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<mods:title id="64A7914F3EE472B988331789EF63553F">An annotated checklist of the chondrichthyans of South Africa</mods:title>
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<taxonomicName id="4C3B4D34953BFFDCCFF75E6FFE102413" ID-CoL="78RL3" authority="Regan, 1906 a" authorityName="Regan" authorityYear="1906" box="[151,489,1122,1149]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="ocellifera">
<emphasis id="B94FEAA5953BFFDCCFF75E6FFE102413" bold="true" box="[151,489,1122,1149]" pageId="82" pageNumber="83">
<emphasis id="B94FEAA5953BFFDCCFF75E6FFEBB2412" bold="true" box="[151,322,1122,1149]" italics="true" pageId="82" pageNumber="83">Raja ocellifera</emphasis>
<bibRefCitation id="EFAA4B46953BFFDCCE285E6FFE102413" author="Regan" box="[328,489,1122,1148]" firstAuthor="Regan" pageId="82" pageNumber="83" pagination="1 - 6" refId="ref111992" refString="Regan, C. T. (1906 a) Descriptions of new or little-known fishes from the coast of Natal. Annals of the Natal Government Museum, 1 (1), 1 - 6." type="journal article" year="1906" yearSuffix="a">Regan, 1906a</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8B8436B7953BFFDCCFF75E8BFEB324CF" blockId="82.[151,1437,1122,2013]" box="[151,330,1158,1184]" pageId="82" pageNumber="83">Twineyed Skate</paragraph>
</subSubSection>
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<paragraph id="8B8436B7953BFFDCCFF75EA6FAE92487" blockId="82.[151,1437,1122,2013]" pageId="82" pageNumber="83">
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<treatmentCitation id="0A9A10A6953BFFDCCFF75EA6FDF424AB" author="Regan, C. T." box="[151,525,1194,1221]" page="2" pageId="82" pageNumber="83" year="1906">
<taxonomicName id="4C3B4D34953BFFDCCFF75EA6FDF424AB" ID-CoL="78RL3" authority="Regan, 1906 a: 2" authorityName="Regan" authorityPageNumber="2" authorityYear="1906" box="[151,525,1194,1221]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="ocellifera">
<emphasis id="B94FEAA5953BFFDCCFF75EA6FEBB24AB" box="[151,322,1194,1220]" italics="true" pageId="82" pageNumber="83">Raja ocellifera</emphasis>
<bibRefCitation id="EFAA4B46953BFFDCCE2D5EA6FDF424AB" author="Regan" box="[333,525,1194,1221]" firstAuthor="Regan" page="2" pageId="82" pageNumber="83" pagination="1 - 6" refId="ref111992" refString="Regan, C. T. (1906 a) Descriptions of new or little-known fishes from the coast of Natal. Annals of the Natal Government Museum, 1 (1), 1 - 6." type="journal article" year="1906" yearSuffix="a">Regan, 1906a: 2</bibRefCitation>
</taxonomicName>
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, Pl. 2
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.
<typeStatus id="54808815953BFFDCCD065EA7FD2824AB" box="[614,721,1194,1220]" pageId="82" pageNumber="83" type="syntype">Syntypes</typeStatus>
:
<specimenCode id="DB9D9ECC953BFFDCCDBF5EA6FC7424AA" box="[735,909,1194,1221]" collectionCode="BMNH" pageId="82" pageNumber="83">BMNH 1905.6</specimenCode>
.8.14 (1), NMP (1);
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probable
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<specimenCode id="DB9D9ECC953BFFDCCA285EA6FEDA2486" collectionCode="BMNH" pageId="82" pageNumber="83">BMNH 1895.12</specimenCode>
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.
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27.14 (1). Type locality:
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, northeast of
<collectingMunicipality id="6BE0ACCD953BFFDCCC2D5EC3FC352487" box="[845,972,1230,1256]" pageId="82" pageNumber="83">Bird Island</collectingMunicipality>
,
<collectingRegion id="49FFF855953BFFDCCCB85EC2FB962487" box="[984,1135,1230,1257]" country="South Africa" name="Eastern Cape" pageId="82" pageNumber="83">Eastern Cape</collectingRegion>
,
<collectingCountry id="F32C7627953BFFDCCB1A5EC3FAF52487" box="[1146,1292,1230,1256]" name="South Africa" pageId="82" pageNumber="83">South Africa</collectingCountry>
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.
</paragraph>
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<emphasis id="B94FEAA5953BFFDCCFF75EFFFEA32563" bold="true" box="[151,346,1266,1292]" pageId="82" pageNumber="83">Local synonymy</emphasis>
:
<taxonomicName id="4C3B4D34953BFFDCCE085EFEFDF62563" ID-CoL="78RL3" authority=": Regan, 1906 a: 2" authorityName="Regan" authorityPageNumber="2" authorityYear="1906" box="[360,527,1266,1292]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="ocellifera">
<emphasis id="B94FEAA5953BFFDCCE085EFEFDF62563" box="[360,527,1266,1292]" italics="true" pageId="82" pageNumber="83">Raia ocellifera</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9A10A6953BFFDCCD7E5EFEFCEE2563" box="[542,791,1266,1293]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCD7E5EFEFD2F2563" author="Regan" box="[542,726,1266,1293]" firstAuthor="Regan" page="2" pageId="82" pageNumber="83" pagination="1 - 6" refId="ref111992" refString="Regan, C. T. (1906 a) Descriptions of new or little-known fishes from the coast of Natal. Annals of the Natal Government Museum, 1 (1), 1 - 6." type="journal article" year="1906" yearSuffix="a">Regan, 1906a: 2</bibRefCitation>
, Pl. 2
</treatmentCitation>
;
<treatmentCitation id="0A9A10A6953BFFDCCC435EFEFC0E2563" box="[803,1015,1266,1293]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCC435EFEFC0E2563" author="Regan" box="[803,1015,1266,1293]" firstAuthor="Regan" page="242" pageId="82" pageNumber="83" pagination="241 - 255" refId="ref112080" refString="Regan, C. T. (1908 a) A collection of fishes from the coasts of Natal, Zululand and Cape Colony. Annals of the Natal Government Museum, 1 (3), 241 - 255." type="journal article" year="1908" yearSuffix="a">Regan, 1908a: 242</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9A10A6953BFFDCCB635EFFFB222562" box="[1027,1243,1266,1293]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCB635EFFFB222562" author="Garman" box="[1027,1243,1266,1293]" firstAuthor="Garman" page="365" pageId="82" pageNumber="83" pagination="1 - 515" refId="ref102048" refString="Garman, S. (1913) The Plagiostomia (sharks, skates, and rays). Memoirs of the Museum of Comparative Zoology, Harvard College, 36, 1 - 515." type="journal article" year="1913">Garman, 1913: 365</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F32C7627953BFFDCCB895EFFFA822563" box="[1257,1403,1266,1292]" name="South Africa" pageId="82" pageNumber="83">South Africa</collectingCountry>
, in part);
<bibRefCitation id="EFAA4B46953BFFDCCE695F1BFDB7255F" author="Gilchrist &amp; Thompson" box="[265,590,1302,1328]" firstAuthor="Gilchrist" pageId="82" pageNumber="83" pagination="255 - 290" refId="ref102877" refString="Gilchrist, J. D. F. &amp; Thompson, W. W. (1916) A catalogue of the sea fishes recorded from Natal. Part I. Annals of the Durban Museum, 1 (3), 255 - 290." type="journal article" year="1916">Gilchrist &amp; Thompson, 1916</bibRefCitation>
; 270;
<treatmentCitation id="0A9A10A6953BFFDCCDF25F1AFC2C255E" box="[658,981,1302,1329]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCDF25F1AFC2C255E" author="von Bonde, C. &amp; Swart, D. B." box="[658,981,1302,1329]" pageId="82" pageNumber="83" pagination="1 - 22" refId="ref118185" refString="von Bonde, C. &amp; Swart, D. B. (1923) The Platosomia (skates and rays) collected by the S. S. &quot; Pickle &quot;. University of South Africa, Fisheries and Marine Biological Survey Report, 3 (5), 1 - 22." type="journal article" year="1923">von Bonde &amp; Swart, 1923: 5</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCC815F1BFB56255F" author="Barnard" box="[993,1199,1302,1329]" firstAuthor="Barnard" page="67" pageId="82" pageNumber="83" pagination="1 - 418" refId="ref92438" refString="Barnard, K. H. (1925) A monograph of the marine fishes of South Africa. Part I (Amphioxus, Cyclostomata, Elasmobranchii, and Teleostei-Isospondyli to Heterosomata). Annals of the South African Museum, 21, 1 - 418." type="journal article" year="1925">Barnard, 1925: 67</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCBDA5F1BFA6E255F" author="Fowler" box="[1210,1431,1302,1329]" firstAuthor="Fowler" page="193" pageId="82" pageNumber="83" pagination="187 - 268" refId="ref100995" refString="Fowler, H. W. (1925 b) Fishes from Natal, Zululand and Portuguese East Africa. Proceedings of the Academy of Natural Sciences, Philadelphia, 1925, 77, 187 - 268." type="journal article" year="1925" yearSuffix="b">Fowler, 1925b: 193</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCFA65F36FE4C253A" author="von Bonde, C." box="[198,437,1338,1365]" pageId="82" pageNumber="83" pagination="4 - 109" refId="ref117901" refString="von Bonde, C. (1933) Report no 10 for the year ending December, 1932. Report of Fisheries and Marine Biological Survey for 1932, including lists of fishes, etc., lists of stations of the R. S. Africana, and salinity results obtained during 1932. Union of South Africa, Fisheries and Marine Biological Survey, Report, 10, 1933, 4 - 109." type="journal article" year="1933">von Bonde, 1933: 51</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCEA25F37FD69253B" author="Barnard" box="[450,656,1338,1364]" firstAuthor="Barnard" page="26" pageId="82" pageNumber="83" refId="ref92649" refString="Barnard, K. H. (1947) A Pictorial Guide to South African Fishes. Marine and Freshwater. Maskew Miller Ltd, Cape Town, 226 pp." type="book" year="1947">Barnard, 1947: 26</bibRefCitation>
, fig. 2, pl. 4;
<treatmentCitation id="0A9A10A6953BFFDCCC485F37FBD5253B" box="[808,1068,1338,1364]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCC485F37FC12253B" author="Smith, J. L. B." box="[808,1003,1338,1364]" pageId="82" pageNumber="83" refId="ref114538" refString="Smith, J. L. B. (1949 a) The Sea Fishes of Southern Africa. South Africa Central News Agency Ltd., 550 pp." type="book" year="1949">Smith, 1949a: 66</bibRefCitation>
, pl. 3
</treatmentCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCB595F37FB06253A" author="Smith, J. L. B." box="[1081,1279,1338,1365]" pageId="82" pageNumber="83" refId="ref115644" refString="Smith, J. L. B. (1964) Fishes collected by Dr T. Mortenson off the coast of South Africa in 1929, with an account of the genus Cruriraja Bigelow &amp; Schroeder, 1954 in South Africa. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 126, 283 - 300." type="book chapter" year="1964">Smith, 1964: 285</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCA6B5F37FF1D2517" author="Smith, J. L. B." pageId="82" pageNumber="83" refId="ref115700" refString="Smith, J. L. B. (1965) The Sea Fishes of Southern Africa. 5 th Edition. Central News Agency Ltd., 580 pp." type="book" year="1965">Smith, 1965: 66</bibRefCitation>
, pl. 3.
<taxonomicName id="4C3B4D34953BFFDCCE585F52FD082516" ID-CoL="78RL3" authority=": Thompson, 1914: 158" authorityName="Thompson" authorityPageNumber="158" authorityYear="1914" box="[312,753,1374,1401]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="ocellifera">
<emphasis id="B94FEAA5953BFFDCCE585F52FE182517" box="[312,481,1374,1400]" italics="true" pageId="82" pageNumber="83">Raja ocellifera</emphasis>
:
<treatmentCitation id="0A9A10A6953BFFDCCE925F53FD082516" box="[498,753,1374,1401]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCE925F53FD082516" author="Thompson, W. W." box="[498,753,1374,1401]" pageId="82" pageNumber="83" pagination="132 - 167" refId="ref117072" refString="Thompson, W. W. (1914) Catalogue of the fishes of the Cape Province (Pt 1). Marine Biological Report, Cape Town, 2 (8), 132 - 167." type="journal article" year="1914">Thompson, 1914: 158</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B46953BFFDCCC605F52FC2C2517" author="Norman" box="[768,981,1374,1401]" firstAuthor="Norman" page="42" pageId="82" pageNumber="83" pagination="1 - 58" refId="ref110714" refString="Norman, J. R. (1935) Coast fishes, Part. I. The South Atlantic. ' Discovery' Reports, 12, 1 - 58." type="journal article" year="1935">Norman, 1935: 42</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCC835F53FB412516" author="Fowler" box="[995,1208,1374,1401]" firstAuthor="Fowler" page="375" pageId="82" pageNumber="83" pagination="1 - 879" refId="ref101290" refString="Fowler, H. W. (1941) The fishes of the groups Elasmobranchii, Holocephali, Isospondyli, and Ostarophysi obtained by the U. S. Bureau of Fisheries streamer &quot; Albatross &quot; in 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Bulletin United States National Museum, 100, 1 - 879." type="journal article" year="1941">Fowler, 1941: 375</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCBA75F53FA6E2517" author="Hulley" box="[1223,1431,1374,1400]" firstAuthor="Hulley" page="137" pageId="82" pageNumber="83" pagination="137 - 147" refId="ref104920" refString="Hulley, P. A. (1969) The relationship between Raja miraletus Linnaeus and Raja ocellifera Regan based on a study of the clasper. Annals of the South African Museum, 52 (6), 137 - 147." type="journal article" year="1969">Hulley, 1969: 137</bibRefCitation>
, figs. 13;
<bibRefCitation id="EFAA4B46953BFFDCCE595F8EFDB525F3" author="Seret, B. &amp; Last, P. R." box="[313,588,1410,1437]" pageId="82" pageNumber="83" pagination="674 - 675" refId="ref113146" refString="Seret, B. &amp; Last, P. R. (2016) Giant stingray. Family Plesiobatidae. In: Last, P. R., White, W. T., de Carvalho, M. E., Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.), Rays of the World. Cornell University Press, Ithaca, New York, pp. 674 - 675. https: // doi. org / 10.1071 / 9780643109148" type="book chapter" year="2016">Last &amp; Séret, 2016: 477</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCD395F8EFCB025F3" author="Last" box="[601,841,1410,1437]" etAl="et al." firstAuthor="Last" page="23" pageId="82" pageNumber="83" pagination="11 - 34" refId="ref107873" refString="Last, P. R., Weigmann, S. &amp; Yang, L. (2016 f) Changes to the nomenclature of the skates (Chondrichthyes: Rajiformes). In: Last, P. R. &amp; Yearsley, G. K. (Eds.), Rays of the World: Supplementary Information. CSIRO Australian National Fish Collection, Hobart, pp. 11 - 34." type="book chapter" year="2016" yearSuffix="f">
Last
<emphasis id="B94FEAA5953BFFDCCDF15F8EFD3A25F3" box="[657,707,1410,1436]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
., 2016f: 23
</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCC365F8EFBA025F3" author="Last" box="[854,1113,1410,1437]" etAl="et al." firstAuthor="Last" page="326" pageId="82" pageNumber="83" pagination="204 - 363" refId="ref107642" refString="Last, P. R., Seret, B., Stehmann, M. F. W. &amp; Weigmann, S. (2016 g) Skates. Family Rajidae. In: Last, P. R., White, W. T., de Carvalho, M. E., Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.), Rays of the World. Cornell University Press, Ithaca, New York, pp. 204 - 363. https: // doi. org / 10.1071 / 9780643109148" type="book chapter" year="2016" yearSuffix="g">
Last
<emphasis id="B94FEAA5953BFFDCCCEE5F8EFC3925F3" box="[910,960,1410,1436]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
.,
<quantity id="4CC39B52953BFFDCCCB65F8FFBD925F3" box="[982,1056,1410,1436]" metricMagnitude="0" metricUnit="kg" metricValue="2.016" pageId="82" pageNumber="83" unit="g" value="2016.0">2016g</quantity>
: 326
</bibRefCitation>
, fig. 19.117.
<taxonomicName id="4C3B4D34953BFFDCCB8E5F8EFE3225AE" ID-CoL="6X3LW" authority=": Thompson, 1914: 158" authorityName="Thompson" authorityPageNumber="158" authorityYear="1914" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="miraletus">
<emphasis id="B94FEAA5953BFFDCCB8E5F8EFA6C25F3" box="[1262,1429,1410,1436]" italics="true" pageId="82" pageNumber="83">Raja miraletus</emphasis>
:
<treatmentCitation id="0A9A10A6953BFFDCCFA65FABFE3225AE" box="[198,459,1446,1473]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCFA65FABFE3225AE" author="Thompson, W. W." box="[198,459,1446,1473]" pageId="82" pageNumber="83" pagination="132 - 167" refId="ref117072" refString="Thompson, W. W. (1914) Catalogue of the fishes of the Cape Province (Pt 1). Marine Biological Report, Cape Town, 2 (8), 132 - 167." type="journal article" year="1914">Thompson, 1914: 158</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B46953BFFDCCEBD5FABFD4E25AF" author="Fowler" box="[477,695,1446,1472]" firstAuthor="Fowler" page="114" pageId="82" pageNumber="83" pagination="1 - 605" refId="ref101239" refString="Fowler, H. W. (1936) The marine fishes of West Africa based on the collection of the American Museum Congo expedition, 1909 - 1915. Part 1. Bulletin of the American Museum of Natural History, 70 (1), 1 - 605." type="journal article" year="1936">Fowler, 1936: 114</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCDA95FABFC5825AE" author="Fowler" box="[713,929,1446,1473]" firstAuthor="Fowler" page="375" pageId="82" pageNumber="83" pagination="1 - 879" refId="ref101290" refString="Fowler, H. W. (1941) The fishes of the groups Elasmobranchii, Holocephali, Isospondyli, and Ostarophysi obtained by the U. S. Bureau of Fisheries streamer &quot; Albatross &quot; in 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Bulletin United States National Museum, 100, 1 - 879." type="journal article" year="1941">Fowler, 1941: 375</bibRefCitation>
(in part, including
<taxonomicName id="4C3B4D34953BFFDCCBF05FAAFA8325AF" ID-CoL="6WRL6" authorityName="von Bonde &amp; Swart" authorityYear="1923" box="[1168,1402,1446,1472]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="parcomaculata">
<emphasis id="B94FEAA5953BFFDCCBF05FAAFA8325AF" box="[1168,1402,1446,1472]" italics="true" pageId="82" pageNumber="83">Raia parcomaculata</emphasis>
</taxonomicName>
in synonymy);
<bibRefCitation id="EFAA4B46953BFFDCCE345FC7FDC9258B" author="Wallace, J. H." box="[340,560,1482,1508]" pageId="82" pageNumber="83" pagination="1 - 62" refId="ref118396" refString="Wallace, J. H. (1967 a) The batoid fishes of the east coast of southern Africa. Part III: skates and electric rays. Investigational Report. Oceanographic Research Institute, Durban, 17, 1 - 62." type="journal article" year="1967">Wallace, 1967a: 31</bibRefCitation>
, figs. 1617;
<bibRefCitation id="EFAA4B46953BFFDCCDAE5FC7FC65258B" author="Hulley" box="[718,924,1482,1508]" firstAuthor="Hulley" page="137" pageId="82" pageNumber="83" pagination="137 - 147" refId="ref104920" refString="Hulley, P. A. (1969) The relationship between Raja miraletus Linnaeus and Raja ocellifera Regan based on a study of the clasper. Annals of the South African Museum, 52 (6), 137 - 147." type="journal article" year="1969">Hulley, 1969: 137</bibRefCitation>
, figs, 12c;
<bibRefCitation id="EFAA4B46953BFFDCCB4A5FC7FB00258B" author="Hulley" box="[1066,1273,1482,1508]" firstAuthor="Hulley" page="179" pageId="82" pageNumber="83" pagination="151 - 220" refId="ref104963" refString="Hulley, P. A. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55, 151 - 220." type="journal article" year="1970">Hulley, 1970: 179</bibRefCitation>
, fig. 9, pl. 7b;
<bibRefCitation id="EFAA4B46953BFFDCCFA65FE3FE6A2667" author="Hulley" box="[198,403,1518,1544]" firstAuthor="Hulley" page="86" pageId="82" pageNumber="83" pagination="1 - 103" refId="ref105041" refString="Hulley, P. A. (1972 a) The origin, interrelationships and distribution of southern African Rajidae (Chondrichthyes, Batoidei). Annals of the South African Museum, 60, 1 - 103. https: // doi. org / 10.2307 / 1442499" type="journal article" year="1972" yearSuffix="a">Hulley, 1972a: 86</bibRefCitation>
, figs. 5859;
<bibRefCitation id="EFAA4B46953BFFDCCD515FE3FCF92667" author="Hulley" box="[561,768,1518,1544]" firstAuthor="Hulley" page="123" pageId="82" pageNumber="83" pagination="115 - 128" refId="ref105183" refString="Hulley, P. A. (1986) Family Rajidae, Anacanthobatidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smith's Sea Fishes. Macmillan, Johannesburg, pp. 115 - 128." type="book chapter" year="1986">Hulley, 1986: 123</bibRefCitation>
, fig. 25.14, pl. 6;
<bibRefCitation id="EFAA4B46953BFFDCCCAF5FE3FB3D2667" author="Stehmann, M." box="[975,1220,1518,1545]" pageId="82" pageNumber="83" pagination="1 - 119" refId="ref116339" refString="Stehmann, M. (1995) First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Archiv f ̡ r Fischereiwissenschaft, 43 (1), 1 - 119." type="journal article" year="1995">Stehmann, 1995: 106</bibRefCitation>
;
<treatmentCitation id="0A9A10A6953BFFDCCBB25FE3FF0B2643" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCBB25FE3FF0B2643" author="Compagno" firstAuthor="Compagno" page="116" pageId="82" pageNumber="83" pagination="75 - 120" refId="ref95720" refString="Compagno, L. J. V. (1999) An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa, 54, 75 - 120. https: // doi. org / 10.1080 / 00359199909520406" type="journal article" year="1999">Compagno, 1999: 116</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCF9E5C1EFD942643" author="Heemstra &amp; Heemstra" box="[254,621,1554,1581]" firstAuthor="Heemstra" page="81" pageId="82" pageNumber="83" refId="ref104374" refString="Heemstra, P. C. &amp; Heemstra, E. (2004) Coastal Fishes of Southern Africa. National Inquiry Service Centre and South African Institute for Aquatic Biodiversity, Grahamstown, 488 pp." type="book" year="2004">Heemstra &amp; Heemstra, 2004: 81</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCD195C1FFC2F2643" author="Compagno &amp; Ebert" box="[633,982,1554,1580]" firstAuthor="Compagno" page="139" pageId="82" pageNumber="83" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Compagno &amp; Ebert, 2007: 139</bibRefCitation>
, fig. 7a;
<bibRefCitation id="EFAA4B46953BFFDCCB595C1FFA6E2643" author="Ebert &amp; Compagno" box="[1081,1431,1554,1580]" firstAuthor="Ebert" page="121" pageId="82" pageNumber="83" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Ebert &amp; Compagno, 2007: 121</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCFA65C3BFE40263F" author="Ebert" box="[198,441,1590,1617]" etAl="et al." firstAuthor="Ebert" page="92" pageId="82" pageNumber="83" pagination="81 - 102" refId="ref98910" refString="Ebert, D. A., Compagno, L. J. V. &amp; Cowley, P. D. (2008) Aspects on the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern African waters. ICES Journal of Marine Science, 65, 81 - 102. https: // doi. org / 10.1093 / icesjms / fsm 169" type="journal article" year="2008">
Ebert
<emphasis id="B94FEAA5953BFFDCCE6A5C3AFEC4263F" box="[266,317,1590,1616]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
., 2008: 92
</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCEA65C3BFD71263E" author="NPOA" box="[454,648,1590,1617]" firstAuthor="NPOA" page="54" pageId="82" pageNumber="83" refId="ref110949" refString="NPOA. (2013) National Plan of Action for the Conservation and Management of Sharks (NPOA-Sharks). Department of Agriculture, Forestry and Fisheries (DAFF), Rogge Bay, Cape Town, 63 pp." type="book" year="2013">NPOA, 2013: 54</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCDF55C3BFC4C263F" author="da Silva" box="[661,949,1590,1617]" etAl="et al." firstAuthor="da Silva" page="247" pageId="82" pageNumber="83" pagination="233 - 248" refId="ref96965" refString="da Silva C., Booth, A. J., Dudley, S. F. J., Kerwath, S. E., Lamberth, S. J., Leslie, R. W., McCord, M. E., Sauer, W. H. H. &amp; Zweig, T. (2015) A description and updated overview of the status and management of South Africa's chondrichthyan fisheries. South African Journal of Marine Science, 37, 233 - 248. https: // doi. org / 10.2989 / 1814232 X. 2015.1044471" type="journal article" year="2015">
da Silva
<emphasis id="B94FEAA5953BFFDCCD995C3AFCD2263F" box="[761,811,1590,1616]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
., 2015: 247
</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCCA25C3BFAE9263F" author="Ebert &amp; van Hees" box="[962,1296,1590,1617]" firstAuthor="Ebert" page="147" pageId="82" pageNumber="83" pagination="141 - 156" refId="ref99877" refString="Ebert, D. A. &amp; van Hees, K. E. (2015) Beyond jaws: rediscovering the &quot; Lost Sharks &quot; of southern Africa. African Journal of Marine Science, 37, 141 - 156. https: // doi. org / 10.2989 / 1814232 X. 2015.1048730" type="journal article" year="2015">Ebert &amp; van Hees, 2015: 147</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953BFFDCCA7D5C3BFEC5261A" author="Weigmann, S." pageId="82" pageNumber="83" pagination="837 - 1037" refId="ref118762" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88, 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann, 2016: 955</bibRefCitation>
.
<taxonomicName id="4C3B4D34953BFFDCCE295C56FE08261B" authority=": von Bonde &amp; Swart, 1923: 5" authorityName="von Bonde &amp; Swart" authorityPageNumber="5" authorityYear="1923" box="[329,497,1626,1652]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="miraletus">
<emphasis id="B94FEAA5953BFFDCCE295C56FE08261B" box="[329,497,1626,1652]" italics="true" pageId="82" pageNumber="83">Raia miraletus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9A10A6953BFFDCCD625C56FCA8261A" box="[514,849,1626,1653]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCD625C56FCA8261A" author="von Bonde, C. &amp; Swart, D. B." box="[514,849,1626,1653]" pageId="82" pageNumber="83" pagination="1 - 22" refId="ref118185" refString="von Bonde, C. &amp; Swart, D. B. (1923) The Platosomia (skates and rays) collected by the S. S. &quot; Pickle &quot;. University of South Africa, Fisheries and Marine Biological Survey Report, 3 (5), 1 - 22." type="journal article" year="1923">von Bonde &amp; Swart, 1923: 5</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9A10A6953BFFDCCC005C57FBC9261B" box="[864,1072,1626,1653]" pageId="82" pageNumber="83">
<bibRefCitation id="EFAA4B46953BFFDCCC005C57FBC9261B" author="Barnard" box="[864,1072,1626,1653]" firstAuthor="Barnard" page="68" pageId="82" pageNumber="83" pagination="1 - 418" refId="ref92438" refString="Barnard, K. H. (1925) A monograph of the marine fishes of South Africa. Part I (Amphioxus, Cyclostomata, Elasmobranchii, and Teleostei-Isospondyli to Heterosomata). Annals of the South African Museum, 21, 1 - 418." type="journal article" year="1925">Barnard, 1925: 68</bibRefCitation>
</treatmentCitation>
(in part, not
<typeStatus id="54808815953BFFDCCBAA5C56FAFC261A" box="[1226,1285,1627,1653]" pageId="82" pageNumber="83">types</typeStatus>
of
<taxonomicName id="4C3B4D34953BFFDCCA4F5C56FE9726F7" ID-CoL="ZS3S" baseAuthorityName="von Bonde &amp; Swart" baseAuthorityYear="1923" class="Elasmobranchii" family="Rajidae" genus="Cruriraja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="parcomaculata">
<emphasis id="B94FEAA5953BFFDCCA4F5C56FE9726F7" italics="true" pageId="82" pageNumber="83">Cruriraja parcomaculata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAA4B46953BFFDCCEE65C73FDDC26F7" author="Clark" box="[390,549,1662,1688]" firstAuthor="Clark" page="9" pageId="82" pageNumber="83" pagination="1 - 66" refId="ref95217" refString="Clark, R. S. (1926) Rays and skates. A revision of the European species. Fishery Board for Scotland, Scientific Investigations, 1, 1 - 66." type="journal article" year="1926">Clark, 1926: 9</bibRefCitation>
(
<collectingRegion id="49FFF855953BFFDCCD535C73FD9726F7" box="[563,622,1662,1688]" country="South Africa" name="Western Cape" pageId="82" pageNumber="83">Cape</collectingRegion>
Colony, Mossel Bay).
<taxonomicName id="4C3B4D34953BFFDCCC0D5C72FA6E26F7" ID-CoL="6X3LW" authority=": Compagno et al., 1989: 96" authorityName="Compagno" authorityPageNumber="96" authorityYear="1989" box="[877,1431,1662,1689]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="82" pageNumber="83" phylum="Chordata" rank="species" species="miraletus" subGenus="Raja">
<emphasis id="B94FEAA5953BFFDCCC0D5C72FC5B26F7" box="[877,930,1663,1688]" italics="true" pageId="82" pageNumber="83">Raja</emphasis>
(
<emphasis id="B94FEAA5953BFFDCCCD15C72FC1F26F7" box="[945,998,1663,1688]" italics="true" pageId="82" pageNumber="83">Raja</emphasis>
)
<emphasis id="B94FEAA5953BFFDCCC965C73FB9926F7" box="[1014,1120,1662,1688]" italics="true" pageId="82" pageNumber="83">miraletus</emphasis>
:
<bibRefCitation id="EFAA4B46953BFFDCCB0E5C73FA6E26F7" author="Compagno" box="[1134,1431,1662,1689]" etAl="et al." firstAuthor="Compagno" page="96" pageId="82" pageNumber="83" refId="ref96138" refString="Compagno, L. J. V., Ebert, D. A. &amp; Smale, M. J. (1989) Guide to the Sharks and Rays of Southern Africa. Struik Publishers, Cape Town, 158 pp." type="book" year="1989">
Compagno
<emphasis id="B94FEAA5953BFFDCCB8F5C72FAE626F7" box="[1263,1311,1662,1688]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
., 1989: 96
</bibRefCitation>
</taxonomicName>
, pl.;
<bibRefCitation id="EFAA4B46953BFFDCCF915CAFFDD426D3" author="Compagno" box="[241,557,1698,1725]" etAl="et al." firstAuthor="Compagno" page="102" pageId="82" pageNumber="83" pagination="43 - 139" refId="ref96066" refString="Compagno, L. J. V., Ebert, D. A. &amp; Cowley, P. D. (1991) Distribution of offshore demersal cartilaginous fishes (class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11, 43 - 139. https: // doi. org / 10.2989 / 025776191784287664" type="journal article" year="1991">
Compagno
<emphasis id="B94FEAA5953BFFDCCE135CAEFE5C26D3" box="[371,421,1698,1724]" italics="true" pageId="82" pageNumber="83">et al</emphasis>
., 1991: 102
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321653C953BFFDCCFF75CCBFAF927D7" pageId="82" pageNumber="83" type="materials_examined">
<paragraph id="8B8436B7953BFFDCCFF75CCBFAD7271E" blockId="82.[151,1437,1122,2013]" pageId="82" pageNumber="83">
<materialsCitation id="3B533CEA953BFFDCCFF75CCBFC9C268E" ID-GBIF-Occurrence="3072574774" box="[151,869,1734,1761]" collectionCode="BMNH" country="South Africa" location="South Africa" pageId="82" pageNumber="83" specimenCode="BMNH 1905.6" specimenCount="1" typeStatus="syntype">
<emphasis id="B94FEAA5953BFFDCCFF75CCBFDE5268F" bold="true" box="[151,540,1734,1760]" pageId="82" pageNumber="83">
<collectingCountry id="F32C7627953BFFDCCFF75CCBFEC1268F" box="[151,312,1734,1760]" name="South Africa" pageId="82" pageNumber="83">South Africa</collectingCountry>
voucher material
</emphasis>
:
<typeStatus id="54808815953BFFDCCD545CCBFD66268F" box="[564,671,1734,1760]" pageId="82" pageNumber="83" type="syntype">Syntypes</typeStatus>
:
<specimenCode id="DB9D9ECC953BFFDCCDD25CCAFC9C268E" box="[690,869,1734,1761]" collectionCode="BMNH" pageId="82" pageNumber="83">BMNH 1905.6</specimenCode>
</materialsCitation>
.8.14 (1),
<collectionCode id="ED2AAE72953BFFDCCC825CCAFBE7268E" box="[994,1054,1735,1761]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="82" pageNumber="83">NMP</collectionCode>
(1);
<materialsCitation id="3B533CEA953BFFDCCB065CCBFEDA276A" ID-GBIF-Occurrence="3072574681" collectionCode="BMNH" country="South Africa" location="Probable" pageId="82" pageNumber="83" specimenCode="BMNH 1895.12" specimenCount="1" typeStatus="syntype">
<location id="8EE4606C953BFFDCCB065CCBFB33268F" LSID="urn:lsid:plazi:treatment:039287A1953BFFDDCFF75E6FFDEB2096:8EE4606C953BFFDCCB065CCBFB33268F" box="[1126,1226,1734,1760]" country="South Africa" name="Probable" pageId="82" pageNumber="83">Probable</location>
<typeStatus id="54808815953BFFDCCBB95CCAFACD268E" box="[1241,1332,1735,1761]" pageId="82" pageNumber="83" type="syntype">syntype</typeStatus>
:
<specimenCode id="DB9D9ECC953BFFDCCA285CCAFEDA276A" collectionCode="BMNH" pageId="82" pageNumber="83">BMNH 1895.12</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B533CEA953BFFDCCE465CE7FAD3271E" ID-GBIF-Occurrence="3072574518" collectionCode="SAIAB" country="South Africa" location="Non-types" pageId="82" pageNumber="83" specimenCode="SAIAB 7842, SAIAB 8067, SAIAB 8270, SAIAB 11148, SAIAB 11149, SAIAB 11150, SAIAB 12007, SAIAB 12031, SAIAB 12194, SAIAB 13313, SAIAB 14621, SAIAB 16518, SAIAB 17227, SAIAB 17414, SAIAB 26903, SAIAB 26904, SAIAB 26905, SAIAB 26906, SAIAB 26907, SAIAB 27152, SAIAB 27153, SAIAB 27154, SAIAB 27155, SAIAB 27156, SAIAB 44240, SAIAB 44250" specimenCount="1" typeStatus="syntype">
27.14 (1).
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:
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,
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.
</paragraph>
<paragraph id="8B8436B7953BFFDCCFF75D77FAF927D7" blockId="82.[151,1437,1122,2013]" pageId="82" pageNumber="83">
<emphasis id="B94FEAA5953BFFDCCFF75D77FE2A27FB" bold="true" box="[151,467,1914,1940]" pageId="82" pageNumber="83">South African distribution</emphasis>
: Possibly endemic to
<collectingCountry id="F32C7627953BFFDCCDAA5D77FCA227FB" box="[714,859,1914,1940]" name="South Africa" pageId="82" pageNumber="83">South Africa</collectingCountry>
. Confirmed from
<collectingRegion id="49FFF855953BFFDCCB435D76FB4627FB" box="[1059,1215,1914,1941]" country="South Africa" name="Western Cape" pageId="82" pageNumber="83">Western Cape</collectingRegion>
to Port Alfred (EC) and from
<taxonomicName id="4C3B4D34953BFFDCCE535D93FE7E27D7" box="[307,391,1950,1976]" genus="Durban" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="82" pageNumber="126" phylum="Mollusca" rank="genus">Durban</taxonomicName>
to Richards Bay (KZN). Records outside of
<collectingCountry id="F32C7627953BFFDCCC1C5D93FBF227D7" box="[892,1035,1950,1976]" name="South Africa" pageId="82" pageNumber="83">South Africa</collectingCountry>
require confirmation.
</paragraph>
</subSubSection>
<subSubSection id="C321653C953BFFDDCFF75DCFFDEB2096" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="discussion">
<paragraph id="8B8436B7953BFFDDCFF75DCFFADC20BA" blockId="82.[151,1437,1122,2013]" lastBlockId="83.[151,1436,151,249]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<emphasis id="B94FEAA5953BFFDCCFF75DCFFEFD27B3" bold="true" box="[151,260,1986,2012]" pageId="82" pageNumber="83">Remarks</emphasis>
: Recent molecular and morphological data reveal that the South African form is distinct from other similar species in the Eastern Atlantic. The reported range from southern
<collectingCountry id="F32C7627953AFFDDCCCD5A9AFBF420DE" box="[941,1037,151,177]" name="Namibia" pageId="83" pageNumber="84">Namibia</collectingCountry>
to KZN, and possibly extending to
<collectingCountry id="F32C7627953AFFDDCFA65AB6FEEB20BA" box="[198,274,187,213]" name="Kenya" pageId="83" pageNumber="84">Kenya</collectingCountry>
, requires confirmation as records from north of
<collectingCountry id="F32C7627953AFFDDCC4B5AB6FC4320BA" box="[811,954,187,213]" name="South Africa" pageId="83" pageNumber="84">South Africa</collectingCountry>
may represent different species.
</paragraph>
<paragraph id="8B8436B7953AFFDDCFF75AD3FDEB2096" blockId="83.[151,1436,151,249]" box="[151,530,222,249]" pageId="83" pageNumber="84">
<emphasis id="B94FEAA5953AFFDDCFF75AD3FE7D2096" bold="true" box="[151,388,222,249]" pageId="83" pageNumber="84">Conservation status</emphasis>
: EN (2020).
</paragraph>
</subSubSection>
</treatment>
</document>