622 lines
61 KiB
XML
622 lines
61 KiB
XML
<document ID-CLB-Dataset="3828" ID-DOI="http://dx.doi.org/10.3897/zookeys.1061.70420" ID-GBIF-Dataset="74154259-34a6-4153-93ce-447ac1a90a27" ID-PMC="PMC8505391" ID-Pensoft-Pub="1313-2970-1061-87" ID-Pensoft-UUID="AF38CEA80126576FA4C341BB14489336" ID-PubMed="34707454" ID-ZooBank="F2701BACD2E14F97A0BDDC8B09966247" ModsDocID="1313-2970-1061-87" checkinTime="1633431410146" checkinUser="pensoft" docAuthor="Shi, Jing-Song, Liu, Jin-Cheng, Giri, Rohit, Owens, John Benjamin, Santra, Vishal, Kuttalam, Sourish, Selvan, Melvin, Guo, Ke-Ji & Malhotra, Anita" docDate="2021" docId="01E6E10AADF759CAA589BD6DFEDF3CAA" docLanguage="en" docName="ZooKeys 1061: 87-108" docOrigin="ZooKeys 1061" docPubDate="2021-10-04" docSource="http://dx.doi.org/10.3897/zookeys.1061.70420" docTitle="Gloydius lipipengi Shi, Liu & Malhotra 2021, sp. nov." docType="treatment" docUuid="6DF30D06-937B-470B-AFE4-D4CABEAF7DAB" docUuidSource="ZooBank" docVersion="6" id="AF38CEA80126576FA4C341BB14489336" lastPageNumber="87" masterDocId="AF38CEA80126576FA4C341BB14489336" masterDocTitle="Molecular phylogenetic analysis of the genus Gloydius (Squamata, Viperidae, Crotalinae), with description of two new alpine species from Qinghai-Tibet Plateau, China" masterLastPageNumber="108" masterPageNumber="87" pageNumber="87" updateTime="1732846796274" updateUser="ExternalLinkService">
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<mods:title id="7644E26EFED47806FC8465FAB73C0F93">Molecular phylogenetic analysis of the genus Gloydius (Squamata, Viperidae, Crotalinae), with description of two new alpine species from Qinghai-Tibet Plateau, China</mods:title>
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<mods:namePart id="4B2B96BDA3A6A7B6000A18B72B363EAC">Shi, Jing-Song</mods:namePart>
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<mods:affiliation id="5D47E3FEC7D830380EA980D686468CA0">Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China & Institute of Herpetology, Shenyang Normal University, Shenyang 110034, China</mods:affiliation>
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<mods:nameIdentifier id="192974818298B54DD095536D3CAB106C" type="email">shijingsong@ivpp.ac.cn</mods:nameIdentifier>
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<mods:namePart id="6C82CA8A62D022970A3E68D259F3F4B8">Liu, Jin-Cheng</mods:namePart>
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<mods:affiliation id="B9907840A6D253DCE9E9012C866E3E86">Institute of Herpetology, Shenyang Normal University, Shenyang 110034, China</mods:affiliation>
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<mods:namePart id="2FA61838E1F6CA3BA6A23982ED367836">Giri, Rohit</mods:namePart>
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<mods:affiliation id="4376576843741BF85DAC9FE21552DDF8">Department of Zoology, Prithvi Narayan Campus, Bhimkalipatan- 1, Pokhara 33700, Nepal</mods:affiliation>
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</mods:role>
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<mods:namePart id="CDFBC475CF59A1E90B6B7DE92AC319B8">Owens, John Benjamin</mods:namePart>
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<mods:nameIdentifier id="72AEB8C79CA4D877CA5B509092604661" type="ORCID">https://orcid.org/0000-0003-0298-451X</mods:nameIdentifier>
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<mods:affiliation id="D432E46BB4FED05984151583B22B36E2">Molecular Ecology and Evolution at Bangor, School of Natural Sciences, Bangor University, Bangor, Gwynedd LL 57 2 UW, UK & Captive and Field Herpetology, Wales, UK</mods:affiliation>
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</mods:name>
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<mods:namePart id="56E116F838D1D39D9E593DCE3FD8F519">Santra, Vishal</mods:namePart>
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<mods:affiliation id="7C908BE5FD33E624E6291F06EB9FF05C">Captive and Field Herpetology, Wales, UK & Society for Nature Conservation, Research and Community Engagement (CONCERN), Nalikul, Hooghly, West Bengal, India</mods:affiliation>
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<mods:namePart id="02C50015A96D6F62AB66FB3A6776AE20">Kuttalam, Sourish</mods:namePart>
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<mods:nameIdentifier id="690F4C6998C191D381CF11D8E96896C3" type="ORCID">https://orcid.org/0000-0003-1072-1699</mods:nameIdentifier>
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<mods:affiliation id="4162C6E6C7EF4EFD1831BD3B2B054A8D">Captive and Field Herpetology, Wales, UK & Society for Nature Conservation, Research and Community Engagement (CONCERN), Nalikul, Hooghly, West Bengal, India</mods:affiliation>
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<mods:namePart id="97DB2DB91B8F4FE9A15CBA4C85F3E699">Selvan, Melvin</mods:namePart>
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<mods:affiliation id="E818F2631A7F11E3E066CE789BF20AF0">Endangered Wildlife Trust, Dindigull, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="58621B52BC12D88E9D4D504D260CE0EA">Guo, Ke-Ji</mods:namePart>
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<mods:affiliation id="A04EE12B6B5D19806EB9E9C9A56EBEB7">Central South Inventory and Planning Institute of National Forestry and Grassland Administration, College of Life Sciences and Technology, Central South University of Forestry and Technology, Changsha, Hunan 410004, China</mods:affiliation>
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<mods:namePart id="2B3A185B81B6707DE5A92E191A9BE673">Malhotra, Anita</mods:namePart>
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<mods:nameIdentifier id="0CCFEB7F865D3B77DA1566F34F61F3ED" type="ORCID">https://orcid.org/0000-0002-1738-9046</mods:nameIdentifier>
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<mods:affiliation id="310E845E9B7436ED16466C807673FE33">Molecular Ecology and Evolution at Bangor, School of Natural Sciences, Bangor University, Bangor, Gwynedd LL 57 2 UW, UK</mods:affiliation>
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<mods:nameIdentifier id="409EAF8E8EC264249D21CBACAA8E681E" type="email">a.malhotra@bangor.ac.uk</mods:nameIdentifier>
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<mods:typeOfResource id="7AAE5747885DB39C17F938B3DA8B77CC">text</mods:typeOfResource>
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<mods:title id="640D57BAAB2655A3DBD67CEACB6CD2FB">ZooKeys</mods:title>
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<mods:part id="D82EA598D2806ECFE25848FFEF0348A6">
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<mods:date id="C11AA059B0748F0F59568A8F26F6A1DA">2021</mods:date>
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<mods:number id="7C098EE557591BE545475FB7234314A6">2021-10-04</mods:number>
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<mods:number id="E55864056E845F01902788FFC5252495">1061</mods:number>
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<mods:start id="9C038EE0878095F9163A1CC4A7002496">87</mods:start>
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<mods:url id="8931D1B734F8FA6AB81E531AF4ACFD08">http://dx.doi.org/10.3897/zookeys.1061.70420</mods:url>
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<mods:classification id="037F3B368EBCF953AE0A3494C7A64A56">journal article</mods:classification>
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<mods:identifier id="C230F24DF7E423A389EDF5E5DC972EBC" type="DOI">http://dx.doi.org/10.3897/zookeys.1061.70420</mods:identifier>
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<mods:identifier id="B98123216D56545E0980A4821A6B11D8" type="Pensoft-Pub">1313-2970-1061-87</mods:identifier>
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<treatment id="01E6E10AADF759CAA589BD6DFEDF3CAA" ID-GBIF-Taxon="189470255" LSID="urn:lsid:zoobank.org:act:6DF30D06-937B-470B-AFE4-D4CABEAF7DAB" httpUri="http://treatment.plazi.org/id/01E6E10AADF759CAA589BD6DFEDF3CAA" lastPageNumber="87" pageId="0" pageNumber="87" scope_class="Squamata" scope_family="Viperidae">
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<subSubSection id="81553C9A9521DAD3AE6621F43AA3E8CC" pageId="0" pageNumber="87" type="nomenclature">
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<paragraph id="F7F6949A399DFE70F7FD5E08544B2B6F" pageId="0" pageNumber="87">
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<taxonomicName id="B8EEA176045D650B97AAAE82F63AC64C" ID-CoL="8SJNQ" LSID="http://zoobank.org/6DF30D06-937B-470B-AFE4-D4CABEAF7DAB" authority="Shi, Liu & Malhotra" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi" status="sp. nov.">Gloydius lipipengi Shi, Liu & Malhotra</taxonomicName>
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||
<taxonomicNameLabel id="C7088AF9AEF818CA5A7E089247402C5F" pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="FBF34E74EC859272AABD46457F6A85A3" pageId="0" pageNumber="87" type="etymology">
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<paragraph id="FD2BA30D58B18C726CAAAE21F051C951" pageId="0" pageNumber="87">Etymology.</paragraph>
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<paragraph id="B6E706ADF13D67495D11B3D96F15B574" pageId="0" pageNumber="87">
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The specific epithet of the new species from Tibet is dedicated to the senior
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<normalizedToken id="25936C92A622C18C1D4868A1BEF47A20" originalValue="author’s">author's</normalizedToken>
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<normalizedToken id="605CA14D4AF86B4C16668234501EF521" originalValue="Master’s">Master's</normalizedToken>
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||
supervisor, Professor Pi-Peng Li (Institute of Herpetology, Shenyang Normal University) on
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||
<normalizedToken id="EB437DA0C07A6B322439CA240DDBF31E" originalValue="Li’s">Li's</normalizedToken>
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sixtieth birthday. Prof. Li has devoted himself to the study of the herpetological diversity of the Qinghai-Tibet Plateau. The senior author became an Asian pit viper enthusiast and professional herpetological researcher under his instruction. The common name of
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<taxonomicName id="5B0BBC474138C26F5D92486044B55323" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
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<emphasis id="3C06D2F845E890DCFCA7491763335892" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
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</taxonomicName>
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||
sp. nov. is suggested as "Nujiang pit viper" in English, and
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||
<normalizedToken id="033BC4FD97164BC722B9A8FFCAAEA062" originalValue="“Nù">"Nu</normalizedToken>
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<normalizedToken id="A0A3F10D64869E67C53D23A04F074DA3" originalValue="Jiāng">Jiang</normalizedToken>
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<normalizedToken id="E9CB961B502A0E528C6898B13F185FA7" originalValue="Fù">Fu</normalizedToken>
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(
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||
<normalizedToken id="2490170E88B254F16D671F2BC803F131" originalValue="怒江蝮)”">怒江蝮)"</normalizedToken>
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in Chinese.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="B061E00E9D9D98A274FD9BFF45D0E975" pageId="0" pageNumber="87" type="type specimen">
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<paragraph id="0ABB1E7A51BB68DF6A7342F0B2AC58DD" pageId="0" pageNumber="87">Type specimen.</paragraph>
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<paragraph id="9C5FAA59487DE2B754CD8F5F7B5FECA9" pageId="0" pageNumber="87">
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<taxonomicName id="0A408CA1F5E3C387287E2E0E144BBC0E" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
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<emphasis id="2E2DA207F4F91552826ED7FAA91A81A9" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
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</taxonomicName>
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sp. nov., holotype. IVPP OV2720 (G2, Figs
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<figureCitation id="15B9104B8BFB5BA3519D4B456C276F72" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gloydius lipipengi sp. nov. (A, B IVPP OV 2720, holotype) and Gloydius swild sp. nov. (C IVPP OV 2725, holotype, D IVPP OV, 2726, paratype) in life, not to scale." figureDoi="10.3897/zookeys.1061.70420.figure1" httpUri="https://binary.pensoft.net/fig/595356" pageId="0" pageNumber="87">1</figureCitation>
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-
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||
<figureCitation id="1AE793FD1F1BEF4A325D13A80B997C59" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Color rendered three-dimensional model of Gloydius lipipengi sp. nov. (holotype, IVPP OV 2720) A dorsal view B palatal view, mandibles not shown C lateral view. Abbreviations: bo, basioccipital; bs, basisphenoid; col, columella; cp, compound bone; d, dentary; ecp, ectopterygoid; exo, exoccipital; f, frontal; na, nasal; ma, maxilla; p, parietal; pcr, prearticular crest of compound bone; pfr, prefrontal; pmx, premaxilla; po, postorbital; pp, palatine process of maxilla; pro, prootic; psp, parasphenoid rostrum; pt, pterygoid; sac, surangular crest of compound bone; spm; septomaxilla; so, supraoccipital; sp, splenial; st, supratemporal; v, vomer. Conducted by Ye-Mao Hou and Jingsong Shi." figureDoi="10.3897/zookeys.1061.70420.figure4" httpUri="https://binary.pensoft.net/fig/595359" pageId="0" pageNumber="87">4</figureCitation>
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), adult male, collected from Muza Village, Zayu, Nyingchi Prefecture, Tibet (
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<geoCoordinate id="E1CD9B7B37015DDF959E783806786344" degrees="28.54" direction="north" orientation="latitude" precision="555" value="28.54">28.54°N</geoCoordinate>
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,
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<geoCoordinate id="D1A288B69F31AC157EC6B0A15A3B33A6" degrees="98.23" direction="east" orientation="longitude" precision="555" value="98.23">98.23°E</geoCoordinate>
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, 2883 m), by Jin-Cheng Liu, on 8 September 2014.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="731744ADB377353E9E28718A28002914" pageId="0" pageNumber="87" type="diagnosis">
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||
<paragraph id="9CCFEEA6D55688AFB9A973E8D85A8870" pageId="0" pageNumber="87">Diagnosis.</paragraph>
|
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<paragraph id="DF7386C28E48D4882CFE3AA918F432C1" pageId="0" pageNumber="87">
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The specimens of the new species, IVPP OV 2720, IVPP OV 2725 and IVPP OV 2726 were identified as the member of the genus
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<taxonomicName id="F2796A169CF6CCDE79EFD2024C027E70" authorityName="Hoge & Romano-Hoge" authorityYear="1981" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius" order="Squamata" pageId="0" pageNumber="87" rank="genus">
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||
<emphasis id="9AE5C65C4933FB77BBC1ABFED30AC3E0" italics="true" pageId="0" pageNumber="87">Gloydius</emphasis>
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</taxonomicName>
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based on the small body size, bilateral pits, and divided subcaudal scales (Hoge and Romano-Hoge 1981).
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</paragraph>
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<paragraph id="9915D121835400CEEBC512CC451AD64C" pageId="0" pageNumber="87">
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||
<taxonomicName id="BC1900F42DA57876B2906F2773FAA3FA" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
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<emphasis id="8BDF266EDF3E63FE960F6A4872FD1D57" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
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</taxonomicName>
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||
sp. nov. differs from other congeneric species in the following characteristics: i) third supralabial scale not touching the orbit; ii) a pair of prominent black markings on the occiput; iii) black-bordered greyish cheek stripe extending from the posterior margin of orbit (not separated by the postoculars) to the ventral surface of the neck; iv) black irregular annular crossbands on the mid-body; iv) two rows of black blotches on the ventral side; v) 23-21-15 circum-body scales; vi) 165 ventral scales; and vii) 46 subcaudal scales.
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</paragraph>
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<paragraph id="85BD78CB5B5344B6BDD3AAA672B3AA64" pageId="0" pageNumber="87">
|
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Table
|
||
<tableCitation id="8F902D73720DE9F662F560BFEFFDAA5C" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Brief morphological comparisons between Gloydius lipipengi sp. nov., G. swild sp. nov. and other congeneric species." httpUri="http://table.plazi.org/id/BD83D965C796E96243DDBDE40DE25ED9" pageId="0" pageNumber="87" tableUuid="BD83D965C796E96243DDBDE40DE25ED9">5</tableCitation>
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provides a brief summary of the differences between
|
||
<taxonomicName id="4543FA286F63D1EB5D9DBDB277255EFC" lsidName="G. lipipengi" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="DFA0EB9003ABD4D3C36A89011A4A55A5" italics="true" pageId="0" pageNumber="87">G. lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="E5C85873044773A2C35FBBC9BEC67F2E" lsidName="G. swild" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="E6082E19B2B324ECF2C9A5245EBAFE3E" italics="true" pageId="0" pageNumber="87">G. swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and other congeneric species.
|
||
</paragraph>
|
||
<caption id="BD83D965C796E96243DDBDE40DE25ED9" ID-Table-UUID="BD83D965C796E96243DDBDE40DE25ED9" httpUri="http://table.plazi.org/id/BD83D965C796E96243DDBDE40DE25ED9" pageId="0" pageNumber="87" start="Table 5" startId="T5">
|
||
<paragraph id="DB417E782F5019BF6BD3AB7B063640CF" pageId="0" pageNumber="87">
|
||
<emphasis id="B21EC29B7524FA58F4488DA35666B14D" bold="true" pageId="0" pageNumber="87">Table 5.</emphasis>
|
||
Brief morphological comparisons between
|
||
<taxonomicName id="349FD049EB3A8392EB01BD23360E8ED2" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="9017B7608AA8CB764287F0B394DAF431" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="180403A0C6B2932ECFD0DBBF62FD45BB" lsidName="G. swild" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="F19E02EE16497A18AB567450464AE521" italics="true" pageId="0" pageNumber="87">G. swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and other congeneric species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8A2673470801E423DA25A7D5451D4FDB" pageId="0" pageNumber="87">
|
||
<table id="A74610062D4DD88021D2722285E22D13" pageId="0" pageNumber="87">
|
||
<tr id="4335BECB473F9E91B243DA4036F74240" pageId="0" pageNumber="87">
|
||
<th id="CDC5DACF5F25CF20A26DE9BA2C866BBB" colspan="1" pageId="0" pageNumber="87" rowspan="1">Species</th>
|
||
<th id="49DCBF24A7A1476507E11645DF6D53F8" colspan="1" pageId="0" pageNumber="87" rowspan="1">Dorsal head</th>
|
||
<th id="3454EE9098ECC31620BA31669E53F368" colspan="1" pageId="0" pageNumber="87" rowspan="1">Spots on supralabials</th>
|
||
<th id="2E01AF92B84A70CF8E051461E1C58AF0" colspan="1" pageId="0" pageNumber="87" rowspan="1">Canthus rostralis</th>
|
||
<th id="169DE0F79CE7DD7E3C7BB0BED3EE95FE" colspan="1" pageId="0" pageNumber="87" rowspan="1">Background coloration</th>
|
||
<th id="EC0392C26621C118F12DC2B24CB9FECA" colspan="1" pageId="0" pageNumber="87" rowspan="1">Dorsal color patterns</th>
|
||
</tr>
|
||
<tr id="E6024A5930DA86A4E58C50F5C82CD450" pageId="0" pageNumber="87">
|
||
<td id="3F630C146BBFEEC91A7B4CCE12FF5AEE" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="5E3FDEBD0344D8B24ED75478936EE790" lsidName="G. lipipengi" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="DCF29FAA449419578E71745E279FD1B2" italics="true" pageId="0" pageNumber="87">G. lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</td>
|
||
<td id="8B910812CAFA97934EBC107EFC37AFBB" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="DEBCD2D2CF47E630707D46A3D06F6E3B" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown discrete spots</td>
|
||
<td id="D0FAB5DEF64ED33E59EF7EB3DFF02D09" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="281FEE71F546E17768FCCBB23CDDA0C6" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown</td>
|
||
<td id="4760F6926B29AFB8F0545DAEFBDDC2FF" colspan="1" pageId="0" pageNumber="87" rowspan="1">irregular large black interlaced patches</td>
|
||
</tr>
|
||
<tr id="89A0721F5728C892B864C260C95FADDC" pageId="0" pageNumber="87">
|
||
<td id="7670A79BBE240331ADFF21577628E63F" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="85E290DB1FC802206DFB9D3ACCC9BD9D" lsidName="G. swild" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="24A97418AB48CC122C88AFB6076B1864" italics="true" pageId="0" pageNumber="87">G. swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</td>
|
||
<td id="E5A16CF95A17414B5DEC4ECB5CE9F464" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="E45FCD6002B30B4B7ECEA8048DDE692E" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown discrete spots</td>
|
||
<td id="BC4FE0857313406371992022B3C1F5D5" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="2B4FD55A5A746E83EF6EAA95F80B2A4D" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown or blueish-grey</td>
|
||
<td id="EB2A4B1EEE431BE9D222DD63609DB9EB" colspan="1" pageId="0" pageNumber="87" rowspan="1">irregular zigzag dark brown markings</td>
|
||
</tr>
|
||
<tr id="6ECD71D75A41ACBB66B0A51F046EB34C" pageId="0" pageNumber="87">
|
||
<td id="B7D24CCF3FADA0CE73E59FAD08998A6F" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="138E23434F6791F74FCCCE84FB371D1C" lsidName="G. angusticeps" pageId="0" pageNumber="87" rank="species" species="angusticeps">
|
||
<emphasis id="B921808D61988A02A52C73EF0400A32F" italics="true" pageId="0" pageNumber="87">G. angusticeps</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="C3CE18E67EDBF10555A773992E08F364" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="4B75DF15442F9B624C1973BA0B33254F" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown, vaporous spots</td>
|
||
<td id="E7B52B0833674B1CD43D0FC8209E5C2E" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="D88A964956D91D5FC6917A2C95915E05" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown or yellowish brown</td>
|
||
<td id="7BAA9A2A9A6622E17BB894AD90C36F06" colspan="1" pageId="0" pageNumber="87" rowspan="1">dark brown small blotches, large patcher or stripes</td>
|
||
</tr>
|
||
<tr id="112BEE24ADC0AA0F5CE74A298CF89C29" pageId="0" pageNumber="87">
|
||
<td id="E60D6E8D65C0F825BF1A0A84D7068BC8" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="29BE35C4E8771EC20F535D7A10370D48" lsidName="G. himalayanus" pageId="0" pageNumber="87" rank="species" species="himalayanus">
|
||
<emphasis id="7B6B6F12AD10DBBDFAF4D8BDE5892C6B" italics="true" pageId="0" pageNumber="87">G. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="897B9FA3366298883F88928440B6C30C" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="CD9E15B04CFD29267275F7CD3F2BEB78" colspan="1" pageId="0" pageNumber="87" rowspan="1">one triangular spot between the third and fourth supralabial</td>
|
||
<td id="6B119EEF525EB9648C08803EF29B16A0" colspan="1" pageId="0" pageNumber="87" rowspan="1">very conspicuous</td>
|
||
<td id="46243E06B231884A42623B6A3DBCF3FB" colspan="1" pageId="0" pageNumber="87" rowspan="1">reddish brown or light grey</td>
|
||
<td id="29ED94D457E616C582BAD03632D9FC4B" colspan="1" pageId="0" pageNumber="87" rowspan="1">brown, reddish brown or dark grey small blotches or large black bordered patches</td>
|
||
</tr>
|
||
<tr id="B8E80B31FB9CA4D12C4F2EE788E40569" pageId="0" pageNumber="87">
|
||
<td id="20F6B45DEE318476BC9BB0AB2AF45FE6" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="BF685B5784667D1333995C344E33215C" lsidName="G. huangi" pageId="0" pageNumber="87" rank="species" species="huangi">
|
||
<emphasis id="4BAE508366AAB6FD20D22E9CA48780A8" italics="true" pageId="0" pageNumber="87">G. huangi</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="CBED4AF4B5449E23ACCD119672F5A1CF" colspan="1" pageId="0" pageNumber="87" rowspan="1">rounded</td>
|
||
<td id="1A2141152ADE483E4BE57B5B12E5178A" colspan="1" pageId="0" pageNumber="87" rowspan="1">yellowish brown small spots</td>
|
||
<td id="C8B97110F59803915C5D70DDF6BC4EA9" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="3AA86AD0ADC9724B4A585CA571528C15" colspan="1" pageId="0" pageNumber="87" rowspan="1">light or pale greyish yellow</td>
|
||
<td id="0DFE80D7D0AD4DE4EB9157BA06688D74" colspan="1" pageId="0" pageNumber="87" rowspan="1">regular large greyish brown patches</td>
|
||
</tr>
|
||
<tr id="DCF5C1E9F07265D9D13B667E809D1CA9" pageId="0" pageNumber="87">
|
||
<td id="2FF5B7B3F37D2B0041BF3E52A2F380F8" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="B48429347CAF592A06D12FF3BA534B54" lsidName="G. liupanensis" pageId="0" pageNumber="87" rank="species" species="liupanensis">
|
||
<emphasis id="7B6AC1CE571745E45055380D4A1379F1" italics="true" pageId="0" pageNumber="87">G. liupanensis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="CC87A7D90BD24BF89499CA8957E41981" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="5830E8882C58A486589B47DCD22963AE" colspan="1" pageId="0" pageNumber="87" rowspan="1">none</td>
|
||
<td id="77B35ECC817F1399B0109EFC333013F6" colspan="1" pageId="0" pageNumber="87" rowspan="1">conspicuous</td>
|
||
<td id="BF1B119780C12E6CCFAA92DA09CE4952" colspan="1" pageId="0" pageNumber="87" rowspan="1">light reddish brown or yellowish brown</td>
|
||
<td id="C12965C8348331FD27AB2F41AC35233F" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
similar with
|
||
<taxonomicName id="7E919714813B671B912C7DFEBE1E6C42" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius qinlingensis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="qinlingensis">Gloydius qinlingensis</taxonomicName>
|
||
, with white stripe on the body side
|
||
</td>
|
||
</tr>
|
||
<tr id="C552753C47C051D739484307EF578B71" pageId="0" pageNumber="87">
|
||
<td id="2C3E012B410B9A3DDA6C4E4A8AE380FB" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="ABE1CCF510D6E8C0C80776C0C8E2EC9D" lsidName="G. monticola" pageId="0" pageNumber="87" rank="species" species="monticola">
|
||
<emphasis id="7FBA5135D2F7EE369FE8A2DB101A3B0A" italics="true" pageId="0" pageNumber="87">G. monticola</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="330E53CD0A577F3372346C0620006297" colspan="1" pageId="0" pageNumber="87" rowspan="1">rounded</td>
|
||
<td id="E9227FF20CF2594EE531F25DDBA67399" colspan="1" pageId="0" pageNumber="87" rowspan="1">whitish borders of the labials along the mouth line</td>
|
||
<td id="ADF44B4E0D87444EC7ADB7335F970FA8" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="F139833F144C46FD8388F2F079A1AD09" colspan="1" pageId="0" pageNumber="87" rowspan="1">greyish brown, greenish brown or reddish brown</td>
|
||
<td id="383B7A583D488F7B53B99866AE8696A8" colspan="1" pageId="0" pageNumber="87" rowspan="1">dark grey or dark brown blotches or zigzag stripes</td>
|
||
</tr>
|
||
<tr id="6D41C75C1828608E09A13ED3134AA272" pageId="0" pageNumber="87">
|
||
<td id="6B1845D0E76CE928175F3B4174335D01" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="90A6D4B11C4A31F5031F24300DF60C0B" lsidName="G. qinlingensis" pageId="0" pageNumber="87" rank="species" species="qinlingensis">
|
||
<emphasis id="E38C238D971C989FD914C157C1A1AAB0" italics="true" pageId="0" pageNumber="87">G. qinlingensis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="07E50A06655A974EA3481AE7C1650767" colspan="1" pageId="0" pageNumber="87" rowspan="1">triangular</td>
|
||
<td id="6AB9DD1F6176889DF0F1E4C2912F9FC9" colspan="1" pageId="0" pageNumber="87" rowspan="1">none</td>
|
||
<td id="6AA31CCE186EA09F27B2C6999D3D778E" colspan="1" pageId="0" pageNumber="87" rowspan="1">conspicuous</td>
|
||
<td id="15B8B1DEFFEC64D4CD604AF34C72B22D" colspan="1" pageId="0" pageNumber="87" rowspan="1">light reddish brown or yellowish brown</td>
|
||
<td id="DDC0D191733824B8B5EF5390C95CDC30" colspan="1" pageId="0" pageNumber="87" rowspan="1">two columns of irregular yellowish brown or dark brown</td>
|
||
</tr>
|
||
<tr id="A591AEFE5FAB5417A53C25ACFDF6C4C5" pageId="0" pageNumber="87">
|
||
<td id="A819E456109B76751C6A31050E1E106B" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="37159AC18B58A94E59E8502258119A06" lsidName="G. rubromaculatus" pageId="0" pageNumber="87" rank="species" species="rubromaculatus">
|
||
<emphasis id="79EAD980E4CC37EE8EBB83995B032ED7" italics="true" pageId="0" pageNumber="87">G. rubromaculatus</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="EE51F0B619DAA55006275372600CC769" colspan="1" pageId="0" pageNumber="87" rowspan="1">rounded</td>
|
||
<td id="F95DED004430326D5410270987FD00C2" colspan="1" pageId="0" pageNumber="87" rowspan="1">irregular small black spots</td>
|
||
<td id="C058D9CD7C7E62D846D2EC026F5F416A" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="EA85B5FFC5BA768C011CBB57E7602888" colspan="1" pageId="0" pageNumber="87" rowspan="1">khaki or yellowish brown</td>
|
||
<td id="EC44481DB12771387C49213672B98716" colspan="1" pageId="0" pageNumber="87" rowspan="1">regular large (or discrete small) scarlet or brownish yellow patches or stripes</td>
|
||
</tr>
|
||
<tr id="76D5CC9CCFEF301E731D7E10E45F132D" pageId="0" pageNumber="87">
|
||
<td id="6F69A25D143F07D86EAF7DC33DF0682A" colspan="1" pageId="0" pageNumber="87" rowspan="1">
|
||
<taxonomicName id="A7C0ED8D83F48D4FB7E5530400BDC93E" lsidName="G. strauchi" pageId="0" pageNumber="87" rank="species" species="strauchi">
|
||
<emphasis id="D036B4C9DE009059735DB0C1EA30A7F1" italics="true" pageId="0" pageNumber="87">G. strauchi</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="9EB726481192DB0ED8020AA0F60EBA77" colspan="1" pageId="0" pageNumber="87" rowspan="1">rounded</td>
|
||
<td id="B261A8E6412F4BF025ED6884F8F3BA56" colspan="1" pageId="0" pageNumber="87" rowspan="1">large brown between the second, third and fourth or none</td>
|
||
<td id="5F58065AA3C9B3E37C334943F3669631" colspan="1" pageId="0" pageNumber="87" rowspan="1">inconspicuous</td>
|
||
<td id="390593DE4E9C17C4D5639039893D4FFD" colspan="1" pageId="0" pageNumber="87" rowspan="1">greenish brown, yellowish brown or nut-brown</td>
|
||
<td id="0894B17CE8C4C4EEBAC4831FA2DC8C9C" colspan="1" pageId="0" pageNumber="87" rowspan="1">patches, transverse crossbands or four longitudinal zigzag strips</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="C3ADFFB3DA7BF12964923278AF515755" pageId="0" pageNumber="87">
|
||
<taxonomicName id="175E51BE2460ED8CCB7C1ABBD751F5F7" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="C7FEA9B0CBE7294102EA234877411604" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="9AC5DBCA7AFEAADF830DA90F99EA1DB3" lsidName="G. swild" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="E069FD7721A03EBBEF576037AEF0AC25" italics="true" pageId="0" pageNumber="87">G. swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be differentiated from the species in the
|
||
<taxonomicName id="8B60855FF547581B2BCBC7195C0A6D25" lsidName="G. blomhoffii" pageId="0" pageNumber="87" rank="species" species="blomhoffii">
|
||
<emphasis id="A0DD8EA607373A69D3340F553BE1B409" italics="true" pageId="0" pageNumber="87">G. blomhoffii</emphasis>
|
||
</taxonomicName>
|
||
complex by having three palatine teeth (versus four palatine teeth), from the
|
||
<taxonomicName id="0D88ADC5957B570B124CB9C93D0F612A" lsidName="G. halys" pageId="0" pageNumber="87" rank="species" species="halys">
|
||
<emphasis id="763D4BC7828116CC6358491E70D586DD" italics="true" pageId="0" pageNumber="87">G. halys</emphasis>
|
||
</taxonomicName>
|
||
complex by having 21 rows of mid-body dorsal scales (versus 22 or 23 rows).
|
||
<taxonomicName id="A717FB65E206DEFDE98F00B7E1778BEF" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="5FDB414CFC6CAA882DB23471592A8546" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from other species in
|
||
<taxonomicName id="E089833E5071F1CF1C3A3B3A5AEB9098" lsidName="G. strauchi" pageId="0" pageNumber="87" rank="species" species="strauchi">
|
||
<emphasis id="A85640AEA690386319D55A998AD5B3CA" italics="true" pageId="0" pageNumber="87">G. strauchi</emphasis>
|
||
</taxonomicName>
|
||
complex by the third supralabial scale not touching the orbit, from
|
||
<taxonomicName id="1F1F356311559DF3E6DEA28581543BEF" lsidName="G. strauchi" pageId="0" pageNumber="87" rank="species" species="strauchi">
|
||
<emphasis id="1BFF454D26DB8F7CA9A2BDA2DC834745" italics="true" pageId="0" pageNumber="87">G. strauchi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="37FE0E71BD4A0D82BB3998C13CEC3A0B" lsidName="G. huangi" pageId="0" pageNumber="87" rank="species" species="huangi">
|
||
<emphasis id="2AFD3C00FCF883CFBB777A91CB9E2F15" italics="true" pageId="0" pageNumber="87">G. huangi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3C598E0A62DFE23C6C79CD254152E8F7" lsidName="G. rubromaculatus" pageId="0" pageNumber="87" rank="species" species="rubromaculatus">
|
||
<emphasis id="1DE0C586274ECC1E8BA7A840543803CD" italics="true" pageId="0" pageNumber="87">G. rubromaculatus</emphasis>
|
||
</taxonomicName>
|
||
by having large irregular black markings on the back (versus four irregular longitudinal stripes or discrete blotches in
|
||
<taxonomicName id="FE88F25375845E9BA47A7511B999C864" lsidName="G. strauchi" pageId="0" pageNumber="87" rank="species" species="strauchi">
|
||
<emphasis id="0354A72CA7C9FFD3F721B2E77B1F0294" italics="true" pageId="0" pageNumber="87">G. strauchi</emphasis>
|
||
</taxonomicName>
|
||
, complete dark brown patches in
|
||
<taxonomicName id="D033B8671D83DBD722101A2A655B064A" lsidName="G. huangi" pageId="0" pageNumber="87" rank="species" species="huangi">
|
||
<emphasis id="4626E073AAB884908D78ABA9D099E67C" italics="true" pageId="0" pageNumber="87">G. huangi</emphasis>
|
||
</taxonomicName>
|
||
, and large red crossbands in
|
||
<taxonomicName id="F971E1D968F00CF1403E7D6FA6CEA898" lsidName="G. rubromaculatus" pageId="0" pageNumber="87" rank="species" species="rubromaculatus">
|
||
<emphasis id="EAD4C77D3C7BB85BD4C8E29AC0928278" italics="true" pageId="0" pageNumber="87">G. rubromaculatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4C482189ACCF8AAB3E1DA9209B31F7BF" DOI="https://doi.org/10.1670/18-126" author="Wang, K" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="87" pagination="224 - 236" refId="B22" refString="Wang, K, Ren, J, Dong, W, Jiang, K, Shi, J, Siler, CD, Che, J, 2019. A New Species of Plateau Pit Viper (Reptilia: Serpentes: Gloydius) from the Upper Lancang (= Mekong) Valley in the Hengduan Mountain Region, Tibet, China. Journal of Herpetology 53 (3): 224 - 236, DOI: https://doi.org/10.1670/18-126" title="A New Species of Plateau Pit Viper (Reptilia: Serpentes: Gloydius) from the Upper Lancang (= Mekong) Valley in the Hengduan Mountain Region, Tibet, China." url="https://doi.org/10.1670/18-126" volume="53" year="2019">Wang et al. 2019</bibRefCitation>
|
||
), from
|
||
<taxonomicName id="46A8BEFDA7FACE02D16361D9621470FD" lsidName="G. monticola" pageId="0" pageNumber="87" rank="species" species="monticola">
|
||
<emphasis id="CF8EE7D1F8E003D1AD5349A2DF2845C6" italics="true" pageId="0" pageNumber="87">G. monticola</emphasis>
|
||
</taxonomicName>
|
||
by having seven supralabials (versus always six supralabials) and more subcaudal scales (46 pairs versus always fewer than 30 pairs), from
|
||
<taxonomicName id="4DD5CDCA9EFD5505C5E13A8E6FF04CBE" lsidName="G. qinlingensis" pageId="0" pageNumber="87" rank="species" species="qinlingensis">
|
||
<emphasis id="6E904C88D78EEF6F23AB18E27529BA15" italics="true" pageId="0" pageNumber="87">G. qinlingensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3FF070A732EF6C6408AF87590F19943" lsidName="G. liupanensis" pageId="0" pageNumber="87" rank="species" species="liupanensis">
|
||
<emphasis id="2A349EC84D72FBFF107404813A5E77AA" italics="true" pageId="0" pageNumber="87">G. liupanensis</emphasis>
|
||
</taxonomicName>
|
||
by its greyish brown body colour (versus yellowish-brown body colour) and lacking a lateral white line on each lateral side (versus possessing a lateral white line on each side).
|
||
<taxonomicName id="1BA3AB796271E05FC51A3EFD615F4F13" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="6C66DFC000992699078829756DCBE6AC" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be differentiated from
|
||
<taxonomicName id="4D49B411BFD3AAF29246C0BD56E3E830" lsidName="G. himalayanus" pageId="0" pageNumber="87" rank="species" species="himalayanus">
|
||
<emphasis id="22E3365FDE0837862190CC8852F25E9B" italics="true" pageId="0" pageNumber="87">G. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
by possessing an indistinct canthus rostralis (versus very distinct canthus rostralis; Gloyd and Conant 1990).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="528C73E2FEC54290D026563BC1C2A34A" pageId="0" pageNumber="87" type="description">
|
||
<paragraph id="E3DAF24C057D6940162F861FA22BC64D" pageId="0" pageNumber="87">Description of the holotype.</paragraph>
|
||
<paragraph id="BDAF7A0459F110EE34A591D02D7C3654" pageId="0" pageNumber="87">
|
||
IVPP OV 2720, adult male, a slender pit viper with a total length of 628.2 mm (SVL 540.6 mm and TL 87.6 mm), preserved in 75% ethanol with its left hemipenes partially extruded (Figs
|
||
<figureCitation id="6B77533F87E3448CBB78F9A176F0E3A4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gloydius lipipengi sp. nov. (A, B IVPP OV 2720, holotype) and Gloydius swild sp. nov. (C IVPP OV 2725, holotype, D IVPP OV, 2726, paratype) in life, not to scale." figureDoi="10.3897/zookeys.1061.70420.figure1" httpUri="https://binary.pensoft.net/fig/595356" pageId="0" pageNumber="87">1</figureCitation>
|
||
,
|
||
<figureCitation id="87FB7020CC6E083FB436C3E5D3ACB9A0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Gloydius lipipengi sp. nov. (IVPP OV 2720) in preservative A dorsal view B ventral view." figureDoi="10.3897/zookeys.1061.70420.figure2" httpUri="https://binary.pensoft.net/fig/595357" pageId="0" pageNumber="87">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="7F88C41146980C3A993C5E24E130F8FC" doi="10.3897/zookeys.1061.70420.figure1" httpUri="https://binary.pensoft.net/fig/595356" pageId="0" pageNumber="87" start="Figure 1" startId="F1">
|
||
<paragraph id="01570EC14A9D8182646962161D4913D1" pageId="0" pageNumber="87">
|
||
<emphasis id="A878BE2B3095D2DC8F83AB1173FE1257" bold="true" pageId="0" pageNumber="87">Figure 1.</emphasis>
|
||
<taxonomicName id="0014F648D28EA9391D9FF3669CD6A9B4" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="2C57E59703A0B81FFE140CD64D325B0D" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis id="4D5E480C1EAAF602D49EC2EC95A62D59" bold="true" pageId="0" pageNumber="87">A, B</emphasis>
|
||
IVPP OV 2720, holotype) and
|
||
<taxonomicName id="E29807DE8383899762AC89856D920857" authorityName="Shi & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius swild" order="Squamata" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="264222BB687AD30B957FFFF1E18DE3E4" italics="true" pageId="0" pageNumber="87">Gloydius swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis id="4AF0F5284165340CCD3CC046A509BF2C" bold="true" pageId="0" pageNumber="87">C</emphasis>
|
||
IVPP OV 2725, holotype,
|
||
<emphasis id="7F6341247FEB9DE20E96409A108F2020" bold="true" pageId="0" pageNumber="87">D</emphasis>
|
||
IVPP OV, 2726, paratype) in life, not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CC375BE2D3FABCDAAECF9E501A7750DD" doi="10.3897/zookeys.1061.70420.figure2" httpUri="https://binary.pensoft.net/fig/595357" pageId="0" pageNumber="87" start="Figure 2" startId="F2">
|
||
<paragraph id="28A0FF68779F6FDC066AACC0A89B7240" pageId="0" pageNumber="87">
|
||
<emphasis id="6ACDFFC0E2FD2E4B6522A9DF86218C59" bold="true" pageId="0" pageNumber="87">Figure 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="AC99A275D3E1C116B59AC165D36CFDD6" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="C28A0AB1C5C2C98EAAFC88D5B380B578" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (IVPP OV 2720) in preservative
|
||
<emphasis id="8E77E008DF5C456DA62E65326F7ABE1F" bold="true" pageId="0" pageNumber="87">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="A82617A5C4AFDAAC08556BC59BA41027" bold="true" pageId="0" pageNumber="87">B</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="892C35CCE21B1FE6E29510A2518D3401" pageId="0" pageNumber="87">The head is slender and triangular shaped in dorsal view, distinct from the neck. Canthus rostralis are not distinct. The head is 25.2 mm in length, 13.2 mm in width and 8.2 mm in depth.</paragraph>
|
||
<paragraph id="0A45485B061B8E340E8760E1D8F4C26A" pageId="0" pageNumber="87">
|
||
<emphasis id="663A2B080247C153418453F31654E569" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="76C8FF8A9E49634B93BA4E0F859E1E47" italics="true" pageId="0" pageNumber="87">Scalation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Rostral scale slightly up-turned, visible from dorsal view; nasal divided, anterior part larger; seven supralabials on both sides: second smallest, not reaching the pit; third highest, not touching the bottom of orbit (separated by one small subocular); fourth longest, not touching the orbit; three preoculars, two postoculars, inferior one touching the top of the third supralabials, forming the bottom margin of the orbit; two rows of temporals (2+4); infralabials 10 on left side while 11 on right, first pair in contact behind the mental; second, third and fourth pairs meet on the chin shield; chin shield is rhomboidal in shape, the posterior chin shield comprises two pairs of scales, forming the mental groove (Fig.
|
||
<figureCitation id="DA2F59E12AB3F8C940C4EC87CB155F4B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Head squamation of Gloydius lipipengi sp. nov. (Holotype, IVPP OV 2720: A lateral view B dorsal view C ventral view) and G. swild sp. nov. (Holotype, IVPP OV 2725: D lateral view E dorsal view F ventral view). Scale bar: 10 mm." figureDoi="10.3897/zookeys.1061.70420.figure3" httpUri="https://binary.pensoft.net/fig/595358" pageId="0" pageNumber="87">3A-C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="E58D405036EC6E643CAEF046FBB68701" doi="10.3897/zookeys.1061.70420.figure3" httpUri="https://binary.pensoft.net/fig/595358" pageId="0" pageNumber="87" start="Figure 3" startId="F3">
|
||
<paragraph id="3030BC1C1BD370A6ABB0C4106291C962" pageId="0" pageNumber="87">
|
||
<emphasis id="155FBAA9066DC48282D332EB9FC74EE4" bold="true" pageId="0" pageNumber="87">Figure 3.</emphasis>
|
||
Head squamation of
|
||
<taxonomicName id="B27DBA29FA23E9E0A4DB56DE953DC9C5" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="DFD130BA7C1478F320A3E8C3D43C127E" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Holotype, IVPP OV 2720:
|
||
<emphasis id="36193E76E213FB346E5749008B343D38" bold="true" pageId="0" pageNumber="87">A</emphasis>
|
||
lateral view
|
||
<emphasis id="83B4E4857EA2E8AA690C3B91D96BB2D6" bold="true" pageId="0" pageNumber="87">B</emphasis>
|
||
dorsal view
|
||
<emphasis id="CD4A46D8979CD9527A5691E317586C9E" bold="true" pageId="0" pageNumber="87">C</emphasis>
|
||
ventral view) and
|
||
<taxonomicName id="89E2D17A270DB93E60E8180B941A2231" lsidName="G. swild" pageId="0" pageNumber="87" rank="species" species="swild">
|
||
<emphasis id="D5B5159ED60633F5DFDA94D049882E19" italics="true" pageId="0" pageNumber="87">G. swild</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Holotype, IVPP OV 2725:
|
||
<emphasis id="9A14B61B63ACF663FEF5A9DA6A734B8C" bold="true" pageId="0" pageNumber="87">D</emphasis>
|
||
lateral view
|
||
<emphasis id="FDE6C3EF7F2CB35ABC8D48E77644E57F" bold="true" pageId="0" pageNumber="87">E</emphasis>
|
||
dorsal view
|
||
<emphasis id="D0972ED0AD11C531058D9B18E7634DEA" bold="true" pageId="0" pageNumber="87">F</emphasis>
|
||
ventral view). Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6DFA0BCF1EECD96A10BBE27AA8C384EE" pageId="0" pageNumber="87">Dorsal scales in 23-21-15 rows (reducing from 19 to 18 posteriorly at ventral 94-96), keeled except for the first scale row bordering the ventral scales; ventral scales 165 (excluding four preventral scales); anal plate single; subcaudals 46, in pairs</paragraph>
|
||
<paragraph id="1B8C2C99637D86A886D239A6611FDC1F" pageId="0" pageNumber="87">
|
||
<emphasis id="EC9CB262F55893D0AA1F518D60D2D8D1" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="67951D5EC7A8F445CB07930769DA5EEE" italics="true" pageId="0" pageNumber="87">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Eye dark brown on the upper half while black on the bottom half, pupil black, vertical with light yellow margins; postorbital stripe wide, greyish brown and black bordered on the lower edge, extending from the posterior orbit to the ventral surface of the neck; supralabials and infralabials greyish brown, scattered with very small irregularly sized black blotches. One black triangular mark on the anterodorsal head, covering the caudomedial part of prefrontals. One bold black M-shaped mark on the dorsomedial head, covering the caudal part of lateral frontals, the lateral part of parietals, merged with the postorbital stripe at the largest temporal scale (but not covering the upper postorbital). The upper postorbital white while the top part of the bottom postorbital is black (covered by the postorbital stripe).
|
||
</paragraph>
|
||
<paragraph id="5F09CCF082EC426ED32D70E30038073E" pageId="0" pageNumber="87">
|
||
The body coloration is dark greyish brown, with two rows of irregular black annular crossbands on the mid-body, each covering 20 or more scales, separated by a gap of two row scale vertically, extending laterally to one or two dorsal scales from the ventrals. Ventral scales light grey, with two large black blotches on each side, clustered into two ventral stripes. The tip of tail is similar to the main body in coloration (Figs
|
||
<figureCitation id="71588003BDD5AF23752FC68CEEC8452F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gloydius lipipengi sp. nov. (A, B IVPP OV 2720, holotype) and Gloydius swild sp. nov. (C IVPP OV 2725, holotype, D IVPP OV, 2726, paratype) in life, not to scale." figureDoi="10.3897/zookeys.1061.70420.figure1" httpUri="https://binary.pensoft.net/fig/595356" pageId="0" pageNumber="87">1</figureCitation>
|
||
,
|
||
<figureCitation id="69E8282C20A7F18C9B23D688332EA97C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Gloydius lipipengi sp. nov. (IVPP OV 2720) in preservative A dorsal view B ventral view." figureDoi="10.3897/zookeys.1061.70420.figure2" httpUri="https://binary.pensoft.net/fig/595357" pageId="0" pageNumber="87">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4F634D870BBBC0960248DB5A918A2055" pageId="0" pageNumber="87">
|
||
<emphasis id="426E54720C08C41E8EA62495A6FD82D0" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="70C7319C22D3B958C5816927703C34CD" italics="true" pageId="0" pageNumber="87">Skull</emphasis>
|
||
.
|
||
</emphasis>
|
||
The description of the skull of
|
||
<taxonomicName id="8E6D04B7050ECB75B8E6DE24292DB91F" lsidName="G. lipipengi" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="48C27BFBA7EABB2409DD4F26DFBAD1A5" italics="true" pageId="0" pageNumber="87">G. lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is based on the 3D-reconstructed model of the holotype.
|
||
</paragraph>
|
||
<paragraph id="C98757B458083C51AA7E28AC4D231B2E" pageId="0" pageNumber="87">
|
||
<emphasis id="44A467FDF348470E8D178C863E2E780A" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="25B9D17EA9CB01EB469CC0C7470A4F3D" italics="true" pageId="0" pageNumber="87">Snout</emphasis>
|
||
.
|
||
</emphasis>
|
||
The premaxilla has bifurcated transverse process on each side. The anterior margin of the premaxilla is blunt. The dorsal tip of the ascending process of premaxilla is triangular in lateral view, not reaching the anterior tip of nasals. The horizontal laminae of the nasals are scutiform in dorsal view. The septomaxillae have prominent dorsolateral processes, nearly meeting the horizontal laminae of the nasals.
|
||
</paragraph>
|
||
<paragraph id="5D5B59C976E65046BCEAE48986DE787D" pageId="0" pageNumber="87">
|
||
<emphasis id="DC7DDBDC758EA0FF023365569335C9C6" bold="true" pageId="0" pageNumber="87">
|
||
<taxonomicName id="DE3F78C8437E4F90034FFB579E23D220" genus="Braincase" lsidName="Braincase" pageId="0" pageNumber="87" rank="genus">
|
||
<emphasis id="25E89CD72AD7A3C2F6E194237B358410" italics="true" pageId="0" pageNumber="87">Braincase</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The parietal is roughly T-shaped in dorsal view. The anterolateral part of the parietal bulges prominently laterally while the dorsoposterior part tapers medially. The postorbital processes of the parietal are prominent. The frontals are squared. The lateral margin of frontals concaved obviously on each side, forming the dorsal edge of orbit. The prefrontal has an elongate blunt lateral process, posterolaterally pointed. The lacrimal foramen perforates the medial lamina of the prefrontal. The prefrontal-frontal join surface is waved in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="5BAA8749BDA3AA5662E81CC1EA05024A" pageId="0" pageNumber="87">The postorbital is relatively small and cashew-shaped, the top of the postorbital does not reach the posterolateral end of frontal. The basisphenoid is spearhead in shape, narrow anteriorly and expanded laterally. The supraoccipital is longitudinally compressed, occupies almost two thirds the total width of the otic region.</paragraph>
|
||
<paragraph id="5674F6C714F4F0AEA3ABBA2D55D396DF" pageId="0" pageNumber="87">
|
||
<emphasis id="454888B1E076E3BBBFFAC13A9736F57E" bold="true" pageId="0" pageNumber="87">
|
||
<taxonomicName id="6936BDE496A7C8D804E7D3ACF9EE28BE" genus="Palatomaxillary" lsidName="Palatomaxillary apparatus" pageId="0" pageNumber="87" rank="species" species="apparatus">
|
||
<emphasis id="0C0E5A54A127C42BAFC7025D3B05DFDB" italics="true" pageId="0" pageNumber="87">Palatomaxillary apparatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The fang is relatively short and curved, roughly the same length of the maxilla, one third the length of ectopterygoid, attached with seven or eight replacement fangs on each side. The palatine bears three teeth. The ectopterygoid is flat and widened at the anterior part. The pterygoid is slender, the dentigerous process of pterygoid is straight, bearing 12/11 teeth (left/right), occupies almost half the total width of the pterygoid, the posterior portion of pterygoid is medially expanded.
|
||
</paragraph>
|
||
<paragraph id="17CA37C8E23BAFA5E99512B45EC70DA3" pageId="0" pageNumber="87">
|
||
<emphasis id="A7A78243F4A73021D3BF4378EE5A70E8" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="9A09FFC5A9EFEE718C296E88A78209AA" italics="true" pageId="0" pageNumber="87">Suspensorium and mandible</emphasis>
|
||
.
|
||
</emphasis>
|
||
The supratemporal is slender, has a lateral process, anterolaterally pointed, lies in front of the supratemporal-quadrate articulation. The quadrate is straight, slender, and enlarged on both ends. The mandible is slender and moderately curved. The prearticular crest of the compound bone is prominent while the surangular crest is slightly concaved. The dental bone bears 11/12 teeth (left/right); the dentary teeth are perpendicular to the dentary bone, decreasing in size at the third tooth. The posterior tip of ventral process of dentary extends farther posteriorly than the dorsal process.
|
||
</paragraph>
|
||
<paragraph id="A550593BBD120A2294DF45B15E627451" pageId="0" pageNumber="87">
|
||
<emphasis id="840CC79ED2CBE63358C701D69C2E4035" bold="true" pageId="0" pageNumber="87">
|
||
<emphasis id="04C153993C6BEF6727712640D78AE48E" italics="true" pageId="0" pageNumber="87">Dentition</emphasis>
|
||
.
|
||
</emphasis>
|
||
Palatine: 3/3, pterygoid: 12/11, dentary: 11/12.
|
||
</paragraph>
|
||
<paragraph id="1D8A16B5F311FE183B838B90345EDB66" pageId="0" pageNumber="87">
|
||
<emphasis id="1C21203292FE7ADC9B820D30EF220361" bold="true" pageId="0" pageNumber="87">
|
||
<taxonomicName id="65B2C3EE20F4E7A04DD28C1A54D7D6ED" genus="Hemipenes" lsidName="Hemipenes" pageId="0" pageNumber="87" rank="genus">
|
||
<emphasis id="CB37FAF079465A6028D1034155E5EA84" italics="true" pageId="0" pageNumber="87">Hemipenes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The hemipenes of
|
||
<taxonomicName id="6F1DD2C714430EF5D4340579B68A2CE0" lsidName="G. lipipengi" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="EE12551488285A44FD272A5677870D42" italics="true" pageId="0" pageNumber="87">G. lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are generally similar to those of
|
||
<taxonomicName id="70C9826A67560FBA36D4E6C850F26853" lsidName="G. rubromaculatus" pageId="0" pageNumber="87" rank="species" species="rubromaculatus">
|
||
<emphasis id="C3D4F55680C3A5E84AF1B9847FF76675" italics="true" pageId="0" pageNumber="87">G. rubromaculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CD75AD7E904CBB5012D89FA38B39E648" lsidName="G. huangi" pageId="0" pageNumber="87" rank="species" species="huangi">
|
||
<emphasis id="5C00FD2852FD2F6B51442E8B0E6CE621" italics="true" pageId="0" pageNumber="87">G. huangi</emphasis>
|
||
</taxonomicName>
|
||
but differ by the possession of longer and stronger spines, seven or eight subcaudals in length, and forked for two subcaudals. Small and stubby spines range from the basal to the distal side of the organ, without any conspicuously enlarged spines (versus 3-5 enlarged spines on the base in the
|
||
<taxonomicName id="1769AFE787483A461369CAA5A4DB1918" lsidName="G. halys" pageId="0" pageNumber="87" rank="species" species="halys">
|
||
<emphasis id="65FD720D3EBC964C985CC733B0E08608" italics="true" pageId="0" pageNumber="87">G. halys</emphasis>
|
||
</taxonomicName>
|
||
complex;
|
||
<bibRefCitation id="CDBB2BDF23E16A7397DBB60F164CBC81" DOI="https://doi.org/10.5358/hsj1972.14.1_23" author="Gloyd, HK" journalOrPublisher="Society for the Study of Amphibians and Reptiles, USA" pageId="0" pageNumber="87" refId="B7" refString="Gloyd, HK, 1990. Snakes of the Agkistrodon Complex: A Monographic Review. Society for the Study of Amphibians and Reptiles, USA, DOI: https://doi.org/10.5358/hsj1972.14.1_23" title="Snakes of the Agkistrodon Complex: A Monographic Review." url="https://doi.org/10.5358/hsj1972.14.1_23" year="1990">Gloyd 1990</bibRefCitation>
|
||
). The spines gradually increase in length distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F358AFA3B5D70205EA7D862237E82F9C" pageId="0" pageNumber="87" type="distribution">
|
||
<paragraph id="E7C390DDE899F7971C30E046776AFF18" pageId="0" pageNumber="87">Distribution and ecology.</paragraph>
|
||
<paragraph id="2748FBBE6FF08A323CDF59202BFDDFDF" pageId="0" pageNumber="87">
|
||
At present,
|
||
<taxonomicName id="674C3EFF259BA2996BDDC00060CFB590" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="5906431A34BE4C9179A3DF6560E11181" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has only been reported from the type locality, Muza village, Zayu, Tibet, China (Fig.
|
||
<figureCitation id="26049CED3F5C6FA2A08857D069684774" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Type localities of Gloydius lipipengi sp. nov. (red triangles) and G. swild sp. nov. (black triangles), with the collection localities of some other congeneric species." figureDoi="10.3897/zookeys.1061.70420.figure6" httpUri="https://binary.pensoft.net/fig/596192" pageId="0" pageNumber="87">6</figureCitation>
|
||
). The specimen was collected at 09:00 h on leaf litter in forest near the hot, dry valley on the lower reaches of the Nujiang River (Fig.
|
||
<figureCitation id="3192040AD6B5509373819F3334BF78E7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. The habitat of Gloydius lipipengi sp. nov. (A Muza Village, Zaty, Tibet, type locality of G. lipipengi sp. nov. B the landscape of the Nujiang River, 15 km from the type locality) and Gloydius swild sp. nov. (C Heishui, Sichuan) A and B Photographs by Jin-Cheng Liu." figureDoi="10.3897/zookeys.1061.70420.figure7" httpUri="https://binary.pensoft.net/fig/595362" pageId="0" pageNumber="87">7</figureCitation>
|
||
).
|
||
<taxonomicName id="5D5D3D46E5B0A3AB5A00B1E207D435A2" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="7E0358B2FFA6B165A02B15FE06678643" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. accepted pink mice in captivity.
|
||
</paragraph>
|
||
<caption id="50AFAEE39ABF1C65489CDC9628C32987" doi="10.3897/zookeys.1061.70420.figure4" httpUri="https://binary.pensoft.net/fig/595359" pageId="0" pageNumber="87" start="Figure 4" startId="F4">
|
||
<paragraph id="E406B0BA97BBE7FD809FF649AEF9A256" pageId="0" pageNumber="87">
|
||
<emphasis id="2E74444FECAF4C39B662EAE9A9F1C2AF" bold="true" pageId="0" pageNumber="87">Figure 4.</emphasis>
|
||
Color rendered three-dimensional model of
|
||
<taxonomicName id="F71D9AA4D49B69A252706AE1188ED13A" authorityName="Shi, Liu & Malhotra" authorityYear="2021" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius lipipengi" order="Squamata" pageId="0" pageNumber="87" rank="species" species="lipipengi">
|
||
<emphasis id="D5BBF249D406CE7D1B0B94F3F458BE67" italics="true" pageId="0" pageNumber="87">Gloydius lipipengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (holotype, IVPP OV 2720)
|
||
<emphasis id="21D17DBB2A698050923B735EE90CAAEF" bold="true" pageId="0" pageNumber="87">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="4F4708FF2C34983B8D6C73832C5EFC07" bold="true" pageId="0" pageNumber="87">B</emphasis>
|
||
palatal view, mandibles not shown
|
||
<emphasis id="1920B3FDB9705B166B385817F4EC7748" bold="true" pageId="0" pageNumber="87">C</emphasis>
|
||
lateral view. Abbreviations: bo, basioccipital; bs, basisphenoid; col, columella; cp, compound bone; d, dentary; ecp, ectopterygoid; exo, exoccipital; f, frontal; na, nasal; ma, maxilla; p, parietal; pcr, prearticular crest of compound bone; pfr, prefrontal; pmx, premaxilla; po, postorbital; pp, palatine process of maxilla; pro, prootic; psp, parasphenoid rostrum; pt, pterygoid; sac, surangular crest of compound bone; spm; septomaxilla; so, supraoccipital; sp, splenial; st, supratemporal; v, vomer. Conducted by Ye-Mao Hou and Jingsong Shi.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="19685DBDEA1EB3ED94860C72BA4F760C" doi="10.3897/zookeys.1061.70420.figure5" httpUri="https://binary.pensoft.net/fig/595360" pageId="0" pageNumber="87" start="Figure 5" startId="F5">
|
||
<paragraph id="A25135B9CD2692B7AF62C695F6D6CEEC" pageId="0" pageNumber="87">
|
||
<emphasis id="2C700F083E8A167C1CA75E63931742B4" bold="true" pageId="0" pageNumber="87">Figure 5.</emphasis>
|
||
Bayesian inferenced tree of the genus
|
||
<taxonomicName id="1B6B32BCB8CA07A338F12C81A071FBA9" authorityName="Hoge & Romano-Hoge" authorityYear="1981" family="Viperidae" genus="Gloydius" kingdom="Animalia" lsidName="Gloydius" order="Squamata" pageId="0" pageNumber="87" rank="genus">
|
||
<emphasis id="5366116F4A197F177A089340DC90E370" italics="true" pageId="0" pageNumber="87">Gloydius</emphasis>
|
||
</taxonomicName>
|
||
, along with some relative genus of the family
|
||
<taxonomicName id="18C0346DB5A889BEBC8C323C74933532" class="Reptilia" family="Viperidae" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="family">Viperidae</taxonomicName>
|
||
, based on 12S, 16S, ND4, and cytb sequences, with the maximum likelihood bootstrap supports (left, regular) and Bayesian posterior probabilities (right, italic) displayed on the nodes (those <50% are displayed as
|
||
<normalizedToken id="237CA67FE430E71D58E3AD71D2094E8E" originalValue="“-”">"-"</normalizedToken>
|
||
). Holotypes are marked with asterisks.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |