treatments-xml/data/03/88/5D/03885D14FF87894D1A82F8AEECC6FE1D.xml
2024-06-21 12:22:17 +02:00

1180 lines
152 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="F4ADC9D2810B1E808DC730B80E020A7D" ID="10.11646/zootaxa.4161.2.1" ID-DOI="10.11646/zootaxa.4161.2.1" ID-GBIF-Dataset="03838f4b-82c0-42a0-9f6c-a8feb4f210a6" ID-ISSN="1175-5326" ID-Zenodo-Dep="263909" ID-ZooBank="B36A20C2-A9DA-4F35-89A3-2153F88673BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473055639834" checkinUser="plazi" docAuthor="Surugiu, Victor" docDate="2016" docId="03885D14FF87894D1A82F8AEECC6FE1D" docLanguage="en" docName="zootaxa.4161.2.1.pdf" docOrigin="Zootaxa 4161 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scolelepis (Scolelepis) neglecta Surugiu, 2016, new species" docType="treatment" docUuid="53D8862C-E62F-4B65-8E7F-22DFF8384A11" docUuidSource="ZooBank" docVersion="7" lastPageNumber="168" masterDocId="FFB1256CFF8D895C1A15FFE2EF14FF95" masterDocTitle="On the taxonomic status of the European Scolelepis (Scolelepis) squamata (Polychaeta: Spionidae), with description of a new species from southern Europe" masterLastPageNumber="176" masterPageNumber="151" pageNumber="161" updateTime="1698383940617" updateUser="plazi">
<mods:mods id="5BEC0BF10604BB015B1BA9CFD71FA701" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6C10E0F8189D998CABB9A052E215D222">
<mods:title id="195AD6FAA475F905BC7879B7F52B510F">On the taxonomic status of the European Scolelepis (Scolelepis) squamata (Polychaeta: Spionidae), with description of a new species from southern Europe</mods:title>
</mods:titleInfo>
<mods:name id="8691D4D415A39C98B5A93357A5285092" type="personal">
<mods:role id="481BAA713FC76691F9E18A8F6DF0FE17">
<mods:roleTerm id="74735AF444BB058C95F020DB31FAF605">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8FBF14F8602AD080C12F6132D092AA0C">Surugiu, Victor</mods:namePart>
</mods:name>
<mods:typeOfResource id="9D77614B1CA63C672EEC843AD609D44C">text</mods:typeOfResource>
<mods:relatedItem id="4856ECEEBDF22C88AAAC7126C444066A" type="host">
<mods:titleInfo id="1E1E77379B9ABC9B1A70614F15506608">
<mods:title id="51D2D953BE200A568B4E1CD6E4E4B234">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5479D093503EF2155E015EED20FFBFB7">
<mods:date id="D98B313E5D19A8BA023D708A3D0AB45F">2016</mods:date>
<mods:detail id="89956BCCE35E9ADEF0EE509DA6256BAF" type="volume">
<mods:number id="BACBDAD17C990003BEB4D123A28E9994">4161</mods:number>
</mods:detail>
<mods:detail id="047323F5B7571E653137E898D94A71F8" type="issue">
<mods:number id="677A12B6C9CF527B3B3317A47A7BD664">2</mods:number>
</mods:detail>
<mods:extent id="139AB7E7722E8B70A44B2B14E4044D09" unit="page">
<mods:start id="85D44517B0BA1405EB09D273A97B4ED4">151</mods:start>
<mods:end id="DA390F5C87682DADA0997BC81978CC6E">176</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B037612411AC39E11728A565B5BC2453">journal article</mods:classification>
<mods:identifier id="EDE347151684EBE7BCF15EB896D9264F" type="DOI">10.11646/zootaxa.4161.2.1</mods:identifier>
<mods:identifier id="04055016AE104399E75C018E5F3C2B17" type="GBIF-Dataset">03838f4b-82c0-42a0-9f6c-a8feb4f210a6</mods:identifier>
<mods:identifier id="11D34D03B33EDFE222C5454734CFB8D7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6C9C78BA81578E0454C8ADD43393A7D3" type="Zenodo-Dep">263909</mods:identifier>
<mods:identifier id="9CD6156FEC74D8D5202B008DEEDF3C88" type="ZooBank">B36A20C2-A9DA-4F35-89A3-2153F88673BC</mods:identifier>
</mods:mods>
<treatment id="03885D14FF87894D1A82F8AEECC6FE1D" ID-DOI="http://doi.org/10.5281/zenodo.5613791" ID-GBIF-Taxon="124409960" ID-Zenodo-Dep="5613791" LSID="urn:lsid:zoobank.org:act:53D8862C-E62F-4B65-8E7F-22DFF8384A11" httpUri="http://treatment.plazi.org/id/03885D14FF87894D1A82F8AEECC6FE1D" lastPageId="17" lastPageNumber="168" pageId="10" pageNumber="161">
<subSubSection id="C33BBF89FF8789561A82F8AEEE08F81C" pageId="10" pageNumber="161" type="nomenclature">
<paragraph id="8B9EEC02FF8789561A82F8AEEDB9F8F2" blockId="10.[151,685,1868,1929]" box="[151,685,1868,1895]" pageId="10" pageNumber="161">
<heading id="D0D65B6EFF8789561A82F8AEEDB9F8F2" bold="true" box="[151,685,1868,1895]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<emphasis id="B9553010FF8789561A82F8AEEDB9F8F2" bold="true" box="[151,685,1868,1895]" pageId="10" pageNumber="161">
<taxonomicName id="4C219781FF8789561A82F8AEED1AF8F3" ID-CoL="4VRN6" box="[151,526,1868,1895]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="10" pageNumber="161" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8789561A82F8AEEE04F8F3" bold="true" box="[151,272,1868,1894]" italics="true" pageId="10" pageNumber="161">Scolelepis</emphasis>
(
<emphasis id="B9553010FF8789561B35F8AEEE8CF8F3" bold="true" box="[288,408,1868,1894]" italics="true" pageId="10" pageNumber="161">Scolelepis</emphasis>
)
<emphasis id="B9553010FF8789561BBFF8AEED1AF8F3" bold="true" box="[426,526,1868,1894]" italics="true" pageId="10" pageNumber="161">neglecta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2668D6BFF8789561808F8AFEDB9F8F2" box="[541,685,1869,1895]" pageId="10" pageNumber="161" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9EEC02FF8789561A82F892EE08F81C" blockId="10.[151,685,1868,1929]" box="[151,284,1904,1929]" pageId="10" pageNumber="161">
<figureCitation id="131AF087FF8789561A82F892EE08F81C" box="[151,284,1904,1929]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="13.[151,250,1881,1903]" captionStartId-1="14.[151,250,1821,1843]" captionStartId-2="15.[151,250,1189,1211]" captionStartId-3="17.[151,250,1279,1301]" captionTargetBox-0="[230,1339,196,1830]" captionTargetBox-1="[260,1340,214,1782]" captionTargetBox-2="[151,1436,193,1167]" captionTargetBox-3="[188,1364,460,1240]" captionTargetId-0="figure@13.[223,1342,193,1839]" captionTargetId-1="figure@14.[240,1341,193,1794]" captionTargetId-2="figure@15.[151,1436,193,1167]" captionTargetId-3="figure@17.[176,1375,447,1248]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="17" captionText-0="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." captionText-1="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." captionText-2="FIGURE 7. Scolelepis (Scolelepis) neglecta sp. nov., SEM micrographs. A F, paratype (MNCN 3855). A, distal tip of left palp with elevated lobes, frontal-lateral view (arrowheads indicate faint splitting of lateral rows into two smaller lobes); B, distal section of right palp, frontal view, showing elevated lobes; C, basal part of right palp, lateral view (arrowhead shows smooth basal palpal sheath); D, chaetigers 13 25 (from top right to bottom left), ventro-lateral view (arrowheads indicate prechaetal notopodial lamellae; arrow indicates ventral median groove); E, chaetigers 16 21 (from top right to bottom left), dorso-lateral view (arrowheads indicate lateral ciliated organs); F, chaetigers 29 32 (from right to left), lateral view (note ventrally directed inferior part of notopodial postchaetal lamella and dorsally directed superior part of neuropodial postchaetal lamella). Scale bars: A, B = 50 µm; C = 100 µm; D = 1 mm; E = 0.5 mm; F = 0.2 mm." captionText-3="FIGURE 8. Scolelepis (Scolelepis) neglecta sp. nov., relationship between the body-size (referring to the maximum anterior width) and the first chaetiger on which a notch is present in the neuropodial postchaetal lamellae (triangles, holotype empty symbol, y = 13.28 + 10.106 x; r s = 0.931; P &lt;0.0001; n = 36), and between the body-size (referring to the maximum anterior width) and the first chaetiger on which neuropodial hooded hooks are present (circles, holotype empty symbol, y = 15.673 + 11.97 x; r s = 0.846; P &lt;0.0001; n = 36)." httpUri-0="https://zenodo.org/record/263914/files/figure.png" httpUri-1="https://zenodo.org/record/263915/files/figure.png" httpUri-2="https://zenodo.org/record/263916/files/figure.png" httpUri-3="https://zenodo.org/record/263917/files/figure.png" pageId="10" pageNumber="161">Figures 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8789571A82F85AEB96FF59" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="reference_group">
<paragraph id="8B9EEC02FF8789561A82F85AEDF0F879" blockId="10.[151,1253,1976,2028]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="AB31CB2CFF8789561A82F85AEDF0F879" pageId="10" pageNumber="161">
<taxonomicName id="4C219781FF8789561A82F85AEE85F85B" box="[151,401,1976,1998]" class="Polychaeta" family="Spionidae" genus="Scolecolepis" higherTaxonomySource="CoL" kingdom="Animalia" order="Spionida" pageId="10" pageNumber="161" phylum="Annelida" rank="species" species="squammata">
<emphasis id="B9553010FF8789561A82F85AEE85F85B" box="[151,401,1976,1998]" italics="true" pageId="10" pageNumber="161">Scolecolepis squammata</emphasis>
</taxonomicName>
[sic].—
<treatmentCitation id="0A80CA13FF8789561BC8F85AED81F85B" author="Dorjes" box="[477,661,1976,1998]" page="213" pageId="10" pageNumber="161" year="1971">
<bibRefCitation id="EFB091F3FF8789561BC8F85AED77F85B" author="Dorjes" box="[477,611,1976,1998]" pageId="10" pageNumber="161" refString="Dorjes, J. (1971) Der Golf von Gaeta (Tyrrhenisches Meer). IV. Das Makrobenthos und seine kustenparallele Zonierung. Senckenbergiana maritima, 3, 203 - 246." type="journal article" year="1971">Dörjes 1971</bibRefCitation>
: 213
</treatmentCitation>
, 215, fig. 4. Partim. Not Abildgaard,
<emphasis id="B9553010FF8789561E1AF85BEB37F85B" box="[1039,1059,1977,1998]" italics="true" pageId="10" pageNumber="161">in</emphasis>
<bibRefCitation id="EFB091F3FF8789561E3CF85AEBF5F85B" author="Muller" box="[1065,1249,1976,1998]" pageId="10" pageNumber="161" refString="Muller, O. F., Abildgaard, P. C., Vahl, M., Holten, J. S. &amp; Rathke, J. (1806) Zoologiae Danica seu Animalium Daniae et Norvegiae rariorum ac minus notorum. Descriptiones et historia. Volume IV. Volume Atlas. N. Christensen: Havniae [Copenhagen], 1 - 2, 46 pp., pls. 121 - 160." type="book chapter" year="1806">O.F. Müller, 1806</bibRefCitation>
.
<taxonomicName id="4C219781FF8789561A82F834EE4FF879" ID-CoL="4754Y" box="[151,347,2006,2028]" class="Polychaeta" family="Spionidae" genus="Nerinides" kingdom="Animalia" order="Spionida" pageId="10" pageNumber="161" phylum="Annelida" rank="species" species="cantabra">
<emphasis id="B9553010FF8789561A82F834EE4FF879" box="[151,347,2006,2028]" italics="true" pageId="10" pageNumber="161">Nerinides cantabra</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A80CA13FF8789561B6DF834ED25F879" author="Vieitez" box="[376,561,2006,2028]" page="240" pageId="10" pageNumber="161" year="1976">
<bibRefCitation id="EFB091F3FF8789561B6DF834EEEBF879" author="Vieitez" box="[376,511,2006,2028]" pageId="10" pageNumber="161" refString="Vieitez, J. M. (1976) Ecologia de Poliquetos y moluscos de la playa de Meira (Ria de Vigo). I. Estudio de las comunidades. Investigacion Pesquera, 40 (1), 223 - 248." type="journal article" year="1976">Viéitez 1976</bibRefCitation>
: 240
</treatmentCitation>
. Not
<bibRefCitation id="EFB091F3FF8789561872F834EDF5F879" author="Rioja" box="[615,737,2006,2028]" pageId="10" pageNumber="161" refString="Rioja, E. (1918) Adiciones a la fauna de anelidos del Cantabrico. Revista de la Real Academia de Ciencias Exactas, Fisicas y Naturales de Madrid, 17, 54 - 79." type="journal article" year="1918">
<collectingRegion id="49E522E0FF8789561872F834EDB4F879" box="[615,672,2006,2028]" country="Spain" name="La Rioja" pageId="10" pageNumber="161">Rioja</collectingRegion>
, 1918
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9EEC02FF8689571A82FF7AECD0FF3B" blockId="11.[151,1154,152,205]" box="[151,964,152,174]" pageId="11" pageNumber="162">
<treatmentCitationGroup id="AB31CB2CFF8689571A82FF7AECD0FF3B" box="[151,964,152,174]" pageId="11" pageNumber="162">
<taxonomicName id="4C219781FF8689571A82FF7AEED7FF3B" ID-CoL="4P2BL" box="[151,451,152,174]" class="Polychaeta" family="Spionidae" genus="Pseudomalacoceros" kingdom="Animalia" order="Spionida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="cantabra">
<emphasis id="B9553010FF8689571A82FF7AEED7FF3B" box="[151,451,152,174]" italics="true" pageId="11" pageNumber="162">Pseudomalacoceros cantabra</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A80CA13FF8689571BF4FF7AED8DFF3B" author="Vieitez" box="[481,665,152,174]" page="247" pageId="11" pageNumber="162" year="1981">
<bibRefCitation id="EFB091F3FF8689571BF4FF7AED73FF3B" author="Vieitez" box="[481,615,152,174]" pageId="11" pageNumber="162" refString="Vieitez, J. M. (1981) Estudio de las comunidades bentonicas de dos playas de las rias de Pontevedra y Vigo (Galicia, Espana). Boletin. Instituto Espanol de Oceanografia, 6 (331), 242 - 258." type="journal article" year="1981">Viéitez 1981</bibRefCitation>
: 247
</treatmentCitation>
, 254, tab. 1. Not
<bibRefCitation id="EFB091F3FF8689571952FF7AECD5FF3B" author="Rioja" box="[839,961,152,174]" pageId="11" pageNumber="162" refString="Rioja, E. (1918) Adiciones a la fauna de anelidos del Cantabrico. Revista de la Real Academia de Ciencias Exactas, Fisicas y Naturales de Madrid, 17, 54 - 79." type="journal article" year="1918">
<collectingRegion id="49E522E0FF8689571952FF7AEC94FF3B" box="[839,896,152,174]" country="Spain" name="La Rioja" pageId="11" pageNumber="162">Rioja</collectingRegion>
, 1918
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9EEC02FF8689571A82FF54EB96FF59" blockId="11.[151,1154,152,205]" box="[151,1154,182,205]" pageId="11" pageNumber="162">
<treatmentCitationGroup id="AB31CB2CFF8689571A82FF54EB96FF59" box="[151,1154,182,205]" pageId="11" pageNumber="162">
<taxonomicName id="4C219781FF8689571A82FF54EE72FF58" ID-CoL="4VRNB" box="[151,358,182,205]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF8689571A82FF54EE72FF58" box="[151,358,182,205]" italics="true" pageId="11" pageNumber="162">Scolelepis squamata</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A80CA13FF8689571B96FF55ED2BFF59" author="Parapar" box="[387,575,182,204]" page="188" pageId="11" pageNumber="162" year="1991">
<bibRefCitation id="EFB091F3FF8689571B96FF55ED04FF59" author="Parapar" box="[387,528,182,204]" pageId="11" pageNumber="162" refString="Parapar, J. (1991) Anelidos Poliquetos bentonicos de la ria de Ferrol (Galicia). PhD-thesis, University of Santiago de Compostela, 1104 pp." type="book" year="1991">Parapar 1991</bibRefCitation>
: 188
</treatmentCitation>
189, 990. Partim. Not Abildgaard,
<emphasis id="B9553010FF86895719B9FF5AECD4FF58" box="[940,960,184,205]" italics="true" pageId="11" pageNumber="162">in</emphasis>
<bibRefCitation id="EFB091F3FF86895719D3FF54EB6AFF59" author="Muller" box="[966,1150,182,204]" pageId="11" pageNumber="162" refString="Muller, O. F., Abildgaard, P. C., Vahl, M., Holten, J. S. &amp; Rathke, J. (1806) Zoologiae Danica seu Animalium Daniae et Norvegiae rariorum ac minus notorum. Descriptiones et historia. Volume IV. Volume Atlas. N. Christensen: Havniae [Copenhagen], 1 - 2, 46 pp., pls. 121 - 160." type="book chapter" year="1806">O.F. Müller, 1806</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8689571A82FF1BECA6FC9F" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8B9EEC02FF8689571A82FF1BEEABFECF" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<materialsCitation id="3B49E65FFF8689571A82FF1BEEABFECF" ID-GBIF-Occurrence="1303591705" collectionCode="ATLANTIC, OCEAN, MNCN" collectorName="R. Acuna Castroviejo" country="Spain" location="Cabo Vidio" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="2" stateProvince="Cantabria" typeStatus="holotype">
<emphasis id="B9553010FF8689571A82FF1BECD1FE87" bold="true" box="[151,965,249,274]" pageId="11" pageNumber="162">
Material examined.
<typeStatus id="549A52A0FF8689571B9CFF1BEED0FE87" box="[393,452,249,274]" pageId="11" pageNumber="162">Type</typeStatus>
material.
<typeStatus id="549A52A0FF868957182AFF1BEDA4FE87" box="[575,688,249,274]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED3074C7FF86895718A8FF1BECABFE87" box="[701,959,249,274]" pageId="11" pageNumber="162">ATLANTIC OCEAN</collectionCode>
:
</emphasis>
<collectingCountry id="F336AC92FF86895719D8FF1BEB34FE84" box="[973,1056,249,273]" name="Spain" pageId="11" pageNumber="162">SPAIN</collectingCountry>
:
<collectingRegion id="49E522E0FF8689571E3EFF1BEB8FFE87" box="[1067,1179,249,274]" country="Spain" name="Cantabria" pageId="11" pageNumber="162">Cantábria</collectingRegion>
,
<collectingRegion id="49E522E0FF8689571EB2FF1BEA13FE87" box="[1191,1287,249,274]" country="Spain" name="Asturias" pageId="11" pageNumber="162">Asturias</collectingRegion>
, sedimentary area between
<location id="8EFEBAD9FF8689571B26FEFFEEA0FEA3" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF8689571B26FEFFEEA0FEA3" box="[307,436,285,310]" country="Spain" name="Cabo Vidio" pageId="11" pageNumber="162" stateProvince="Cantabria">Cabo Vidio</location>
and Cabo de Peñas, st. 6 D 27, 25 m depth, fine sand, 1998, leg.
<collectorName id="26D489D4FF8689571EB4FEFCEA8CFEA3" box="[1185,1432,285,310]" pageId="11" pageNumber="162">R. Acuña Castroviejo</collectorName>
,
<emphasis id="B9553010FF8689571A82FEA3EE60FECF" bold="true" box="[151,372,321,346]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571A82FEA3EE26FECF" box="[151,306,321,346]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571A82FEA3EFFAFECF" box="[151,238,321,346]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/3852
</emphasis>
(1 af).
</materialsCitation>
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2FE87EE38FDC3" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<materialsCitation id="3B49E65FFF8689571AD2FE87EA17FE34" ID-GBIF-Occurrence="1303591708" collectionCode="MNCN" elevation="26" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="1" typeStatus="paratype">
<emphasis id="B9553010FF8689571AD2FE87EE5CFEEB" bold="true" box="[199,328,357,382]" pageId="11" pageNumber="162">
<typeStatus id="549A52A0FF8689571AD2FE87EE57FEEB" box="[199,323,357,382]" pageId="11" pageNumber="162" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as holotype, st. 2 D 59,
<elevation id="000C0B31FF86895718F9FE87EC2AFEE8" box="[748,830,357,382]" metricMagnitude="1" metricUnit="m" metricValue="2.56" pageId="11" pageNumber="162" unit="m" value="25.6">
<quantity id="4CD941E7FF86895718F9FE87EC2AFEE8" box="[748,830,357,382]" metricMagnitude="1" metricUnit="m" metricValue="2.56" pageId="11" pageNumber="162" unit="m" value="25.6">25.6 m</quantity>
</elevation>
depth,
<emphasis id="B9553010FF8689571982FE87EB6CFEEB" bold="true" box="[919,1144,357,382]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571982FE87EB23FEEB" box="[919,1079,357,382]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571982FE87ECFAFEEB" box="[919,1006,357,382]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/3851
</emphasis>
(1 af); st. D 84,
<emphasis id="B9553010FF8689571F50FE87EE01FE37" bold="true" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571F50FE87EFC0FE37" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571F50FE87EA88FEEB" box="[1349,1436,357,382]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/3854
</emphasis>
(1 af); st. 7 D 53, 32 m depth,
<emphasis id="B9553010FF8689571861FE6BEC46FE37" bold="true" box="[628,850,393,418]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571861FE6BEC04FE37" box="[628,784,393,418]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571861FE6BEDDFFE37" box="[628,715,393,418]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/3855
</emphasis>
partim (1 af on SEM stub); st. 4 D 72
</materialsCitation>
,
<quantity id="4CD941E7FF8689571F1AFE6BEA5BFE34" box="[1295,1359,393,417]" metricMagnitude="0" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="162" unit="m" value="8.4">8.4 m</quantity>
depth,
<emphasis id="B9553010FF8689571A82FE4FEE67FE53" bold="true" box="[151,371,429,454]" pageId="11" pageNumber="162">MNCN 16.01/3856</emphasis>
(1 af); st. 9 D 16, 28 m depth,
<emphasis id="B9553010FF86895718D2FE4FECB7FE53" bold="true" box="[711,931,429,454]" pageId="11" pageNumber="162">MNCN 16.01/3857</emphasis>
(1 af); st. 8 D 18, 32 m depth,
<emphasis id="B9553010FF8689571EEDFE4FEFDBFE7F" bold="true" pageId="11" pageNumber="162">MNCN 16.01/ 3858</emphasis>
(1 af, 1 mf); st. 7 D 53, 32 m depth,
<emphasis id="B9553010FF8689571864FE33EC5AFE7F" bold="true" box="[625,846,465,490]" pageId="11" pageNumber="162">MNCN 16.01/3859</emphasis>
(2 rehydrated af); st. 8 D 19,
<emphasis id="B9553010FF8689571EB5FE33EA68FE7F" bold="true" box="[1184,1404,465,490]" pageId="11" pageNumber="162">MNCN 16.01/3860</emphasis>
(2 af);
<materialsCitation id="3B49E65FFF8689571AD2FE17EA83FD9B" ID-GBIF-Occurrence="1303591703" box="[199,1431,501,526]" collectingDate="1975-10" collectionCode="MNCN" collectorName="Ria de Vigo &amp; J. M. Vieitez" county="Pontevedra" location="Meira" municipality="Ria de Vigo" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="1" stateProvince="Galicia" typeStatus="paratype">
<collectingRegion id="49E522E0FF8689571AD2FE17EE0FFD9B" box="[199,283,501,526]" country="Spain" name="Galicia" pageId="11" pageNumber="162">Galicia</collectingRegion>
,
<collectingCounty id="62FF948EFF8689571B3CFE17EEBEFD9B" box="[297,426,501,526]" pageId="11" pageNumber="162">Pontevedra</collectingCounty>
,
<collectorName id="26D489D4FF8689571BADFE14ED52FD9B" box="[440,582,501,526]" pageId="11" pageNumber="162">
<collectingMunicipality id="6BFA7678FF8689571BADFE14ED52FD9B" box="[440,582,501,526]" pageId="11" pageNumber="162">Ría de Vigo</collectingMunicipality>
</collectorName>
, beach of
<location id="8EFEBAD9FF86895718D5FE17EC12FD9B" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF86895718D5FE17EC12FD9B" box="[704,774,501,526]" county="Pontevedra" municipality="Ria de Vigo" name="Meira" pageId="11" pageNumber="162" stateProvince="Galicia">Meira</location>
,
<collectingDate id="EFDB332AFF8689571906FE14EC95FD9B" box="[787,897,501,526]" pageId="11" pageNumber="162" value="1975-10">
<date id="FF9FCAC2FF8689571906FE14EC95FD9B" box="[787,897,501,526]" pageId="11" pageNumber="162" value="1975-10">Oct 1975</date>
</collectingDate>
, leg.
<collectorName id="26D489D4FF86895719D4FE14EB5BFD9B" box="[961,1103,501,526]" pageId="11" pageNumber="162">J.M. Viéitez</collectorName>
,
<emphasis id="B9553010FF8689571E49FE17EA5FFD9B" bold="true" box="[1116,1355,501,526]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571E49FE17EBEFFD9B" box="[1116,1275,501,526]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571E49FE17EBA7FD9B" box="[1116,1203,501,526]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/17364
</emphasis>
(2 af)
</materialsCitation>
;
<materialsCitation id="3B49E65FFF8689571A82FDFBEE38FDC3" ID-GBIF-Occurrence="1303591704" collectingDate="1988-01-22" collectionCode="MNCN" collectorName="Ria de Ferrol &amp; J. Parapar" county="Coruna" location="Laxe" municipality="Ria de Ferrol" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="1" stateProvince="Galicia" typeStatus="paratype">
<collectingRegion id="49E522E0FF8689571A82FDFBEFFFFDA7" box="[151,235,537,562]" country="Spain" name="Galicia" pageId="11" pageNumber="162">Galicia</collectingRegion>
,
<collectingCounty id="62FF948EFF8689571AECFDFBEE5AFDA4" box="[249,334,537,561]" pageId="11" pageNumber="162">Coruña</collectingCounty>
,
<collectorName id="26D489D4FF8689571B49FDFBEEECFDA7" box="[348,504,537,562]" pageId="11" pageNumber="162">
<collectingMunicipality id="6BFA7678FF8689571B49FDFBEEECFDA7" box="[348,504,537,562]" pageId="11" pageNumber="162">Ría de Ferrol</collectingMunicipality>
</collectorName>
,
<location id="8EFEBAD9FF8689571812FDF8ED56FDA4" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF8689571812FDF8ED56FDA4" box="[519,578,538,561]" county="Coruna" municipality="Ria de Ferrol" name="Laxe" pageId="11" pageNumber="162" stateProvince="Galicia">Laxe</location>
, intertidal, coarse sand,
<collectingDate id="EFDB332AFF8689571949FDFBECF8FDA4" box="[860,1004,537,561]" pageId="11" pageNumber="162" value="1988-01-22">
<date id="FF9FCAC2FF8689571949FDFBECF8FDA4" box="[860,1004,537,561]" pageId="11" pageNumber="162" value="1988-01-22">22 Jan 1988</date>
</collectingDate>
, leg.
<collectorName id="26D489D4FF8689571E38FDF8EBB5FDA4" box="[1069,1185,538,561]" pageId="11" pageNumber="162">J. Parapar</collectorName>
,
<emphasis id="B9553010FF8689571EBBFDFBEA88FDA7" bold="true" box="[1198,1436,537,562]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571EBBFDFBEA59FDA7" box="[1198,1357,537,562]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571EBBFDFBEA11FDA7" box="[1198,1285,537,562]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/12568
</emphasis>
partim (4 af).
</materialsCitation>
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2FD83EA25FD57" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<materialsCitation id="3B49E65FFF8689571AD2FD83ECB5FD0B" ID-GBIF-Occurrence="1303591702" collectingDate="1891-10" collectionCode="ATLANTIC, OCEAN, MNCN" collectorName="Estacion Biologica" country="Spain" county="Non-type" location="Cantabrian" municipality="Gulf of Biscaya" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="1">
<emphasis id="B9553010FF8689571AD2FD83EDA1FDEF" bold="true" box="[199,693,609,634]" pageId="11" pageNumber="162">
<collectingCounty id="62FF948EFF8689571AD2FD83EE21FDEF" box="[199,309,609,634]" pageId="11" pageNumber="162">Non-type</collectingCounty>
material.
<collectionCode id="ED3074C7FF8689571BBAFD83EDA4FDEF" box="[431,688,609,634]" pageId="11" pageNumber="162">ATLANTIC OCEAN</collectionCode>
:
</emphasis>
<collectingCountry id="F336AC92FF86895718A9FD80EC1BFDEF" box="[700,783,610,634]" name="Spain" pageId="11" pageNumber="162">SPAIN</collectingCountry>
:
<collectingMunicipality id="6BFA7678FF868957190CFD83ECD9FDEF" box="[793,973,609,634]" pageId="11" pageNumber="162">Gulf of Biscaya</collectingMunicipality>
,
<location id="8EFEBAD9FF86895719C2FD83EB46FDEF" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF86895719C2FD83EB46FDEF" box="[983,1106,609,634]" country="Spain" county="Non-type" municipality="Gulf of Biscaya" name="Cantabrian" pageId="11" pageNumber="162">Cantabrian</location>
coast,
<location id="8EFEBAD9FF8689571EB5FD83EA06FDEF" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF8689571EB5FD83EA06FDEF" box="[1184,1298,609,634]" country="Spain" county="Non-type" municipality="Gulf of Biscaya" name="Santander" pageId="11" pageNumber="162">Santander</location>
,
<location id="8EFEBAD9FF8689571F08FD80EA8CFDEF" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF8689571F08FD80EA8CFDEF" box="[1309,1432,609,634]" country="Spain" county="Non-type" municipality="Gulf of Biscaya" name="Pena Vieja" pageId="11" pageNumber="162">Peña Vieja</location>
,
<collectingDate id="EFDB332AFF8689571A82FD64EE17FD0B" box="[151,259,646,670]" pageId="11" pageNumber="162" value="1891-10">
<date id="FF9FCAC2FF8689571A82FD64EE17FD0B" box="[151,259,646,670]" pageId="11" pageNumber="162" value="1891-10">Oct 1891</date>
</collectingDate>
, coll.
<collectorName id="26D489D4FF8689571B5EFD67ED31FD0B" box="[331,549,645,670]" pageId="11" pageNumber="162">Estación Biológica</collectorName>
,
<emphasis id="B9553010FF8689571827FD67EC17FD0B" bold="true" box="[562,771,645,670]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571827FD67EDDBFD0B" box="[562,719,645,670]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571827FD67ED9DFD0B" box="[562,649,645,670]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/370
</emphasis>
partim (2 af)
</materialsCitation>
;
<materialsCitation id="3B49E65FFF86895719A5FD67EA25FD57" ID-GBIF-Occurrence="1303591706" collectingDate="1972-08" collectionCode="MNCN" collectorName="Ria de Vigo &amp; Playa de Meira &amp; J. M. Vieitez" country="Spain" county="Pontevedra" latitude="42.284443" location="Playa de Meira" longLatPrecision="19" longitude="-8.718888" municipality="Ria de Vigo" pageId="11" pageNumber="162" specimenCode="MNCN 16.01" specimenCount="1" stateProvince="Galicia">
<collectingRegion id="49E522E0FF86895719A5FD67EB10FD0B" box="[944,1028,645,670]" country="Spain" name="Galicia" pageId="11" pageNumber="162">Galicia</collectingRegion>
,
<collectingCounty id="62FF948EFF8689571E04FD67EB86FD0B" box="[1041,1170,645,670]" pageId="11" pageNumber="162">Pontevedra</collectingCounty>
,
<collectorName id="26D489D4FF8689571E8AFD64EA38FD0B" box="[1183,1324,645,670]" pageId="11" pageNumber="162">
<collectingMunicipality id="6BFA7678FF8689571E8AFD64EA38FD0B" box="[1183,1324,645,670]" pageId="11" pageNumber="162">Ría de Vigo</collectingMunicipality>
</collectorName>
,
<collectorName id="26D489D4FF8689571F2FFD67EFC9FD57" pageId="11" pageNumber="162">
<location id="8EFEBAD9FF8689571F2FFD67EFC9FD57" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF8689571F2FFD67EFC9FD57" country="Spain" county="Pontevedra" latitude="42.284443" longLatPrecision="19" longitude="-8.718888" municipality="Ria de Vigo" name="Playa de Meira" pageId="11" pageNumber="162" stateProvince="Galicia">Playa de Meira</location>
</collectorName>
, St. 13,
<geoCoordinate id="EE158AC5FF8689571B23FD48EEA9FD57" box="[310,445,682,706]" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="15" value="42.284443">42°17'04&quot;N</geoCoordinate>
,
<geoCoordinate id="EE158AC5FF8689571BDDFD48ED50FD57" box="[456,580,682,706]" direction="west" orientation="longitude" pageId="11" pageNumber="162" precision="15" value="-8.718888">8°43'08&quot;W</geoCoordinate>
,
<collectingDate id="EFDB332AFF8689571845FD48EDD6FD57" box="[592,706,682,706]" pageId="11" pageNumber="162" value="1972-08">
<date id="FF9FCAC2FF8689571845FD48EDD6FD57" box="[592,706,682,706]" pageId="11" pageNumber="162" value="1972-08">Aug 1972</date>
</collectingDate>
, leg.
<collectorName id="26D489D4FF86895718E8FD48EC9CFD57" box="[765,904,681,706]" pageId="11" pageNumber="162">J.M. Viéitez</collectorName>
,
<emphasis id="B9553010FF8689571986FD4BEB76FD57" bold="true" box="[915,1122,681,706]" pageId="11" pageNumber="162">
<specimenCode id="DB874479FF8689571986FD4BEB3AFD57" box="[915,1070,681,706]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571986FD4BECFEFD57" box="[915,1002,681,706]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="162">MNCN</collectionCode>
16.01
</specimenCode>
/373
</emphasis>
(1 dehydrated af).
</materialsCitation>
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2FD2FECA6FC9F" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<emphasis id="B9553010FF8689571AD2FD2FED19FD73" bold="true" box="[199,525,717,742]" pageId="11" pageNumber="162">
MEDITERRANEAN
<collectionCode id="ED3074C7FF8689571BDBFD2FED1CFD73" box="[462,520,717,742]" pageId="11" pageNumber="162">SEA</collectionCode>
:
</emphasis>
TYRRHENIAN
<collectionCode id="ED3074C7FF86895718C3FD2CEC1BFD73" box="[726,783,718,742]" pageId="11" pageNumber="162">SEA</collectionCode>
:
<collectingCountry id="F336AC92FF8689571908FD2CEC7AFD73" box="[797,878,718,742]" name="Italy" pageId="11" pageNumber="162">ITALY</collectingCountry>
:
<collectingRegion id="49E522E0FF868957196FFD2FECFBFD73" box="[890,1007,717,742]" country="Italy" name="Campania" pageId="11" pageNumber="162">Campania</collectingRegion>
, N. of Napoli, Gulf of Gaeta, Licola,
<date id="FF9FCAC2FF8689571A82FD13EE03FC9F" box="[151,279,753,778]" pageId="11" pageNumber="162" value="1966" valueMax="1968">19661968</date>
, leg. J. Dörjes,
<emphasis id="B9553010FF8689571BD1FD13ED59FC9F" bold="true" box="[452,589,753,778]" pageId="11" pageNumber="162">
<collectionCode id="ED3074C7FF8689571BD1FD13EEEBFC9F" box="[452,511,753,778]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="162">SMF</collectionCode>
12914
</emphasis>
partim (~30 af, juveniles only).
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8689571AD2FCF7EB57FCC7" pageId="11" pageNumber="162" type="description">
<paragraph id="8B9EEC02FF8689571AD2FCF7EB57FCC7" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<emphasis id="B9553010FF8689571AD2FCF7EE65FCBB" bold="true" box="[199,369,789,814]" pageId="11" pageNumber="162">BLACK SEA:</emphasis>
<collectingCountry id="F336AC92FF8689571B6DFCF4ED16FCBB" box="[376,514,790,814]" name="Romania" pageId="11" pageNumber="162">ROMANIA</collectingCountry>
: Năvodari—Digul de Sud,
<geoCoordinate id="EE158AC5FF8689571926FCF7ECDBFCBB" box="[819,975,789,814]" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="1" value="44.323723">44°19'25.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE158AC5FF86895719CFFCF4EB67FCBB" box="[986,1139,790,814]" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="1" value="28.639278">28°38'21.4&quot;E</geoCoordinate>
,
<quantity id="4CD941E7FF8689571E6BFCF7EBE5FCB8" box="[1150,1265,789,814]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="162" unit="m" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 m</quantity>
, fine sand with shells,
<date id="FF9FCAC2FF8689571AF1FCD8EE64FCC7" box="[228,368,826,850]" pageId="11" pageNumber="162" value="2014-06-29">29 Jun 2014</date>
, leg. V. Surugiu,
<emphasis id="B9553010FF8689571824FCDBEC31FCC7" bold="true" box="[561,805,825,850]" pageId="11" pageNumber="162">MNINGA PLY 0 0 59</emphasis>
(1 af, in good condition).
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8689571AD2FCBFEC77FB03" pageId="11" pageNumber="162" type="diagnosis">
<paragraph id="8B9EEC02FF8689571AD2FCBFEC77FB03" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<emphasis id="B9553010FF8689571AD2FCBFEE55FCE3" bold="true" box="[199,321,861,886]" pageId="11" pageNumber="162">Diagnosis.</emphasis>
Prostomium anteriorly with medial portion elongated into an acute point and with rounded anterolateral lobes; prostomium with transverse furrow at level of palp insertions; caruncle subtriangular, slightly inflated, attached to dorsum. Occipital antenna absent. Peristomium short, separated from prostomium by distinct groove, forming well-developed dorso-lateral wings. Palps short, with smooth basal sheaths and with two widely spaced longitudinal bands of elevated lobes bearing transverse rows of cilia. Low dorsal, segmentally arranged, transverse ciliated bands present throughout body. Chaetiger 1 with foliaceous notopodial postchaetal lamellae, with chaetae in both rami. Anterior chaetigers with obtuse, subtriangular prechaetal notopodial lamellae. Branchial tips always free from notopodial postchaetal lamellae. Bidentate neuropodial hooded hooks from chaetiger 1949; shafts of hooks strongly curved and slightly constricted at bend.
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8689571AD2FB43EA96FB2C" box="[199,1410,1185,1210]" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8B9EEC02FF8689571AD2FB43EA96FB2C" blockId="11.[151,1437,249,2038]" box="[199,1410,1185,1210]" pageId="11" pageNumber="162">
<materialsCitation id="3B49E65FFF8689571AD2FB43EA96FB2C" ID-GBIF-Occurrence="1303591707" box="[199,1410,1185,1210]" country="Spain" elevation="25" location="Cabo Vidio" pageId="11" pageNumber="162" specimenCount="25" stateProvince="Cantabria" typeStatus="holotype">
<emphasis id="B9553010FF8689571AD2FB43EE71FB2F" bold="true" box="[199,357,1185,1210]" pageId="11" pageNumber="162">
<typeStatus id="549A52A0FF8689571AD2FB43EE16FB2F" box="[199,258,1185,1210]" pageId="11" pageNumber="162">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F336AC92FF8689571B79FB40EEA9FB2F" box="[364,445,1186,1210]" name="Spain" pageId="11" pageNumber="162">SPAIN</collectingCountry>
,
<collectingRegion id="49E522E0FF8689571BDDFB43ED2CFB2F" box="[456,568,1185,1210]" country="Spain" name="Cantabria" pageId="11" pageNumber="162">Cantabria</collectingRegion>
,
<collectingRegion id="49E522E0FF8689571856FB43EDB7FB2F" box="[579,675,1185,1210]" country="Spain" name="Asturias" pageId="11" pageNumber="162">Asturias</collectingRegion>
, sedimentary area between
<location id="8EFEBAD9FF86895719CDFB43EB4CFB2F" LSID="urn:lsid:plazi:treatment:03885D14FF87894D1A82F8AEECC6FE1D:8EFEBAD9FF86895719CDFB43EB4CFB2F" box="[984,1112,1185,1210]" country="Spain" name="Cabo Vidio" pageId="11" pageNumber="162" stateProvince="Cantabria">Cabo Vidio</location>
and Cabo de Peñas,
<specimenCount id="9D27278BFF8689571F57FB43EA96FB2C" box="[1346,1410,1185,1210]" pageId="11" pageNumber="162" type="generic">
<elevation id="000C0B31FF8689571F57FB43EA6AFB2C" box="[1346,1406,1185,1210]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="162" unit="m" value="25.0">
<quantity id="4CD941E7FF8689571F57FB43EA6AFB2C" box="[1346,1406,1185,1210]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="162" unit="m" value="25.0">25 m</quantity>
</elevation>
.
</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF8689501AD2FB27EED5F82C" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162" type="description">
<paragraph id="8B9EEC02FF8689571AD2FB27EEF0FADF" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
<emphasis id="B9553010FF8689571AD2FB27EE4DFB4B" bold="true" box="[199,345,1221,1246]" pageId="11" pageNumber="162">Description.</emphasis>
All specimens incomplete; longest anterior fragment
<quantity id="4CD941E7FF86895719D6FB27EB00FB48" box="[963,1044,1221,1246]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="11" pageNumber="162" unit="mm" value="51.0">51 mm</quantity>
for 93 chaetigers and widest
<quantity id="4CD941E7FF8689571F7EFB27EFD2FA94" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="11" pageNumber="162" unit="mm" value="2.75">2.75 mm</quantity>
.
<typeStatus id="549A52A0FF8689571AC2FB0BEE2BFA97" box="[215,319,1257,1282]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
<quantity id="4CD941E7FF8689571B58FB08EEB7FA94" box="[333,419,1258,1282]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="162" unit="mm" value="24.0">24 mm</quantity>
long,
<quantity id="4CD941E7FF8689571BE2FB08ED75FA94" box="[503,609,1258,1282]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="11" pageNumber="162" unit="mm" value="1.42">1.42 mm</quantity>
wide (at chaetiger 15) for 61+ chaetigers, widest anteriorly (around chaetigers 721), then gradually tapering to posterior end; body suboval in cross section. Colour in alcohol yellowbrown, without pigmentation.
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2FAB7EA1FFA6B" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
Prostomium anteriorly trilobate, medially acuminate, antero-laterally rounded, posteriorly extended as a short, bluntly rounded, attached, slightly raised caruncle, extending to posterior margin of chaetiger 1 (
<figureCitation id="131AF087FF8689571EE1FA9BEA51FA07" box="[1268,1349,1401,1426]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Figs. 5</figureCitation>
A, 6A), with transverse constriction in posterior third. Occipital antenna absent. Eyes usually absent, but four faded, subcuticular eyes in a trapezoidal arrangement observed in a few specimens. Nuchal organs a J-shaped inconspicuous structure, on posterior lateral margins of caruncle, just behind bases of palps (
<figureCitation id="131AF087FF8689571EB6FA07EBF2FA6B" box="[1187,1254,1509,1534]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Fig. 5</figureCitation>
A).
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2F9EBEC2BF8F3" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
Peristomium short, distinctly separated from prostomium by a shallow groove, forming well-developed dorsolateral wings, indistinctly separated from chaetiger 1 (
<figureCitation id="131AF087FF8689571947F9CFEC8CF9D3" box="[850,920,1581,1606]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Fig. 5</figureCitation>
AB). Eversible proboscis sac-like, inflated. Palps deciduous, present only in six examined specimens; when present, palps very short, tapering (
<figureCitation id="131AF087FF8689571EE9F9B3EA5EF9FF" box="[1276,1354,1617,1642]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Figs. 5</figureCitation>
B, 6B), reaching at most to chaetiger 5; ciliation of palps consisting of two frontal longitudinal distinctly separated bands of transverse rows of short, non-motile cilia, both on elevated 12-µm-tall lobes of approximately equal width, those on medial side approximately 1414.3 µm wide, those on lateral side subdivided into two closely applied rows of approximately 7 µm wide each (14 µm in total with a distance less than 1 µm between them) (
<figureCitation id="131AF087FF8689571EA6F903EBE1F96F" box="[1203,1269,1761,1786]" captionStart="FIGURE 7" captionStartId="15.[151,250,1189,1211]" captionTargetBox="[151,1436,193,1167]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 7. Scolelepis (Scolelepis) neglecta sp. nov., SEM micrographs. A F, paratype (MNCN 3855). A, distal tip of left palp with elevated lobes, frontal-lateral view (arrowheads indicate faint splitting of lateral rows into two smaller lobes); B, distal section of right palp, frontal view, showing elevated lobes; C, basal part of right palp, lateral view (arrowhead shows smooth basal palpal sheath); D, chaetigers 13 25 (from top right to bottom left), ventro-lateral view (arrowheads indicate prechaetal notopodial lamellae; arrow indicates ventral median groove); E, chaetigers 16 21 (from top right to bottom left), dorso-lateral view (arrowheads indicate lateral ciliated organs); F, chaetigers 29 32 (from right to left), lateral view (note ventrally directed inferior part of notopodial postchaetal lamella and dorsally directed superior part of neuropodial postchaetal lamella). Scale bars: A, B = 50 µm; C = 100 µm; D = 1 mm; E = 0.5 mm; F = 0.2 mm." httpUri="https://zenodo.org/record/263916/files/figure.png" pageId="11" pageNumber="162">Fig. 7</figureCitation>
AB). Ratio of long to short rows approximately 1.201.25 with a distance of 1115 µm separating them. Palp sheaths rounded, short, smooth, tightly adhering to lateral and abfrontal bases of palps (
<figureCitation id="131AF087FF86895719CDF8CBEB34F8D7" box="[984,1056,1833,1858]" captionStart="FIGURE 7" captionStartId="15.[151,250,1189,1211]" captionTargetBox="[151,1436,193,1167]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 7. Scolelepis (Scolelepis) neglecta sp. nov., SEM micrographs. A F, paratype (MNCN 3855). A, distal tip of left palp with elevated lobes, frontal-lateral view (arrowheads indicate faint splitting of lateral rows into two smaller lobes); B, distal section of right palp, frontal view, showing elevated lobes; C, basal part of right palp, lateral view (arrowhead shows smooth basal palpal sheath); D, chaetigers 13 25 (from top right to bottom left), ventro-lateral view (arrowheads indicate prechaetal notopodial lamellae; arrow indicates ventral median groove); E, chaetigers 16 21 (from top right to bottom left), dorso-lateral view (arrowheads indicate lateral ciliated organs); F, chaetigers 29 32 (from right to left), lateral view (note ventrally directed inferior part of notopodial postchaetal lamella and dorsally directed superior part of neuropodial postchaetal lamella). Scale bars: A, B = 50 µm; C = 100 µm; D = 1 mm; E = 0.5 mm; F = 0.2 mm." httpUri="https://zenodo.org/record/263916/files/figure.png" pageId="11" pageNumber="162">Fig. 7</figureCitation>
C).
<typeStatus id="549A52A0FF8689571E58F8CBEBA1F8D7" box="[1101,1205,1833,1858]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
with only left palp present (fell off during the study), reaching back chaetiger 4.
</paragraph>
<paragraph id="8B9EEC02FF8689571AD2F893ECBCF863" blockId="11.[151,1437,249,2038]" pageId="11" pageNumber="162">
Branchiae from chaetiger 2 to end of fragments, best developed by chaetigers 520; ciliation along inner edge of branchiae; tips of branchiae acuminate, glandular, devoid of cilia. In anterior chaetigers, branchiae fused to notopodial postchaetal lamellae for approximately half of their length (
<figureCitation id="131AF087FF86895719AAF85BEB18F847" box="[959,1036,1977,2002]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Figs. 5</figureCitation>
A, B, D, E, 7E); on last chaetigers, branchiae fused only basally to postchaetal lamellae (
<figureCitation id="131AF087FF86895718F8F83FEC2EF863" box="[749,826,2013,2038]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="11" pageNumber="162">Figs. 5</figureCitation>
FH, 7F).
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FF75EC6CFEF0" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Chaetiger 1 well developed, capillary chaetae present in both rami, fewer than in following chaetigers; notopodial lamellae elongate, digitiform, tapered subdistally; neuropodial lamellae shorter, acute-ovate (
<figureCitation id="131AF087FF8189501F22FF5EEA6EFF40" box="[1335,1402,188,213]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B C); notopodial capillaries with indistinct double rows and a dorsal superior tuft of thin and long capillaries numbering up to 10, longer than ventral ones and as long as, or slightly shorter than those of dorsal superior tuft on following chaetigers; neuropodial capillaries more numerous, arranged in two distinct rows, 79 per row, plus a ventral inferior tuft, all shorter than those of following chaetigers.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FE8DED32FDA9" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Dorsal transverse ciliation as primary ciliary bands on mid-part of each chaetiger (nototrochs), continuous with ciliation of branchiae; additional short row of cilia on outer edge of branchiae between tips of notopodial lamellae and subdistal portion of branchiae. In anterior chaetigers (to about chaetiger 2630) secondary dorsal transverse rows of cilia located on anterior part of chaetigers also present (
<figureCitation id="131AF087FF8189501976FE3EECB1FE60" box="[867,933,476,501]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A). In middle and posterior chaetigers dorsal surface smooth with low transverse folds uniting branchiae (from about chaetiger 25). Ventral surface smooth with a shallow median groove (
<figureCitation id="131AF087FF8189501BAAFDC6ED16FDA9" box="[447,514,548,573]" captionStart="FIGURE 7" captionStartId="15.[151,250,1189,1211]" captionTargetBox="[151,1436,193,1167]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 7. Scolelepis (Scolelepis) neglecta sp. nov., SEM micrographs. A F, paratype (MNCN 3855). A, distal tip of left palp with elevated lobes, frontal-lateral view (arrowheads indicate faint splitting of lateral rows into two smaller lobes); B, distal section of right palp, frontal view, showing elevated lobes; C, basal part of right palp, lateral view (arrowhead shows smooth basal palpal sheath); D, chaetigers 13 25 (from top right to bottom left), ventro-lateral view (arrowheads indicate prechaetal notopodial lamellae; arrow indicates ventral median groove); E, chaetigers 16 21 (from top right to bottom left), dorso-lateral view (arrowheads indicate lateral ciliated organs); F, chaetigers 29 32 (from right to left), lateral view (note ventrally directed inferior part of notopodial postchaetal lamella and dorsally directed superior part of neuropodial postchaetal lamella). Scale bars: A, B = 50 µm; C = 100 µm; D = 1 mm; E = 0.5 mm; F = 0.2 mm." httpUri="https://zenodo.org/record/263916/files/figure.png" pageId="12" pageNumber="163">Fig. 7</figureCitation>
D).
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FDA5EC71FC80" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Notopodial postchaetal lamellae well-developed from chaetiger 2 (
<figureCitation id="131AF087FF81895019C9FDA5EB37FDF5" box="[988,1059,583,608]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AB, D), on anterior chaetigers elongated, narrow, attached to branchiae, with outer margin smooth, entire, with free tips acuminate (
<figureCitation id="131AF087FF8189501EEEFD8EEA2FFD10" box="[1275,1339,620,645]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
DE), in middle and posterior chaetigers becoming gradually shorter, wider and attached only at base of branchiae, with lower portion directed ventrally towards neuropodial postchaetal lamella (
<figureCitation id="131AF087FF81895019CBFD56EB3FFD58" box="[990,1067,692,717]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Figs. 5</figureCitation>
FH, 7F). Notopodial prechaetal lamellae subtriangular, with stubby (subobtuse) tip, best developed from chaetiger 2 to around chaetiger 29 (
<figureCitation id="131AF087FF8189501F7AFD35EFB1FC80" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B, 7D), then becoming lower and broader inconspicuous folds.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FCFDEC2CFBCD" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Neuropodial postchaetal lamellae in anteriormost 25 chaetigers acute, with a small median knob (mammilate), as long as wide, in following chaetigers becoming progressively more rounded, semicircular, entire, wider than long (
<figureCitation id="131AF087FF8189501B75FC85EEB2FC15" box="[352,422,871,896]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B, D); slight notch developing in inferior 1/3 of lamella on chaetigers 1441, in adults (usually wider than
<quantity id="4CD941E7FF8189501B91FC6EEEF4FC31" box="[388,480,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="mm" value="1.5">1.5 mm</quantity>
) on chaetigers 3541 (
<figureCitation id="131AF087FF81895018E7FC6EEC57FC30" box="[754,835,908,933]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Figs. 5</figureCitation>
F, 8); notch becoming deeper in middle chaetigers, dividing lamella into two separate lobes, upper lobe being more than twice as large as lower subtriangular lobe, lower lobe located at level of ventral inferior bundle of capillaries (
<figureCitation id="131AF087FF81895019B1FC36ECF3FC78" box="[932,999,980,1005]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
FH). In posteriormost chaetigers gap between lobes wider, setting lobes further apart, upper lobe becoming narrow, rounded, with upper portion elongated and directed towards ventral portion of notopodial postchaetal lamella, lower part reduced to triangular cirrus (
<figureCitation id="131AF087FF8189501AF3FBDDEE33FBCD" box="[230,295,1087,1112]" captionStart="FIGURE 5" captionStartId="13.[151,250,1881,1903]" captionTargetBox="[230,1339,196,1830]" captionTargetId="figure@13.[223,1342,193,1839]" captionTargetPageId="13" captionText="FIGURE 5. Scolelepis (Scolelepis) neglecta sp. nov. A, B, holotype (MNCN 3852); C H, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: A H = 1 mm." httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
GH). Neuropodial prechaetal lamellae absent.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FB86ED04FAA5" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Anterior chaetigers with capillaries only, arranged in three distinct groups in both noto- and neuropodia. Notopodial capillary chaetae elongate, narrow, arranged in double rows (711 per row) and a dorsal superior tuft of 210 longer (up to ~twice the branchiae length) and thinner capillaries; capillaries of anterior row unilimbate, with fibrose cores, of approximately same length and width as those of posterior row; capillaries of posterior row unilimbate, with uniformly granulated cores (
<figureCitation id="131AF087FF81895018BEFB16EDE7FA98" box="[683,755,1268,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
C). Number and length of notopodial capillaries gradually decreasing towards posterior end.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FADEED42FA71" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Neuropodial capillaries similar in morphology to those of notopodia, although shorter and broader, with narrow limbation, with granulated cores (when viewed in direct light) or appearing fibrous (when viewed in reflected light), also arranged in double vertical rows (616 per row) plus a ventral inferior bundle of 14 long, unilimbate capillaries in position of sabre chaetae (
<figureCitation id="131AF087FF81895018CAFA45EC30FA55" box="[735,804,1447,1472]" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
E); capillaries of anterior row (
<figureCitation id="131AF087FF8189501E92FA45EBDFFA55" box="[1159,1227,1447,1472]" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
D) slightly shorter and broader than those of posterior row.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2FA0DEB55F928" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
Neuropodial hooded hooks first present in posterior row of chaetigers 1949, in adults in chaetigers 4049 (
<figureCitation id="131AF087FF8189501A8AF9F6EFF7F9B9" box="[159,227,1556,1581]" captionStart="FIGURE 8" captionStartId="17.[151,250,1279,1301]" captionTargetBox="[188,1364,460,1240]" captionTargetId="figure@17.[176,1375,447,1248]" captionTargetPageId="17" captionText="FIGURE 8. Scolelepis (Scolelepis) neglecta sp. nov., relationship between the body-size (referring to the maximum anterior width) and the first chaetiger on which a notch is present in the neuropodial postchaetal lamellae (triangles, holotype empty symbol, y = 13.28 + 10.106 x; r s = 0.931; P &lt;0.0001; n = 36), and between the body-size (referring to the maximum anterior width) and the first chaetiger on which neuropodial hooded hooks are present (circles, holotype empty symbol, y = 15.673 + 11.97 x; r s = 0.846; P &lt;0.0001; n = 36)." httpUri="https://zenodo.org/record/263917/files/figure.png" pageId="12" pageNumber="163">Fig. 8</figureCitation>
), up to 13 per fascicle, accompanied in middle and posterior chaetigers by 04 alternating slender capillary chaetae and by 07 shorter slender limbate capillaries in ventral inferior tuft. Hooded hooks with bluntly rounded main fang surmounted by a smaller bluntly rounded apical tooth, angle between main tooth and shaft 8595°; shaft long, with small constriction and bend just below hood insertion (
<figureCitation id="131AF087FF818950199CF99DECD9F90D" box="[905,973,1663,1688]" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
FG). Hood elongate, more than 5 times length of main fang, with apical-rostral slit-like opening. Ventral sabre setae absent.
</paragraph>
<paragraph id="8B9EEC02FF8189501AD2F925ED93F975" blockId="12.[151,1437,151,1977]" box="[199,647,1735,1760]" pageId="12" pageNumber="163">Notopodial hooded hooks not observed.</paragraph>
<paragraph id="8B9EEC02FF8189501AD2F90EEEBFF890" blockId="12.[151,1437,151,1977]" box="[199,427,1772,1797]" pageId="12" pageNumber="163">Pygidium unknown.</paragraph>
<paragraph id="8B9EEC02FF8189501AD2F8EDEED5F82C" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="163">
<emphasis id="B9553010FF8189501AD2F8EDED5DF8BD" bold="true" box="[199,585,1807,1832]" pageId="12" pageNumber="163">Methyl green staining pattern.</emphasis>
Most intense blue staining on peristomial wings, on margins of both notopodial and neuropodial postchaetal lamellae, on tips of notopodial prechaetal lamellae, on tips of branchiae and on superior margins of basal sheaths of palps (
<figureCitation id="131AF087FF81895018E3F8B5EC29F8E5" box="[758,829,1879,1904]" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
AB). Prostomium and peristomium with uniformly dispersed speckles forming indistinct longitudinal stripes. Ventral side uniformly speckled on each chaetiger, with unclear transversal stripes.
</paragraph>
</subSubSection>
<caption id="DF5EBC8AFF8089511A82F8BBEB15F87C" httpUri="https://zenodo.org/record/263914/files/figure.png" pageId="13" pageNumber="164" targetBox="[230,1339,196,1830]" targetPageId="13">
<paragraph id="8B9EEC02FF8089511A82F8BBEB15F87C" blockId="13.[151,1436,1881,2025]" pageId="13" pageNumber="164">
<emphasis id="B9553010FF8089511A82F8BBEE03F8FB" bold="true" box="[151,279,1881,1903]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
<taxonomicName id="4C219781FF8089511B34F8BBED7FF8FA" box="[289,619,1881,1903]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8089511B34F8BBEE93F8FA" box="[289,391,1881,1903]" italics="true" pageId="13" pageNumber="164">Scolelepis</emphasis>
(
<emphasis id="B9553010FF8089511B8EF8BBED15F8FA" box="[411,513,1881,1903]" italics="true" pageId="13" pageNumber="164">Scolelepis</emphasis>
)
<emphasis id="B9553010FF8089511800F8BBED7FF8FA" box="[533,619,1881,1903]" italics="true" pageId="13" pageNumber="164">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF8089511860F8BBEDDEF8FB" box="[629,714,1881,1902]" pageId="13" pageNumber="164" rank="species">
<emphasis id="B9553010FF8089511860F8BBEDDEF8FB" bold="true" box="[629,714,1881,1902]" pageId="13" pageNumber="164">sp. nov.</emphasis>
</taxonomicNameLabel>
A, B, holotype (MNCN 3852); CH, paratype (MNCN 12568). A, anterior end, dorsal view (palps missing, left notopodial lamella and branchia of chaetiger 5 missing); B, anterior end, lateral view (left palp present only); C, parapodium of chaetiger 1, anterior view; D, parapodium of chaetiger 2, anterior view; E, parapodium of chaetiger 15, anterior view; F, parapodium of chaetiger 42, posterior view; G, parapodium of chaetiger 50, anterior view; H, parapodium of chaetiger 56, anterior view. Scale bars: AH = 1 mm.
</paragraph>
</caption>
<caption id="DF5EBC8AFF8389521A82F8FFEE14F87E" httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="14" pageNumber="165" targetBox="[260,1340,214,1782]" targetPageId="14">
<paragraph id="8B9EEC02FF8389521A82F8FFEE14F87E" blockId="14.[151,1436,1821,2027]" pageId="14" pageNumber="165">
<emphasis id="B9553010FF8389521A82F8FFEE07F8A7" bold="true" box="[151,275,1821,1843]" pageId="14" pageNumber="165">FIGURE 6.</emphasis>
<taxonomicName id="4C219781FF8389521B0FF8FFED48F8A6" box="[282,604,1821,1843]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="14" pageNumber="165" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8389521B0FF8FFEE94F8A6" box="[282,384,1821,1843]" italics="true" pageId="14" pageNumber="165">Scolelepis</emphasis>
(
<emphasis id="B9553010FF8389521B85F8FFEEE3F8A6" box="[400,503,1821,1843]" italics="true" pageId="14" pageNumber="165">Scolelepis</emphasis>
)
<emphasis id="B9553010FF8389521813F8FFED48F8A6" box="[518,604,1821,1843]" italics="true" pageId="14" pageNumber="165">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF8389521876F8FFEDA0F8A7" box="[611,692,1821,1842]" pageId="14" pageNumber="165" rank="species">
<emphasis id="B9553010FF8389521876F8FFEDA0F8A7" bold="true" box="[611,692,1821,1842]" pageId="14" pageNumber="165">sp. nov.</emphasis>
</taxonomicNameLabel>
, A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, CG, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: AB = 1 mm; CI = 0.1 mm.
</paragraph>
</caption>
<caption id="DF5EBC8AFF8289531A82FB47EC71FA07" httpUri="https://zenodo.org/record/263916/files/figure.png" pageId="15" pageNumber="166" targetBox="[151,1436,193,1167]" targetPageId="15">
<paragraph id="8B9EEC02FF8289531A82FB47EC71FA07" blockId="15.[151,1436,1189,1426]" pageId="15" pageNumber="166">
<emphasis id="B9553010FF8289531A82FB47EE06FB2F" bold="true" box="[151,274,1189,1211]" pageId="15" pageNumber="166">FIGURE 7.</emphasis>
<taxonomicName id="4C219781FF8289531B0DFB47ED43FB2E" box="[280,599,1189,1211]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8289531B0DFB47EE6BFB2E" box="[280,383,1189,1211]" italics="true" pageId="15" pageNumber="166">Scolelepis</emphasis>
(
<emphasis id="B9553010FF8289531B99FB47EEE7FB2E" box="[396,499,1189,1211]" italics="true" pageId="15" pageNumber="166">Scolelepis</emphasis>
)
<emphasis id="B9553010FF8289531814FB47ED43FB2E" box="[513,599,1189,1211]" italics="true" pageId="15" pageNumber="166">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF8289531848FB47EDB9FB2F" box="[605,685,1189,1210]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B9553010FF8289531848FB47EDB9FB2F" bold="true" box="[605,685,1189,1210]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
, SEM micrographs. AF, paratype (MNCN 3855). A, distal tip of left palp with elevated lobes, frontal-lateral view (arrowheads indicate faint splitting of lateral rows into two smaller lobes); B, distal section of right palp, frontal view, showing elevated lobes; C, basal part of right palp, lateral view (arrowhead shows smooth basal palpal sheath); D, chaetigers 1325 (from top right to bottom left), ventro-lateral view (arrowheads indicate prechaetal notopodial lamellae; arrow indicates ventral median groove); E, chaetigers 1621 (from top right to bottom left), dorso-lateral view (arrowheads indicate lateral ciliated organs); F, chaetigers 2932 (from right to left), lateral view (note ventrally directed inferior part of notopodial postchaetal lamella and dorsally directed superior part of neuropodial postchaetal lamella). Scale bars: A, B = 50 µm; C = 100 µm; D = 1 mm; E = 0.5 mm; F = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C33BBF89FF82894C1AD2FA5BEA00FDF5" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166" type="discussion">
<paragraph id="8B9EEC02FF8289531AD2FA5BEFF5F93F" blockId="15.[151,1436,1465,2030]" pageId="15" pageNumber="166">
<emphasis id="B9553010FF8289531AD2FA5BEE2FFA47" bold="true" box="[199,315,1465,1490]" pageId="15" pageNumber="166">Remarks.</emphasis>
<taxonomicName id="4C219781FF8289531B57FA5BED5CFA44" box="[322,584,1465,1489]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8289531B57FA5BEEA0FA44" box="[322,436,1465,1489]" italics="true" pageId="15" pageNumber="166">Scolelepis</emphasis>
(
<emphasis id="B9553010FF8289531BD1FA5BEECDFA45" box="[452,473,1465,1488]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531BFCFA5BED5CFA44" box="[489,584,1465,1489]" italics="true" pageId="15" pageNumber="166">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF828953185AFA5BEDBDFA47" box="[591,681,1465,1490]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B9553010FF828953185AFA5BEDBDFA47" bold="true" box="[591,681,1465,1490]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="4C219781FF82895319AEFA5BEA83FA44" authority="Hartmann-Schroder, 1981" authorityName="Hartmann-Schroder" authorityYear="1981" box="[955,1431,1465,1490]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="kudenovi" subGenus="Scolelepis">
<emphasis id="B9553010FF82895319AEFA5BECC4FA45" box="[955,976,1465,1488]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF82895319F5FA5BECE1FA45" box="[992,1013,1465,1488]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531E10FA5BEB7EFA44" box="[1029,1130,1465,1489]" italics="true" pageId="15" pageNumber="166">kudenovi</emphasis>
<bibRefCitation id="EFB091F3FF8289531E67FA5BEA83FA44" author="Hartmann-Schroder" box="[1138,1431,1465,1490]" pageId="15" pageNumber="166" refString="Hartmann-Schroder, G. (1981) Zur Kenntnis des Eulitorals der australischen Kusten unter besonderer Berucksichtigung der Polychaeten und Ostracoden. Teil 6: Die Polychaeten der tropisch-subtropischen Westkuste Australiens (zwischen Exmouth im Norden und Cervantes im Suden). Mitteilungen aus dem Hamburger Zoologischen Museum und Institut, 78, 19 - 96." type="journal article" year="1981">Hartmann-Schröder, 1981</bibRefCitation>
</taxonomicName>
, described from
<collectingRegion id="49E522E0FF8289531B5CFA3CED02FA60" box="[329,534,1500,1525]" country="Australia" name="Western Australia" pageId="15" pageNumber="166">
<collectingRegion id="49E522E0FF8289531B5CFA3CEEB0FA60" box="[329,420,1502,1525]" country="American Samoa" name="Western" pageId="15" pageNumber="166">Western</collectingRegion>
<collectingCountry id="F336AC92FF8289531BB8FA3EED02FA60" box="[429,534,1500,1525]" name="Australia" pageId="15" pageNumber="166">Australia</collectingCountry>
</collectingRegion>
, and
<taxonomicName id="4C219781FF8289531841FA3FEC86FA60" authority="Zhou, 2014" authorityName="Zhou" authorityYear="2014" box="[596,914,1500,1525]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF8289531841FA3FED7DFA61" box="[596,617,1501,1524]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF828953186FFA3FED9BFA61" box="[634,655,1501,1524]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF82895318B4FA3FEC11FA60" box="[673,773,1501,1525]" italics="true" pageId="15" pageNumber="166">angulata</emphasis>
<bibRefCitation id="EFB091F3FF8289531918FA3EEC86FA60" author="Zhou" box="[781,914,1500,1525]" pageId="15" pageNumber="166" refString="Zhou, J. (2014) A new species of Scolelepis (Polychaeta: Spionidae) from Chinese seas. Raffles Bulletin of Zoology, 62, 490 - 495." type="journal article" year="2014">Zhou, 2014</bibRefCitation>
</taxonomicName>
, recently described from the South
<collectingCountry id="F336AC92FF8289531F3DFA3EEA7FFA60" box="[1320,1387,1500,1525]" name="China" pageId="15" pageNumber="166">China</collectingCountry>
Sea and the Yellow Sea. The three species are morphologically similar in the following characteristics: the shape of the prostomium (somewhat truncate, with a conical medial tip and subdistal lateral angles anteriorly and with a transversal constriction in the posterior third), the caruncle extending to the posterior margin of chaetiger 1, short palps (never exceeding chaetiger 5), the presence of anterior prechaetal notopodial lamellae, and bidentate hooded hooks.
</paragraph>
<paragraph id="8B9EEC02FF8289531AD2F957EE79F833" blockId="15.[151,1436,1465,2030]" pageId="15" pageNumber="166">
With regard to palp ciliation pattern, the new species is somewhat similar to
<taxonomicName id="4C219781FF8289531E0BF957EA6BF95B" authority="Dauer, 1985" authorityName="Dauer" authorityYear="1985" box="[1054,1407,1717,1742]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="hutchingsae" subGenus="Scolelepis">
<emphasis id="B9553010FF8289531E0BF957EB27F959" box="[1054,1075,1717,1740]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF8289531E56F957EB4CF959" box="[1091,1112,1717,1740]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531E7DF957EBFBF958" box="[1128,1263,1717,1741]" italics="true" pageId="15" pageNumber="166">hutchingsae</emphasis>
<bibRefCitation id="EFB091F3FF8289531EE3F954EA6BF95B" author="Dauer" box="[1270,1407,1717,1742]" pageId="15" pageNumber="166" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">
Dauer,
<quantity id="4CD941E7FF8289531F52F957EA6BF95B" box="[1351,1407,1717,1742]" metricMagnitude="1" metricUnit="m" metricValue="5.0419" pageId="15" pageNumber="166" unit="in" value="1985.0">1985</quantity>
</bibRefCitation>
</taxonomicName>
in having distinctly separated long and short rows, with the short rows on elevated lobes (
<bibRefCitation id="EFB091F3FF8289531E65F93BEA09F964" author="Williams" box="[1136,1309,1753,1778]" pageId="15" pageNumber="166" refString="Williams, J. D. (2007) New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on the palp ciliation patterns of the genus Scolelepis. Zootaxa, 1459, 1 - 35." type="journal article" year="2007">Williams 2007</bibRefCitation>
). However,
<taxonomicName id="4C219781FF8289531A82F91FEE5CF880" box="[151,328,1789,1813]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8289531A82F91FEFB8F881" box="[151,172,1789,1812]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF8289531AD5F91FEFC1F881" box="[192,213,1789,1812]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531AFCF91FEE5CF880" box="[233,328,1789,1813]" italics="true" pageId="15" pageNumber="166">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF8289531B47F91FEEBBF883" box="[338,431,1789,1814]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B9553010FF8289531B47F91FEEBBF883" bold="true" box="[338,431,1789,1814]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
presents a new pattern in having both rows on elevated lobes. Thus, for those eleven species of
<taxonomicName id="4C219781FF8289531B0BF8C3EE9BF8AC" box="[286,399,1825,1849]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="genus">
<emphasis id="B9553010FF8289531B0BF8C3EE9BF8AC" box="[286,399,1825,1849]" italics="true" pageId="15" pageNumber="166">Scolelepis</emphasis>
</taxonomicName>
that have been so far examined with SEM for the palp ciliary pattern, six distinct morphological
<typeStatus id="549A52A0FF8289531B50F8A4EE94F8CB" box="[325,384,1862,1886]" pageId="15" pageNumber="166">types</typeStatus>
have been recognized (
<bibRefCitation id="EFB091F3FF8289531882F8A7EC50F8C8" author="Williams" box="[663,836,1861,1886]" pageId="15" pageNumber="166" refString="Williams, J. D. (2007) New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on the palp ciliation patterns of the genus Scolelepis. Zootaxa, 1459, 1 - 35." type="journal article" year="2007">Williams 2007</bibRefCitation>
;
<bibRefCitation id="EFB091F3FF8289531941F8A7ECC1F8C8" author="Zhou" box="[852,981,1861,1886]" pageId="15" pageNumber="166" refString="Zhou, J. (2014) A new species of Scolelepis (Polychaeta: Spionidae) from Chinese seas. Raffles Bulletin of Zoology, 62, 490 - 495." type="journal article" year="2014">Zhou 2014</bibRefCitation>
; present study). It is likely that further investigation of palp morphology for other species will greatly increase the number of morphological
<typeStatus id="549A52A0FF8289531F54F888EA68F817" box="[1345,1404,1898,1922]" pageId="15" pageNumber="166">types</typeStatus>
to which they belong.
</paragraph>
<paragraph id="8B9EEC02FF82894C1AD2F853EB56FED5" blockId="15.[151,1436,1465,2030]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">
The new species is similar to the Australian specimens of
<taxonomicName id="4C219781FF828953194FF853EB1FF85C" box="[858,1035,1969,1993]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="kudenovi" subGenus="Scolelepis">
<emphasis id="B9553010FF828953194FF853EC7BF85D" box="[858,879,1969,1992]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF8289531995F853EC80F85D" box="[896,916,1969,1992]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF82895319B3F853EB1FF85C" box="[934,1035,1969,1993]" italics="true" pageId="15" pageNumber="166">kudenovi</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C219781FF8289531E9BF853EA2EF85C" box="[1166,1338,1969,1993]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF8289531E9BF853EBB0F85D" box="[1166,1188,1969,1992]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF8289531EA1F853EBDEF85D" box="[1204,1226,1969,1992]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531ECEF853EA2EF85C" box="[1243,1338,1969,1993]" italics="true" pageId="15" pageNumber="166">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF8289531F57F853EA88F85F" box="[1346,1436,1969,1994]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B9553010FF8289531F57F853EA88F85F" bold="true" box="[1346,1436,1969,1994]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C219781FF8289531B38F837EEF3F878" box="[301,487,2005,2029]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="kudenovi" subGenus="Scolelepis">
<emphasis id="B9553010FF8289531B38F837EE57F879" box="[301,323,2005,2028]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
(
<emphasis id="B9553010FF8289531B4DF837EE79F879" box="[344,365,2005,2028]" italics="true" pageId="15" pageNumber="166">S.</emphasis>
)
<emphasis id="B9553010FF8289531B97F837EEF3F878" box="[386,487,2005,2029]" italics="true" pageId="15" pageNumber="166">kudenovi</emphasis>
</taxonomicName>
by the shape of subdistal anterolateral lobes on prostomium, which are broadly rounded in the former and bluntly tapered in the latter, the presence of well-developed peristomial wings in the former, and the shape of the neuropodial hooded hooks, which have smaller, bluntly rounded teeth instead of sharp teeth with an upright apical tooth. Also, specimens of
<taxonomicName id="4C219781FF9D894C18FBFF02EC88FF6D" box="[750,924,224,248]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="kudenovi" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18FBFF02EC10FF62" box="[750,772,224,247]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1906FF02EC3CFF62" box="[787,808,224,247]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1922FF02EC88FF6D" box="[823,924,224,248]" italics="true" pageId="16" pageNumber="167">kudenovi</emphasis>
</taxonomicName>
appear to be much more slender than those of
<taxonomicName id="4C219781FF9D894C1A82FEE6EE57FE89" box="[151,323,260,284]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1A82FEE6EFB8FE8E" box="[151,172,260,283]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1AA8FEE6EFC6FE8E" box="[189,210,260,283]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1AF1FEE6EE57FE89" box="[228,323,260,284]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1B5FFEE6EEB1FE88" box="[330,421,260,285]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1B5FFEE6EEB1FE88" bold="true" box="[330,421,260,285]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<tableCitation id="C6A3D9B9FF9D894C1BA1FEE6ED1DFE89" box="[436,521,260,285]" captionStart="TABLE 1" captionStartId="18.[119,201,608,629]" captionTargetBox="[119,1883,664,1542]" captionTargetPageId="18" captionText="TABLE 1. Taxonomic characters of selected species in the genus Scolelepis (Scolelepis)." httpUri="http://table.plazi.org/id/DF5EBC8AFF9F894E1A62FD82ECC6FDE0" pageId="16" pageNumber="167" tableUuid="DF5EBC8AFF9F894E1A62FD82ECC6FDE0">Table 1</tableCitation>
). Specimens of
<taxonomicName id="4C219781FF9D894C18D6FEE6EC61FE89" box="[707,885,260,284]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="kudenovi" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18D6FEE6EDCCFE8E" box="[707,728,260,283]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C18FCFEE6EDEAFE8E" box="[745,766,260,283]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1905FEE6EC61FE89" box="[784,885,260,284]" italics="true" pageId="16" pageNumber="167">kudenovi</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="EFB091F3FF9D894C1E13FEE6EBA2FE89" author="Imajima" box="[1030,1206,260,285]" pageId="16" pageNumber="167" refString="Imajima, M. (1992) Spionidae (Annelida, Polychaeta) from Japan, 8. The genus Scolelepis. Bulletin of the National Science Museum Tokyo (Japan). Series A. Zoology, 18, 1 - 34." type="journal article" year="1992">Imajima (1992)</bibRefCitation>
from
<collectingCountry id="F336AC92FF9D894C1EE8FEE7EA29FE89" box="[1277,1341,261,284]" name="Japan" pageId="16" pageNumber="167">Japan</collectingCountry>
seem to belong to a species distinct from the Australian material (Meißner &amp; Götting 2015).
</paragraph>
<paragraph id="8B9EEC02FF9D894C1AD2FEAEEA00FDF5" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
<taxonomicName id="4C219781FF9D894C1AD2FEAEEEC1FEF1" box="[199,469,332,356]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1AD2FEAEEE2DFEF1" box="[199,313,332,356]" italics="true" pageId="16" pageNumber="167">Scolelepis</emphasis>
(
<emphasis id="B9553010FF9D894C1B59FEAEEE76FEF6" box="[332,354,332,355]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1B63FEAEEEC1FEF1" box="[374,469,332,356]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1BCAFEAEED29FEF0" box="[479,573,332,357]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1BCAFEAEED29FEF0" bold="true" box="[479,573,332,357]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C219781FF9D894C18C9FEAEEC87FEF1" box="[732,915,332,356]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18C9FEAEEDE5FEF6" box="[732,753,332,355]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1910FEAEEC0FFEF6" box="[773,795,332,355]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C193AFEAEEC87FEF1" box="[815,915,332,356]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
in the shape of the prostomium, the lack of pigmentation on the posterior caruncle, the shape of the peristomium, which has a smooth contour in
<taxonomicName id="4C219781FF9D894C1F46FE92EFE2FE39" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1F46FE92EA7CFE12" box="[1363,1384,368,391]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1F68FE92EA87FE12" box="[1405,1427,368,391]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1A82FE76EFE2FE39" box="[151,246,404,428]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1B15FE76EE49FE38" box="[256,349,404,429]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1B15FE76EE49FE38" bold="true" box="[256,349,404,429]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and a wavy contour in
<taxonomicName id="4C219781FF9D894C186DFE76EC38FE39" box="[632,812,404,428]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C186DFE76ED99FE3E" box="[632,653,404,427]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C18B5FE76EDA2FE3E" box="[672,694,404,427]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C18DCFE76EC38FE39" box="[713,812,404,428]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
, the shape of the neuropodial postchaetal lamellae of anteriormost chaetigers, which are ovate in
<taxonomicName id="4C219781FF9D894C18B5FE5AEC40FE45" box="[672,852,440,464]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18B5FE5AEDA1FE5A" box="[672,693,440,463]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C18DEFE5AEDF4FE5A" box="[715,736,440,463]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C18E0FE5AEC40FE45" box="[757,852,440,464]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1974FE55ECD4FE45" box="[865,960,439,464]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1974FE55ECD4FE45" bold="true" box="[865,960,439,464]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and rectangular in
<taxonomicName id="4C219781FF9D894C1EB8FE5AEA72FE45" box="[1197,1382,440,464]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1EB8FE5AEBD6FE5A" box="[1197,1218,440,463]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1ECDFE5AEBF9FE5A" box="[1240,1261,440,463]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1F16FE5AEA72FE45" box="[1283,1382,440,464]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
, the extension of branchiae, which are present on at least 79 chaetigers (the longest anterior fragment), while in
<taxonomicName id="4C219781FF9D894C1F42FE3EEFEFFD8D" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1F42FE3EEA79FE66" box="[1367,1389,476,499]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1F68FE3EEA87FE66" box="[1405,1427,476,499]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1A82FDE2EFEFFD8D" box="[151,251,512,536]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
the branchiae are limited to first 4351 chaetigers, and by ciliation on palps (absent in
<taxonomicName id="4C219781FF9D894C1ECEFDE2EA98FD8D" box="[1243,1420,512,536]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1ECEFDE2EBE4FD82" box="[1243,1264,512,535]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1F17FDE2EA03FD82" box="[1282,1303,512,535]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1F3DFDE2EA98FD8D" box="[1320,1420,512,536]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
). There are also differences in the habitat preferences:
<taxonomicName id="4C219781FF9D894C1916FDC6ECACFDA9" box="[771,952,548,572]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="angulata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1916FDC6EC0CFDAE" box="[771,792,548,571]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1939FDC6EC55FDAE" box="[812,833,548,571]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1941FDC6ECACFDA9" box="[852,952,548,572]" italics="true" pageId="16" pageNumber="167">angulata</emphasis>
</taxonomicName>
mainly inhabits tidal flats with saltmarsh cordgrass (
<taxonomicName id="4C219781FF9D894C1B07FDAAEEEFFDF5" box="[274,507,584,608]" class="Liliopsida" family="Poaceae" genus="Spartina" kingdom="Plantae" order="Poales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="species" species="alterniflora">
<emphasis id="B9553010FF9D894C1B07FDAAEEEFFDF5" box="[274,507,584,608]" italics="true" pageId="16" pageNumber="167">Spartina alterniflora</emphasis>
</taxonomicName>
), whilst
<taxonomicName id="4C219781FF9D894C184BFDAAEC13FDF5" box="[606,775,584,608]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C184BFDAAED67FDCA" box="[606,627,584,607]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1896FDAAED8CFDCA" box="[643,664,584,607]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C18BDFDAAEC13FDF5" box="[680,775,584,608]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C191BFDA5EC7CFDF5" box="[782,872,583,608]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C191BFDA5EC7CFDF5" bold="true" box="[782,872,583,608]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
mainly inhabits fine sublittoral sands.
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF9D894C1AD2FD8EECDEFA98" pageId="16" pageNumber="167" type="reference_group">
<paragraph id="8B9EEC02FF9D894C1AD2FD8EECF9FCC9" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
<taxonomicName id="4C219781FF9D894C1AD2FD8EEEC3FD11" ID-CoL="4VRN6" box="[199,471,620,644]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1AD2FD8EEE2DFD11" box="[199,313,620,644]" italics="true" pageId="16" pageNumber="167">Scolelepis</emphasis>
(
<emphasis id="B9553010FF9D894C1B5BFD8EEE77FD16" box="[334,355,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1B6DFD8EEEC3FD11" box="[376,471,620,644]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1BF6FD8EED56FD10" box="[483,578,620,645]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1BF6FD8EED56FD10" bold="true" box="[483,578,620,645]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
has been confused in the past with
<taxonomicName id="4C219781FF9D894C19ECFD8EEBAFFD11" ID-CoL="4VRNB" box="[1017,1211,620,644]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C19ECFD8EEB1AFD16" box="[1017,1038,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1E31FD8EEB2CFD16" box="[1060,1080,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1E5BFD8FEBAFFD11" box="[1102,1211,621,644]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1EDBFD8EEA60FD11" ID-CoL="73X4V" box="[1230,1396,620,644]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="mesnili" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1EDBFD8EEBF7FD16" box="[1230,1251,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1EEDFD8EEA1AFD16" box="[1272,1294,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1F36FD8EEA60FD11" box="[1315,1396,620,644]" italics="true" pageId="16" pageNumber="167">mesnili</emphasis>
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1F92FD8EEE9CFD3D" ID-CoL="4P2BL" class="Polychaeta" family="Spionidae" genus="Pseudomalacoceros" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="cantabra" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1F92FD8EEA88FD16" box="[1415,1436,620,643]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1AB5FD72EE06FD3D" box="[160,274,656,680]" italics="true" pageId="16" pageNumber="167">Scolelepis</emphasis>
)
<emphasis id="B9553010FF9D894C1B36FD72EE9CFD3D" box="[291,392,656,680]" italics="true" pageId="16" pageNumber="167">cantabra</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C219781FF9D894C1BD3FD72EC32FD3D" ID-CoL="36VN5" authority="Harman, 1951" authorityName="Harman" authorityYear="1951" box="[454,806,655,680]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="B9553010FF9D894C1BD3FD72ED6FFD32" box="[454,635,656,679]" italics="true" pageId="16" pageNumber="167">Dispio uncinata</emphasis>
Harman, 1951
</taxonomicName>
. Thus, individuals from
<collectingRegion id="49E522E0FF9D894C1E2EFD6DEBBCFD3D" box="[1083,1192,655,680]" country="Spain" name="Cantabria" pageId="16" pageNumber="167">Cantabria</collectingRegion>
were identified by G. San Martín either as
<taxonomicName id="4C219781FF9D894C1B6AFD57ED26FD59" ID-CoL="36VN5" box="[383,562,693,716]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="B9553010FF9D894C1B6AFD57ED26FD59" box="[383,562,693,716]" italics="true" pageId="16" pageNumber="167">Dispio uncinata</emphasis>
</taxonomicName>
and subsequently revised by O. Díaz as
<taxonomicName id="4C219781FF9D894C19E3FD56EBD4FD59" ID-CoL="73X4V" box="[1014,1216,692,716]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="mesnili">
<emphasis id="B9553010FF9D894C19E3FD56EBD4FD59" box="[1014,1216,692,716]" italics="true" pageId="16" pageNumber="167">Scolelepis mesnili</emphasis>
</taxonomicName>
(unpublished study by
<collectingRegion id="49E522E0FF9D894C1AAEFD3AEE13FD65" box="[187,263,728,752]" country="France" name="Centre" pageId="16" pageNumber="167">Centro</collectingRegion>
de Investigaciones Submarinas, 1998, Guillermo San Martín, pers. comm.,
<date id="FF9FCAC2FF9D894C1E73FD3AEBC3FD65" box="[1126,1239,727,752]" pageId="16" pageNumber="167" value="2015-12">Dec 2015</date>
), or as
<taxonomicName id="4C219781FF9D894C1F3FFD3AEE10FC81" ID-CoL="4VRNB" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF9D894C1F3FFD3AEE10FC81" italics="true" pageId="16" pageNumber="167">Scolelepis squamata</emphasis>
</taxonomicName>
by G. San Martín and J.M. Viéitez. The material from
<collectingRegion id="49E522E0FF9D894C196DFD1EECDDFC80" box="[888,969,764,789]" country="Spain" name="Galicia" pageId="16" pageNumber="167">Galicia</collectingRegion>
was identified as
<taxonomicName id="4C219781FF9D894C1E8DFD1EEA63FC81" ID-CoL="4VRMP" box="[1176,1399,764,788]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="cantabra">
<emphasis id="B9553010FF9D894C1E8DFD1EEA63FC81" box="[1176,1399,764,788]" italics="true" pageId="16" pageNumber="167">Scolelepis cantabra</emphasis>
</taxonomicName>
by J.M.
<bibRefCitation id="EFB091F3FF9D894C1AC0FCFDEE61FCAD" author="Vieitez" box="[213,373,799,824]" pageId="16" pageNumber="167" refString="Vieitez, J. M. (1976) Ecologia de Poliquetos y moluscos de la playa de Meira (Ria de Vigo). I. Estudio de las comunidades. Investigacion Pesquera, 40 (1), 223 - 248." type="journal article" year="1976">Viéitez (1976</bibRefCitation>
,
<bibRefCitation id="EFB091F3FF9D894C1B93FCC2EEABFCAD" author="Vieitez" box="[390,447,800,824]" pageId="16" pageNumber="167" refString="Vieitez, J. M. (1981) Estudio de las comunidades bentonicas de dos playas de las rias de Pontevedra y Vigo (Galicia, Espana). Boletin. Instituto Espanol de Oceanografia, 6 (331), 242 - 258." type="journal article" year="1981">1981</bibRefCitation>
) or as
<taxonomicName id="4C219781FF9D894C1809FCC2EC13FCAD" ID-CoL="4VRNB" box="[540,775,800,824]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF9D894C1809FCC2EC13FCAD" box="[540,775,800,824]" italics="true" pageId="16" pageNumber="167">Scolelepis squamata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB091F3FF9D894C1929FCC2ECF8FCAD" author="Parapar" box="[828,1004,800,824]" pageId="16" pageNumber="167" refString="Parapar, J. (1991) Anelidos Poliquetos bentonicos de la ria de Ferrol (Galicia). PhD-thesis, University of Santiago de Compostela, 1104 pp." type="book" year="1991">Parapar (1991)</bibRefCitation>
. Also, the material from Licola, the Mediterranean Sea, was identified as
<taxonomicName id="4C219781FF9D894C182EFCA6EC34FCC9" ID-CoL="4VRNB" box="[571,800,836,860]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF9D894C182EFCA6EC34FCC9" box="[571,800,836,860]" italics="true" pageId="16" pageNumber="167">Scolelepis squamata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB091F3FF9D894C195FFCA6ECFEFCC9" author="Dorjes" box="[842,1002,836,861]" pageId="16" pageNumber="167" refString="Dorjes, J. (1971) Der Golf von Gaeta (Tyrrhenisches Meer). IV. Das Makrobenthos und seine kustenparallele Zonierung. Senckenbergiana maritima, 3, 203 - 246." type="journal article" year="1971">Dörjes (1971)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9EEC02FF9D894C1AD2FC85ECDEFA98" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
The new species differs from
<taxonomicName id="4C219781FF9D894C1808FC8AEDCDFC15" ID-CoL="4VRNB" box="[541,729,872,896]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1808FC8AED27FCEA" box="[541,563,872,895]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1850FC8AED4EFCEA" box="[581,602,872,895]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1879FC8BEDCDFC15" box="[620,729,873,896]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
by the shape of the prostomium (with anterolateral lobes and with a transverse furrow in front of the caruncle in
<taxonomicName id="4C219781FF9D894C18F9FC6EEC8EFC31" ID-CoL="4VRN6" box="[748,922,908,932]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18F9FC6EEC14FC36" box="[748,768,908,931]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1906FC6EEC3CFC36" box="[787,808,908,931]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C192EFC6EEC8EFC31" box="[827,922,908,932]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C19B6FC6EECEBFC30" box="[931,1023,908,933]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C19B6FC6EECEBFC30" bold="true" box="[931,1023,908,933]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and without anterolateral lobes and without a transverse furrow in front of the caruncle in
<taxonomicName id="4C219781FF9D894C190EFC52ECCDFC5D" ID-CoL="4VRNB" box="[795,985,944,968]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C190EFC52EC24FC52" box="[795,816,944,967]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1951FC52EC4CFC52" box="[836,856,944,967]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1979FC53ECCDFC5D" box="[876,985,945,968]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
), by having a shorter peristomium, by having well-developed lateral peristomial wings which are separated from the prostomium by a distinct grove, by the shape of basal sheaths (smooth in
<taxonomicName id="4C219781FF9D894C1840FC1AEC11FB85" ID-CoL="4VRN6" box="[597,773,1016,1040]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1840FC1AED7EFB9A" box="[597,618,1016,1039]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1868FC1AED86FB9A" box="[637,658,1016,1039]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C18B3FC1AEC11FB85" box="[678,773,1016,1040]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C191AFC15EC78FB85" box="[783,876,1015,1040]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C191AFC15EC78FB85" bold="true" box="[783,876,1015,1040]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and slightly rugose in
<taxonomicName id="4C219781FF9D894C1E6EFC1AEA2DFB85" ID-CoL="4VRNB" box="[1147,1337,1016,1040]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1E6EFC1AEB84FB9A" box="[1147,1168,1016,1039]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1EB1FC1AEBADFB9A" box="[1188,1209,1016,1039]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1ED9FC1BEA2DFB85" box="[1228,1337,1017,1040]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
), by the ciliation pattern of palps, by the chaetiger 1 having much longer notopodial postchaetal lamellae and capillaries, by the shape of anterior postchaetal lamellae (with entire, smooth margins in
<taxonomicName id="4C219781FF9D894C19C1FBA2EB6AFBCD" ID-CoL="4VRN6" box="[980,1150,1088,1112]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C19C1FBA2ECFDFBC2" box="[980,1001,1088,1111]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C19EFFBA2EB1BFBC2" box="[1018,1039,1088,1111]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1E0AFBA2EB6AFBCD" box="[1055,1150,1088,1112]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1E90FBDDEBCAFBCD" box="[1157,1246,1087,1112]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1E90FBDDEBCAFBCD" bold="true" box="[1157,1246,1087,1112]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and with folded, notched margins in
<taxonomicName id="4C219781FF9D894C1B66FB86ED3EFBE9" ID-CoL="4VRNB" box="[371,554,1124,1148]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1B66FB86EE9CFBEE" box="[371,392,1124,1147]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1B8DFB86EEB9FBEE" box="[408,429,1124,1147]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1BA8FB87ED3EFBE9" box="[445,554,1125,1148]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
), and by the presence of a distinct prechaetal lamellae. Also, in
<taxonomicName id="4C219781FF9D894C1EE6FB86EA88FBE9" ID-CoL="4VRN6" box="[1267,1436,1124,1148]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1EE6FB86EA1CFBEE" box="[1267,1288,1124,1147]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1F0DFB86EA39FBEE" box="[1304,1325,1124,1147]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1F28FB86EA88FBE9" box="[1341,1436,1124,1148]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1A82FB65EFE7FB35" box="[151,243,1159,1184]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1A82FB65EFE7FB35" bold="true" box="[151,243,1159,1184]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
, the branchiae on anterior chaetigers are less fused to the notopodial postchaetal lamellae (about 1/2 as against to 2/
<quantity id="4CD941E7FF9D894C1B37FB4EEE5FFB50" box="[290,331,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="167" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C219781FF9D894C1B47FB4EED1EFB51" ID-CoL="4VRNB" box="[338,522,1196,1220]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1B47FB4EEE73FB56" box="[338,359,1196,1219]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1B6DFB4EEE99FB56" box="[376,397,1196,1219]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1B88FB4FED1EFB51" box="[413,522,1197,1220]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
). Palps are usually deciduous in
<taxonomicName id="4C219781FF9D894C1969FB4EEB32FB51" ID-CoL="4VRN6" box="[892,1062,1196,1220]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1969FB4EEC86FB56" box="[892,914,1196,1219]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C19B7FB4EECA3FB56" box="[930,951,1196,1219]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C19D2FB4EEB32FB51" box="[967,1062,1196,1220]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1E38FB4EEB93FB50" box="[1069,1159,1196,1221]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1E38FB4EEB93FB50" bold="true" box="[1069,1159,1196,1221]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
and non-deciduous in
<taxonomicName id="4C219781FF9D894C1F92FB4EEE27FB7D" ID-CoL="4VRNB" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1F92FB4EEA88FB56" box="[1415,1436,1196,1219]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1AB5FB32EFA1FB72" box="[160,181,1232,1255]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1AD3FB33EE27FB7D" box="[198,307,1233,1256]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
. The eggs in
<taxonomicName id="4C219781FF9D894C1BDBFB32ED93FB7D" ID-CoL="4VRNB" box="[462,647,1232,1256]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="squamata" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1BDBFB32EEF7FB72" box="[462,483,1232,1255]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C1BE1FB32ED1DFB72" box="[500,521,1232,1255]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C180FFB33ED93FB7D" box="[538,647,1233,1256]" italics="true" pageId="16" pageNumber="167">squamata</emphasis>
</taxonomicName>
are characterized by 12 peripheral vesicles (
<bibRefCitation id="EFB091F3FF9D894C1E6BFB32EA1BFB7D" author="Joyner" box="[1150,1295,1231,1256]" pageId="16" pageNumber="167" refString="Joyner, A. (1962) Reproduction and larval life of Nerine cirratulus (Delle Chiaje) Family Spionidae. Proceedings of the Zoological Society of London, 138 (4), 655 - 666. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1962. tb 05338. x" type="journal article" year="1962">Joyner 1962</bibRefCitation>
), whereas in
<taxonomicName id="4C219781FF9D894C1A82FB16EE89FA99" ID-CoL="4VRN6" box="[151,413,1268,1292]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1A82FB16EE1DFA99" box="[151,265,1268,1292]" italics="true" pageId="16" pageNumber="167">Scolelepis</emphasis>
(
<emphasis id="B9553010FF9D894C1B0CFB16EE3AFA9E" box="[281,302,1268,1291]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1B2BFB16EE89FA99" box="[318,413,1268,1292]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1BB1FB16EEE9FA98" box="[420,509,1268,1293]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1BB1FB16EEE9FA98" bold="true" box="[420,509,1268,1293]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
the eggs have 2731 peripheral vesicles.
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF9D894C1AD2FAF5ECB0F9C5" pageId="16" pageNumber="167" type="description">
<paragraph id="8B9EEC02FF9D894C1AD2FAF5EDBAFAED" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
The new species can be easily distinguished from
<taxonomicName id="4C219781FF9D894C18E1FAFAECB6FAA5" box="[756,930,1304,1328]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="cantabra" subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C18E1FAFAEC1CFABA" box="[756,776,1304,1327]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
(
<emphasis id="B9553010FF9D894C190DFAFAEC3AFABA" box="[792,814,1304,1327]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1928FAFAECB6FAA5" box="[829,930,1304,1328]" italics="true" pageId="16" pageNumber="167">cantabra</emphasis>
</taxonomicName>
by the absence of occipital antenna (which is present in
<taxonomicName id="4C219781FF9D894C1B19FADEED99FAC1" authority="Maciolek (1987)" authorityName="Maciolek" authorityYear="1987" box="[268,653,1340,1365]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="subSpecies" species="cantabra" subSpecies="fide">
<emphasis id="B9553010FF9D894C1B19FADEEEABFAC1" box="[268,447,1340,1364]" italics="true" pageId="16" pageNumber="167">S. cantabra fide</emphasis>
<bibRefCitation id="EFB091F3FF9D894C1BD0FADEED99FAC1" author="Maciolek" box="[453,653,1340,1365]" pageId="16" pageNumber="167" refString="Maciolek. N. J. (1987) New species and records of Scolelepis (Polychaeta: Spionidae) from the east coast of North America. Proceedings of the Biological Society of Washington, 7, 16 - 40." type="journal article" year="1987">Maciolek (1987))</bibRefCitation>
</taxonomicName>
, by the presence of notochaetae in chaetiger 1, and by the presence of a notch in the neuropodial postchaetal lamellae.
</paragraph>
<paragraph id="8B9EEC02FF9D894C1AD2FA66ECB0F9C5" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
<taxonomicName id="4C219781FF9D894C1AD2FA66EEDAFA09" box="[199,462,1412,1436]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9D894C1AD2FA66EE2DFA09" box="[199,313,1412,1436]" italics="true" pageId="16" pageNumber="167">Scolelepis</emphasis>
(
<emphasis id="B9553010FF9D894C1B5FFA66EE4BFA0E" box="[330,351,1412,1435]" italics="true" pageId="16" pageNumber="167">S.</emphasis>
)
<emphasis id="B9553010FF9D894C1B7AFA66EEDAFA09" box="[367,462,1412,1436]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9D894C1BC3FA66ED25FA08" box="[470,561,1412,1437]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B9553010FF9D894C1BC3FA66ED25FA08" bold="true" box="[470,561,1412,1437]" pageId="16" pageNumber="167">sp. nov.</emphasis>
</taxonomicNameLabel>
has been confused also with
<taxonomicName id="4C219781FF9D894C196BFA67EB20FA09" box="[894,1076,1413,1436]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="B9553010FF9D894C196BFA67EB20FA09" box="[894,1076,1413,1436]" italics="true" pageId="16" pageNumber="167">Dispio uncinata</emphasis>
</taxonomicName>
because of the large notopodial postchaetal lamellae on chaetiger 1, which resembles a lamella joined with branchia. At first glance it was difficult to discern whether on chaetiger 1 there is a large lamella or a small branchia completely fused to the notopodial lamella. However, the vascular core of the branchia, which can be detected on chaetiger 2, was not observed on chaetiger 1. Also, anterior notochaetae are very long, resembling those of
<taxonomicName id="4C219781FF9D894C19E4F9F7EBB3F9B9" box="[1009,1191,1557,1580]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="B9553010FF9D894C19E4F9F7EBB3F9B9" box="[1009,1191,1557,1580]" italics="true" pageId="16" pageNumber="167">Dispio uncinata</emphasis>
</taxonomicName>
. There are, however, major differences in most other characters between these two species.
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF9D894C1AD2F9BEED12F848" pageId="16" pageNumber="167" type="etymology">
<paragraph id="8B9EEC02FF9D894C1AD2F9BEECE0F928" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
<emphasis id="B9553010FF9D894C1AD2F9BEEE5AF9E0" bold="true" box="[199,334,1628,1653]" pageId="16" pageNumber="167">Etymology.</emphasis>
The species name “
<taxonomicName id="4C219781FF9D894C1854F9BEEDB4F9E1" box="[577,672,1628,1652]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta">
<emphasis id="B9553010FF9D894C1854F9BEEDB4F9E1" box="[577,672,1628,1652]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
” (derived from the Latin nominative singular feminine adjective
<taxonomicName id="4C219781FF9D894C1A82F962EFE2F90D" box="[151,246,1664,1688]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="neglecta">
<emphasis id="B9553010FF9D894C1A82F962EFE2F90D" box="[151,246,1664,1688]" italics="true" pageId="16" pageNumber="167">neglecta</emphasis>
</taxonomicName>
meaning disregarded, neglected, or ignored) refers to the fact that the species has in the past been misidentified and overlooked, without being recognized as a distinct species.
</paragraph>
<paragraph id="8B9EEC02FF9D894C1AD2F925ED12F848" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="167">
<emphasis id="B9553010FF9D894C1AD2F925EEA9F975" bold="true" box="[199,445,1735,1760]" pageId="16" pageNumber="167">Habitat and ecology.</emphasis>
The species inhabits fine sublittoral sands with an admixture of shells at depths ranging from
<quantity id="4CD941E7FF9D894C1AC1F90EEE42F891" box="[212,342,1772,1797]" metricMagnitude="1" metricUnit="m" metricValue="1.625" metricValueMax="3.2" metricValueMin="0.05" pageId="16" pageNumber="167" unit="m" value="16.25" valueMax="32.0" valueMin="0.5">
0.5 to
<specimenCount id="9D27278BFF9D894C1B0EF90EEE42F891" box="[283,342,1772,1796]" pageId="16" pageNumber="167" type="generic">32 m</specimenCount>
</quantity>
. Short palps could indicate that this species is dependent on habitats rich in organic matter (
<bibRefCitation id="EFB091F3FF9D894C1F43F90EEFC0F8BD" author="Dauer" pageId="16" pageNumber="167" refString="Dauer, D. M. (2000) Functional morphology and feeding behavior of Spio setosa (Polychaeta: Spionidae). Bulletin of Marine Science, 67 (1), 269 - 275." type="journal article" year="2000">Dauer 2000</bibRefCitation>
). It usually occurs together with
<taxonomicName id="4C219781FF9D894C1879F8F2EB10F8BD" authority="Baird, 1870" authorityName="Baird" authorityYear="1870" box="[620,1028,1807,1832]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="bilineata">
<emphasis id="B9553010FF9D894C1879F8F2EC42F8BD" box="[620,854,1808,1832]" italics="true" pageId="16" pageNumber="167">Aponuphis bilineata</emphasis>
(Baird, 1870)
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1E02F8F2EA8CF8BD" authority="Orsted, 1843" authorityName="Orsted" authorityYear="1843" box="[1047,1432,1807,1832]" class="Polychaeta" family="Goniadidae" genus="Goniada" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="maculata">
<emphasis id="B9553010FF9D894C1E02F8F2EBFBF8BD" box="[1047,1263,1808,1832]" italics="true" pageId="16" pageNumber="167">Goniada maculata</emphasis>
Örsted, 1843
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1A82F8D6ED0FF8D8" authority="Ehlers, 1868" authorityName="Ehlers" authorityYear="1868" box="[151,539,1844,1869]" class="Polychaeta" family="Lumbrineridae" genus="Hilbigneris" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="B9553010FF9D894C1A82F8D6EE66F8D9" box="[151,370,1844,1868]" italics="true" pageId="16" pageNumber="167">Hilbigneris gracilis</emphasis>
(Ehlers, 1868)
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1832F8D6ECF0F8D8" authority="F. Muller, 1858" authorityName="F. Muller" authorityYear="1858" box="[551,996,1844,1869]" class="Polychaeta" family="Magelonidae" genus="Magelona" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="papillicornis">
<emphasis id="B9553010FF9D894C1832F8D6EC3AF8D9" box="[551,814,1844,1868]" italics="true" pageId="16" pageNumber="167">Magelona papillicornis</emphasis>
F. Müller, 1858
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C19FAF8D6EA23F8D8" authority="Ehlers, 1868" authorityName="Ehlers" authorityYear="1868" box="[1007,1335,1844,1869]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="cirrosa">
<emphasis id="B9553010FF9D894C19FAF8D6EBB4F8D9" box="[1007,1184,1844,1868]" italics="true" pageId="16" pageNumber="167">Nephtys cirrosa</emphasis>
Ehlers, 1868
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1F57F8D6ED40F8E5" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="hombergii">
<emphasis id="B9553010FF9D894C1F57F8D6EE1DF8E5" italics="true" pageId="16" pageNumber="167">Nephtys hombergii</emphasis>
Savigny
<emphasis id="B9553010FF9D894C1B6AF8BAEE81F8FA" box="[383,405,1880,1903]" italics="true" pageId="16" pageNumber="167">in</emphasis>
Lamarck, 1818
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1870F8BAECF3F8E5" authority="Sars, 1835" authorityName="Sars" authorityYear="1835" box="[613,999,1879,1904]" class="Polychaeta" family="Onuphidae" genus="Nothria" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="conchylega">
<emphasis id="B9553010FF9D894C1870F8BAEC5EF8E5" box="[613,842,1880,1904]" italics="true" pageId="16" pageNumber="167">Nothria conchylega</emphasis>
(Sars, 1835)
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C19ECF8BAEE57F801" authority="Audouin &amp; Milne Edwards, 1833" authorityName="Audouin &amp; Milne Edwards" authorityYear="1833" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9553010FF9D894C19ECF8BAEBAEF8FA" box="[1017,1210,1880,1904]" italics="true" pageId="16" pageNumber="167">Onuphis eremita</emphasis>
Audouin &amp; Milne Edwards, 1833
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C1B45F89EEC15F801" authority="Delle Chiaje, 1844" authorityName="Delle Chiaje" authorityYear="1844" box="[336,769,1916,1941]" class="Polychaeta" family="Oweniidae" genus="Owenia" kingdom="Animalia" order="Sabellida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="fusiformis">
<emphasis id="B9553010FF9D894C1B45F89EED34F801" box="[336,544,1916,1940]" italics="true" pageId="16" pageNumber="167">Owenia fusiformis</emphasis>
Delle Chiaje, 1844
</taxonomicName>
,
<taxonomicName id="4C219781FF9D894C191BF89EEFC6F82D" authority="Audouin &amp; Milne Edwards in Cuvier, 1830" authorityName="Audouin &amp; Milne Edwards in Cuvier" authorityYear="1830" class="Polychaeta" family="Sigalionidae" genus="Sigalion" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="mathildae">
<emphasis id="B9553010FF9D894C191BF89EECF0F801" box="[782,996,1916,1940]" italics="true" pageId="16" pageNumber="167">Sigalion mathildae</emphasis>
Audouin &amp; Milne Edwards
<emphasis id="B9553010FF9D894C1F3EF89FEA55F801" box="[1323,1345,1917,1940]" italics="true" pageId="16" pageNumber="167">in</emphasis>
Cuvier, 1830
</taxonomicName>
, and
<taxonomicName id="4C219781FF9D894C1B09F842EDBBF82C" authority="Ehlers, 1864" authorityName="Ehlers" authorityYear="1864" box="[284,687,1952,1977]" class="Polychaeta" family="Sigalionidae" genus="Sthenelais" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="limicola">
<emphasis id="B9553010FF9D894C1B09F842EEEDF82D" box="[284,505,1952,1976]" italics="true" pageId="16" pageNumber="167">Sthenelais limicola</emphasis>
(Ehlers, 1864)
</taxonomicName>
(
<bibRefCitation id="EFB091F3FF9D894C18D0F842EC48F82D" author="Dorjes" box="[709,860,1952,1977]" pageId="16" pageNumber="167" refString="Dorjes, J. (1971) Der Golf von Gaeta (Tyrrhenisches Meer). IV. Das Makrobenthos und seine kustenparallele Zonierung. Senckenbergiana maritima, 3, 203 - 246." type="journal article" year="1971">Dörjes 1971</bibRefCitation>
;
<bibRefCitation id="EFB091F3FF9D894C197AF842EB1CF82C" author="Vieitez" box="[879,1032,1952,1977]" pageId="16" pageNumber="167" refString="Vieitez, J. M. (1976) Ecologia de Poliquetos y moluscos de la playa de Meira (Ria de Vigo). I. Estudio de las comunidades. Investigacion Pesquera, 40 (1), 223 - 248." type="journal article" year="1976">Viéitez 1976</bibRefCitation>
,
<bibRefCitation id="EFB091F3FF9D894C1E0EF842EB42F82D" author="Vieitez" box="[1051,1110,1952,1976]" pageId="16" pageNumber="167" refString="Vieitez, J. M. (1981) Estudio de las comunidades bentonicas de dos playas de las rias de Pontevedra y Vigo (Galicia, Espana). Boletin. Instituto Espanol de Oceanografia, 6 (331), 242 - 258." type="journal article" year="1981">1981</bibRefCitation>
;
<collectingRegion id="49E522E0FF9D894C1E7FF842EBA2F82D" box="[1130,1206,1952,1976]" country="France" name="Centre" pageId="16" pageNumber="167">Centro</collectingRegion>
de Investigaciones Submarinas, unpublished study).
</paragraph>
</subSubSection>
<subSubSection id="C33BBF89FF9C894D1AD2FF75ECC6FE1D" pageId="17" pageNumber="168" type="description">
<paragraph id="8B9EEC02FF9C894D1AD2FF75EFF5FE88" blockId="17.[151,1437,151,392]" pageId="17" pageNumber="168">
<emphasis id="B9553010FF9C894D1AD2FF75EE67FF25" bold="true" box="[199,371,151,176]" pageId="17" pageNumber="168">Reproduction.</emphasis>
Oocytes were observed in the coelomic cavity from chaetiger 40 to the end of fragments. The eggs are ellipsoid, flattened disks, 200230 µm × 150170 µm in diameter. They have a thick papillated vitelline membrane, 2731 peripheral vesicles, opaque yolky cytoplasm, and a centrally positioned hyaline nucleus (
<figureCitation id="131AF087FF9C894D1F7AFF3DEFB1FE88" captionStart="FIGURE 6" captionStartId="14.[151,250,1821,1843]" captionTargetBox="[260,1340,214,1782]" captionTargetId="figure@14.[240,1341,193,1794]" captionTargetPageId="14" captionText="FIGURE 6. Scolelepis (Scolelepis) neglecta sp. nov., A, B, LM micrographs of formaldehyde-fixed specimens stained with a solution of methyl green in alcohol, H, I, LM micrographs. A, C G, holotype (MNCN 3852); B, paratype (MNCN 3860); H, I, paratype (MNCN 12568). A, anterior end, dorsal view; B, anterior end, lateral view; C, capillary notochaeta from posterior row of chaetiger 9; D, capillary neurochaeta from anterior row of chaetiger 13; E, ventral inferior capillary from chaetiger 33. F, neuropodial hooded hook from chaetiger 50, lateral view; G, neuropodial hooded hook from chaetiger 59, front-on view; H, egg envelope ornamentation; I, coelomic egg with thick papillate membrane and peripheral vesicles. Scale bars: A B = 1 mm; C I = 0.1 mm." httpUri="https://zenodo.org/record/263915/files/figure.png" pageId="17" pageNumber="168">Fig. 6</figureCitation>
HI).
</paragraph>
<paragraph id="8B9EEC02FF9C894D1AD2FEC5ECC6FE1D" blockId="17.[151,1437,151,392]" pageId="17" pageNumber="168">
<emphasis id="B9553010FF9C894D1AD2FEC5EE4DFED5" bold="true" box="[199,345,295,320]" pageId="17" pageNumber="168">Occurrence.</emphasis>
<taxonomicName id="4C219781FF9C894D1B75FECAED73FED5" box="[352,615,296,320]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="neglecta" status="sp. nov." subGenus="Scolelepis">
<emphasis id="B9553010FF9C894D1B75FECAEEC6FED5" box="[352,466,296,320]" italics="true" pageId="17" pageNumber="168">Scolelepis</emphasis>
(
<emphasis id="B9553010FF9C894D1BF7FECAEEE3FEAA" box="[482,503,296,319]" italics="true" pageId="17" pageNumber="168">S.</emphasis>
)
<emphasis id="B9553010FF9C894D181DFECAED73FED5" box="[520,615,296,320]" italics="true" pageId="17" pageNumber="168">neglecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2668D6BFF9C894D187BFEC5EDD3FED5" box="[622,711,295,320]" pageId="17" pageNumber="168" rank="species">
<emphasis id="B9553010FF9C894D187BFEC5EDD3FED5" bold="true" box="[622,711,295,320]" pageId="17" pageNumber="168">sp. nov.</emphasis>
</taxonomicNameLabel>
is known so far only from the Cantabrian and Galician coasts of
<collectingCountry id="F336AC92FF9C894D1A82FEAEEFC3FEF0" box="[151,215,332,357]" name="Spain" pageId="17" pageNumber="168">Spain</collectingCountry>
(
<bibRefCitation id="EFB091F3FF9C894D1AF0FEAEEE63FEF0" author="Vieitez" box="[229,375,332,357]" pageId="17" pageNumber="168" refString="Vieitez, J. M. (1976) Ecologia de Poliquetos y moluscos de la playa de Meira (Ria de Vigo). I. Estudio de las comunidades. Investigacion Pesquera, 40 (1), 223 - 248." type="journal article" year="1976">Viéitez 1976</bibRefCitation>
,
<bibRefCitation id="EFB091F3FF9C894D1B97FEAEEEA8FEF1" author="Vieitez" box="[386,444,332,356]" pageId="17" pageNumber="168" refString="Vieitez, J. M. (1981) Estudio de las comunidades bentonicas de dos playas de las rias de Pontevedra y Vigo (Galicia, Espana). Boletin. Instituto Espanol de Oceanografia, 6 (331), 242 - 258." type="journal article" year="1981">1981</bibRefCitation>
, as
<taxonomicName id="4C219781FF9C894D1BF0FEAEEDD6FEF1" box="[485,706,332,356]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="cantabra">
<emphasis id="B9553010FF9C894D1BF0FEAEEDD6FEF1" box="[485,706,332,356]" italics="true" pageId="17" pageNumber="168">Scolelepis cantabra</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB091F3FF9C894D18C4FEAFEC73FEF1" author="Parapar" box="[721,871,332,356]" pageId="17" pageNumber="168" refString="Parapar, J. (1991) Anelidos Poliquetos bentonicos de la ria de Ferrol (Galicia). PhD-thesis, University of Santiago de Compostela, 1104 pp." type="book" year="1991">Parapar 1991</bibRefCitation>
, as
<taxonomicName id="4C219781FF9C894D1984FEAEEB62FEF1" box="[913,1142,332,356]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF9C894D1984FEAEEB62FEF1" box="[913,1142,332,356]" italics="true" pageId="17" pageNumber="168">Scolelepis squamata</emphasis>
</taxonomicName>
), from the Tyrrhenian Sea (
<bibRefCitation id="EFB091F3FF9C894D1A8BFE8DED28FE1D" author="Dorjes" box="[158,572,367,392]" pageId="17" pageNumber="168" refString="Dorjes, J. (1971) Der Golf von Gaeta (Tyrrhenisches Meer). IV. Das Makrobenthos und seine kustenparallele Zonierung. Senckenbergiana maritima, 3, 203 - 246." type="journal article" year="1971">
Dörjes 1971, as
<taxonomicName id="4C219781FF9C894D1B43FE92ED28FE1D" box="[342,572,368,392]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B9553010FF9C894D1B43FE92ED28FE1D" box="[342,572,368,392]" italics="true" pageId="17" pageNumber="168">Scolelepis squamata</emphasis>
</taxonomicName>
</bibRefCitation>
), and the Black Sea (present study).
</paragraph>
</subSubSection>
</treatment>
</document>