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<document id="407E3E486C4AB7BC94153E7CBCD4EAC9" ID-CLB-Dataset="31016" ID-DOI="10.5852/ejt.2018.394" ID-GBIF-Dataset="54a11801-d3d3-4b76-9f6c-7a9c27677825" ID-Zenodo-Dep="1146158" ID-ZooBank="94028C61-FAC5-4A21-BF2D-A75BBF3CCC4D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1515747489213" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2018" docId="038D2864FFBBFFBBFDDBFBFB8BCBFD0E" docLanguage="en" docName="ejt-394_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 394" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helicochetus Attems 1909" docType="treatment" docVersion="19" lastPageNumber="11" masterDocId="FFB4501CFFB2FFB1FF94FFB0892EFF8E" masterDocTitle="A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="29" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547161722869" originalUpdateUser="yanell" pageNumber="10" updateTime="1703024528746" updateUser="carolina">
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<mods:title id="6B8C7B8C27B4947800763C188E2855FD">A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:title id="9CDFAED1ADBF295E4D84FA5DD7EA1915">European Journal of Taxonomy</mods:title>
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<mods:date id="A022BB649E7FAABC0B0B646CCA511A18">2018</mods:date>
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<treatment id="038D2864FFBBFFBBFDDBFBFB8BCBFD0E" ID-DOI="http://doi.org/10.5281/zenodo.5613980" ID-GBIF-Taxon="140251436" ID-Zenodo-Dep="5613980" LSID="urn:lsid:plazi:treatment:038D2864FFBBFFBBFDDBFBFB8BCBFD0E" httpUri="http://treatment.plazi.org/id/038D2864FFBBFFBBFDDBFBFB8BCBFD0E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C33ECAF9FFBBFFB8FDDBFBFB8ACBFBEB" box="[591,997,1098,1125]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9B9972FFBBFFB8FDDBFBFB8ACBFBEB" blockId="9.[591,997,1098,1125]" box="[591,997,1098,1125]" pageId="9" pageNumber="10">
<heading id="D0D32E1EFFBBFFB8FDDBFBFB8ACBFBEB" box="[591,997,1098,1125]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
Genus
<taxonomicName id="4C24E2F1FFBBFFB8FD35FBFA8ACBFBEB" ID-CoL="7PJNB" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[673,997,1098,1125]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9504560FFBBFFB8FD35FBFA8A11FBEB" bold="true" box="[673,831,1098,1125]" italics="true" pageId="9" pageNumber="10">Helicochetus</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FCD3FBFB8ACBFBEB" author="Attems" box="[839,997,1099,1125]" firstAuthor="Attems" journalOrPublisher="Zoologischer Anzeiger" pageId="9" pageNumber="10" pagination="156 - 159" part="34" publicationUrl="https://biodiversitylibrary.org/page/30144159" refId="ref8995" refString="Attems C. 1909 a. Zur Systematik der Spirostreptoidea. Zoologischer Anzeiger 34 (5): 156 - 159. Available from https: // biodiversitylibrary. org / page / 30144159 [accessed 27 Nov. 2017]." title="Zur Systematik der Spirostreptoidea" type="journal article" year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33ECAF9FFBBFFB8FF29FB3C8B64FB29" box="[189,586,1164,1191]" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8B9B9972FFBBFFB8FF29FB3C8B64FB29" blockId="9.[189,586,1164,1191]" box="[189,586,1164,1191]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB34BE5CFFBBFFB8FF29FB3C8B64FB29" box="[189,586,1164,1191]" pageId="9" pageNumber="10">
<treatmentCitation id="0A85BF63FFBBFFB8FF29FB3C8B6BFB29" author="Attems" box="[189,581,1164,1191]" page="158" pageId="9" pageNumber="10" year="1909">
<taxonomicName id="4C24E2F1FFBBFFB8FF29FB3C8B6BFB29" ID-CoL="7PJNB" authority="Attems, 1909 a: 158" authorityName="Attems" authorityPageNumber="158" authorityYear="1909" box="[189,581,1164,1191]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9504560FFBBFFB8FF29FB3C8878FB28" box="[189,342,1164,1190]" italics="true" pageId="9" pageNumber="10">Helicochetus</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FEC8FB3C8B6BFB29" author="Attems C." box="[348,581,1164,1191]" firstAuthor="Attems" journalOrPublisher="Zoologischer Anzeiger" page="158" pageId="9" pageNumber="10" pagination="156 - 159" part="34" publicationUrl="https://biodiversitylibrary.org/page/30144159" refId="ref8995" refString="Attems C. 1909 a. Zur Systematik der Spirostreptoidea. Zoologischer Anzeiger 34 (5): 156 - 159. Available from https: // biodiversitylibrary. org / page / 30144159 [accessed 27 Nov. 2017]." title="Zur Systematik der Spirostreptoidea" type="journal article" year="1909" yearSuffix="a">Attems, 1909a: 158</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33ECAF9FFBBFFB8FF29FB7D8D5BFA98" pageId="9" pageNumber="10" type="type_taxon">
<paragraph id="8B9B9972FFBBFFB8FF29FB7D8876FB69" blockId="9.[189,344,1229,1255]" box="[189,344,1229,1255]" pageId="9" pageNumber="10">
<heading id="D0D32E1EFFBBFFB8FF29FB7D8876FB69" bold="true" box="[189,344,1229,1255]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9504560FFBBFFB8FF29FB7D8876FB69" bold="true" box="[189,344,1229,1255]" pageId="9" pageNumber="10">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B9972FFBBFFB8FF29FB4C8D5BFA98" blockId="9.[189,1141,1276,1303]" box="[189,1141,1276,1303]" pageId="9" pageNumber="10">
<taxonomicName id="4C24E2F1FFBBFFB8FF29FB4C8B5BFA99" authority="Peters" authorityName="Peters" authorityYear="1855" box="[189,629,1276,1303]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B9504560FFBBFFB8FF29FB4C88F5FA98" box="[189,475,1276,1302]" italics="true" pageId="9" pageNumber="10">Spirostreptus dimidiatus</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FE76FB4C8B5BFA99" author="Peters W. C. H." box="[482,629,1276,1303]" firstAuthor="Peters" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin" pageId="9" pageNumber="10" pagination="75 - 85" part="14" refId="ref10261" refString="Peters W. C. H. 1855. Uber die Myriapoden im allgemeinen und insbesondere uber die in Mossambique beobachteten Arten. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin 14: 75 - 85." title="Uber die Myriapoden im allgemeinen und insbesondere uber die in Mossambique beobachteten Arten" type="journal article" year="1855">Peters, 1855</bibRefCitation>
</taxonomicName>
, by original designation. See further below.
</paragraph>
</subSubSection>
<subSubSection id="C33ECAF9FFBBFFBBFF29FA8D8BCBFD0E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8B9B9972FFBBFFB8FF29FA8D88F6FAD9" blockId="9.[189,1399,1341,1626]" box="[189,472,1341,1367]" pageId="9" pageNumber="10">
<heading id="D0D32E1EFFBBFFB8FF29FA8D88F6FAD9" bold="true" box="[189,472,1341,1367]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9504560FFBBFFB8FF29FA8D88F6FAD9" bold="true" box="[189,472,1341,1367]" pageId="9" pageNumber="10">Other included species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B9972FFBBFFB8FF29FADC8A5AF9D7" blockId="9.[189,1399,1341,1626]" pageId="9" pageNumber="10">
<taxonomicName id="4C24E2F1FFBBFFB8FF29FADC8BFBFA09" ID-CoL="3K82C" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" box="[189,725,1388,1415]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B9504560FFBBFFB8FF29FADC88E9FA08" box="[189,455,1388,1414]" italics="true" pageId="9" pageNumber="10">Helicochetus aberrans</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FE5BFADC8B70FA08" author="Kraus O." box="[463,606,1388,1414]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="9" pageNumber="10" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus, 1966</bibRefCitation>
(
<collectingCountry id="F333D9E2FFBBFFB8FDF9FADC8BE3FA09" box="[621,717,1388,1415]" name="Zambia" pageId="9" pageNumber="10">Zambia</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C24E2F1FFBBFFB8FD75FADD8D0FFA09" ID-CoL="6LJ74" authority="Kraus, 1957" authorityName="Kraus" authorityYear="1957" box="[737,1057,1388,1415]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="digititarsus">
<emphasis id="B9504560FFBBFFB8FD75FADD8AA5FA08" box="[737,907,1388,1414]" italics="true" pageId="9" pageNumber="10">H. digititarsus</emphasis>
Kraus, 1957
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FBA4FADC8820FA27" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FF29FA3F8820FA27" box="[189,270,1423,1449]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
,
<collectingCountry id="F333D9E2FFBBFFB8FE8DFA3F88A9FA24" box="[281,391,1423,1450]" name="Tanzania" pageId="9" pageNumber="10">Tanzania</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FE0EFA208BEBFA24" ID-CoL="3K82G" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[410,709,1423,1450]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="electricus">
<emphasis id="B9504560FFBBFFB8FE0EFA208B01FA27" box="[410,559,1423,1449]" italics="true" pageId="9" pageNumber="10">H. electricus</emphasis>
Kraus, 1958
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FD40FA3F8D5DFA27" box="[724,1139,1423,1449]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FBB4FA3F8D5DFA27" box="[1056,1139,1423,1449]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FB12FA208841FA42" ID-CoL="3K82K" authority="(Pocock, 1896) (Kenya)" baseAuthorityName="Pocock" baseAuthorityYear="1896" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gregorii">
<emphasis id="B9504560FFBBFFB8FB12FA208C26FA27" box="[1158,1288,1424,1449]" italics="true" pageId="9" pageNumber="10">H. gregorii</emphasis>
(
<bibRefCitation id="EFB5E483FFBBFFB8FA83FA3F89D7FA43" author="Pocock R. I." firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History, Series 6" pageId="9" pageNumber="10" pagination="425 - 444" part="17" publicationUrl="https://doi.org/10.1080/00222939608680394" refId="ref10362" refString="Pocock R. I. 1896. On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History, Series 6, 17: 425 - 444. https: // doi. org / 10.1080 / 00222939608680394" title="On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa" type="journal article" year="1896">Pocock, 1896</bibRefCitation>
) (
<collectingCountry id="F333D9E2FFBBFFB8FE81FA028849FA42" box="[277,359,1458,1484]" name="Kenya" pageId="9" pageNumber="10">Kenya</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C24E2F1FFBBFFB8FEEAFA038B88FA43" ID-CoL="3K82L" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[382,678,1458,1485]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="inversus">
<emphasis id="B9504560FFBBFFB8FEEAFA038B26FA42" box="[382,520,1459,1484]" italics="true" pageId="9" pageNumber="10">H. inversus</emphasis>
Kraus, 1958
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FD2CFA028D49FA42" box="[696,1127,1458,1484]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FB80FA028D49FA42" box="[1044,1127,1458,1484]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FBEAFA0389D6FA7E" ID-CoL="3K82M" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="involutus">
<emphasis id="B9504560FFBBFFB8FBEAFA038C3CFA42" box="[1150,1298,1458,1484]" italics="true" pageId="9" pageNumber="10">H. involutus</emphasis>
Attems, 1935
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FE9EFA658B99FA61" box="[266,695,1493,1519]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FDF0FA658B99FA61" box="[612,695,1493,1519]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FD5AFA668D3AFA7E" ID-CoL="3K82N" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" box="[718,1044,1493,1520]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="laciniatus">
<emphasis id="B9504560FFBBFFB8FD5AFA668A47FA61" box="[718,873,1493,1519]" italics="true" pageId="9" pageNumber="10">H. laciniatus</emphasis>
Attems, 1935
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FBB2FA65883EF99D" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FF29FA49883EF99D" box="[189,272,1529,1555]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FEBCFA498B49F99D" ID-CoL="3K82P" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[296,615,1529,1555]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="levifolius">
<emphasis id="B9504560FFBBFFB8FEBCFA498895F99D" box="[296,443,1529,1555]" italics="true" pageId="9" pageNumber="10">H. levifolius</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FE51FA498B49F99D" author="Attems C." box="[453,615,1529,1555]" firstAuthor="Attems" journalOrPublisher="Zoologica (Stuttgart)" pageId="9" pageNumber="10" pagination="1 - 235" part="25" refId="ref9083" refString="Attems C. 1914. Afrikanische Spirostreptiden nebst Uberblick uber die Spirostreptiden orbis terrarum. Zoologica (Stuttgart) 25: 1 - 235." title="Afrikanische Spirostreptiden nebst Uberblick uber die Spirostreptiden orbis terrarum" type="journal article" year="1914">Attems, 1914</bibRefCitation>
</taxonomicName>
(”Sambesi”,
<collectingCountry id="F333D9E2FFBBFFB8FC98FA488A82F99D" box="[780,940,1528,1555]" name="Mozambique" pageId="9" pageNumber="10">Mozambique</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FC57FA498C5CF99D" ID-CoL="3K82Q" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[963,1394,1528,1555]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="monodon">
<emphasis id="B9504560FFBBFFB8FC57FA498D76F99D" box="[963,1112,1529,1555]" italics="true" pageId="9" pageNumber="10">H. monodon</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FBF7FA498DD9F99C" author="Kraus O." box="[1123,1271,1528,1555]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="9" pageNumber="10" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960</bibRefCitation>
(
<collectingCountry id="F333D9E2FFBBFFB8FA9EFA488C44F99D" box="[1290,1386,1528,1555]" name="Zambia" pageId="9" pageNumber="10">Zambia</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C24E2F1FFBBFFB8FF29F9AD88F1F9B8" ID-CoL="3K82R" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[189,479,1564,1590]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutaba">
<emphasis id="B9504560FFBBFFB8FF29F9AD8810F9B8" box="[189,318,1564,1590]" italics="true" pageId="9" pageNumber="10">H. mutaba</emphasis>
<bibRefCitation id="EFB5E483FFBBFFB8FEDEF9AC88F1F9B8" author="Kraus O." box="[330,479,1564,1590]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="9" pageNumber="10" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FE67F9AC8A86F9B8" box="[499,936,1564,1590]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FCC1F9AC8A86F9B8" box="[853,936,1564,1590]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
),
<taxonomicName id="4C24E2F1FFBBFFB8FC55F9AD8C41F9B9" ID-CoL="3K82S" authority="(Carl, 1909) (Tanzania)" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[961,1391,1564,1591]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pococki">
<emphasis id="B9504560FFBBFFB8FC55F9AD8D6BF9B8" box="[961,1093,1564,1590]" italics="true" pageId="9" pageNumber="10">H. pococki</emphasis>
(Carl, 1909) (
<collectingCountry id="F333D9E2FFBBFFB8FB6EF9AC8C49F9B9" box="[1274,1383,1564,1591]" name="Tanzania" pageId="9" pageNumber="10">Tanzania</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C24E2F1FFBBFFB8FF29F9F08899F9D4" ID-CoL="3K82T" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[189,439,1599,1626]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9504560FFBBFFB8FF29F9F0880FF9D7" box="[189,289,1600,1625]" italics="true" pageId="9" pageNumber="10">H. rarus</emphasis>
Kraus, 1958
</taxonomicName>
(
<collectingCountry id="F333D9E2FFBBFFB8FE52F98F8A46F9D7" box="[454,872,1599,1625]" name="Democratic Republic of the Congo" pageId="9" pageNumber="10">
Democratic Republic of the
<collectingRegion id="49E05790FFBBFFB8FC81F98F8A46F9D7" box="[789,872,1599,1625]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="9" pageNumber="10">Congo</collectingRegion>
</collectingCountry>
).
</paragraph>
<paragraph id="8B9B9972FFBBFFB8FF29F9318D72F914" blockId="9.[189,1116,1664,1691]" box="[189,1116,1664,1691]" pageId="9" pageNumber="10">
<bibRefCitation id="EFB5E483FFBBFFB8FF29F9318876F915" author="Kraus O." box="[189,344,1664,1691]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="9" pageNumber="10" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus (1960)</bibRefCitation>
provided an identification key to the species, except
<emphasis id="B9504560FFBBFFB8FC53F9318D72F914" box="[967,1116,1664,1690]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C24E2F1FFBBFFB8FC53F9318D77F914" ID-CoL="3K82C" authorityName="Kraus" authorityYear="1966" box="[967,1113,1664,1690]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aberrans">H. aberrans</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B9B9972FFBBFFB8FF29F9728801F952" blockId="9.[189,303,1730,1756]" box="[189,303,1730,1756]" pageId="9" pageNumber="10">
<heading id="D0D32E1EFFBBFFB8FF29F9728801F952" bold="true" box="[189,303,1730,1756]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9504560FFBBFFB8FF29F9728801F952" bold="true" box="[189,303,1730,1756]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B9972FFBBFFB8FF29F9408A42F8DF" blockId="9.[189,1399,1776,1873]" pageId="9" pageNumber="10">
The material from the Udzungwa Mts includes specimens of two species of
<emphasis id="B9504560FFBBFFB8FBCCF9408C2EF884" box="[1112,1280,1776,1802]" italics="true" pageId="9" pageNumber="10">Helichochetus</emphasis>
. Separate descriptive notes are given for each species, but two characters shared by both species may prove to be genus-characteristic and are therefore commented on here.
</paragraph>
<paragraph id="8B9B9972FFBBFFBBFF29F8C988B5FE5B" blockId="9.[189,1399,1912,2009]" lastBlockId="10.[189,1399,267,469]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
LIMBUS (
<figureCitation id="131F85F7FFBBFFB8FE8AF8C8889BF81D" box="[286,437,1912,1939]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, C, E</figureCitation>
).
<bibRefCitation id="EFB5E483FFBBFFB8FE53F8C98B71F81D" author="Kraus O." box="[455,607,1912,1939]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="9" pageNumber="10" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus (1960)</bibRefCitation>
characterized the limbus of
<emphasis id="B9504560FFBBFFB8FC09F8C88D6BF81C" box="[925,1093,1912,1938]" italics="true" pageId="9" pageNumber="10">Helichochetus</emphasis>
(translated from German): “Limbus divided by rounded indentations into processes which each carry three to six nail-like points. Limbus very broad, ca. 10 × as broad as the processes, with a silky sheen due to the characteristic surface sculpture (exception
<taxonomicName id="4C24E2F1FFB8FFBBFE2CFEBC8B7CFEAB" authorityName="Attems" authorityYear="1935" box="[440,594,267,293]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="laciniatus">
<emphasis id="B9504560FFB8FFBBFE2CFEBC8B7CFEAB" box="[440,594,267,293]" italics="true" pageId="10" pageNumber="11">H. laciniatus</emphasis>
</taxonomicName>
)”. The limbus of the two studied species, especially
<taxonomicName id="4C24E2F1FFB8FFBBFB47FEBC8C7CFEAB" authorityName="Kraus" authorityYear="1960" box="[1235,1362,267,293]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mutaba">
<emphasis id="B9504560FFB8FFBBFB47FEBC8C7CFEAB" box="[1235,1362,267,293]" italics="true" pageId="10" pageNumber="11">H. mutaba</emphasis>
</taxonomicName>
, is indeed very broad (
<figureCitation id="131F85F7FFB8FFBBFE0FFE9E8B39FEC7" box="[411,535,302,329]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, C</figureCitation>
), and the processes and nail-like points agree fully with Kraus’ description. It is not clear from his publications (
<bibRefCitation id="EFB5E483FFB8FFBBFDE3FEE28A2BFEE2" author="Kraus O." box="[631,773,338,364]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="10" pageNumber="11" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
,
<bibRefCitation id="EFB5E483FFB8FFBBFC87FEE28A61FEE3" author="Kraus O." box="[787,847,338,365]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="10" pageNumber="11" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">1966</bibRefCitation>
) what he meant by the “characteristic surface structure”. The SEM images (
<figureCitation id="131F85F7FFB8FFBBFDB0FEC58BE4FE01" box="[548,714,373,399]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, C, E</figureCitation>
) provide no clue. There is, however, another remarkable detail, visible in
<figureCitation id="131F85F7FFB8FFBBFEEBFE2888F4FE3C" box="[383,474,408,434]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="10" pageNumber="11">Fig. 4E</figureCitation>
: under the limbus proper, there is a “sub-limbus” consisting of rounded, almost semicircular lobes.
</paragraph>
<paragraph id="8B9B9972FFB8FFBBFF29FE4D8BCBFD0E" blockId="10.[189,1399,509,641]" pageId="10" pageNumber="11">
CYTOSCUTE DENTICLES (
<figureCitation id="131F85F7FFB8FFBBFE53FE4D8B45FD99" box="[455,619,509,535]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="10" pageNumber="11">Fig. 4B, D, F</figureCitation>
). The “cytoscutes”, i.e., cuticular “cells” each corresponding to a hypodermis cell (
<bibRefCitation id="EFB5E483FFB8FFBBFE19FD908B4CFDB4" author="Fusco G. &amp; Brena C. &amp; Minelli A." box="[397,610,544,570]" etAl="et al." firstAuthor="Fusco" journalOrPublisher="Zoologischer Anzeiger" pageId="10" pageNumber="11" pagination="91 - 102" part="239" refId="ref10009" refString="Fusco G., Brena C. &amp; Minelli A. 2000. Cellular processes in the growth of lithobiomorph centipedes (Chilopoda: Lithobiomorpha). A cuticular view. Zoologischer Anzeiger 239: 91 - 102." title="Cellular processes in the growth of lithobiomorph centipedes (Chilopoda: Lithobiomorpha). A cuticular view" type="journal article" year="2000">
Fusco
<emphasis id="B9504560FFB8FFBBFE4BFD918B32FDB4" box="[479,540,544,570]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2000
</bibRefCitation>
) very often carry a row of tiny denticles along one edge. In both studied species of
<taxonomicName id="4C24E2F1FFB8FFBBFE0DFDF38B1CFDD3" authorityName="Attems" authorityYear="1909" box="[409,562,579,605]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9504560FFB8FFBBFE0DFDF38B1CFDD3" box="[409,562,579,605]" italics="true" pageId="10" pageNumber="11">Helicochetus</emphasis>
</taxonomicName>
, these denticles are much larger than normal and point away from the cuticular surface rather than lying parallel to it.
</paragraph>
</subSubSection>
</treatment>
</document>
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