB1054C8CAADFFDF9FD49" ID-CoL="63722" ID-ENA="540434" authorityName="Townes" authorityYear="1969" box="[447,600,705,732]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B97D5B20FFFCB1054C8CAADFFDF9FD49" bold="true" box="[447,600,705,732]" italics="true" pageId="9" pageNumber="580">Hylophasma</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFFCB1054DA4AB19FA3DF897" pageId="9" pageNumber="580" type="key">
<keyStep id="30FD9F97FFFCB1054DA4AB19FA3DFC7C" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4AB19FA3DFCE1" blockId="9.[151,1437,775,1002]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AB19FA3DFCE1" pageId="9" pageNumber="580">
1. Occipital carina ventrally interrupted, its lower end projecting into a small but distinct tooth. Hind wing with posterior section of AA (brachiella) present. Head predominantly white with occiput, frons and vertex black. Mesoscutum with mark centrally and lateral margins posteriorly white.Antenna of both sexes with white ring occupying at least five flagellomeres. The
<taxonomicName id="4C09FCB1FFFCB1054877AB5FFA3DFCC3" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1348,1436,833,854]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="cavigena">
<emphasis id="B97D5B20FFFCB1054877AB5FFA3DFCC3" bold="true" box="[1348,1436,833,854]" italics="true" pageId="9" pageNumber="580">cavigena</emphasis>
</taxonomicName>
species-group.
<collectingCountry id="F31EC7A2FFFCB1054C40AB41FE1DFCE0" box="[371,444,863,885]" name="Mexico" pageId="9" pageNumber="580">Mexico</collectingCountry>
................................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4AB63FA3DFC7C" blockId="9.[151,1437,775,1002]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AB63FA3DFC7C" pageId="9" pageNumber="580">
- Occipital carina complete, joining to hypostomal carina ventrally. Hind wing with posterior section of AA (brachiella) usually absent. Head black, clypeus sometimes pale. Mesoscutum usually completely black, without white spot centrally, sometimes with reddish markings. Antenna of both sexes with white ring occupying at most five flagellomeres. The
<taxonomicName id="4C09FCB1FFFCB1054835ABA8FAE7FC59" authorityName="Townes" authorityYear="1962" box="[1286,1350,950,972]" class="Insecta" family="Ichneumonidae" genus="Hemisphragia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B97D5B20FFFCB1054835ABA8FAE7FC59" bold="true" box="[1286,1350,950,972]" italics="true" pageId="9" pageNumber="580">debilis</emphasis>
</taxonomicName>
species-group.............................................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD9F97FFFCB1054DA4AC11FA3DFBE8" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4AC11FA3DFBD7" blockId="9.[151,1436,1039,1149]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AC11FA3DFBD7" pageId="9" pageNumber="580">
2. Gena ventrally with a deep concavity just behind the lower corner of the mandible...................................................................................................
<emphasis id="B97D5B20FFFCB1054ECEAC32FBAEFBD4" bold="true" box="[1021,1039,1068,1089]" pageId="9" pageNumber="580">3.</emphasis>
<taxonomicName id="4C09FCB1FFFCB1054926AC32FA3DFBD7" ID-CoL="3NGHQ" authority="Kasparyan &amp; Ruiz-Cancino" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1045,1436,1068,1090]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="cavigena">
<emphasis id="B97D5B20FFFCB1054926AC32FB2DFBD4" bold="true" box="[1045,1164,1068,1089]" italics="true" pageId="9" pageNumber="580">H. cavigena</emphasis>
Kasparyan &amp; Ruíz-Cancino
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4AC54FA3DFBE8" blockId="9.[151,1436,1039,1149]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AC54FA3DFBE8" pageId="9" pageNumber="580">
- Gena ventrally evenly rounded between lower corner of mandible and hypostomal carina, without concavity..............................................................................
<emphasis id="B97D5B20FFFCB1054933AC79FBB3FBE9" bold="true" box="[1024,1042,1127,1148]" pageId="9" pageNumber="580">1.</emphasis>
<taxonomicName id="4C09FCB1FFFCB105492BAC79FA3DFBE8" ID-CoL="3NGHP" authority="Kasparyan &amp; Ruiz-Cancino" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1048,1436,1126,1149]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="altacima">
<emphasis id="B97D5B20FFFCB105492BAC79FB2AFBE9" bold="true" box="[1048,1163,1126,1148]" italics="true" pageId="9" pageNumber="580">H. altacima</emphasis>
Kasparyan &amp; Ruíz-Cancino
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD9F97FFFCB1054DA4ACBFFA3DFA9A" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4ACBFFA3DFB41" blockId="9.[151,1437,1185,1295]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4ACBFFA3DFB41" pageId="9" pageNumber="580">3. Mesosoma predominantly black except for rufous collar, tegulae, small whitish marks in hind corner of pronotum and sometimes at apex of scutellum................................................................................... 4</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4ACC3FA3DFA9A" blockId="9.[151,1437,1185,1295]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4ACC3FA3DFA9A" pageId="9" pageNumber="580">
- Mesosoma predominantly black with metapleuron and propodeum reddish; in
<taxonomicName id="4C09FCB1FFFCB1054EFDACC3FBE2FB67" ID-CoL="3NGHS" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[974,1091,1245,1266]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFCB1054EFDACC3FBE2FB67" box="[974,1091,1245,1266]" italics="true" pageId="9" pageNumber="580">H. nigriceps</emphasis>
</taxonomicName>
speculum and scutellum sometimes white.............................................................................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD9F97FFFCB1054DA4AD2AFA3DFA49" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4AD2AFA3DFA11" blockId="9.[151,1437,1332,1501]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AD2AFA3DFA11" pageId="9" pageNumber="580">
4. Hind wing with posterior section of AA (brachiella) distinct. Abdomen of female fulvoferruginous, tergites more or less in basal half brownish. In male tergites dark brown, first five tergites in their apical 0.5± brownish fulvous; tergites 6 and 7 entirely dark brown.
<collectingCountry id="F31EC7A2FFFCB1054C18AD71FEFBFA11" box="[299,346,1391,1412]" name="United States of America" pageId="9" pageNumber="580">USA</collectingCountry>
.......................................................................
<taxonomicName id="4C09FCB1FFFCB105498BAD71FA3DFA11" authority="Townes, 1970" authorityName="Townes" authorityYear="1970" box="[1208,1436,1390,1412]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="debils">
<emphasis id="B97D5B20FFFCB105498BAD71FAB1FA11" bold="true" box="[1208,1296,1390,1412]" italics="true" pageId="9" pageNumber="580">H. debils</emphasis>
<bibRefCitation id="EF98FAC3FFFCB1054825AD71FA3DFA11" author="Townes, H. K." box="[1302,1436,1391,1412]" pageId="9" pageNumber="580" pagination="1 - 537" refId="ref7309" refString="Townes, H. K. (1970) The genera of Ichneumonidae. Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes, 1970</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4AD93FA3DFA49" blockId="9.[151,1437,1332,1501]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AD93FA3DFA49" pageId="9" pageNumber="580">
- Hind wing without posterior section of AA (brachiella). Abdomen of female blackish with hind margin of tergites
<quantity id="4CF12AD7FFFCB1054812AD92FAFFFA37" box="[1313,1374,1420,1442]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.778" metricValueMin="0.508" pageId="9" pageNumber="580" unit="in" value="4.5" valueMax="7.0" valueMin="2.0">2–7 in</quantity>
apical 0.2–0.35 pale brownish (
<figureCitation id="13329BB7FFFCB1054CE0ADB7FDBDFA2A" box="[467,540,1449,1471]" captionStart="FIGURES 12–17" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 12–17. Hylophasma aterrima sp. nov., ♀, holotype (12–15) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="9" pageNumber="580">Fig. 12</figureCitation>
). In male tergites dark brown with rather narrow pale rufous apical band widened in middle part anteriorly; tergite 7 pale rufous with brownish its basal 0.4 (
<figureCitation id="13329BB7FFFCB1054E7FADD9FC35FA48" box="[844,916,1479,1501]" captionStart="FIGURES 12–17" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 12–17. Hylophasma aterrima sp. nov., ♀, holotype (12–15) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="9" pageNumber="580">Fig. 16</figureCitation>
).
<collectingCountry id="F31EC7A2FFFCB1054E96ADD9FC4FFA48" box="[933,1006,1479,1501]" name="Mexico" pageId="9" pageNumber="580">Mexico</collectingCountry>
................
<emphasis id="B97D5B20FFFCB105498DADD9FA3DFA49" bold="true" box="[1214,1436,1479,1500]" pageId="9" pageNumber="580">
2.
<taxonomicName id="4C09FCB1FFFCB10549E5ADD9FAE8FA49" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[1238,1353,1479,1500]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="aterrima" status="sp. nov.">
<emphasis id="B97D5B20FFFCB10549E5ADD9FAE8FA49" bold="true" box="[1238,1353,1479,1500]" italics="true" pageId="9" pageNumber="580">H. aterrima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EE65BFFFCB1054863ADD9FA3DFA49" box="[1360,1436,1479,1500]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD9F97FFFCB1054DA4AE1FFA3DF918" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4AE1FFA3DF9FA" blockId="9.[151,1437,1537,1677]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AE1FFA3DF9FA" pageId="9" pageNumber="580">
5. Pronotum entirely black. [Mesopleuron black except for reddish lower posterior corner of mesepimeron. Antenna black with flagellomeres 7–10 white ventrally and two basal ones reddish dorsally; scape reddish. Body black with metapleuron, propodeum completely and hind half metasomal tergites 1–3 reddish; legs reddish with fore and mid coxae and trochanters white, and hind tibia and all tarsus brownish. Brachiella absent.]
<collectingCountry id="F31EC7A2FFFCB1054E62AE47FC2BF9FA" box="[849,906,1625,1647]" name="China" pageId="9" pageNumber="580">China</collectingCountry>
.....................
<taxonomicName id="4C09FCB1FFFCB10549ABAE44FA3DF9FA" authority="Sheng et al., 2019" authorityName="Sheng" authorityYear="2019" box="[1176,1436,1625,1647]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="luica">
<emphasis id="B97D5B20FFFCB10549ABAE44FB49F9FA" bold="true" box="[1176,1256,1625,1647]" italics="true" pageId="9" pageNumber="580">H. luica</emphasis>
Sheng
<emphasis id="B97D5B20FFFCB1054803AE44FAFAF9FB" box="[1328,1371,1625,1647]" italics="true" pageId="9" pageNumber="580">et al</emphasis>
., 2019
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4AE69FA3DF918" blockId="9.[151,1437,1537,1677]" box="[151,1436,1655,1677]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AE69FA3DF918" box="[151,1436,1655,1677]" pageId="9" pageNumber="580">
- Pronotum partly or entirely white.
<collectingCountry id="F31EC7A2FFFCB1054F1CAE69FDD9F918" box="[559,632,1655,1677]" name="Mexico" pageId="9" pageNumber="580">Mexico</collectingCountry>
................................................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD9F97FFFCB1054DA4AEAFFA3DF897" pageId="9" pageNumber="580">
<paragraph id="8BB68732FFFCB1054DA4AEAFFA3DF952" blockId="9.[151,1436,1713,1794]" box="[151,1436,1713,1735]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AEAFFA3DF952" box="[151,1436,1713,1735]" pageId="9" pageNumber="580">
6. Pronotum completely white. Mesopleuron pale reddish with speculum white....
<emphasis id="B97D5B20FFFCB1054EC9AEAFFBADF953" bold="true" box="[1018,1036,1713,1734]" pageId="9" pageNumber="580">4.</emphasis>
<taxonomicName id="4C09FCB1FFFCB1054921AEACFA3DF952" ID-CoL="3NGHS" authority="Kasparyan &amp; Ruiz-Cancino" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1042,1436,1713,1735]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFCB1054921AEACFB2DF953" bold="true" box="[1042,1164,1713,1735]" italics="true" pageId="9" pageNumber="580">H. nigriceps</emphasis>
Kasparyan &amp; Ruíz-Cancino
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB68732FFFCB1054DA4AED1FA3DF897" blockId="9.[151,1436,1713,1794]" pageId="9" pageNumber="580">
<keyLead id="30F82407FFFCB1054DA4AED1FA3DF897" pageId="9" pageNumber="580">
- Pronotum partly black, with front margin entirely white. Mesopleuron black except for pale subtegular ridge and mesepimeron.......................................................
<emphasis id="B97D5B20FFFCB105493BAEF2FBBBF894" bold="true" box="[1032,1050,1772,1793]" pageId="9" pageNumber="580">5.</emphasis>
<taxonomicName id="4C09FCB1FFFCB1054913AEF2FA3DF897" ID-CoL="3NGHT" authority="Kasparyan &amp; Ruiz-Cancino" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1056,1436,1772,1794]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B97D5B20FFFCB1054913AEF2FB2AF897" bold="true" box="[1056,1163,1772,1794]" italics="true" pageId="9" pageNumber="580">H. pulchra</emphasis>
Kasparyan &amp; Ruíz-Cancino
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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