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<document id="00629AB069E7860410FC05269D1170B9" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia &amp; Kessler, Michael" docDate="2011" docId="0380F37CFFE37F0A99A6E1F1D452FAA5" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus nudus Nakai 1950" docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="39" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B6E16D3509691CCCCA71B00552BA25AA">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
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<mods:namePart id="A14E58B6DD217C35626DC36AB8BD9930">Gonzales, Jasivia</mods:namePart>
<mods:affiliation id="05E69989DF725E508BCBDAC8E5F732CC">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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<mods:namePart id="1CC80E4D005A6B415645927E8743B6EB">Kessler, Michael</mods:namePart>
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<mods:title id="439C7CE76D39C2C42E659D83BFF8471B">Phytotaxa</mods:title>
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<mods:date id="43F97FAC68FEDA4E5B89ADD3E3FEEFF3">2011</mods:date>
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<treatmentCitation id="0A88647BFFE37F0A99A6E1F1D586FD02" author="Nakai, T." box="[151,660,654,680]" page=")" pageId="38" pageNumber="39" year="1950">
<taxonomicName id="4C2939E9FFE37F0A99A6E1F1D586FD02" ID-CoL="52HWT" authority="(Moritz) Nakai (1950: 23)" authorityName="Nakai" authorityPageNumber="23" authorityYear="1950" baseAuthorityName="Moritz" box="[151,660,654,680]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="nudus">
<emphasis id="B95D9E78FFE37F0A99A6E1F1D648FD02" bold="true" box="[151,346,654,680]" italics="true" pageId="38" pageNumber="39">Sticherus nudus</emphasis>
(Moritz)
<bibRefCitation id="EFB83F9BFFE37F0A98FDE1F1D586FD02" author="Nakai, T." box="[460,660,654,680]" pageId="38" pageNumber="39" pagination="1 - 71" refId="ref32966" refString="Nakai, T. (1950) A new classification of Gleicheniales. Bulletin of the National Science Museum 29: 1 - 71." type="journal article" year="1950">Nakai (1950: 23)</bibRefCitation>
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<treatmentCitationGroup id="AB396544FFE37F0A99A6E1A3D410FD59" box="[151,770,732,755]" pageId="38" pageNumber="39">
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<emphasis id="B95D9E78FFE37F0A99A6E1A3D62DFD59" box="[151,319,732,755]" italics="true" pageId="38" pageNumber="39">Mertensia nuda</emphasis>
Moritz
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(in
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<bibRefCitation id="EFB83F9BFFE37F0A9888E1A3D595FD59" author="Reichardt, H. W." box="[441,647,732,755]" pageId="38" pageNumber="39" pagination="21 - 48" refId="ref33475" refString="Reichardt, H. W. (1859) Uber die Gefassbundel-Vertheilung im Stamme und Stipes der Farne. Ein Beitrag zur anatomischen und systematischen Kenntniss dieser Familie. Akademie der Wissenschaften in Wien, Mathematisch- Naturwissenschaftliche Klasse, Denkschriften 17: 21 - 48, plates 1 - 3." type="journal article" year="1859">Reichardt 1859: 28</bibRefCitation>
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, t 1, f 14).
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<emphasis id="B95D9E78FFE37F0A99F7E184D639FCBF" bold="true" box="[198,299,763,789]" pageId="38" pageNumber="39">Type:—</emphasis>
<materialsCitation id="3B414837FFE37F0A981AE183D4D9FCBC" ID-GBIF-Occurrence="3131398390" box="[299,971,764,790]" collectionCode="W" country="Venezuela" location="Caracas" municipality="Moritz" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F33E02FAFFE37F0A981AE183D6C8FCBC" box="[299,474,764,790]" name="Venezuela" pageId="38" pageNumber="39">VENEZUELA</collectingCountry>
.
<location id="8EF614B1FFE37F0A98D5E183D556FCBC" LSID="urn:lsid:plazi:treatment:0380F37CFFE37F0A99A6E1F1D452FAA5:8EF614B1FFE37F0A98D5E183D556FCBC" box="[484,580,764,790]" country="Venezuela" municipality="Moritz" name="Caracas" pageId="38" pageNumber="39">Caracas</location>
, 1856,
<emphasis id="B95D9E78FFE37F0A9BA8E183D408FCBF" box="[665,794,764,789]" italics="true" pageId="38" pageNumber="39">
<collectingMunicipality id="6BF2D810FFE37F0A9BA8E183D5F5FCBF" box="[665,743,764,789]" pageId="38" pageNumber="39">Moritz</collectingMunicipality>
452
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(
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!)
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.
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<subSubSection id="C33311E1FFE37F0A99F7E05DD452FAA5" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8B96426AFFE37F0A99F7E05DD556FCC9" blockId="38.[151,1437,732,1295]" pageId="38" pageNumber="39">
<emphasis id="B95D9E78FFE37F0A99F7E05DD530FC96" bold="true" box="[198,546,802,828]" pageId="38" pageNumber="39">Distribution and ecology:—</emphasis>
Andes in
<collectingCountry id="F33E02FAFFE37F0A9BA2E05DD414FC96" box="[659,774,802,828]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
and
<collectingCountry id="F33E02FAFFE37F0A9A71E05DD4AEFC96" box="[832,956,802,828]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
. It forms dense colonies in open habitats in montane forests, at
<quantity id="4CD1EF8FFFE37F0A98ABE036D52DFCC9" box="[410,575,841,867]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.7" metricValueMin="1.6" pageId="38" pageNumber="39" unit="m" value="2150.0" valueMax="2700.0" valueMin="1600.0">16002700 m</quantity>
.
</paragraph>
<paragraph id="8B96426AFFE37F0A99F7E010D452FAA5" blockId="38.[151,1437,732,1295]" pageId="38" pageNumber="39">
<emphasis id="B95D9E78FFE37F0A99F7E010D621FC23" bold="true" box="[198,307,879,905]" pageId="38" pageNumber="39">Notes:—</emphasis>
<taxonomicName id="4C2939E9FFE37F0A9802E00FD6E0FC23" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Moritz" box="[307,498,880,905]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="nudus">
<emphasis id="B95D9E78FFE37F0A9802E00FD6E0FC23" box="[307,498,880,905]" italics="true" pageId="38" pageNumber="39">Sticherus nudus</emphasis>
</taxonomicName>
belongs to a group of four confusingly similar Mesoamerican and Andean species, including
<taxonomicName id="4C2939E9FFE37F0A984DE0E8D571FC1A" authorityName="Ostergaard &amp; Ollgaard" authorityYear="2001" box="[380,611,919,944]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="brevitomentosus">
<emphasis id="B95D9E78FFE37F0A984DE0E8D571FC1A" box="[380,611,919,944]" italics="true" pageId="38" pageNumber="39">S. brevitomentosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2939E9FFE37F0A9B45E0E8D5F1FC1A" authorityName="J. Gonzales" authorityYear="2011" box="[628,739,919,944]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="B95D9E78FFE37F0A9B45E0E8D5F1FC1A" box="[628,739,919,944]" italics="true" pageId="38" pageNumber="39">S. ovatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2939E9FFE37F0A9A18E0E8D4D9FC1A" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Christ" box="[809,971,919,944]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="pteridellus">
<emphasis id="B95D9E78FFE37F0A9A18E0E8D4D9FC1A" box="[809,971,919,944]" italics="true" pageId="38" pageNumber="39">S. pteridellus</emphasis>
</taxonomicName>
. These all have ovate segments, nonpectinate proximal branches, approximate, usually flat segments, generally rather sparsely scaly branches, ovate-lanceolate to ovate scales that appear papyraceous and dull whitish to brown when dry, and finally, they also lack aphlebiae. Among these species,
<taxonomicName id="4C2939E9FFE37F0A9BB6E775D5FBFB89" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Moritz" box="[647,745,1034,1059]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="nudus">
<emphasis id="B95D9E78FFE37F0A9BB6E775D5FBFB89" box="[647,745,1034,1059]" italics="true" pageId="38" pageNumber="39">S. nudus</emphasis>
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is characterized by abaxially protruding ultimate branches, 1
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branches longer than the 2
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ones, arched branches, petioles and rachises much thicker than the pinna branches, thick rhizomes with scattered, rigid scales, cartilaginous, undulate segment margins, abaxially glabrous segments with raised, parallel veinlets, sparse, reduced, mixed ovate and linear scales restricted to the midveins and sometimes around the inframedial sori, and ovate to rounded branch and bud scales with usually glandular apices. The arched branches are reminiscent of
<taxonomicName id="4C2939E9FFE37F0A9A94E7B0D343FB42" authorityName="Ostergaard &amp; Ollgaard" authorityYear="2001" box="[933,1105,1231,1256]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="aurantiacus">
<emphasis id="B95D9E78FFE37F0A9A94E7B0D343FB42" box="[933,1105,1231,1256]" italics="true" pageId="38" pageNumber="39">S. aurantiacus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2939E9FFE37F0A9DBDE7B0D233FB42" authorityName="Smith" authorityYear="1990" box="[1164,1313,1231,1256]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="tepuiensis">
<emphasis id="B95D9E78FFE37F0A9DBDE7B0D233FB42" box="[1164,1313,1231,1256]" italics="true" pageId="38" pageNumber="39">S. tepuiensis</emphasis>
</taxonomicName>
, but these species are densely scaly, with narrowly lanceolate scales.
</paragraph>
</subSubSection>
</treatment>
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