184 lines
20 KiB
XML
184 lines
20 KiB
XML
<document id="FAE798255F25CE596E03A759D844973D" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. & Gordon, Dennis P." docDate="2018" docId="03EB8789FFE74467A1E177DDFD87FC12" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Collura Jullien 1882" docType="treatment" docVersion="3" lastPageNumber="1672" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of ‘ onychocellid’ cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1671" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="FFCBE7CE490A6B376D41C8BA89EFE9BC">
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<mods:title id="93CB23AF644748FD301BAEC8AD6065BB">Synopsis of ‘ onychocellid’ cheilostome bryozoan genera</mods:title>
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<mods:namePart id="92409DA67FE4FF8394E15638D2658973">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="9604B31ACA409C2B146ABE2693A4D043">Martha, Silviu O.</mods:namePart>
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<mods:namePart id="2A6509DBBF931257939C17230452C11E">Gordon, Dennis P.</mods:namePart>
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<mods:title id="FA3CC9D39A5D3F5872F060222708FDDA">Journal of Natural History</mods:title>
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<mods:part id="34445701ECAE80D494EA0BEA86000745">
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<mods:date id="A82D92D7F260E56B32CC93E662DB02A0">2018</mods:date>
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<mods:detail id="487BACCEBCB28F77C05B1AA2D996A4EE" type="series">
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<mods:title id="16F1D84BDA83CC81FC1FD3DF8C84F464">J. Nat. Hist.</mods:title>
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<mods:number id="A1253AC17957FDE7EF6F30E7810EC3A2">2018-07-03</mods:number>
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<mods:number id="8C89DC27616C8239ECC5AF3FD76010EE">52</mods:number>
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<mods:number id="3BD0E3E96159D3282A92B2A1F7E82A90">25 - 26</mods:number>
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<mods:url id="B0CEF94F3BC524635173BE0EA33DE932">http://dx.doi.org/10.1080/00222933.2018.1481235</mods:url>
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<subSubSection id="C3586514FFE74478A1E177DDFCA7FC0D" box="[493,821,922,947]" pageId="15" pageNumber="1671" type="nomenclature">
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<paragraph id="8BFD369FFFE74478A1E177DDFCA7FC0D" blockId="15.[493,821,922,982]" box="[493,821,922,947]" pageId="15" pageNumber="1671">
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<heading id="D0B581F3FFE74478A1E177DDFCA7FC0D" box="[493,821,922,947]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="1671" reason="2">
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Genus
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<taxonomicName id="4C424D1CFFE74478A24D77DDFCA7FC0D" ID-CoL="85BGM" authority="Jullien, 1882" authorityName="Jullien" authorityYear="1882" box="[577,821,922,947]" class="Gymnolaemata" family="Onychocellidae" genus="Collura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="1671" phylum="Bryozoa" rank="genus">
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<emphasis id="B936EA8DFFE74478A24D77DDFD0BFC0D" bold="true" box="[577,665,922,947]" italics="true" pageId="15" pageNumber="1671">Collura</emphasis>
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<bibRefCitation id="EFD34B6EFFE74478A2AE77DDFCA7FC0D" author="Jullien J" box="[674,821,922,947]" pageId="15" pageNumber="1671" pagination="271 - 285" refId="ref26644" refString="Jullien J. 1882. Note sur une nouvelle division des bryozoaires cheilostomiens. Bull Soc Zool Fr. 6: 271 - 285." type="journal article" year="1882">Jullien, 1882</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3586514FFE74478A25577FAFD59FC68" box="[601,715,957,982]" pageId="15" pageNumber="1671" type="description">
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||
<paragraph id="8BFD369FFFE74478A25577FAFD59FC68" blockId="15.[493,821,922,982]" box="[601,715,957,982]" pageId="15" pageNumber="1671">
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(
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<figureCitation id="13792A1AFFE74478A26D77FAFD50FC68" box="[609,706,957,982]" captionStart="Figure 6" captionStartId="15.[160,225,1677,1700]" captionTargetBox="[162,1152,1168,1646]" captionTargetId="figure-277@15.[162,1153,1168,1646]" captionTargetPageId="15" captionText="Figure 6. Collura athulia (d’Orbigny); lectotype, MNHN R61508; Cretaceous, Senonian (probably Maastrichtian), Néhou, Manche, Normandy, France. (a) Branch fragment; (b) autozooids with grooves around perimeter of the opesiae. Scale bars: a = 500 µm; b = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747155" httpUri="https://zenodo.org/record/4747155/files/figure.png" pageId="15" pageNumber="1671">Figure 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFE74478A0AC7044FB11FBDF" pageId="15" pageNumber="1671" type="type_taxon">
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<paragraph id="8BFD369FFFE74478A0AC7044FEA6FBA2" blockId="15.[160,1155,1027,1121]" box="[160,308,1027,1052]" pageId="15" pageNumber="1671">
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<emphasis id="B936EA8DFFE74478A0AC7044FEA6FBA2" bold="true" box="[160,308,1027,1052]" italics="true" pageId="15" pageNumber="1671">
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<typeStatus id="54F9883DFFE74478A0AC7044FF4AFBA2" box="[160,216,1027,1052]" pageId="15" pageNumber="1671">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFE74478A0AC7061FB11FBDF" blockId="15.[160,1155,1027,1121]" pageId="15" pageNumber="1671">
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<emphasis id="B936EA8DFFE74478A0AC7061FF61FB80" box="[160,243,1062,1086]" italics="true" pageId="15" pageNumber="1671">Eschara</emphasis>
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<taxonomicName id="4C424D1CFFE74478A0F77061FD93FB80" ID-CoL="86FWH" authority="d' Orbigny, 1851" authorityName="d' Orbigny" authorityYear="1851" box="[251,513,1062,1086]" class="Gymnolaemata" family="Onychocellidae" genus="Collura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="1671" phylum="Bryozoa" rank="species" species="athulia">
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<emphasis id="B936EA8DFFE74478A0F77061FED4FB80" box="[251,326,1062,1086]" italics="true" pageId="15" pageNumber="1671">athulia</emphasis>
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d’ Orbigny, 1851
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</taxonomicName>
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, by subsequent designation.
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<materialsCitation id="3B2A3CC2FFE74478A3447061FC9BFBDF" country="France" location="Normandy" pageId="15" pageNumber="1671" specimenCount="1" stateProvince="Haute-Normandie" typeStatus="holotype">
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Cretaceous, Senonian [probably Maastrichtian], Néhou, Manche,
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<collectingRegion id="4986F87DFFE74478A237700EFD20FBDF" box="[571,690,1097,1121]" country="France" name="Haute-Normandie" pageId="15" pageNumber="1671">Normandy</collectingRegion>
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,
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<collectingCountry id="F355760FFFE74478A2B1700EFC9BFBDF" box="[701,777,1097,1121]" name="France" pageId="15" pageNumber="1671">France</collectingCountry>
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</materialsCitation>
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.
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<bibRefCitation id="EFD34B6EFFE74478A318700EFC35FBDF" author="Jullien J" box="[788,935,1097,1121]" pageId="15" pageNumber="1671" pagination="271 - 285" refId="ref26644" refString="Jullien J. 1882. Note sur une nouvelle division des bryozoaires cheilostomiens. Bull Soc Zool Fr. 6: 271 - 285." type="journal article" year="1882">Jullien (1882)</bibRefCitation>
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listed three species
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF3D6617FFE74478A0AC72CAFCFEF961" ID-DOI="http://doi.org/10.5281/zenodo.4747155" ID-Zenodo-Dep="4747155" httpUri="https://zenodo.org/record/4747155/files/figure.png" pageId="15" pageNumber="1671" startId="15.[160,225,1677,1700]" targetBox="[162,1152,1168,1646]" targetPageId="15">
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||
<paragraph id="8BFD369FFFE74478A0AC72CAFCFEF961" blockId="15.[160,1155,1677,1759]" pageId="15" pageNumber="1671">
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||
<emphasis id="B936EA8DFFE74478A0AC72CAFE92F91A" bold="true" box="[160,256,1677,1700]" pageId="15" pageNumber="1671">Figure 6.</emphasis>
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||
<taxonomicName id="4C424D1CFFE74478A10572CAFE0EF91A" authorityName="d' Orbigny" authorityYear="1851" box="[265,412,1677,1700]" class="Gymnolaemata" family="Onychocellidae" genus="Collura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="1671" phylum="Bryozoa" rank="species" species="athulia">
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<emphasis id="B936EA8DFFE74478A10572CAFE0EF91A" box="[265,412,1677,1700]" italics="true" pageId="15" pageNumber="1671">Collura athulia</emphasis>
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</taxonomicName>
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(d
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<emphasis id="B936EA8DFFE74478A1B172CAFE50F91A" box="[445,450,1677,1700]" italics="true" pageId="15" pageNumber="1671">’</emphasis>
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Orbigny); lectotype, MNHN R61508; Cretaceous, Senonian (probably Maastrichtian), Néhou, Manche, Normandy, France. (a) Branch fragment; (b) autozooids with grooves around perimeter of the opesiae. Scale bars: a = 500 µm; b = 200 µm.
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</paragraph>
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</caption>
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<subSubSection id="C3586514FFF84467A0AC74C9FBE4FF77" pageId="16" pageNumber="1672" type="reference_group">
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||
<paragraph id="8BFD369FFFF84467A0AC74C9FBE4FF77" blockId="16.[160,1155,142,201]" pageId="16" pageNumber="1672">
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||
<treatmentCitationGroup id="AB5211B1FFF84467A0AC74C9FBE4FF77" pageId="16" pageNumber="1672">
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under
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<taxonomicName id="4C424D1CFFF84467A0EB74C9FEA1FF18" ID-CoL="85BGM" authorityName="Jullien" authorityYear="1882" box="[231,307,142,166]" class="Gymnolaemata" family="Onychocellidae" genus="Collura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="1672" phylum="Bryozoa" rank="genus">
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<emphasis id="B936EA8DFFF84467A0EB74C9FEA1FF18" box="[231,307,142,166]" italics="true" pageId="16" pageNumber="1672">Collura</emphasis>
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</taxonomicName>
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but failed to choose a
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<typeStatus id="54F9883DFFF84467A22074C9FDCFFF18" box="[556,605,142,166]" pageId="16" pageNumber="1672">type</typeStatus>
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||
species. The first of these three species was stated by
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<treatmentCitation id="0AE3108EFFF84467A0CE74F6FE07FF77" author="Bassler RS" box="[194,405,177,201]" page="77" pageId="16" pageNumber="1672" year="1935">
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||
<bibRefCitation id="EFD34B6EFFF84467A0CE74F6FEC0FF77" author="Bassler RS" box="[194,338,177,201]" pageId="16" pageNumber="1672" refId="ref23629" refString="Bassler RS. 1935. Bryozoa (generum et genotyporum index et bibliographia). Den Haag: W. Junk." type="book" year="1935">Bassler (1935</bibRefCitation>
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, p. 77
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</treatmentCitation>
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||
) to be the
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<typeStatus id="54F9883DFFF84467A20374F6FDD2FF77" box="[527,576,177,201]" pageId="16" pageNumber="1672">type</typeStatus>
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species, which may represent the first designation.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3586514FFF84467A0AC74BEFBECFE01" pageId="16" pageNumber="1672" type="diagnosis">
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<paragraph id="8BFD369FFFF84467A0AC74BEFE86FEAC" blockId="16.[160,1156,249,447]" box="[160,276,249,274]" pageId="16" pageNumber="1672">
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||
<emphasis id="B936EA8DFFF84467A0AC74BEFE86FEAC" bold="true" box="[160,276,249,274]" italics="true" pageId="16" pageNumber="1672">Diagnosis</emphasis>
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||
</paragraph>
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<paragraph id="8BFD369FFFF84467A0AC755AFBECFE01" blockId="16.[160,1156,249,447]" pageId="16" pageNumber="1672">
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Colony erect, vincularian (
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<figureCitation id="13792A1AFFF84467A1DC755AFDA3FE8B" box="[464,561,285,309]" captionStart="Figure 6" captionStartId="15.[160,225,1677,1700]" captionTargetBox="[162,1152,1168,1646]" captionTargetId="figure-277@15.[162,1153,1168,1646]" captionTargetPageId="15" captionText="Figure 6. Collura athulia (d’Orbigny); lectotype, MNHN R61508; Cretaceous, Senonian (probably Maastrichtian), Néhou, Manche, Normandy, France. (a) Branch fragment; (b) autozooids with grooves around perimeter of the opesiae. Scale bars: a = 500 µm; b = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747155" httpUri="https://zenodo.org/record/4747155/files/figure.png" pageId="16" pageNumber="1672">Figure 6</figureCitation>
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||
(a)). Autozooids subhexagonal to diamond shaped; zooidal boundaries broadly grooved due to voids in calcification around the proximal cryptocyst (
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||
<figureCitation id="13792A1AFFF84467A12A7525FE15FEC4" box="[294,391,354,378]" captionStart="Figure 6" captionStartId="15.[160,225,1677,1700]" captionTargetBox="[162,1152,1168,1646]" captionTargetId="figure-277@15.[162,1153,1168,1646]" captionTargetPageId="15" captionText="Figure 6. Collura athulia (d’Orbigny); lectotype, MNHN R61508; Cretaceous, Senonian (probably Maastrichtian), Néhou, Manche, Normandy, France. (a) Branch fragment; (b) autozooids with grooves around perimeter of the opesiae. Scale bars: a = 500 µm; b = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747155" httpUri="https://zenodo.org/record/4747155/files/figure.png" pageId="16" pageNumber="1672">Figure 6</figureCitation>
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(b)), the remaining cryptocyst having an inverted teardrop shape. Opesiae terminal, occupying about half of the frontal surface, longitudinally ovoidal, lacking opesiular indentations or lateral constrictions. Ovicells and avicularia unknown.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3586514FFF84467A0AC75A8FD87FC12" pageId="16" pageNumber="1672" type="discussion">
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||
<paragraph id="8BFD369FFFF84467A0AC75A8FE94FDB6" blockId="16.[160,1156,495,832]" box="[160,262,495,520]" pageId="16" pageNumber="1672">
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||
<emphasis id="B936EA8DFFF84467A0AC75A8FE94FDB6" bold="true" box="[160,262,495,520]" italics="true" pageId="16" pageNumber="1672">Remarks</emphasis>
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||
</paragraph>
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||
<paragraph id="8BFD369FFFF84467A0AC7654FBD4FCFE" blockId="16.[160,1156,495,832]" pageId="16" pageNumber="1672">
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||
Prud’
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<bibRefCitation id="EFD34B6EFFF84467A0D57654FEE7FD95" author="Homme J" box="[217,373,531,555]" pageId="16" pageNumber="1672" pagination="212 - 218" refId="ref27775" refString="Prud' Homme J. 1968. Revision de quelques types de la collection d' Orbigny (Cretace) conservee au Museum d' Histoire Naturelle de Paris. Atti Soc Ital Sci Nat Museo Civ Nat Milano. 108: 212 - 218." type="journal article" year="1968">Homme (1968</bibRefCitation>
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||
, p. 214, pl. 1, fig. 3) doubted the validity of this genus, commenting that d’ Orbigny’ s
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<typeStatus id="54F9883DFFF84467A1287671FEC7FDF0" box="[292,341,566,590]" pageId="16" pageNumber="1672">type</typeStatus>
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||
material of the
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<typeStatus id="54F9883DFFF84467A2047671FDABFDF0" box="[520,569,566,590]" pageId="16" pageNumber="1672">type</typeStatus>
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species was ‘represented by a fragment of colony in which the particular form of the zooids appears to result from an alteration identical to that which often affects colonies of
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<emphasis id="B936EA8DFFF84467A1FE763CFDD7FD2D" box="[498,581,635,659]" italics="true" pageId="16" pageNumber="1672">Eschara</emphasis>
|
||
[
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||
<taxonomicName id="4C424D1CFFF84467A25E763CFD46FD2D" authorityName="Jullien" authorityYear="1882" box="[594,724,635,659]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="1672" phylum="Bryozoa" rank="genus">
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||
<emphasis id="B936EA8DFFF84467A25E763CFD46FD2D" box="[594,724,635,659]" italics="true" pageId="16" pageNumber="1672">Onychocella</emphasis>
|
||
</taxonomicName>
|
||
]
|
||
<emphasis id="B936EA8DFFF84467A2ED763CFCBEFD2D" box="[737,812,635,659]" italics="true" pageId="16" pageNumber="1672">royana</emphasis>
|
||
, among others’. She appears to have been referring to the voids in the cryptocyst, to judge from subsequent scanning electron microscopy of this material (
|
||
<figureCitation id="13792A1AFFF84467A24B7687FD31FD66" box="[583,675,704,728]" captionStart="Figure 6" captionStartId="15.[160,225,1677,1700]" captionTargetBox="[162,1152,1168,1646]" captionTargetId="figure-277@15.[162,1153,1168,1646]" captionTargetPageId="15" captionText="Figure 6. Collura athulia (d’Orbigny); lectotype, MNHN R61508; Cretaceous, Senonian (probably Maastrichtian), Néhou, Manche, Normandy, France. (a) Branch fragment; (b) autozooids with grooves around perimeter of the opesiae. Scale bars: a = 500 µm; b = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747155" httpUri="https://zenodo.org/record/4747155/files/figure.png" pageId="16" pageNumber="1672">Figure 6</figureCitation>
|
||
). The genus
|
||
<taxonomicName id="4C424D1CFFF84467A3357687FC17FD66" authorityName="Jullien" authorityYear="1882" box="[825,901,704,728]" class="Gymnolaemata" family="Onychocellidae" genus="Collura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="1672" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFF84467A3357687FC17FD66" box="[825,901,704,728]" italics="true" pageId="16" pageNumber="1672">Collura</emphasis>
|
||
</taxonomicName>
|
||
has fallen into disuse, although it was listed as a valid genus of
|
||
<taxonomicName id="4C424D1CFFF84467A26C76A4FC9BFD45" authorityName="Jullien" authorityYear="1882" box="[608,777,739,763]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="1672" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFD34B6EFFF84467A33C76A4FBD3FD45" author="Gordon DP & Taylor PD" box="[816,1089,739,763]" pageId="16" pageNumber="1672" pagination="83 - 97" refId="ref26071" refString="Gordon DP, Taylor PD 2005. The cheilostomatous genera of Alcide d' Orbigny - nomenclatural and taxonomic status. In: Moyano HI, Wyse Jackson PN, editors. Bryozoan Studies 2004. Proceedings of the 13 th International Bryozoology Association conference, Concepcion / Chile; January 11 - 16 2004; p. 83 - 97. London: CRC Press." type="journal article" year="2005">Gordon and Taylor (2005</bibRefCitation>
|
||
, table 2). The absence of avicularia in the
|
||
<typeStatus id="54F9883DFFF84467A2387742FDF7FCA3" box="[564,613,773,797]" pageId="16" pageNumber="1672">type</typeStatus>
|
||
material, combined with the lack of conspecific colonies that might have these polymorphs, hampers a clear definition of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFF84467A0AC7737FF79FC37" blockId="16.[160,533,880,940]" box="[160,235,880,905]" pageId="16" pageNumber="1672">
|
||
<emphasis id="B936EA8DFFF84467A0AC7737FF79FC37" bold="true" box="[160,235,880,905]" italics="true" pageId="16" pageNumber="1672">Range</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFF84467A0AC77D3FD87FC12" blockId="16.[160,533,880,940]" box="[160,533,916,940]" pageId="16" pageNumber="1672">Late Cretaceous [?Maastrichtian].</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |