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<document id="C240801D6C942A76E732C189AB7E0DB7" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFEA7D70ACE4D63EFE689B9C" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus dorsalis Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="EDC8D353-BE43-4D79-8BA4-E23D0FD57CE7" docUuidSource="ZooBank" docVersion="8" lastPageNumber="25" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="22" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C3AAB0714E5D84DAB4049C7E5493A413">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="E1EB04D06538F14734CB85B5F195C1B5">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="7C4A6CE45C4C773587DBBA6668943449">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="454534C936A3673A92D94C97A319A716">Golovatch, Sergei I.</mods:namePart>
<mods:nameIdentifier id="F7776570B543C4AD9BDF47AAE66B59A6" type="LSID">71532F45-BDD5-415D-BC54-86256E5D5D4A</mods:nameIdentifier>
<mods:affiliation id="CAC4291010C7FF4F6EFBC3B0D27C720A">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="4A011545958454B291DC0F7CDDFB0709" type="email">sgolovatch@yandex.ru&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="5B243801394CF9EB2D9B228FE744AB11">Panha, Somsak</mods:namePart>
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<mods:affiliation id="3F0BFEB10F52FA1540CA45AA952874A4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:title id="6198072D1D2FA58A8840A9904B8977C4">European Journal of Taxonomy</mods:title>
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<mods:date id="B0015FD248D1E39F033D19DD0E88B5E6">2016</mods:date>
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<mods:number id="65264BA22B1CE0003E3354CD855D4B98">2016-05-10</mods:number>
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<treatment id="03EB5A00FFEA7D70ACE4D63EFE689B9C" ID-DOI="http://doi.org/10.5281/zenodo.3852408" ID-GBIF-Taxon="164246091" ID-Zenodo-Dep="3852408" LSID="urn:lsid:zoobank.org:act:EDC8D353-BE43-4D79-8BA4-E23D0FD57CE7" httpUri="http://treatment.plazi.org/id/03EB5A00FFEA7D70ACE4D63EFE689B9C" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="C358B89DFFEA7D7DACE4D63EFB1E9A89" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BFDEB16FFEA7D7DACE4D63EFC7F9AAD" blockId="21.[356,1231,623,720]" box="[646,942,623,650]" pageId="21" pageNumber="22">
<heading id="D0B55C7AFFEA7D7DACE4D63EFC7F9AAD" box="[646,942,623,650]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="4C429095FFEA7D7DACE4D63EFC9A9AAE" ID-CoL="7CZN4" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[646,843,623,650]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="dorsalis" status="sp. nov.">
<emphasis id="B9363704FFEA7D7DACE4D63EFC9A9AAE" bold="true" box="[646,843,623,650]" italics="true" pageId="21" pageNumber="22">Tylopus dorsalis</emphasis>
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<taxonomicNameLabel id="A2058A7FFFEA7D7DAD30D621FC7F9AAD" box="[850,942,624,650]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C358B89DFFEA7D7DACA0D6E7FCA09AF4" box="[706,881,694,720]" pageId="21" pageNumber="22" type="description">
<paragraph id="8BFDEB16FFEA7D7DACA0D6E7FCA09AF4" blockId="21.[356,1231,623,720]" box="[706,881,694,720]" pageId="21" pageNumber="22">
<figureCitation id="1379F793FFEA7D7DACA0D6E7FC969AF4" box="[706,839,694,720]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="22.[189,232,1893,1919]" captionStartId-1="23.[189,232,1964,1990]" captionTargetBox-0="[189,1398,592,1841]" captionTargetBox-1="[314,1249,559,1910]" captionTargetId-0="figure@22.[189,1398,592,1841]" captionTargetId-1="figure@23.[310,1255,559,1920]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." captionText-1="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi-0="http://doi.org/10.5281/zenodo.3837907" figureDoi-1="http://doi.org/10.5281/zenodo.3837909" httpUri-0="https://zenodo.org/record/3837907/files/figure.png" httpUri-1="https://zenodo.org/record/3837909/files/figure.png" pageId="21" pageNumber="22">
Figs 16
<emphasis id="B9363704FFEA7D7DAD7BD6E7FCF99AF4" bold="true" box="[793,808,694,720]" pageId="21" pageNumber="22">–</emphasis>
17
</figureCitation>
,
<figureCitation id="1379F793FFEA7D7DAD36D6E7FCA09AF4" box="[852,881,694,720]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="21" pageNumber="22">26</figureCitation>
</paragraph>
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<subSubSection id="C358B89DFFEA7D7DAEDFD6A6FD879B8D" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BFDEB16FFEA7D7DAEDFD6A6FEE79B35" blockId="21.[189,310,759,785]" box="[189,310,759,785]" pageId="21" pageNumber="22">
<heading id="D0B55C7AFFEA7D7DAEDFD6A6FEE79B35" bold="true" box="[189,310,759,785]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9363704FFEA7D7DAEDFD6A6FEE79B35" bold="true" box="[189,310,759,785]" pageId="21" pageNumber="22">Diagnosis</emphasis>
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The live coloration is similar to that of
<taxonomicName id="4C429095FFEA7D7DACECD776FCC69B64" authorityName="Likhitrakarn, Golovatch &amp; Panha" authorityYear="2014" box="[654,791,807,832]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="B9363704FFEA7D7DACECD776FCC69B64" box="[654,791,807,832]" italics="true" pageId="21" pageNumber="22">T. trigonum</emphasis>
</taxonomicName>
, but the new species differs in the head and collum being light brown (versus blackish), and the pattern not contrasting (versus contrasting); the gonopod structure is similar to that of
<taxonomicName id="4C429095FFEA7D7DAC68D73CFC8F9BA2" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[522,862,876,902]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B9363704FFEA7D7DAC68D73CFDF19BA2" box="[522,544,877,902]" italics="true" pageId="21" pageNumber="22">T.</emphasis>
<emphasis id="B9363704FFEA7D7DAC44D73CFD429BA2" box="[550,659,877,902]" italics="true" pageId="21" pageNumber="22">crassipes</emphasis>
<bibRefCitation id="EFD396E7FFEA7D7DACF8D73DFC8F9BA2" author="Golovatch S. I." box="[666,862,876,902]" pageId="21" pageNumber="22" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
, but the new species differs in process
<emphasis id="B9363704FFEA7D7DAB42D73DFAFE9BA2" bold="true" box="[1312,1327,876,902]" pageId="21" pageNumber="22">x</emphasis>
being longer and its tip rounded dorsally.
</paragraph>
</subSubSection>
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<heading id="D0B55C7AFFEA7D7DAEDFD780FE959BCF" bold="true" box="[189,324,977,1003]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9363704FFEA7D7DAEDFD780FE959BCF" bold="true" box="[189,324,977,1003]" pageId="21" pageNumber="22">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFEA7D7DAEDFD051FC289C3D" blockId="21.[189,1017,1023,1050]" box="[189,1017,1023,1050]" pageId="21" pageNumber="22">
To emphasize process
<emphasis id="B9363704FFEA7D7DAFA7D7AEFE059C3D" bold="true" box="[453,468,1023,1049]" pageId="21" pageNumber="22">x</emphasis>
on the solenophore arising dorsally; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFEA7D7DAEDFD010FD269D10" pageId="21" pageNumber="22" type="materials_examined">
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<heading id="D0B55C7AFFEA7D7DAEDFD010FE7B9C7F" bold="true" box="[189,426,1089,1115]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9363704FFEA7D7DAEDFD010FE7B9C7F" bold="true" box="[189,426,1089,1115]" pageId="21" pageNumber="22">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B2AE14BFFEA7D7DAE88D03EFBD39CF4" ID-GBIF-Occurrence="2621711303" collectingDate="2014-10-16" collectionCode="R, CUMZ" collectorName="C. Sutcharit &amp; Srisonchai" country="Laos" elevation="1125" latitude="21.625" location="Boun Neua" longLatPrecision="21" longitude="101.93194" municipality="Phuthalang forest" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Phongsali" typeStatus="holotype">
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<emphasis id="B9363704FFEA7D7DAE88D03EFE8D9CAD" bold="true" box="[234,348,1135,1161]" pageId="21" pageNumber="22">Holotype</emphasis>
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: Ƌ,
<collectingMunicipality id="6B99716CFFEA7D7DAF24D0C3FDC99C88" box="[326,536,1170,1196]" pageId="21" pageNumber="22">Phuthalang forest</collectingMunicipality>
protected area,
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,
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,
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21°37'30&quot;
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,
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101°55'55&quot;
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, ca
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a.s.l.
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,
<collectingDate id="EFB8343EFFEA7D7DAF03D0E7FE2C9CF4" box="[353,509,1206,1232]" pageId="21" pageNumber="22" value="2014-10-16">
<date id="FFFCCDD6FFEA7D7DAF03D0E7FE2C9CF4" box="[353,509,1206,1232]" pageId="21" pageNumber="22" value="2014-10-16">16 Oct. 2014</date>
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, leg.
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. Sutcharit
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&amp;
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.
<collectorName id="26B78EC0FFEA7D7DAD6DD0E7FC5A9CF4" box="[783,907,1206,1232]" pageId="21" pageNumber="22">Srisonchai</collectorName>
(
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).
</paragraph>
</materialsCitation>
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<emphasis id="B9363704FFEA7D7DAE88D0A6FE8D9D35" bold="true" box="[234,348,1271,1297]" pageId="21" pageNumber="22">Paratype</emphasis>
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</paragraph>
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<collectingCountry id="F355AB86FFEA7D7DAEDFD14BFEDF9D10" box="[189,270,1306,1332]" name="Laos" pageId="21" pageNumber="22">
<collectionCode id="ED5373D3FFEA7D7DAEDFD14BFEDF9D10" box="[189,270,1306,1332]" pageId="21" pageNumber="22">LAOS</collectionCode>
</collectingCountry>
:
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, same data as for
<typeStatus id="54F955B4FFEA7D7DAC7BD14BFD519D10" box="[537,640,1306,1332]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
(
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).
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFEA7D70AEDFD10DFE689B9C" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22" type="description">
<paragraph id="8BFDEB16FFEA7D7DAEDFD10DFE9F9D52" blockId="21.[189,334,1372,1398]" box="[189,334,1372,1398]" pageId="21" pageNumber="22">
<heading id="D0B55C7AFFEA7D7DAEDFD10DFE9F9D52" bold="true" box="[189,334,1372,1398]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9363704FFEA7D7DAEDFD10DFE9F9D52" bold="true" box="[189,334,1372,1398]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFEA7D7DAEDFD1DAFCBB9EBF" blockId="21.[189,1399,1418,1691]" pageId="21" pageNumber="22">
MEASUREMENTS AND COLOUR. Length 22.1 (Ƌ) or
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(
<specimenCount id="9D44209FFFEA7D7DAD1FD1DBFC419D80" box="[893,912,1418,1444]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
), width of midbody pro- and metazonae 2.5 and
<quantity id="4CBA46F3FFEA7D7DAF42D1FCFEAC9DEC" box="[288,381,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="21" pageNumber="22" unit="mm" value="3.6">3.6 mm</quantity>
(Ƌ) or 1.97 and
<quantity id="4CBA46F3FFEA7D7DAC2CD1FCFD689DEC" box="[590,697,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="21" pageNumber="22" unit="mm" value="2.55">2.55 mm</quantity>
(
<specimenCount id="9D44209FFFEA7D7DACACD1FFFD309DEC" box="[718,737,1454,1480]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
), respectively. Live coloration light brown (
<figureCitation id="1379F793FFEA7D7DAA9BD1FCFAB99DEC" box="[1273,1384,1453,1480]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
); paraterga, legs and epiproct light yellow-brown, head and collum light brown, antennomere 7 blackish, collum and following terga each with a light brown triangle and a dark brown collar covering both pro- and metazonae. Coloration of alcohol material faded after one year of preservation to light brown, with a pattern of a dark brown band in pre-sulcus area on metazonae and a pair of paramedian, longitudinal, brown stripes dorsally both on pro- and metaterga; paraterga light red-brown, head brown, antennae, legs, venter and epiproct light yellow-brown (
<figureCitation id="1379F793FFEA7D7DACB4D2D1FCFB9EBF" box="[726,810,1664,1691]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="21" pageNumber="22">Fig. 16</figureCitation>
B–J).
</paragraph>
<paragraph id="8BFDEB16FFEA7D7DAEDFD292FCC69FA8" blockId="21.[189,1399,1730,1933]" pageId="21" pageNumber="22">
HEAD. Clypeolabral region densely, vertex sparsely, setose; epicranial suture distinct. Antennae rather short (
<figureCitation id="1379F793FFEA7D7DAF65D2B4FEA29F24" box="[263,371,1765,1792]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
), extending behind body segment 3 (Ƌ) or 2 (
<specimenCount id="9D44209FFFEA7D7DADEAD2B4FC4A9EDB" box="[904,923,1765,1791]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
) when stretched dorsally. In width, head &lt;segment 3 &lt;2 = 4 &lt;collum &lt;segments 5–17 (Ƌ,
<specimenCount id="9D44209FFFEA7D7DAD4AD358FCEC9F07" box="[808,829,1801,1827]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 3+3 anterior, 1+1 intermediate and 2+2 posterior; a small lateral denticle at about anterior 1/3 (
<figureCitation id="1379F793FFEA7D7DAC1BD31EFCDF9F4D" box="[633,782,1871,1897]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="21" pageNumber="22">Fig. 16B, D</figureCitation>
); caudal corner of paraterga very broadly rounded, declined ventrad, not surpassing rear tergal margin.
</paragraph>
<paragraph id="8BFDEB16FFE97D70AEDFD55DFB2099DD" blockId="22.[189,1399,267,540]" lastBlockId="24.[189,1399,267,505]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
BODY. Tegument smooth and shining, prozonae finely shagreened, metaterga smooth and leathery, posterior halves rugulose, surface below paraterga microgranulate (
<figureCitation id="1379F793FFE97D7EADA3D57FFBC0996C" box="[961,1041,302,328]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="22" pageNumber="23">Fig. 16</figureCitation>
B–F, H). Postcollum metaterga with two transverse rows of setae: 2+
<quantity id="4CBA46F3FFE97D7EAC1BD503FD769948" box="[633,679,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="22" pageNumber="23" unit="in" value="2.0">2 in</quantity>
anterior (pre-sulcus) row, also 2+
<quantity id="4CBA46F3FFE97D7EAA5AD503FBB79948" box="[1080,1126,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="22" pageNumber="23" unit="in" value="2.0">2 in</quantity>
posterior (post-sulcus) one. Tergal setae long, strong, slender, about 2/3 of metatergal length.Axial line traceable both on pro- and metazonae. Paraterga well-developed (
<figureCitation id="1379F793FFE97D7EACE0D5C9FCFC9997" box="[642,813,408,435]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="22" pageNumber="23">Fig. 16B, D, F</figureCitation>
), especially so in Ƌ, set rather high (at upper 1/3 of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, extending increasingly behind rear tergal margin, posterior edge slightly oblique. Calluses on paraterga delimited by a sulcus only dorsally. Paratega 2 broad, anterior edge angular, lateral edge with three evident incisions. Lateral edge of following paraterga with a clear incision in anterior 1/3 (
<figureCitation id="1379F793FFE87D7FAB6AD55DFF07996E" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="23" pageNumber="24">Fig. 16B, D</figureCitation>
). Paraterga 14–19 with caudal corner extending increasingly behind, with tip evidently curved mesad (
<figureCitation id="1379F793FFE87D7FAEA4D502FEFF994A" box="[198,302,339,366]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="23" pageNumber="24">Fig. 16F</figureCitation>
). Ozopores distinct, lateral, lying in an ovoid groove at about 1/3 of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (
<figureCitation id="1379F793FFE87D7FADACD527FBAC99B4" box="[974,1149,374,400]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="23" pageNumber="24">Fig. 16B, D, F</figureCitation>
), slightly incomplete on segment 3, complete on segments 4–18, narrow, line-shaped, deep, not reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazona evident, wide, rather deep, clearly ribbed at bottom down to base of paraterga (
<figureCitation id="1379F793FFE87D7FAC3ED58EFCC099DE" box="[604,785,479,506]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="23" pageNumber="24">
Fig. 16B, D
<emphasis id="B9363704FFE87D7FAC8BD5B1FD2999DE" bold="true" box="[745,760,480,506]" pageId="23" pageNumber="24">–</emphasis>
G
</figureCitation>
). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2–7, reduced and remaining only a sharp caudal tooth on segment 8, reduced to a small, delicately denticulate, caudal crest on segments 9–15, retained as a minute tooth on segment 16, thereafter missing (Ƌ), or with complete crests and a sharp caudal tooth on segments 2–4, remaining only an evident caudal tooth on segments 5–12, a rounded caudal crest on segments 13–17, thereafter missing (
<specimenCount id="9D44209FFFE77D70AF48D5C9FEEC9996" box="[298,317,408,434]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) (
<figureCitation id="1379F793FFE77D70AF3AD5C9FDD79997" box="[344,518,408,435]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="24" pageNumber="25">Fig. 16C, E, H</figureCitation>
). Epiproct (
<figureCitation id="1379F793FFE77D70ACECD5C9FD0E9997" box="[654,735,408,435]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="24" pageNumber="25">Fig. 16</figureCitation>
E–G) conical, flattened dorsoventrally, with two evident, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae evident, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (
<figureCitation id="1379F793FFE77D70AA14D58FFB3599DD" box="[1142,1252,478,505]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="24" pageNumber="25">Fig. 16G</figureCitation>
).
</paragraph>
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<paragraph id="8BFDEB16FFE97D7EAEDFD334FCDD9FCD" blockId="22.[189,1399,1893,2025]" pageId="22" pageNumber="23">
<emphasis id="B9363704FFE97D7EAEDFD334FEC89F5B" bold="true" box="[189,281,1893,1919]" pageId="22" pageNumber="23">Fig. 16.</emphasis>
<taxonomicName id="4C429095FFE97D7EAF42D334FE339F5B" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[288,482,1893,1919]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dorsalis" status="sp. nov.">
<emphasis id="B9363704FFE97D7EAF42D334FE339F5B" box="[288,482,1893,1919]" italics="true" pageId="22" pageNumber="23">Tylopus dorsalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE97D7EAF8AD337FD979FA4" box="[488,582,1894,1920]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE97D7EAC89D334FCD19F5B" bold="true" box="[747,768,1893,1919]" pageId="22" pageNumber="23">A</emphasis>
. Habitus, live coloration.
<emphasis id="B9363704FFE97D7EAA50D334FBB89F5B" bold="true" box="[1074,1129,1893,1919]" pageId="22" pageNumber="23">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE97D7EAC1FD3D8FD659F87" bold="true" box="[637,692,1929,1955]" pageId="22" pageNumber="23">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE97D7EAEDFD3FDFF259FE2" bold="true" box="[189,244,1964,1990]" pageId="22" pageNumber="23">F–H</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively.
<emphasis id="B9363704FFE97D7EAA55D3FDFBB19FE2" bold="true" box="[1079,1120,1964,1990]" pageId="22" pageNumber="23">I–J</emphasis>
. Sternal cones between coxae 4, caudal and sublateral views, respectively.
</paragraph>
</caption>
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<paragraph id="8BFDEB16FFE87D7FAEDFD3FDFBA69FCE" blockId="23.[189,1399,1964,2026]" pageId="23" pageNumber="24">
<emphasis id="B9363704FFE87D7FAEDFD3FDFECD9FE2" bold="true" box="[189,284,1964,1990]" pageId="23" pageNumber="24">Fig. 17.</emphasis>
<taxonomicName id="4C429095FFE87D7FAF44D3FDFE389FE2" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[294,489,1964,1990]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dorsalis" status="sp. nov.">
<emphasis id="B9363704FFE87D7FAF44D3FDFE389FE2" box="[294,489,1964,1990]" italics="true" pageId="23" pageNumber="24">Tylopus dorsalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE87D7FAF91D3FDFD829FE2" box="[499,595,1964,1990]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE87D7FAD62D3FDFCE89FE2" bold="true" box="[768,825,1964,1990]" pageId="23" pageNumber="24">A–D</emphasis>
. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE77D70AEDFD671FA8A9ACE" blockId="24.[189,1399,544,746]" pageId="24" pageNumber="25">
STERNA. Separsely setose, without modifications; cross-impressions shallow; an entire, linguiform, sternal lobe between Ƌ coxae 4 (
<figureCitation id="1379F793FFE77D70AC2BD612FD179A7A" box="[585,710,579,606]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="24" pageNumber="25">
Fig.
<date id="FFFCCDD6FFE77D70ACE0D612FD789A79" box="[642,681,579,605]" pageId="24" pageNumber="25" value="1916-01">16I</date>
<emphasis id="B9363704FFE77D70ACCBD612FD699A79" bold="true" box="[681,696,579,605]" pageId="24" pageNumber="25">–</emphasis>
J
</figureCitation>
). Legs long and slender, midbody ones ca 1.3–1.4 (Ƌ) or 0.9–1.1 (
<specimenCount id="9D44209FFFE77D70AF48D637FEEC9AA4" box="[298,317,614,640]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) as long as body height; Ƌ femora 6 bulged and microtuberculate ventrally (
<figureCitation id="1379F793FFE77D70AABFD637FA989AA4" box="[1245,1353,614,640]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="24" pageNumber="25">Fig. 17E</figureCitation>
); Ƌ femora 7 also bulged ventrally (
<figureCitation id="1379F793FFE77D70AC23D6D8FD7A9A80" box="[577,683,649,676]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="24" pageNumber="25">Fig. 17F</figureCitation>
), but following femora not swollen; legs of segments 7–16 each with femur, postfemur, tibia and tarsus carrying a small adenostyle (tubercle) medially (
<figureCitation id="1379F793FFE77D70AA9ED6FCFAB99AE3" box="[1276,1384,685,711]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="24" pageNumber="25">Fig. 17G</figureCitation>
), telopodites particularly densely setose ventrally (
<figureCitation id="1379F793FFE77D70AC9ED681FCB99ACE" box="[764,872,720,746]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="24" pageNumber="25">Fig. 17G</figureCitation>
), tarsal brushes present until Ƌ segment 6.
</paragraph>
<paragraph id="8BFDEB16FFE77D70AEDFD743FE689B9C" blockId="24.[189,1399,785,952]" pageId="24" pageNumber="25">
GONOPOD. Rather simple (
<figureCitation id="1379F793FFE77D70AF9AD740FD9E9B08" box="[504,591,785,812]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. A–D. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="24" pageNumber="25">Fig. 17</figureCitation>
A–D); coxa slightly curved caudad, sparsely setose distoventrally. Femorite slightly curved and stout, with an evident mesal groove and a clear distolateral sulcus demarcating a postfemoral part; process
<emphasis id="B9363704FFE77D70ACC6D709FD659B56" bold="true" box="[676,692,856,882]" pageId="24" pageNumber="25">h</emphasis>
rather high, directed anterodorsally, flattened, tip rounded; solenophore (
<emphasis id="B9363704FFE77D70AF3ED72AFE599BB1" bold="true" box="[348,392,891,917]" pageId="24" pageNumber="25">sph</emphasis>
) rather slender, strongly coiled, flattened and evidently bifid, with process
<emphasis id="B9363704FFE77D70AA8AD72AFB269BB1" bold="true" box="[1256,1271,891,917]" pageId="24" pageNumber="25">x</emphasis>
being slim and directed dorsally.
</paragraph>
</subSubSection>
</treatment>
</document>
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