160 lines
18 KiB
XML
160 lines
18 KiB
XML
<document id="F3BBE107AF15E95F11ECFB32A78565D4" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia & Kessler, Michael" docDate="2011" docId="0380F37CFFEF7F0699A6E63ED7E3F833" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus retroflexus Copeland 1947" docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="43" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="90049C53E01C952272450B439BA8F7FC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5F442611D76C8FE3A3F204567536F294">
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<mods:title id="E0C26CF5EB6A20A845642845398CAB42">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
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</mods:titleInfo>
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<mods:name id="F06413F56185C5E57FB079F6184FF4E0" type="personal">
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<mods:role id="65965DA07D417EC8A69A34587B0DC415">
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<mods:roleTerm id="2EC42F69A1FC4BC693F5DDC3452B1AD4">Author</mods:roleTerm>
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<mods:namePart id="75BD646CF5EBBECD8548E563FE27063A">Gonzales, Jasivia</mods:namePart>
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<mods:affiliation id="D8815557A5CA6B9B75D90A91255FB67E">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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</mods:name>
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<mods:name id="E945B19C8234C91F4A8B2841DBD23CE5" type="personal">
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<mods:roleTerm id="9CCE8387939A5A0FF88F1120C24F49E7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A1B39E2D5E42F4D4F4376326BFC7D384">Kessler, Michael</mods:namePart>
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<mods:typeOfResource id="A17C04A001D08D690557D43A7A068DD7">text</mods:typeOfResource>
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<mods:title id="2A78C0D52B7BBEB14D812214329FA8E0">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A6819E04E4DF6AD0FF41B16DB0E31ED3">
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<mods:date id="D0B5188350ED3ED8F92F0061E71BC02D">2011</mods:date>
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<mods:number id="82FBE3092B0987F1F2B6A61DB57BC18C">2011-12-31</mods:number>
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<mods:detail id="7A7D9869EA30ED79F03EBF5822841A71" type="volume">
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<mods:number id="34856A85B0AC170FFFF57C9FF3208618">31</mods:number>
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<mods:start id="62D820E62B6560C43B909F85C3E8B17D">1</mods:start>
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<mods:classification id="C4E79248787C9312F00FA45EF4AE1C16">journal article</mods:classification>
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<mods:identifier id="6CE95BC7136D662E292597F9F88B77DC" type="CLB-Dataset">6085</mods:identifier>
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<mods:identifier id="4BEAD8EAD4C4EB9CABE8DA33B9E4519B" type="DOI">10.11646/phytotaxa.31.1.1</mods:identifier>
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<treatment id="0380F37CFFEF7F0699A6E63ED7E3F833" ID-DOI="http://doi.org/10.5281/zenodo.4894790" ID-GBIF-Taxon="182742280" ID-Zenodo-Dep="4894790" LSID="urn:lsid:plazi:treatment:0380F37CFFEF7F0699A6E63ED7E3F833" httpUri="http://treatment.plazi.org/id/0380F37CFFEF7F0699A6E63ED7E3F833" lastPageNumber="43" pageId="42" pageNumber="43">
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<subSubSection id="C33311E1FFEF7F0699A6E63ED496FAF1" box="[151,900,1345,1371]" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="8B96426AFFEF7F0699A6E63ED496FAF1" blockId="42.[151,900,1345,1371]" box="[151,900,1345,1371]" pageId="42" pageNumber="43">
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<heading id="D0DEF506FFEF7F0699A6E63ED496FAF1" box="[151,900,1345,1371]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="3">
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<treatmentCitationGroup id="AB396544FFEF7F0699A6E63ED496FAF1" box="[151,900,1345,1371]" pageId="42" pageNumber="43">
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<treatmentCitation id="0A88647BFFEF7F0699A6E63ED496FAF1" author="Copeland, E. B." box="[151,900,1345,1371]" page=")" pageId="42" pageNumber="43" year="1947">
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<taxonomicName id="4C2939E9FFEF7F0699A6E63ED496FAF1" ID-CoL="52HXL" authority="(J. Bommer ex Christ) Copeland (1947: 28)" authorityName="Copeland" authorityPageNumber="28" authorityYear="1947" baseAuthorityName="J. Bommer ex Christ" box="[151,900,1345,1371]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="retroflexus">
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<emphasis id="B95D9E78FFEF7F0699A6E63ED69DFAF1" bold="true" box="[151,399,1345,1371]" italics="true" pageId="42" pageNumber="43">Sticherus retroflexus</emphasis>
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(J.Bommer ex Christ)
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<bibRefCitation id="EFB83F9BFFEF7F069BA6E63ED496FAF1" author="Copeland, E. B." box="[663,900,1345,1371]" pageId="42" pageNumber="43" pagination="1 - 247" refId="ref30844" refString="Copeland, E. B. (1947) Genera filicum. The genera of ferns. Annales Cryptogamici et Phytopathologici 5: 1 - 247." type="journal article" year="1947">Copeland (1947: 28)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33311E1FFEF7F0699A6E6F0D24AFA0F" box="[151,1368,1422,1446]" pageId="42" pageNumber="43" type="reference_group">
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<paragraph id="8B96426AFFEF7F0699A6E6F0D24AFA0F" blockId="42.[151,1438,1422,1945]" box="[151,1368,1422,1446]" pageId="42" pageNumber="43">
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<treatmentCitationGroup id="AB396544FFEF7F0699A6E6F0D24AFA0F" box="[151,1368,1422,1446]" pageId="42" pageNumber="43">
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<treatmentCitation id="0A88647BFFEF7F0699A6E6F0D40BFA0F" author="Baker, J." box="[151,793,1422,1446]" page=")" pageId="42" pageNumber="43" year="1887">
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<taxonomicName id="4C2939E9FFEF7F0699A6E6F0D40BFA0F" ID-CoL="3G76L" authority="Baker (1887: 24)" authorityName="Baker" authorityPageNumber="24" authorityYear="1887" box="[151,793,1422,1446]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="variety" species="intermedia" variety="flexuosa">
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<emphasis id="B95D9E78FFEF7F0699A6E6F0D694FA0C" box="[151,390,1423,1446]" italics="true" pageId="42" pageNumber="43">Gleichenia intermedia</emphasis>
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Baker var.
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<emphasis id="B95D9E78FFEF7F069B30E6F0D548FA0C" box="[513,602,1423,1446]" italics="true" pageId="42" pageNumber="43">flexuosa</emphasis>
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<bibRefCitation id="EFB83F9BFFEF7F069B51E6F1D40BFA0F" author="Baker, J." box="[608,793,1422,1445]" pageId="42" pageNumber="43" pagination="24 - 26" refId="ref30363" refString="Baker, J. (1887) Mr. J. J. Cooper's Costa Rica ferns. Journal of Botany British and Foreign 25: 24 - 26." type="journal article" year="1887">Baker (1887: 24)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="0A88647BFFEF7F069A15E6F0D246FA0F" author="Bommer, J. - E. & Christ H." box="[804,1364,1422,1446]" page=")" pageId="42" pageNumber="43" year="1896">
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<taxonomicName id="4C2939E9FFEF7F069A15E6F0D246FA0F" ID-CoL="6KH3M" authority="Bommer & Christ (1896: 657)" authorityName="Bommer & Christ" authorityPageNumber="657" authorityYear="1896" box="[804,1364,1422,1446]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="retroflexa">
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<emphasis id="B95D9E78FFEF7F069A15E6F0D314FA0C" box="[804,1030,1423,1446]" italics="true" pageId="42" pageNumber="43">Gleichenia retroflexa</emphasis>
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<bibRefCitation id="EFB83F9BFFEF7F069D3CE6F1D246FA0F" author="Bommer, J. - E. & Christ H." box="[1037,1364,1422,1445]" pageId="42" pageNumber="43" pagination="657 - 663" refId="ref30424" refString="Bommer, J. - E. & Christ H. (1896) Filices novae. Bulletin de l'Herbier Boissier 10: 657 - 663." type="journal article" year="1896">Bommer & Christ (1896: 657)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33311E1FFEF7F0699F7E6D1D4B4FA63" box="[198,934,1454,1481]" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="8B96426AFFEF7F0699F7E6D1D4B4FA63" blockId="42.[151,1438,1422,1945]" box="[198,934,1454,1481]" pageId="42" pageNumber="43">
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<emphasis id="B95D9E78FFEF7F0699F7E6D1D639FA62" bold="true" box="[198,299,1454,1480]" pageId="42" pageNumber="43">Type:—</emphasis>
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<materialsCitation id="3B414837FFEF7F06981AE6D0D4B1FA63" ID-GBIF-Occurrence="3131398350" box="[299,931,1455,1481]" collectionCode="BR" collectorName="La Palma" country="Costa Rica" location="COSTA RICA" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="F33E02FAFFEF7F06981AE6D0D6C8FA63" box="[299,474,1455,1481]" name="Costa Rica" pageId="42" pageNumber="43">COSTA RICA</collectingCountry>
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.
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<collectorName id="26DC27BCFFEF7F0698D5E6D0D545FA63" box="[484,599,1455,1481]" pageId="42" pageNumber="43">La Palma</collectorName>
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,
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<emphasis id="B95D9E78FFEF7F069B53E6D0D5FDFA62" box="[610,751,1455,1480]" italics="true" pageId="42" pageNumber="43">Pittier 1471</emphasis>
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(
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<typeStatus id="5492FCC8FFEF7F069BCFE6D0D474FA63" box="[766,870,1455,1481]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
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<collectionCode id="ED38DAAFFFEF7F069A5FE6D0D488FA63" box="[878,922,1455,1481]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="42" pageNumber="43">BR</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33311E1FFEF7F0699F7E6AAD7E3F833" pageId="42" pageNumber="43" type="distribution">
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<paragraph id="8B96426AFFEF7F0699F7E6AAD50FF996" blockId="42.[151,1438,1422,1945]" pageId="42" pageNumber="43">
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<emphasis id="B95D9E78FFEF7F0699F7E6AAD534FA45" bold="true" box="[198,550,1493,1519]" pageId="42" pageNumber="43">Distribution and ecology:—</emphasis>
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<collectingCountry id="F33E02FAFFEF7F069B14E6AAD5B2FA45" box="[549,672,1493,1519]" name="Nicaragua" pageId="42" pageNumber="43">Nicaragua</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEF7F069B9EE6AAD426FA45" box="[687,820,1493,1519]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
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, and western
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<collectingCountry id="F33E02FAFFEF7F069AEBE6AAD329FA45" box="[986,1083,1493,1519]" name="Panama" pageId="42" pageNumber="43">Panama</collectingCountry>
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. On steep slopes along forest margins, landslides, and roadbanks, forming thickets in open disturbed sites, often mixed with other species of
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<taxonomicName id="4C2939E9FFEF7F0699A6E55DD65FF996" box="[151,333,1570,1596]" class="Polypodiopsida" family="Gleicheniaceae" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="family">Gleicheniaceae</taxonomicName>
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, at
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<quantity id="4CD1EF8FFFEF7F069844E55DD50BF996" box="[373,537,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.8" metricValueMin="1.1" pageId="42" pageNumber="43" unit="m" value="1950.0" valueMax="2800.0" valueMin="1100.0">1100–2800 m</quantity>
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.
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</paragraph>
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<paragraph id="8B96426AFFEF7F0699F7E536D7E3F833" blockId="42.[151,1438,1422,1945]" pageId="42" pageNumber="43">
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<emphasis id="B95D9E78FFEF7F0699F7E536D621F9C9" bold="true" box="[198,307,1609,1635]" pageId="42" pageNumber="43">Notes:—</emphasis>
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This species belongs to a group of four closely related species, which also include
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<taxonomicName id="4C2939E9FFEF7F069DCFE536D286F9C8" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Rosenst." box="[1278,1428,1609,1634]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="hastulatus">
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<emphasis id="B95D9E78FFEF7F069DCFE536D286F9C8" box="[1278,1428,1609,1634]" italics="true" pageId="42" pageNumber="43">S. hastulatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2939E9FFEF7F0699A6E50FD657F923" authorityName="Chrysler" authorityYear="1944" baseAuthorityName="Christ" box="[151,325,1648,1673]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="orthocladus">
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<emphasis id="B95D9E78FFEF7F0699A6E50FD657F923" box="[151,325,1648,1673]" italics="true" pageId="42" pageNumber="43">S. orthocladus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C2939E9FFEF7F0698BBE50FD525F923" authorityName="Chrysler" authorityYear="1944" baseAuthorityName="Baker" box="[394,567,1648,1673]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="intermedius">
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<emphasis id="B95D9E78FFEF7F0698BBE50FD525F923" box="[394,567,1648,1673]" italics="true" pageId="42" pageNumber="43">S. intermedius</emphasis>
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</taxonomicName>
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. They all have scales restricted to the buds and rhizomes, hastulate, (in some species) short-petiolate segments, accessory segments on the pinna bases, 2- or 3-forked veinlets on the auriculate segment bases, obscure veinlets, abaxial blade papillae in most specimens, slightly to strongly undulate segment margins, and ovate, entire, translucent to slightly matte scales. All four species co-occur in
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<collectingCountry id="F33E02FAFFEF7F0699A6E475D605F88E" box="[151,279,1802,1828]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
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and often
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<taxonomicName id="4C2939E9FFEF7F0698A2E475D50FF88E" box="[403,541,1802,1828]" form="mixed" pageId="42" pageNumber="43" rank="form">form mixed</taxonomicName>
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populations. Morphologically they grade into each other, with
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<taxonomicName id="4C2939E9FFEF7F069C33E474D289F88E" authorityName="Copeland" authorityYear="1947" baseAuthorityName="J. Bommer ex Christ" box="[1282,1435,1803,1828]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="retroflexus">
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<emphasis id="B95D9E78FFEF7F069C33E474D289F88E" box="[1282,1435,1803,1828]" italics="true" pageId="42" pageNumber="43">S. retroflexus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2939E9FFEF7F0699F8E44ED663F8E0" authorityName="Chrysler" authorityYear="1944" baseAuthorityName="Baker" box="[201,369,1841,1866]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="intermedius">
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<emphasis id="B95D9E78FFEF7F0699F8E44ED663F8E0" box="[201,369,1841,1866]" italics="true" pageId="42" pageNumber="43">S. intermedius</emphasis>
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</taxonomicName>
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as the most distinct forms and
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<taxonomicName id="4C2939E9FFEF7F069BD1E44ED464F8E0" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Rosenst." box="[736,886,1841,1866]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="hastulatus">
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<emphasis id="B95D9E78FFEF7F069BD1E44ED464F8E0" box="[736,886,1841,1866]" italics="true" pageId="42" pageNumber="43">S. hastulatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2939E9FFEF7F069A81E44ED348F8E0" authorityName="Chrysler" authorityYear="1944" baseAuthorityName="Christ" box="[944,1114,1841,1866]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="orthocladus">
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<emphasis id="B95D9E78FFEF7F069A81E44ED348F8E0" box="[944,1114,1841,1866]" italics="true" pageId="42" pageNumber="43">S. orthocladus</emphasis>
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</taxonomicName>
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as intermediate forms. It is conceivable that the latter are hybrids between
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<taxonomicName id="4C2939E9FFEF7F069BF1E427D449F8DB" authorityName="Copeland" authorityYear="1947" baseAuthorityName="J. Bommer ex Christ" box="[704,859,1880,1905]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="retroflexus">
|
||
<emphasis id="B95D9E78FFEF7F069BF1E427D449F8DB" box="[704,859,1880,1905]" italics="true" pageId="42" pageNumber="43">S. retroflexus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2939E9FFEF7F069AA5E427D329F8DB" authorityName="Chrysler" authorityYear="1944" baseAuthorityName="Baker" box="[916,1083,1880,1905]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="intermedius">
|
||
<emphasis id="B95D9E78FFEF7F069AA5E427D329F8DB" box="[916,1083,1880,1905]" italics="true" pageId="42" pageNumber="43">S. intermedius</emphasis>
|
||
</taxonomicName>
|
||
, but more detailed studies are needed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |