375 lines
50 KiB
XML
375 lines
50 KiB
XML
<document id="C011EC2843BEB6360EC45FA1F4D5CFC8" ID-CLB-Dataset="31016" ID-DOI="10.5852/ejt.2018.394" ID-GBIF-Dataset="54a11801-d3d3-4b76-9f6c-7a9c27677825" ID-Zenodo-Dep="1146158" ID-ZooBank="94028C61-FAC5-4A21-BF2D-A75BBF3CCC4D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515747489213" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2018" docId="038D2864FFA1FFA7FDD9F98C8D9BF99B" docLanguage="en" docName="ejt-394_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 394" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Spinotarsus fortehamatus Enghoff 2018, sp. nov." docType="treatment" docUuid="B9ACE322-7B35-4879-9DC4-B5F08E666121" docUuidSource="ZooBank" docVersion="17" lastPageNumber="23" masterDocId="FFB4501CFFB2FFB1FF94FFB0892EFF8E" masterDocTitle="A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="29" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547161722869" originalUpdateUser="yanell" pageNumber="20" updateTime="1698513679935" updateUser="ExternalLinkService">
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<mods:mods id="D890B946D59C75C088A5E68D3CB88B3A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CFFC18728B78B64243D61A2FA3D5DD35">
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<mods:title id="F34D6AF7A9BD94CED3889CDCA804149E">A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2BC6E7E5EC46205794292002CB2994A1" type="personal">
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<mods:roleTerm id="7F23E60806B06BDE3FEA3D9E92D6319C">Author</mods:roleTerm>
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<mods:namePart id="56650DC6A843F4CA5270FD3DA3E3E6F6">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="DD0A25CC9D3B9D032A3D0FDC6338ED47">text</mods:typeOfResource>
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<mods:title id="538308EF46F11233B24426C1A701CC3B">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="B641051891967CF14E047F41ACC181CE">
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<mods:date id="7A9AD119C2BC09AE8889665D83AD27A3">2018</mods:date>
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<mods:detail id="FC128CC0D38274A903D2C5C3E6BFFB26" type="pubDate">
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<mods:number id="8458D16BA47B9B1AD20C5786AA0B8A99">2018-01-11</mods:number>
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<mods:detail id="38707473FA799865B52F6648032875FB" type="volume">
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<mods:number id="3E4752ADAC165FF863B32A23B73DD6E7">394</mods:number>
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<mods:classification id="EE9357340843440A760FA161F2A75115">journal article</mods:classification>
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<mods:identifier id="7ADB4EDE505BFFCA0F2D31EC849F98F5" type="DOI">10.5852/ejt.2018.394</mods:identifier>
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<mods:identifier id="E30058B0CCFD8A80C7A208EC9F26C82D" type="Zenodo-Dep">1146158</mods:identifier>
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<mods:identifier id="0BFDAF63D0082EB23CECD61733E3502F" type="ZooBank">94028C61-FAC5-4A21-BF2D-A75BBF3CCC4D</mods:identifier>
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<treatment id="038D2864FFA1FFA7FDD9F98C8D9BF99B" ID-DOI="http://doi.org/10.5281/zenodo.5613994" ID-GBIF-Taxon="140251448" ID-Zenodo-Dep="5613994" LSID="urn:lsid:zoobank.org:act:B9ACE322-7B35-4879-9DC4-B5F08E666121" httpUri="http://treatment.plazi.org/id/038D2864FFA1FFA7FDD9F98C8D9BF99B" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
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<subSubSection id="C33ECAF9FFA1FFA2FDD9F98C8AC9F9D8" box="[589,999,1595,1622]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8B9B9972FFA1FFA2FDD9F98C8AC9F9D8" blockId="19.[370,1217,1595,1692]" box="[589,999,1595,1622]" pageId="19" pageNumber="20">
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<heading id="D0D32E1EFFA1FFA2FDD9F98C8AC9F9D8" box="[589,999,1595,1622]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
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<taxonomicName id="4C24E2F1FFA1FFA2FDD9F98C8AABF9D8" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[589,901,1595,1622]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fortehamatus" status="sp. nov.">
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<emphasis id="B9504560FFA1FFA2FDD9F98C8AABF9D8" bold="true" box="[589,901,1595,1622]" italics="true" pageId="19" pageNumber="20">Spinotarsus fortehamatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A263F81BFFA1FFA2FC18F98C8AC9F9D8" box="[908,999,1596,1622]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33ECAF9FFA1FFA2FEE6F9EF8A12F912" pageId="19" pageNumber="20" type="description">
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||
<paragraph id="8B9B9972FFA1FFA2FEE6F9EF8DEFF9F4" blockId="19.[370,1217,1595,1692]" box="[370,1217,1631,1658]" pageId="19" pageNumber="20">urn:lsid:zoobank.org:act:B9ACE322-7B35-4879-9DC4-B5F08E666121</paragraph>
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<paragraph id="8B9B9972FFA1FFA2FD63F9328A12F912" blockId="19.[370,1217,1595,1692]" box="[759,828,1666,1692]" pageId="19" pageNumber="20">
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<figureCitation id="131F85F7FFA1FFA2FD63F9328A12F912" box="[759,828,1666,1692]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ECAF9FFA1FFA2FF29F9738B3BF8DC" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="8B9B9972FFA1FFA2FF29F9738818F953" blockId="19.[189,310,1731,1757]" box="[189,310,1731,1757]" pageId="19" pageNumber="20">
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||
<heading id="D0D32E1EFFA1FFA2FF29F9738818F953" bold="true" box="[189,310,1731,1757]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
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<emphasis id="B9504560FFA1FFA2FF29F9738818F953" bold="true" box="[189,310,1731,1757]" pageId="19" pageNumber="20">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9B9972FFA1FFA2FF29F9428B3BF8DC" blockId="19.[189,1399,1778,1875]" pageId="19" pageNumber="20">
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A species of
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<taxonomicName id="4C24E2F1FFA1FFA2FEC3F94288CCF882" authorityName="Attems" authorityYear="1909" box="[343,482,1778,1804]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B9504560FFA1FFA2FEC3F94288CCF882" box="[343,482,1778,1804]" italics="true" pageId="19" pageNumber="20">Spinotarsus</emphasis>
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</taxonomicName>
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characterized by the very large, regularly curving lateral metaplical spine, in combination with the presence of a spine at the base of the solenomerite and a row of denticles on the terminal part of the telomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ECAF9FFA1FFA2FF29F8CA8BC1F868" pageId="19" pageNumber="20" type="etymology">
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<paragraph id="8B9B9972FFA1FFA2FF29F8CA886AF81A" blockId="19.[189,1398,1914,2022]" box="[189,324,1914,1940]" pageId="19" pageNumber="20">
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<heading id="D0D32E1EFFA1FFA2FF29F8CA886AF81A" bold="true" box="[189,324,1914,1940]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
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<emphasis id="B9504560FFA1FFA2FF29F8CA886AF81A" bold="true" box="[189,324,1914,1940]" pageId="19" pageNumber="20">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9B9972FFA1FFA2FF29F8188BC1F868" blockId="19.[189,1398,1914,2022]" pageId="19" pageNumber="20">The name is a composite Latin adjective, meaning “with a strong hook” and referring to the strong lateral, hook-shaped spine of the gonopod coxa.</paragraph>
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</subSubSection>
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<subSubSection id="C33ECAF9FFA6FFA5FF29FEBB8DDDFE2F" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8B9B9972FFA6FFA5FF29FEBB8B39FEAB" blockId="20.[189,535,267,293]" box="[189,535,267,293]" pageId="20" pageNumber="21">
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<emphasis id="B9504560FFA6FFA5FF29FEBB88A2FEAB" bold="true" box="[189,396,267,293]" pageId="20" pageNumber="21">Material studied</emphasis>
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(total: 1 ♂)
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</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF7EFE8A8872FEDA" blockId="20.[189,1398,314,418]" box="[234,348,314,340]" pageId="20" pageNumber="21">
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<emphasis id="B9504560FFA6FFA5FF7EFE8A8872FEDA" bold="true" box="[234,348,314,340]" pageId="20" pageNumber="21">Holotype</emphasis>
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</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FED48DDDFE2F" blockId="20.[189,1398,314,418]" pageId="20" pageNumber="21">
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<materialsCitation id="3B4C932FFFA6FFA5FF29FED48DDDFE2F" ID-GBIF-Occurrence="1802834928" collectingDate="2014-02-07" collectionCode="ZMUC" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mountains National Park" elevation="850" latitude="-7.763222" location="Udzungwa Mountains National Park, Sanje Kati camp and plot" longLatPrecision="1" longitude="36.886223" municipality="Udzungwa Mountains" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
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<collectingCountry id="F333D9E2FFA6FFA5FF29FED48879FEF0" box="[189,343,356,382]" name="Tanzania" pageId="20" pageNumber="21">TANZANIA</collectingCountry>
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:
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<specimenCount id="9D2252FBFFA6FFA5FEF1FED58853FEF1" box="[357,381,357,383]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
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,
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<collectingRegion id="49E05790FFA6FFA5FE18FED58B4BFEF0" box="[396,613,356,383]" country="Tanzania" name="Morogoro" pageId="20" pageNumber="21">Morogoro Region</collectingRegion>
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,
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<location id="8EFBCFA9FFA6FFA5FDE0FED48C5CFEF0" LSID="urn:lsid:plazi:treatment:038D2864FFA1FFA7FDD9F98C8D9BF99B:8EFBCFA9FFA6FFA5FDE0FED48C5CFEF0" box="[628,1394,356,383]" country="Tanzania" county="Udzungwa Mountains National Park" latitude="-7.763222" longLatPrecision="1" longitude="36.886223" municipality="Udzungwa Mountains" name="Udzungwa Mountains National Park, Sanje Kati camp and plot" pageId="20" pageNumber="21" stateProvince="Morogoro Region">Udzungwa Mountains National Park, Sanje Kati camp and plot</location>
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,
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<geoCoordinate id="EE10FFB5FFA6FFA5FF29FE36884DFE2F" box="[189,355,390,418]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-7.763222">07°45′47.6′′ S</geoCoordinate>
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,
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||
<geoCoordinate id="EE10FFB5FFA6FFA5FEE4FE368B37FE2C" box="[368,537,390,418]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="36.886223">36°53′10.4′′ E</geoCoordinate>
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,
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<elevation id="00097E41FFA6FFA5FDB1FE378B81FE2F" box="[549,687,391,418]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="21" unit="m" value="850.0">
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<quantity id="4CDC3497FFA6FFA5FDB1FE378B5EFE2C" box="[549,624,391,418]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="21" unit="m" value="850.0">850 m</quantity>
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a.s.l.
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</elevation>
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,
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<date id="FF9ABFB2FFA6FFA5FD28FE388A65FE2F" box="[700,843,391,418]" pageId="20" pageNumber="21" value="2014-02-07">
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<collectingDate id="EFDE465AFFA6FFA5FD28FE388A65FE2F" box="[700,843,391,418]" pageId="20" pageNumber="21" value="2014-02-07">7 Feb. 2014</collectingDate>
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</date>
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,
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<collectorName id="26D1FCA4FFA6FFA5FCC2FE388D67FE2F" box="[854,1097,391,418]" pageId="20" pageNumber="21">J. Malumbres-Olarte</collectorName>
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leg. (
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<collectionCode id="ED3501B7FFA6FFA5FB1FFE378DC9FE2F" box="[1163,1255,391,417]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
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||
).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ECAF9FFA6FFA5FF29FE798817F83B" pageId="20" pageNumber="21" type="description">
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||
<paragraph id="8B9B9972FFA6FFA5FF29FE798860FE6D" blockId="20.[189,334,457,483]" box="[189,334,457,483]" pageId="20" pageNumber="21">
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||
<heading id="D0D32E1EFFA6FFA5FF29FE798860FE6D" bold="true" box="[189,334,457,483]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="B9504560FFA6FFA5FF29FE798860FE6D" bold="true" box="[189,334,457,483]" pageId="20" pageNumber="21">Description</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B9B9972FFA6FFA5FF7EFE488804FD9C" blockId="20.[189,1276,504,573]" box="[234,298,504,530]" pageId="20" pageNumber="21">
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||
<emphasis id="B9504560FFA6FFA5FF7EFE488804FD9C" bold="true" box="[234,298,504,530]" pageId="20" pageNumber="21">Male</emphasis>
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||
</paragraph>
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||
<paragraph id="8B9B9972FFA6FFA5FF29FD928DD2FDB2" blockId="20.[189,1276,504,573]" box="[189,1276,546,573]" pageId="20" pageNumber="21">
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||
SIZE. Length ca
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||
<quantity id="4CDC3497FFA6FFA5FEECFD92889BFDB3" box="[376,437,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="cm" value="6.0">6 cm</quantity>
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, diameter 5.0 mm, 59 podous rings, no apodous rings in front of telson.
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</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FDD48D8FFC84" blockId="20.[189,1398,611,779]" pageId="20" pageNumber="21">COLOUR. Apparently quite well-preserved after 3 years in alcohol: head below antennae yellowish; head above antennae blackish, parietal and interocular furrows contrastingly yellow; antennae brown, collum blackish with lighter margins; body rings light grey, metazona with black sputtering, posterior ⅓ of metazona amber; a yellow dorsal longitudinal stripe on posterior ¼ of body; legs yellow; preanal ring grey, dorsally yellow; anal valves blackish with lighter margin; subanal scale yellow.</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FC828A6DFC1C" blockId="20.[189,1399,818,915]" pageId="20" pageNumber="21">HEAD. Parietal and interocular furrows present; 7 supralabral setae. Antennae reaching to ring 6 when folded back. Eyes reaching to median tangent to antennal socket, each with nine rows of ommatidia, 7 horizontal rows, and a total of ca 38 ommatidia each.</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FC0A8ABFFC5D" blockId="20.[189,913,953,980]" box="[189,913,953,980]" pageId="20" pageNumber="21">COLLUM. Lateral lobes subrectangular, each with two furrows.</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FC4C8D75FBB7" blockId="20.[189,1398,1019,1081]" pageId="20" pageNumber="21">BODY RINGS. Unvaulted; metazona and posterior part of prosoma with numerous short very fine longitudinal furrows; ozopores ca ⅓ of metazona length behind straight suture.</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FBD08C3FFBF5" blockId="20.[189,1297,1119,1147]" box="[189,1297,1119,1147]" pageId="20" pageNumber="21">OZOPORES. Missing from rings 31 (left side), 47 (both sides), 56 (right), 58 (left) and 59 (both).</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FB118A00FB50" blockId="20.[189,1398,1185,1247]" pageId="20" pageNumber="21">ANAL VALVES. Each with a stout dorsal spine and a tiny ventral knob-like spine; mesal margin slightly raised, setiferous tubercles indistinct, not on ravelins.</paragraph>
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<paragraph id="8B9B9972FFA6FFA5FF29FAB68D76FAAE" blockId="20.[189,1112,1286,1313]" box="[189,1112,1286,1313]" pageId="20" pageNumber="21">
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||
LIMBUS (
|
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<figureCitation id="131F85F7FFA6FFA5FEB5FAB68850FAAE" box="[289,382,1286,1312]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9G</figureCitation>
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). With long (more than twice as long as broad), pointed lobes.
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</paragraph>
|
||
<paragraph id="8B9B9972FFA6FFA5FF29FAF78874F99C" blockId="20.[189,1398,1351,1554]" pageId="20" pageNumber="21">
|
||
GONOPOD COXA (
|
||
<figureCitation id="131F85F7FFA6FFA5FEE8FAF788FBFAEF" box="[380,469,1351,1377]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9F</figureCitation>
|
||
). Slightly more than twice as long as broad. Proplica ending in small proplical lobe (
|
||
<emphasis id="B9504560FFA6FFA5FF6BFADA880AFA0A" box="[255,292,1386,1412]" italics="true" pageId="20" pageNumber="21">ppl</emphasis>
|
||
). Metaplica longer than proplica, apically forming a “cucullus” (
|
||
<emphasis id="B9504560FFA6FFA5FB9BFADB8D05FA0A" box="[1039,1067,1387,1412]" italics="true" pageId="20" pageNumber="21">cu</emphasis>
|
||
). Mesal margin of metaplica with a curved lamellar process (
|
||
<emphasis id="B9504560FFA6FFA5FDBAFA3E8B74FA26" box="[558,602,1422,1448]" italics="true" pageId="20" pageNumber="21">mlp</emphasis>
|
||
) just distal to exit point of telopodite (“arculus”). Metaplica laterally with a very large spine (
|
||
<emphasis id="B9504560FFA6FFA5FE41FA028B2AFA45" box="[469,516,1458,1483]" italics="true" pageId="20" pageNumber="21">msp</emphasis>
|
||
) curving distad in a latero-posterior plane. Proplical lobe, lateral metaplical spine and a stripe parallel to lateral metaplical margin from spine to tip of cucullus black (not visible on SEM image).
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA6FFA5FF29F9898806F8C7" blockId="20.[189,1399,1593,1865]" pageId="20" pageNumber="21">
|
||
GONOPOD TELOPODITE (
|
||
<figureCitation id="131F85F7FFA6FFA5FE5DF9898B65F9DD" box="[457,587,1593,1619]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9A–E</figureCitation>
|
||
). Arculus 90°. A very long post-torsal spine (
|
||
<emphasis id="B9504560FFA6FFA5FB1EF9898D82F9DC" box="[1162,1196,1593,1618]" italics="true" pageId="20" pageNumber="21">pts</emphasis>
|
||
) making almost 1½ turns around torsotope (
|
||
<emphasis id="B9504560FFA6FFA5FD95F9ED8B3FF9F8" box="[513,529,1629,1654]" italics="true" pageId="20" pageNumber="21">tt</emphasis>
|
||
) (
|
||
<figureCitation id="131F85F7FFA6FFA5FDBEF9EC8B82F9F8" box="[554,684,1628,1654]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9C–D</figureCitation>
|
||
). Solenomere (
|
||
<emphasis id="B9504560FFA6FFA5FCC8F9EC8AABF9F8" box="[860,901,1628,1654]" italics="true" pageId="20" pageNumber="21">slm</emphasis>
|
||
) as long as telomere, thin, simple, without outgrowths, apically with oblique-longitudinal fluting.A long spine (
|
||
<emphasis id="B9504560FFA6FFA5FC45F9308AC5F917" box="[977,1003,1664,1689]" italics="true" pageId="20" pageNumber="21">ps</emphasis>
|
||
) originating at base of solenomere and projecting at right angles to it (
|
||
<figureCitation id="131F85F7FFA6FFA5FDC6F9128B9EF932" box="[594,688,1698,1724]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
|
||
). Telomere (
|
||
<emphasis id="B9504560FFA6FFA5FCD5F9138A70F932" box="[833,862,1699,1724]" italics="true" pageId="20" pageNumber="21">tm</emphasis>
|
||
) curving in an almost complete circle, basally with large spinose lamella (
|
||
<emphasis id="B9504560FFA6FFA5FD94F9758B0BF951" box="[512,549,1733,1759]" italics="true" pageId="20" pageNumber="21">bla</emphasis>
|
||
) projecting into space delimited by telomere curvature, followed by two smooth lamellae (
|
||
<emphasis id="B9504560FFA6FFA5FE19F95988CFF88D" box="[397,481,1769,1795]" italics="true" pageId="20" pageNumber="21">el1, el2</emphasis>
|
||
) projecting from outside of curvature (
|
||
<figureCitation id="131F85F7FFA6FFA5FC31F9598D2DF88D" box="[933,1027,1769,1795]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
|
||
); distal part of telomere slender, longitudinally folded, one margin with a series of black denticles (
|
||
<emphasis id="B9504560FFA6FFA5FC52F8BC8ACAF8A8" box="[966,996,1804,1830]" italics="true" pageId="20" pageNumber="21">td,</emphasis>
|
||
colour not visible on SEM image;
|
||
<figureCitation id="131F85F7FFA6FFA5FF29F89F8834F8C7" box="[189,282,1839,1865]" captionStart="Fig. 9" captionStartId="21.[189,232,1762,1788]" captionTargetBox="[192,1401,262,1728]" captionTargetId="figure@21.[176,1412,261,1729]" captionTargetPageId="21" captionText="Fig. 9. Spinotarsus fortehamatus sp. nov., holotype. A–E. Right gonopod telopodite.A. Posterior view. B. Basal lamella of telomere, enlarged. C. Anterior view. D. Posterior-mesal view. E. Distal part of solenomere. F. Right gonopod coxa, posterior and slightly mesal view. G. Limbus. Abbreviations: bla = basal lamella of telomere; cu = cucullus; el1, el2 = smooth telomeral lamellae; mlp = metaplical lamellar process; msp= metaplical spine-like process; ppl = proplical lobe; ps = proximal solenomeral spine;pts = post-torsal spine; slm = solenomere; td = telomeral denticles; tm = telomere; tt = torsotope. Scale bars:A, C–D,F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm." httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="20" pageNumber="21">Fig. 9D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA6FFA5FF7EF8C1886BF805" blockId="20.[189,325,1905,1973]" box="[234,325,1905,1931]" pageId="20" pageNumber="21">
|
||
<emphasis id="B9504560FFA6FFA5FF7EF8C1886BF805" bold="true" box="[234,325,1905,1931]" pageId="20" pageNumber="21">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA6FFA5FF29F82B8817F83B" blockId="20.[189,325,1905,1973]" box="[189,313,1947,1973]" pageId="20" pageNumber="21">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5BC9FAFFA7FFA4FF29F9528A88F85E" httpUri="https://zenodo.org/record/1146174/files/figure.png" pageId="21" pageNumber="22" targetBox="[192,1401,262,1728]" targetPageId="21">
|
||
<paragraph id="8B9B9972FFA7FFA4FF29F9528A88F85E" blockId="21.[189,1399,1762,2000]" pageId="21" pageNumber="22">
|
||
<emphasis id="B9504560FFA7FFA4FF29F9528824F972" bold="true" box="[189,266,1762,1788]" pageId="21" pageNumber="22">Fig. 9.</emphasis>
|
||
<taxonomicName id="4C24E2F1FFA7FFA4FE86F9538B11F972" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[274,575,1762,1789]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fortehamatus" status="sp. nov.">
|
||
<emphasis id="B9504560FFA7FFA4FE86F9538B11F972" box="[274,575,1762,1789]" italics="true" pageId="21" pageNumber="22">Spinotarsus fortehamatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A263F81BFFA7FFA4FDD2F9538B8DF973" box="[582,675,1763,1789]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype.
|
||
<emphasis id="B9504560FFA7FFA4FCB0F9528A75F972" bold="true" box="[804,859,1762,1788]" pageId="21" pageNumber="22">A–E</emphasis>
|
||
. Right gonopod telopodite.
|
||
<emphasis id="B9504560FFA7FFA4FB37F9528D96F972" bold="true" box="[1187,1208,1762,1788]" pageId="21" pageNumber="22">A</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="B9504560FFA7FFA4FF29F8B689FFF8AE" bold="true" box="[189,209,1798,1824]" pageId="21" pageNumber="22">B</emphasis>
|
||
. Basal lamella of telomere, enlarged.
|
||
<emphasis id="B9504560FFA7FFA4FD35F8B58B98F891" bold="true" box="[673,694,1797,1823]" pageId="21" pageNumber="22">C</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="B9504560FFA7FFA4FCE9F8B68ABCF8AE" bold="true" box="[893,914,1798,1824]" pageId="21" pageNumber="22">D</emphasis>
|
||
. Posterior-mesal view.
|
||
<emphasis id="B9504560FFA7FFA4FB24F8B68DEAF8AE" bold="true" box="[1200,1220,1798,1824]" pageId="21" pageNumber="22">E</emphasis>
|
||
. Distal part of solenomere.
|
||
<emphasis id="B9504560FFA7FFA4FECCF8998844F8CD" bold="true" box="[344,362,1833,1859]" pageId="21" pageNumber="22">F</emphasis>
|
||
. Right gonopod coxa, posterior and slightly mesal view.
|
||
<emphasis id="B9504560FFA7FFA4FBA6F8988D67F8CC" bold="true" box="[1074,1097,1832,1858]" pageId="21" pageNumber="22">G</emphasis>
|
||
. Limbus. Abbreviations:
|
||
<emphasis id="B9504560FFA7FFA4FF29F8FC89CCF8E8" box="[189,226,1868,1894]" italics="true" pageId="21" pageNumber="22">bla</emphasis>
|
||
= basal lamella of telomere;
|
||
<emphasis id="B9504560FFA7FFA4FDAAF8FD8B74F8E8" box="[574,602,1869,1894]" italics="true" pageId="21" pageNumber="22">cu</emphasis>
|
||
= cucullus;
|
||
<emphasis id="B9504560FFA7FFA4FD79F8FC8A3FF8E8" box="[749,785,1868,1894]" italics="true" pageId="21" pageNumber="22">el1</emphasis>
|
||
,
|
||
<emphasis id="B9504560FFA7FFA4FCB5F8FC8A6AF8E8" box="[801,836,1868,1894]" italics="true" pageId="21" pageNumber="22">el2</emphasis>
|
||
= smooth telomeral lamellae;
|
||
<emphasis id="B9504560FFA7FFA4FB3BF8FC8DF5F8E8" box="[1199,1243,1868,1894]" italics="true" pageId="21" pageNumber="22">mlp</emphasis>
|
||
= metaplical lamellar process;
|
||
<emphasis id="B9504560FFA7FFA4FE1AF8C08893F807" box="[398,445,1904,1929]" italics="true" pageId="21" pageNumber="22">msp</emphasis>
|
||
= metaplical spine-like process;
|
||
<emphasis id="B9504560FFA7FFA4FCAAF8DF8A4DF807" box="[830,867,1903,1929]" italics="true" pageId="21" pageNumber="22">ppl</emphasis>
|
||
= proplical lobe;
|
||
<emphasis id="B9504560FFA7FFA4FBADF8C08D7DF807" box="[1081,1107,1904,1929]" italics="true" pageId="21" pageNumber="22">ps</emphasis>
|
||
= proximal solenomeral spine;
|
||
<emphasis id="B9504560FFA7FFA4FE9EF8238802F822" box="[266,300,1939,1964]" italics="true" pageId="21" pageNumber="22">pts</emphasis>
|
||
= post-torsal spine;
|
||
<emphasis id="B9504560FFA7FFA4FD89F8228B68F822" box="[541,582,1938,1964]" italics="true" pageId="21" pageNumber="22">slm</emphasis>
|
||
= solenomere;
|
||
<emphasis id="B9504560FFA7FFA4FD6FF8228A3CF822" box="[763,786,1938,1964]" italics="true" pageId="21" pageNumber="22">td</emphasis>
|
||
= telomeral denticles;
|
||
<emphasis id="B9504560FFA7FFA4FBB4F8238D13F822" box="[1056,1085,1939,1964]" italics="true" pageId="21" pageNumber="22">tm</emphasis>
|
||
= telomere;
|
||
<emphasis id="B9504560FFA7FFA4FB46F8238DCCF822" box="[1234,1250,1939,1964]" italics="true" pageId="21" pageNumber="22">tt</emphasis>
|
||
= torsotope. Scale bars: A, C–D, F = 0.2 mm; B, E = 0.1 mm; G = 0.01 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33ECAF9FFA4FFA7FF29FEBB8C58FC96" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FEBB8801FEAB" blockId="22.[189,303,267,293]" box="[189,303,267,293]" pageId="22" pageNumber="23">
|
||
<heading id="D0D32E1EFFA4FFA7FF29FEBB8801FEAB" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
|
||
<emphasis id="B9504560FFA4FFA7FF29FEBB8801FEAB" bold="true" box="[189,303,267,293]" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FE898A9DFD59" blockId="22.[189,1399,313,727]" pageId="22" pageNumber="23">
|
||
The basal spinose lamella of the telomere (
|
||
<emphasis id="B9504560FFA4FFA7FD50FE898BC7FEDD" box="[708,745,313,339]" italics="true" pageId="22" pageNumber="23">bla</emphasis>
|
||
) is a key character for
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FB91FE8A8DBEFEDA" authorityName="Attems" authorityYear="1909" box="[1029,1168,314,340]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9504560FFA4FFA7FB91FE8A8DBEFEDA" box="[1029,1168,314,340]" italics="true" pageId="22" pageNumber="23">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
(the name actually refers to the lamella – the telomere was formerly referred to as the tarsus). Many species of
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FB78FEED8C59FEF9" authorityName="Attems" authorityYear="1909" box="[1260,1399,349,375]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9504560FFA4FFA7FB78FEED8C59FEF9" box="[1260,1399,349,375]" italics="true" pageId="22" pageNumber="23">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
possess an additional apomorphy, namely a darkly sclerotized ridge on the posterior surface of the telomere, but such a ridge is absent in
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FD03FE138A7CFE33" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[663,850,419,445]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fortehamatus" status="sp. nov.">
|
||
<emphasis id="B9504560FFA4FFA7FD03FE138A7CFE33" box="[663,850,419,445]" italics="true" pageId="22" pageNumber="23">S. fortehamatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A263F81BFFA4FFA7FCC8FE148A94FE30" box="[860,954,420,446]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
In the most recent key to species of
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FF29FE778866FE6F" authorityName="Attems" authorityYear="1909" box="[189,328,455,481]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9504560FFA4FFA7FF29FE778866FE6F" box="[189,328,455,481]" italics="true" pageId="22" pageNumber="23">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FEC1FE768B7AFE6E" author="Kraus O." box="[341,596,454,480]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" page="110 - 113" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus 1966: 110–113</bibRefCitation>
|
||
), the new species keys out without problems to couplet 60, as follows (couplet texts translated from German and adapted to current terminology, characters of
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FB2BFE5A8C59FD8D" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[1215,1399,489,516]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fortehamatus" status="sp. nov.">
|
||
<emphasis id="B9504560FFA4FFA7FB2BFE5A8C59FD8D" box="[1215,1399,489,516]" italics="true" pageId="22" pageNumber="23">S. fortehamatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A263F81BFFA4FFA7FF29FDBD8834FDA9" box="[189,282,525,551]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in bold): couplet 1: Gonopodal metaplica with
|
||
<emphasis id="B9504560FFA4FFA7FCC9FDBD8AA7FDA9" bold="true" box="[861,905,525,551]" pageId="22" pageNumber="23">one</emphasis>
|
||
or several dark sclerotized lateral spines which stand clearly out in strict oral view → couplet 38:
|
||
<emphasis id="B9504560FFA4FFA7FCF6FD808AB8FDC4" bold="true" box="[866,918,560,586]" pageId="22" pageNumber="23">One</emphasis>
|
||
or more spine(s) (“Postfemoraldorn”) at base of solenomere present → couplet 59:
|
||
<emphasis id="B9504560FFA4FFA7FD39FDE38BCFFDE3" bold="true" box="[685,737,595,621]" pageId="22" pageNumber="23">One</emphasis>
|
||
spine at base of solenomere → couplet 60. “Couplet” 60 is, however, deficient in the key, which can be seen by comparison with the older key by
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FB48FDC788B7FD3A" author="Kraus O." journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="22" pageNumber="23" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus (1960: 122, “couplet” 42)</bibRefCitation>
|
||
. The “couplet” in question is actually a “triplet”, but the third option is lacking in the 1966 key. In translation, “triplet”
|
||
<quantity id="4CDC3497FFA4FFA7FDDCFD0C8BADFD58" box="[584,643,700,726]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="23" unit="in" value="60.0">60 in</quantity>
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FD1FFD0D8A08FD59" author="Kraus O." box="[651,806,700,727]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus (1966)</bibRefCitation>
|
||
should run:
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FD4E8C58FC96" blockId="22.[189,1399,766,898]" box="[189,1398,766,793]" pageId="22" pageNumber="23">
|
||
<key id="FFB5CE43FFA4FFA7FF29FD4E8C58FC96" box="[189,1398,766,793]" pageId="22" pageNumber="23">
|
||
<keyStep id="30D081D7FFA4FFA7FF29FD4E8C58FC96" box="[189,1398,766,793]" pageId="22" pageNumber="23">
|
||
<keyLead id="30D53A47FFA4FFA7FF29FD4E8C58FC96" box="[189,1398,766,793]" pageId="22" pageNumber="23">
|
||
60. Metaplica distally coarsely serrate ...................................................................................
|
||
<emphasis id="B9504560FFA4FFA7FB73FD4E8C58FC96" box="[1255,1398,766,792]" italics="true" pageId="22" pageNumber="23">denticulatus</emphasis>
|
||
</keyLead>
|
||
</keyStep>
|
||
</key>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFA4FFA7FF29FC928C59FC0C" pageId="22" pageNumber="23" type="key">
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FC928C58FCD0" blockId="22.[189,1399,766,898]" pageId="22" pageNumber="23">
|
||
<keyLead id="30D53A47FFA4FFA7FF29FC928C58FCD0" pageId="22" pageNumber="23">– Metaplica distally ending in a slender, spine-like process with a blackened tip; this process inserting apico-mesally...................................................................................................................................61</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA4FFA7FF55FCD88C59FC0C" blockId="22.[189,1399,766,898]" box="[193,1399,871,898]" pageId="22" pageNumber="23">
|
||
<keyLead id="30D53A47FFA4FFA7FF55FCD88C59FC0C" box="[193,1399,871,898]" pageId="22" pageNumber="23">
|
||
–
|
||
<emphasis id="B9504560FFA4FFA7FF7EFCD78B05FC0F" bold="true" box="[234,555,871,897]" pageId="22" pageNumber="23">Tip of metaplica different</emphasis>
|
||
..............................................................................................................65
|
||
</keyLead>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFA4FFA7FF29FC198D9BF99B" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FC198D12FAC8" blockId="22.[189,1399,937,1351]" pageId="22" pageNumber="23">
|
||
Continuing in couplet 65, the new species fulfils the second alternative: Lateral metaplical spine directed ±
|
||
<emphasis id="B9504560FFA4FFA7FF45FC7C880AFC68" bold="true" box="[209,292,972,998]" pageId="22" pageNumber="23">apicad</emphasis>
|
||
→ couplet 68: Metaplica apically
|
||
<emphasis id="B9504560FFA4FFA7FD5DFC7C8A88FC68" bold="true" box="[713,934,972,998]" pageId="22" pageNumber="23">without a process</emphasis>
|
||
or just with a delicate hyaline, spoonshaped appendage → couplet 81: Terminal lamella of telomere with spiny margin and
|
||
<emphasis id="B9504560FFA4FFA7FB21FC5F8DEAFB87" bold="true" box="[1205,1220,1007,1033]" pageId="22" pageNumber="23">1</emphasis>
|
||
–2 longitudinal
|
||
<emphasis id="B9504560FFA4FFA7FF29FBA389C3FBA3" bold="true" box="[189,237,1043,1069]" pageId="22" pageNumber="23">row</emphasis>
|
||
s of denticles on the internal surface (→ couplet 82) vs Terminal lamella of telomere
|
||
<emphasis id="B9504560FFA4FFA7FB7BFBA38836FBDE" bold="true" pageId="22" pageNumber="23">completely smooth</emphasis>
|
||
, internal surface of telomere without obvious spine rows (→ couplet 84). The new species has a smooth margin of the terminal part of the telomere, but it also has a very obvious row of spines on the internal surface of the telomere. It thus fits one criterium of each alternative. In addition to its unique combination of key characters, it also differs from congeners in the shape of the lateral metaplical spine: none of the almost one hundred described species of
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FC15FB738D22FB53" authorityName="Attems" authorityYear="1909" box="[897,1036,1219,1245]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9504560FFA4FFA7FC15FB738D22FB53" box="[897,1036,1219,1245]" italics="true" pageId="22" pageNumber="23">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
have an equally large, equally smoothly curved lateral metaplical spine. Those which come closest in this character, i.e.,
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FB62FB5688EEFAAA" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="demotus">
|
||
<emphasis id="B9504560FFA4FFA7FB62FB568C59FA8E" box="[1270,1399,1254,1280]" italics="true" pageId="22" pageNumber="23">S. demotus</emphasis>
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FF29FABA8863FAAD" author="Kraus O." box="[189,333,1289,1316]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus, 1966</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F333D9E2FFA4FFA7FEC8FAB98896FAAA" box="[348,440,1289,1316]" name="Angola" pageId="22" pageNumber="23">Angola</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FE58FABA8BCAFAAD" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" box="[460,740,1289,1316]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kruegeri">
|
||
<emphasis id="B9504560FFA4FFA7FE58FABA8B63FAAD" box="[460,589,1289,1316]" italics="true" pageId="22" pageNumber="23">S. kruegeri</emphasis>
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FDC0FABA8BCAFAAD" author="Kraus O." box="[596,740,1289,1316]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F333D9E2FFA4FFA7FD67FAB98AA1FAAA" box="[755,911,1289,1316]" name="South Africa" pageId="22" pageNumber="23">
|
||
<collectingRegion id="49E05790FFA4FFA7FD67FAB98A17FAAD" box="[755,825,1289,1315]" country="Tanzania" name="Kusini Unguja" pageId="22" pageNumber="23">South</collectingRegion>
|
||
Africa
|
||
</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FC37FABA8C59FAAA" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" box="[931,1399,1289,1316]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="lanceolatus">
|
||
<emphasis id="B9504560FFA4FFA7FC37FABA8D69FAAD" box="[931,1095,1289,1316]" italics="true" pageId="22" pageNumber="23">S. lanceolatus</emphasis>
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FBDBFABA8DF0FAAD" author="Kraus O." box="[1103,1246,1289,1316]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus, 1966</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F333D9E2FFA4FFA7FB79FAB98C40FAAA" box="[1261,1390,1289,1316]" name="Zimbabwe" pageId="22" pageNumber="23">Zimbabwe</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FF7BFA9D88D0FAC8" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" box="[239,510,1324,1351]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9504560FFA4FFA7FF7BFA9D8849FAC8" box="[239,359,1324,1351]" italics="true" pageId="22" pageNumber="23">S. pusillus</emphasis>
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FEFBFA9D88D0FAC8" author="Kraus O." box="[367,510,1324,1351]" firstAuthor="Kraus" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pageId="22" pageNumber="23" pagination="1 - 143" part="512" refId="ref10157" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." title="Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha)" type="journal article" year="1966">Kraus, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F333D9E2FFA4FFA7FD99FA9C8B87FAC9" box="[525,681,1324,1351]" name="South Africa" pageId="22" pageNumber="23">
|
||
<collectingRegion id="49E05790FFA4FFA7FD99FA9C8B7DFAC8" box="[525,595,1324,1350]" country="Tanzania" name="Kusini Unguja" pageId="22" pageNumber="23">South</collectingRegion>
|
||
Africa
|
||
</collectingCountry>
|
||
), differ clearly in other characters.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA4FFA7FF29FADE8D9BF99B" blockId="22.[189,1399,1390,1557]" pageId="22" pageNumber="23">
|
||
The lack of ozopores on one or both sides of certain body rings is a highly unusual trait. In juliformian millipedes, ozopores are normally present in an uninterrupted series usually starting on body ring 6, rarely on ring 5, exceptionally on ring 3, and continuing to include the last podous ring. Up to now, the only exception to this rule is constituted by two species of the family
|
||
<taxonomicName id="4C24E2F1FFA4FFA7FBDAFA678DD0FA7F" authorityName="Pocock" authorityYear="1903" box="[1102,1278,1495,1521]" class="Diplopoda" family="Mongoliulidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">Mongoliulidae</taxonomicName>
|
||
, in which ozoporeless body rings occur in a regular pattern along the body (
|
||
<bibRefCitation id="EFB5E483FFA4FFA7FC2AFA4A8D87F99B" author="Enghoff" box="[958,1193,1530,1557]" etAl="et al." firstAuthor="Enghoff" journalOrPublisher="Zootaxa" pageId="22" pageNumber="23" pagination="501 - 530" part="4273" publicationUrl="https://doi.org/10.11646/zootaxa.4273.4.3" refId="ref9542" refString="Enghoff H. 2017 a. A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas. Zootaxa 4273 (4): 501 - 530. https: // doi. org / 10.11646 / zootaxa. 4273.4.3" title="A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas" type="journal article" year="2017">
|
||
Enghoff
|
||
<emphasis id="B9504560FFA4FFA7FBB3FA4B8D4CF99A" box="[1063,1122,1530,1556]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |