treatments-xml/data/03/EB/5A/03EB5A00FFD97D44AEDFD55AFAA69E62.xml
2024-06-21 12:22:17 +02:00

1462 lines
185 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="847DF903F21FC9FD174DC9085584FA5F" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFD97D44AEDFD55AFAA69E62" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus Jeekel 1968" docType="key" docVersion="7" lastPageNumber="45" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="39" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="45856CFADB94155ED92ED2D5C9365753" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CBDA61C6B3FCDDA98B35F43BF8F1D445">
<mods:title id="E4CB32E24FCC014077FAA2748D8C75D2">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
</mods:titleInfo>
<mods:name id="6E16A2E997B006D2E9D6E9B8E4371004" type="personal">
<mods:role id="6FDFE7EBAC8FF2E27D6FF2E507360867">
<mods:roleTerm id="8FA1B17856872DE5300D28EF203142EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D29F1D0D41907283BEFF3CF83672D863">Likhitrakarn, Natdanai</mods:namePart>
<mods:nameIdentifier id="2DA8A3FE83875BD1A85F822A586C995D" type="LSID">442E2E6E-1CB3-47B2-9896-1A922CE509E1</mods:nameIdentifier>
<mods:affiliation id="8F34ABFEC8CC939A00F95CC3055F0CCA">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
<mods:nameIdentifier id="E82489E94DC3D1055E24C9E1C9078DAA" type="email">kongerrrr@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="36AA747F7CDBD02245D2580880896794" type="personal">
<mods:role id="BE2C1B0DA1ED2B42F227426CAD2F55FD">
<mods:roleTerm id="662B5259A512871EB95CC3798E8C9671">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B16C90472849D32315B91D207B28D23B">Golovatch, Sergei I.</mods:namePart>
<mods:nameIdentifier id="DDFFBCF3B48177741CF447E713C88015" type="LSID">71532F45-BDD5-415D-BC54-86256E5D5D4A</mods:nameIdentifier>
<mods:affiliation id="85DD3653DBEE2A753139B3C87AEE02F3">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="8734C19158B9CFF8D9A5084EAF2DB701" type="email">sgolovatch@yandex.ru&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name id="12A68BB1385F344AFB0D5B8A8468524E" type="personal">
<mods:role id="7D0049341573ADCC1D3BE9C97D187ADC">
<mods:roleTerm id="D93CFAC33A74AFD50415F228F23137FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6ED4DB254A0221F9A74CFE6A8D44F140">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="D7AD80FEB199DF077E8A86AD15F2013C" type="LSID">AC935098-D901-4F35-A414-4B0D4FE44E79</mods:nameIdentifier>
<mods:affiliation id="F0274D1ABD9B174E2B3893F2F8D17F1C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="CB22D4489E6071F5C459BE77BB54C5BC" type="email">somsak.pan@chula.ac.th&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8AF012050F76B1896C17A774C55D6B8B">text</mods:typeOfResource>
<mods:relatedItem id="5796D77A9B6B4FFB61EDD7AA8681D2B4" type="host">
<mods:titleInfo id="A9D5188BA7B8FEBC77A2BED602323537">
<mods:title id="C9C36F038DA437A9AA6F6A033B4114DB">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="AF136A13063E13FBA5096939CC884341">
<mods:date id="683A06AE18CA6E6F1A754715399E4A72">2016</mods:date>
<mods:detail id="CD14FB848C32EE4D3CC3E74127DCDAAA" type="pubDate">
<mods:number id="88BC5992955BF9130E311AE0BD2B064F">2016-05-10</mods:number>
</mods:detail>
<mods:detail id="75BB60A21D7BB0DFEE4CBC4AD757813E" type="volume">
<mods:number id="DEBD4D1E84C215E39006ADD8ABD5CCED">195</mods:number>
</mods:detail>
<mods:extent id="7568531236FA197D78DE2975A54D3707" unit="page">
<mods:start id="B81DF050C0E9F1B64499E9460148D480">1</mods:start>
<mods:end id="0EA53DFD56A244502C18D3A2C5D12F45">47</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1D1AF00BE411D0A3B956FF1523F1310C">journal article</mods:classification>
<mods:identifier id="AB963F5CD26017DE259C154B4157F1E3" type="CLB-Dataset">21966</mods:identifier>
<mods:identifier id="08701195A60D23899A6655FBB089757F" type="DOI">10.5852/ejt.2016.195</mods:identifier>
<mods:identifier id="FA114FE57AD6F24C709FBD4394720E77" type="GBIF-Dataset">39071d6f-d900-4c81-ace1-dd03875018c5</mods:identifier>
<mods:identifier id="2CAF84362BC2435595191B0B893912D5" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="8770829EA6EF70EB6C723B58F19DC9C5" type="Zenodo-Dep">3837846</mods:identifier>
<mods:identifier id="FBC71AFC8603CB09EBC999AC3555FE37" type="ZooBank">4EEA9AD1-5762-4A93-A189-CF185F64CBAF</mods:identifier>
</mods:mods>
<subSection id="E2CDF2FDFFD97D44AEDFD55AFAA69E62" lastPageId="44" lastPageNumber="45" pageId="38" pageNumber="39" type="key">
<treatment id="03EB5A00FFD97D44AEDFD55AFAA69E62" ID-DOI="http://doi.org/10.5281/zenodo.3852402" ID-GBIF-Taxon="164246085" ID-Zenodo-Dep="3852402" LSID="urn:lsid:plazi:treatment:03EB5A00FFD97D44AEDFD55AFAA69E62" httpUri="http://treatment.plazi.org/id/03EB5A00FFD97D44AEDFD55AFAA69E62" lastPageId="44" lastPageNumber="45" pageId="38" pageNumber="39">
<subSubSection id="C358B89DFFD97D4EAEDFD55AFAA69901" box="[189,1399,266,293]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BFDEB16FFD97D4EAEDFD55AFAA69901" blockId="38.[189,1399,266,293]" box="[189,1399,266,293]" pageId="38" pageNumber="39">
<heading id="D0B55C7AFFD97D4EAEDFD55AFAA69901" box="[189,1399,266,293]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<emphasis id="B9363704FFD97D4EAEDFD55AFAA69901" bold="true" box="[189,1399,266,293]" pageId="38" pageNumber="39">
Key to the known species of
<taxonomicName id="4C429095FFD97D4EAC75D55BFDA79901" ID-CoL="7Q5LS" authorityName="Jeekel" authorityYear="1968" box="[535,630,266,293]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9363704FFD97D4EAC75D55BFDA79901" bold="true" box="[535,630,266,293]" italics="true" pageId="38" pageNumber="39">Tylopus</emphasis>
</taxonomicName>
(based chiefly on Ƌ), modified after
<bibRefCitation id="EFD396E7FFD97D4EAA51D55AFABD9901" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[1075,1388,266,293]" pageId="38" pageNumber="39" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">
Likhitrakarn
<emphasis id="B9363704FFD97D4EAABDD55AFAC09901" bold="true" box="[1247,1297,266,293]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. (2010)
</bibRefCitation>
:
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFD97D44AEDFD56BFAA69E62" lastPageId="44" lastPageNumber="45" pageId="38" pageNumber="39" type="key">
<key id="FFD3BC27FFD97D44AEDFD56BFAA69E62" lastPageId="44" lastPageNumber="45" pageId="38" pageNumber="39">
<paragraph id="8BFDEB16FFD97D4EAEDFD56BFAA69953" blockId="38.[189,1399,313,375]" pageId="38" pageNumber="39">
<keyStep id="30B6F3B3FFD97D4EAEDFD56BFAA69953" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEDFD56BFAA69953" pageId="38" pageNumber="39">
1. Most of Ƌ prefemora evidently swollen laterally (
<figureCitation id="1379F793FFD97D4EAD3AD568FC1D9970" box="[856,972,313,340]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[308,1241,1126,1855]" captionTargetId="figure@18.[302,1247,1119,1870]" captionTargetPageId="18" captionText="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. AB. Lateral and mesal views, respectively. C. Leg 13. Scale bar: AB. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="38" pageNumber="39">Figs 13C</figureCitation>
,
<figureCitation id="1379F793FFD97D4EADBCD56BFBC19970" box="[990,1040,314,340]" captionStart="Fig" captionStartId="32.[189,232,1930,1956]" captionTargetBox="[369,1231,950,1895]" captionTargetId="figure@32.[359,1236,943,1899]" captionTargetPageId="32" captionText="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="38" pageNumber="39">24C</figureCitation>
) ……………………………2 All Ƌ prefemora normal, not bulged laterally (
<figureCitation id="1379F793FFD97D4EAD26D50CFC7F9953" box="[836,942,349,375]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[265,1316,539,1860]" captionTargetId="figure@4.[261,1319,533,1884]" captionTargetPageId="4" captionText="Fig. 2. Tylopus flavolineatus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, caudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837863" httpUri="https://zenodo.org/record/3837863/files/figure.png" pageId="38" pageNumber="39">Figs 2G</figureCitation>
,
<figureCitation id="1379F793FFD97D4EADA0D50CFBCA9953" box="[962,1051,349,375]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="38" pageNumber="39">11F, G</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAA4DD50CFB9B9953" box="[1071,1098,349,375]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. AD. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="38" pageNumber="39">17</figureCitation>
EG, 19C) …………29
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD97D4EAEDFD5CFFAA69A61" pageId="38" pageNumber="39">
<paragraph id="8BFDEB16FFD97D4EAEDFD5CFFAA699DB" blockId="38.[189,1399,414,581]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEDFD5CFFAA699DB" pageId="38" pageNumber="39">
2. Surface of metaterga virtually smooth, at best extremely faintly rugulose in certain places and/ or with a few barely traceable (setigerous) tubercles near caudal margin (setae mostly broken off) (
<figureCitation id="1379F793FFD97D4EAE91D5B5FEA799DB" box="[243,374,484,511]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, subdorsal, ventral and sublateral views, respectively. IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="38" pageNumber="39">Figs 7B, D</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAFE0D5B4FE2C99DB" box="[386,509,485,511]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="38" pageNumber="39">
12A
<emphasis id="B9363704FFD97D4EAFD6D5B4FE1299DB" bold="true" box="[436,451,485,511]" pageId="38" pageNumber="39"></emphasis>
B, D
</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAC6AD5B4FD8F99DB" box="[520,606,485,511]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="38" pageNumber="39">21A, C</figureCitation>
) ……………………………………………………………………3
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD97D4EAEA3D659FAA69A61" blockId="38.[189,1399,414,581]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEA3D659FAA69A61" pageId="38" pageNumber="39">
Surface of metaterga mostly rugulose to coarsely rugose/tuberculate (
<figureCitation id="1379F793FFD97D4EAA23D659FAD39A06" box="[1089,1282,520,546]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="38" pageNumber="39">
Figs 18A
<emphasis id="B9363704FFD97D4EAAD6D659FB129A06" bold="true" box="[1204,1219,520,546]" pageId="38" pageNumber="39"></emphasis>
B, D
</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAB71D659FABC9A06" box="[1299,1389,520,546]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="38" pageNumber="39">23B, D</figureCitation>
) ………………………………………………………………………………………………………9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD97D4EAEDFD63DFAA79AD5" pageId="38" pageNumber="39">
<paragraph id="8BFDEB16FFD97D4EAEDFD63DFAA69A8E" blockId="38.[189,1399,620,753]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEDFD63DFAA69A8E" pageId="38" pageNumber="39">
3. Paraterga moderately developed (
<figureCitation id="1379F793FFD97D4EAC17D63DFD2B9AA2" box="[629,762,620,646]" captionStart="Fig" captionStartId="3.[189,232,1894,1920]" captionTargetBox="[189,1398,906,1846]" captionTargetId="figure@3.[189,1398,906,1846]" captionTargetPageId="3" captionText="Fig. 1. Tylopus flavolineatus sp. nov., holotype, Ƌ (A, CI), paratype, Ƌ (B). AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 10 and 11, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, dorsal and subventral views, respectively. HI. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837856" httpUri="https://zenodo.org/record/3837856/files/figure.png" pageId="38" pageNumber="39">Figs 1A, C</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAD64D63DFC8F9AA2" box="[774,862,620,646]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="38" pageNumber="39">21A, C</figureCitation>
), ratio of Ƌ midbody prozonite to metazonite width below 1:1.2 …………………………………………………………………………………4
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD97D4EAEA3D6E2FAA79AD5" blockId="38.[189,1399,620,753]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEA3D6E2FAA79AD5" pageId="38" pageNumber="39">
Paraterga relatively well-developed (
<figureCitation id="1379F793FFD97D4EACDBD6E2FCB19AE9" box="[697,864,691,717]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="38" pageNumber="39">Figs 10B, D</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAD14D6E2FC069AE9" box="[886,983,691,717]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="38" pageNumber="39">11B, D</figureCitation>
,
<figureCitation id="1379F793FFD97D4EAD8FD6E2FB9E9AE9" box="[1005,1103,691,717]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="38" pageNumber="39">18B, D</figureCitation>
), radio of Ƌ midbody prozonite to metazonite width over 1:1.2 …………………………………………………………5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD97D4EAEDFD746FAA69B9A" pageId="38" pageNumber="39">
<paragraph id="8BFDEB16FFD97D4EAEDFD746FAA69B71" blockId="38.[189,1399,791,959]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEDFD746FAA69B71" pageId="38" pageNumber="39">
4. Transverse sulcus on metaterga starting with segment 5. Calluses without incisions. Gonopod with process
<emphasis id="B9363704FFD97D4EAF26D76AFE859B71" bold="true" box="[324,340,827,853]" pageId="38" pageNumber="39">h</emphasis>
smallanddentiform.
<collectingCountry id="F355AB86FFD97D4EAC5CD76BFD779B70" box="[574,678,826,852]" name="Thailand" pageId="38" pageNumber="39">Thailand</collectingCountry>
………………………
<taxonomicName id="4C429095FFD97D4EADCDD76AFAA69B71" ID-CoL="7CZPJ" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[943,1399,826,853]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="prosperus">
<emphasis id="B9363704FFD97D4EADCDD76AFBEF9B71" bold="true" box="[943,1086,827,853]" italics="true" pageId="38" pageNumber="39">T.prosperus</emphasis>
<bibRefCitation id="EFD396E7FFD97D4EAA22D76BFAA69B71" author="Golovatch S. I. &amp; Enghoff H." box="[1088,1399,826,853]" pageId="38" pageNumber="39" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch&amp;Enghoff,1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD97D4EAEA3D70FFAA69B9A" blockId="38.[189,1399,791,959]" pageId="38" pageNumber="39">
<keyLead id="30B34823FFD97D4EAEA3D70FFAA69B9A" pageId="38" pageNumber="39">
Transverse sulcus on metaterga starting with segment 4. Calluses with faint one or two denticles on poreless and poriferous paraterga, respectively. Gonopod very simple, without processes. Southern
<collectingCountry id="F355AB86FFD97D4EAF38D7F5FE719B9A" box="[346,416,932,958]" name="China" pageId="38" pageNumber="39">China</collectingCountry>
…………………………………………………………
<taxonomicName id="4C429095FFD97D4EAA49D7F4FAA69B9A" ID-CoL="7CZQ9" authority="Golovatch, 2013" authorityName="Golovatch" authorityYear="2013" box="[1067,1399,932,959]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B9363704FFD97D4EAA49D7F4FB7E9B9B" bold="true" box="[1067,1199,932,959]" italics="true" pageId="38" pageNumber="39">T. reductus</emphasis>
<bibRefCitation id="EFD396E7FFD97D4EAAD7D7F5FAA69B9A" author="Golovatch S. I." box="[1205,1399,932,958]" pageId="38" pageNumber="39" pagination="1 - 31" refId="ref19577" refString="Golovatch S. I. 2013. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIII. Arthropoda Selecta 22 (1): 1 - 31." type="journal article" year="2013">Golovatch, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF3DBB9EFFD97D4EAEDFD34CFEC99FC3" ID-DOI="http://doi.org/10.5281/zenodo.3837927" ID-Zenodo-Dep="3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="38" pageNumber="39" startId="38.[189,232,1821,1847]" targetBox="[266,1310,1040,1771]" targetPageId="38">
<paragraph id="8BFDEB16FFD97D4EAEDFD34CFEC99FC3" blockId="38.[189,1399,1821,2023]" pageId="38" pageNumber="39">
<emphasis id="B9363704FFD97D4EAEDFD34CFECD9F13" bold="true" box="[189,284,1821,1847]" pageId="38" pageNumber="39">Fig. 26.</emphasis>
Distribution of the new eight species of
<taxonomicName id="4C429095FFD97D4EAD73D34CFCBD9F13" ID-CoL="7Q5LS" authorityName="Jeekel" authorityYear="1968" box="[785,876,1821,1847]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9363704FFD97D4EAD73D34CFCBD9F13" box="[785,876,1821,1847]" italics="true" pageId="38" pageNumber="39">Tylopus</emphasis>
</taxonomicName>
.
<emphasis id="B9363704FFD97D4EAD1FD34CFBE09F13" bold="true" box="[893,1073,1821,1847]" pageId="38" pageNumber="39">Filled triangle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAA20D34FFB6C9F13" ID-CoL="7CZPX" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1090,1213,1822,1847]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAA20D34FFB6C9F13" box="[1090,1213,1822,1847]" italics="true" pageId="38" pageNumber="39">T. punctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAAA5D34CFAF49F13" box="[1223,1317,1821,1847]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAB4DD34CFED09F7E" bold="true" pageId="38" pageNumber="39">Filled circle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAF76D310FE6D9F7E" ID-CoL="7CZMC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[276,444,1857,1882]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAF76D310FE6D9F7E" box="[276,444,1857,1882]" italics="true" pageId="38" pageNumber="39">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAFA5D310FDF79F7F" box="[455,550,1857,1883]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAC53D311FD349F7E" bold="true" box="[561,741,1856,1882]" pageId="38" pageNumber="39">Crossed circle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAC9AD310FC719F7E" ID-CoL="7CZMC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[760,928,1857,1882]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAC9AD310FC719F7E" box="[760,928,1857,1882]" italics="true" pageId="38" pageNumber="39">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EADCED310FBDC9F7F" box="[940,1037,1857,1883]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C429095FFD97D4EAA7FD310FB4C9F7E" ID-CoL="7CZN4" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1053,1181,1856,1882]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="dorsalis" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAA7FD310FB4C9F7E" box="[1053,1181,1856,1882]" italics="true" pageId="38" pageNumber="39">T. dorsalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAACAD310FAD69F7F" box="[1192,1287,1857,1883]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAB70D311FEC09F59" bold="true" pageId="38" pageNumber="39">Crossed square</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAF46D335FE1D9F59" ID-CoL="7CZMC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[292,460,1892,1917]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAF46D335FE1D9F59" box="[292,460,1892,1917]" italics="true" pageId="38" pageNumber="39">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAFBAD335FDE69F5A" box="[472,567,1892,1918]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C429095FFD97D4EAC20D335FD669F59" ID-CoL="7CZPY" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[578,695,1892,1917]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAC20D335FD669F59" box="[578,695,1892,1917]" italics="true" pageId="38" pageNumber="39">T. retusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EACA0D335FCF09F5A" box="[706,801,1892,1918]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAD4ED332FC119F59" bold="true" box="[812,960,1891,1917]" pageId="38" pageNumber="39">Open circle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EADB1D335FB999F59" ID-CoL="7CZPY" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[979,1096,1892,1917]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EADB1D335FB999F59" box="[979,1096,1892,1917]" italics="true" pageId="38" pageNumber="39">T. retusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAA31D335FB639F5A" box="[1107,1202,1892,1918]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAADFD332FABE9F59" bold="true" box="[1213,1391,1891,1917]" pageId="38" pageNumber="39">Open triangle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAEDFD3D6FD609F84" ID-CoL="7CZMK" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[189,689,1926,1952]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFD97D4EAEDFD3D6FE869F84" box="[189,343,1926,1952]" italics="true" pageId="38" pageNumber="39">T. baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFD97D4EAF02D3D7FD609F84" author="Golovatch S. I. &amp; Enghoff H." box="[352,689,1926,1952]" pageId="38" pageNumber="39" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9363704FFD97D4EACDCD3D7FCB29F85" bold="true" box="[702,867,1926,1953]" pageId="38" pageNumber="39">Filled square</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAD11D3D6FB949F84" ID-CoL="7CZMW" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[883,1093,1926,1952]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAD11D3D6FB949F84" box="[883,1093,1926,1952]" italics="true" pageId="38" pageNumber="39">T. hongkhraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAA2CD3D6FB7B9F85" box="[1102,1194,1927,1953]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAAD1D3D7FABE9F84" bold="true" box="[1203,1391,1926,1952]" pageId="38" pageNumber="39">Open diamond</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAEDFD3FBFEB99FE0" ID-CoL="7CZP9" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[189,360,1962,1988]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAEDFD3FBFEB99FE0" box="[189,360,1962,1988]" italics="true" pageId="38" pageNumber="39">T. thunghaihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAF0DD3FBFE1B9FE0" box="[367,458,1962,1988]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAFB3D3FBFD3A9FE7" bold="true" box="[465,747,1961,1988]" pageId="38" pageNumber="39">Inverted filled triangle</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAC9BD3FBFC7D9FE0" ID-CoL="7CZN5" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[761,940,1962,1988]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="flavolineatus" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAC9BD3FBFC7D9FE0" box="[761,940,1962,1988]" italics="true" pageId="38" pageNumber="39">T. flavolineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EADD1D3FBFBDF9FE0" box="[947,1038,1962,1988]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9363704FFD97D4EAA77D3F8FB649FE0" bold="true" box="[1045,1205,1961,1988]" pageId="38" pageNumber="39">Open square</emphasis>
.
<taxonomicName id="4C429095FFD97D4EAAA6D3FBFAA69FE0" ID-CoL="7CZPF" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1220,1399,1962,1988]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFD97D4EAAA6D3FBFAA69FE0" box="[1220,1399,1962,1988]" italics="true" pageId="38" pageNumber="39">T. moniliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD97D4EAEDFD39CFEC99FC3" box="[189,280,1997,2023]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<keyStep id="30B6F3B3FFD87D4FAEDFD55AFAA699AB" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEDFD55AFAA6996C" blockId="39.[189,1399,267,399]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD55AFAA6996C" pageId="39" pageNumber="40">
5. Calluses without incisions. Gonopod postfemoral lobe
<emphasis id="B9363704FFD87D4FADE1D55DFC5A9902" bold="true" box="[899,907,268,294]" pageId="39" pageNumber="40">l</emphasis>
much broader than long; area basal to
<emphasis id="B9363704FFD87D4FAB0CD55DFAA79902" bold="true" box="[1390,1398,268,294]" pageId="39" pageNumber="40">l</emphasis>
delimited by a distinct cingulum ……………………………………………………………………6
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D503FAA699AB" blockId="39.[189,1399,267,399]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D503FAA699AB" pageId="39" pageNumber="40">
Calluses mostly with 12 incisions. Gonopod postfemoral lobe
<emphasis id="B9363704FFD87D4FADBFD503FC349948" bold="true" box="[989,997,338,364]" pageId="39" pageNumber="40">l</emphasis>
either as long as broad or longer; no cingulum basal to
<emphasis id="B9363704FFD87D4FAF8AD524FE2199AB" bold="true" box="[488,496,373,399]" pageId="39" pageNumber="40">l</emphasis>
………………………………………………………………………………7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD87D4FAEDFD5E7FAA69A1E" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEDFD5E7FAA699D0" blockId="39.[189,1399,438,570]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD5E7FAA699D0" pageId="39" pageNumber="40">
6. Body smaller: length up to ca
<quantity id="4CBA46F3FFD87D4FAC33D5E7FD0299F5" box="[593,723,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.0" metricValueMin="1.9" pageId="39" pageNumber="40" unit="mm" value="24.5" valueMax="30.0" valueMin="19.0">1930 mm</quantity>
. Gonopod with a broadly coiled solenophore and a small finger-shaped process on lobe
<emphasis id="B9363704FFD87D4FAC0CD58BFDA799D0" bold="true" box="[622,630,474,500]" pageId="39" pageNumber="40">l</emphasis>
.
<collectingCountry id="F355AB86FFD87D4FACEFD588FD0299D7" box="[653,723,473,499]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
……………………………
<taxonomicName id="4C429095FFD87D4FAA47D58BFAA699D0" ID-CoL="7CZMX" authority="Golovatch, 2014" authorityName="Golovatch" authorityYear="2014" box="[1061,1399,473,500]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kabaki">
<emphasis id="B9363704FFD87D4FAA47D58BFB4A99D0" bold="true" box="[1061,1179,473,500]" italics="true" pageId="39" pageNumber="40">T. kabaki</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAAC9D588FAA699D0" author="Golovatch S. I." box="[1195,1399,473,500]" pageId="39" pageNumber="40" pagination="1 - 19" refId="ref19610" refString="Golovatch S. I. 2014. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XV. Arthropoda Selecta 23 (1): 1 - 19." type="journal article" year="2014">Golovatch, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D5ACFAA69A1E" blockId="39.[189,1399,438,570]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D5ACFAA69A1E" pageId="39" pageNumber="40">
Body larger: length ca
<quantity id="4CBA46F3FFD87D4FAC66D5ACFD599A33" box="[516,648,509,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="39" pageNumber="40" unit="mm" value="34.0" valueMax="35.0" valueMin="33.0">3335 mm</quantity>
. Gonopod with a slender and coiled solenophore, but without processes on lobe
<emphasis id="B9363704FFD87D4FAFDED671FE159A1E" bold="true" box="[444,452,544,570]" pageId="39" pageNumber="40">l</emphasis>
.
<collectingCountry id="F355AB86FFD87D4FAFB3D671FDE59A1E" box="[465,564,544,570]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
……………………………………………
<taxonomicName id="4C429095FFD87D4FAA4FD671FAA69A1E" ID-CoL="7CZPH" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1069,1399,544,570]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="magicus">
<emphasis id="B9363704FFD87D4FAA4FD671FB7F9A1E" bold="true" box="[1069,1198,544,570]" italics="true" pageId="39" pageNumber="40">T. magicus</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAAD6D671FAA69A1E" author="Golovatch S. I." box="[1204,1399,544,570]" pageId="39" pageNumber="40" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD87D4FAEA3D633FAA69B03" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEA3D633FAA69AE6" blockId="39.[193,1399,609,706]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D633FAA69AE6" pageId="39" pageNumber="40">
7. Metaterga at best only very faintly rugulose near stricture, near sulcus and/or at base of paraterga, with 23 weak, oblong tubercles near rear margin. Ƌ postfemora and tibiae tuberculiferous. Gonopods with process
<emphasis id="B9363704FFD87D4FAFE2D6F9FE5C9AE6" bold="true" box="[384,397,680,706]" pageId="39" pageNumber="40">z</emphasis>
serrate, with 3 spines.
<collectingCountry id="F355AB86FFD87D4FACEDD6F9FD269AE6" box="[655,759,680,706]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
…………
<taxonomicName id="4C429095FFD87D4FAD13D6F9FAA69AE6" ID-CoL="7CZPR" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[881,1399,679,706]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="similirugosus">
<emphasis id="B9363704FFD87D4FAD13D6F9FBE19AE6" bold="true" box="[881,1072,679,706]" italics="true" pageId="39" pageNumber="40">T. similirugosus</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAA57D6F9FAA69AE6" author="Golovatch S. I. &amp; Enghoff H." box="[1077,1399,680,706]" pageId="39" pageNumber="40" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D6BBFAA69B03" blockId="39.[193,1399,745,807]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D6BBFAA69B03" pageId="39" pageNumber="40">
Metatergal surface entirely smooth, polished, without tubercles. Ƌ legs without adenostyles (= tubercles). Gonopods with process
<emphasis id="B9363704FFD87D4FACB7D75CFD339B03" bold="true" box="[725,738,781,807]" pageId="39" pageNumber="40">z</emphasis>
pointed, without spines ……………………………8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD87D4FAEDFD71FFAA69BD1" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEDFD71FFAA79BA8" blockId="39.[189,1399,846,1013]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD71FFAA79BA8" pageId="39" pageNumber="40">
8. Midline wanting. Gonopods with process
<emphasis id="B9363704FFD87D4FAC8AD71FFD299B4C" bold="true" box="[744,760,846,872]" pageId="39" pageNumber="40">h</emphasis>
without additional tooth, while process
<emphasis id="B9363704FFD87D4FAA86D71FFB209B4C" bold="true" box="[1252,1265,846,872]" pageId="39" pageNumber="40">z</emphasis>
very high.
<collectingCountry id="F355AB86FFD87D4FAE88D720FE9C9BAF" box="[234,333,881,907]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
…………………………………………………………………
<taxonomicName id="4C429095FFD87D4FAA53D723FAA79BA8" ID-CoL="7CZPG" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[1073,1398,881,908]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B9363704FFD87D4FAA53D723FB119BA8" bold="true" box="[1073,1216,881,908]" italics="true" pageId="39" pageNumber="40">T. mutilatus</emphasis>
(
<bibRefCitation id="EFD396E7FFD87D4FAAB2D720FABF9BA8" author="Attems C." box="[1232,1390,881,908]" pageId="39" pageNumber="40" pagination="133 - 230" refId="ref19454" refString="Attems C. 1953. Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, N. S. 5 A: 133 - 230." type="book chapter" year="1953">Attems, 1953</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D7C4FAA69BD1" blockId="39.[189,1399,846,1013]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D7C4FAA69BD1" pageId="39" pageNumber="40">
Midline mostly traceable on prozonae and anterior halves of metaterga. Gonopods with process
<emphasis id="B9363704FFD87D4FAB04D7C4FAA79B8B" bold="true" box="[1382,1398,917,943]" pageId="39" pageNumber="40">h</emphasis>
higher with an evident dorsal tooth, while process
<emphasis id="B9363704FFD87D4FAD7ED7E9FCF89BF6" bold="true" box="[796,809,952,978]" pageId="39" pageNumber="40">z</emphasis>
small(
<figureCitation id="1379F793FFD87D4FAD1AD7E6FBD79BF6" box="[888,1030,951,978]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[308,1241,1126,1855]" captionTargetId="figure@18.[302,1247,1119,1870]" captionTargetPageId="18" captionText="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. AB. Lateral and mesal views, respectively. C. Leg 13. Scale bar: AB. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="39" pageNumber="40">
Figs13A
<emphasis id="B9363704FFD87D4FAD80D7E9FC209BF6" bold="true" box="[994,1009,952,978]" pageId="39" pageNumber="40"></emphasis>
B
</figureCitation>
,
<figureCitation id="1379F793FFD87D4FAA6DD7E9FB8A9BF6" box="[1039,1115,951,978]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="19.[187,230,1928,1954]" captionStartId-1="20.[189,232,1856,1882]" captionTargetBox-0="[189,1398,680,1886]" captionTargetBox-1="[189,1398,605,1811]" captionTargetId-0="figure@19.[189,1398,680,1886]" captionTargetId-1="figure@20.[189,1398,605,1811]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Fig. 14. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. AB. Mesal and lateral views, respectively. CF. Distal part, sublateral, submesal, subcaudal and suboral views, respectively. Scale bars = 0.2 mm." captionText-1="Fig. 15. Tylopus acuminatus sp. nov., paratype, Ƌ, from Phufa (AB), paratype, Ƌ, from Phuthalang forest protected area (CD), paratype, Ƌ, from forest near road (E), paratype, Ƌ, from Ban Na Thong (F), paratype, Ƌ, from Phagneung Phoukulom waterfall (GH). AH. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process h. Green colour shows spine z." figureDoi-0="http://doi.org/10.5281/zenodo.3837902" figureDoi-1="http://doi.org/10.5281/zenodo.3837904" httpUri-0="https://zenodo.org/record/3837902/files/figure.png" httpUri-1="https://zenodo.org/record/3837904/files/figure.png" pageId="39" pageNumber="40">
14
<emphasis id="B9363704FFD87D4FAA4ED7E9FBEA9BF6" bold="true" box="[1068,1083,952,978]" pageId="39" pageNumber="40"></emphasis>
15
</figureCitation>
).
<collectingCountry id="F355AB86FFD87D4FAA0FD7E9FB779BF6" box="[1133,1190,952,978]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
………………… ………………………………………………………………………………
<taxonomicName id="4C429095FFD87D4FAA38D78AFADA9BD1" ID-CoL="7CZMC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1114,1291,987,1013]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFD87D4FAA38D78AFADA9BD1" bold="true" box="[1114,1291,987,1013]" italics="true" pageId="39" pageNumber="40">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD87D4FAB75D78AFAA69BD1" box="[1303,1399,987,1013]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD87D4FAEDFD04DFAA69C59" blockId="39.[189,1400,1052,1149]" pageId="39" pageNumber="40">
<keyStep id="30B6F3B3FFD87D4FAEDFD04DFAA69C59" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD04DFAA69C59" pageId="39" pageNumber="40">
9. Body larger:
<quantity id="4CBA46F3FFD87D4FAF1BD04DFE279C13" box="[377,502,1052,1079]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="39" pageNumber="40" unit="mm" value="39.5" valueMax="42.0" valueMin="37.0">3742 mm</quantity>
long, 2.83.8 and 4.15.0 mm wide on pro- and metaterga, respectively …… …………………………………………………………………………………………………10 Body smaller ………………………………………………………………………………………15
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD87D4FAEDFD0F5FAA69D4A" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEDFD0F5FAA69D20" blockId="39.[189,1399,1188,1391]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD0F5FAA69D20" pageId="39" pageNumber="40">
10. Sterna between Ƌ coxae 4 with a pair of separated cones. Gonopod with a short spiniform process
<emphasis id="B9363704FFD87D4FAF2DD096FE8E9CC5" bold="true" box="[335,351,1223,1249]" pageId="39" pageNumber="40">h</emphasis>
, a basally only poorly delimited lobe
<emphasis id="B9363704FFD87D4FAD5DD096FC969CC5" bold="true" box="[831,839,1223,1249]" pageId="39" pageNumber="40">l</emphasis>
and a small lobe-shaped process
<emphasis id="B9363704FFD87D4FAA8CD096FB2A9CC5" bold="true" box="[1262,1275,1223,1249]" pageId="39" pageNumber="40">z</emphasis>
.
<collectingCountry id="F355AB86FFD87D4FAB6CD096FAA79CC5" box="[1294,1398,1223,1249]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
…………………………………
<taxonomicName id="4C429095FFD87D4FAC0AD0BAFAA69D20" ID-CoL="7CZN2" authority="Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010" authorityName="Likhitrakarn, Golovatch, Prateepasen &amp; Panha" authorityYear="2010" box="[616,1399,1258,1285]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B9363704FFD87D4FAC0AD0BAFD349D21" bold="true" box="[616,741,1258,1285]" italics="true" pageId="39" pageNumber="40">T. grandis</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAC90D0BBFAA69D20" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[754,1399,1258,1285]" pageId="39" pageNumber="40" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D15FFAA69D4A" blockId="39.[189,1399,1188,1391]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D15FFAA69D4A" pageId="39" pageNumber="40">
Sterna between Ƌ coxae 4 with a slightly notched sternal lobe. Gonopod with a rather long process
<emphasis id="B9363704FFD87D4FAE88D160FF2B9D6F" bold="true" box="[234,250,1329,1355]" pageId="39" pageNumber="40">h</emphasis>
carrying two branches, plus a basally evidently delimited lobe
<emphasis id="B9363704FFD87D4FAD96D160FC2D9D6F" bold="true" box="[1012,1020,1329,1355]" pageId="39" pageNumber="40">l</emphasis>
and a large, serrated process
<emphasis id="B9363704FFD87D4FAB00D160FABE9D6F" bold="true" box="[1378,1391,1329,1355]" pageId="39" pageNumber="40">z</emphasis>
.
<collectingCountry id="F355AB86FFD87D4FAE88D105FE9C9D4A" box="[234,333,1364,1390]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
………………………………………………………………
<taxonomicName id="4C429095FFD87D4FAA75D105FAA69D4A" ID-CoL="7CZPN" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[1047,1399,1364,1391]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="phanluongi">
<emphasis id="B9363704FFD87D4FAA75D105FB159D4B" bold="true" box="[1047,1220,1364,1391]" italics="true" pageId="39" pageNumber="40">T. phanluongi</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAAACD104FAA69D4A" author="Nguyen A. D." box="[1230,1399,1364,1391]" pageId="39" pageNumber="40" pagination="289 - 311" refId="ref20147" refString="Nguyen A. D. 2012. Tylopus millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology 60 (2): 289 - 311." type="journal article" year="2012">Nguyen, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD87D4FAEDFD1C7FAA69DD2" blockId="39.[189,1399,1429,1526]" pageId="39" pageNumber="40">
<keyStep id="30B6F3B3FFD87D4FAEDFD1C7FAA69DD2" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD1C7FAA69DD2" pageId="39" pageNumber="40">
11. Both processes
<emphasis id="B9363704FFD87D4FAFC6D1C7FE659D94" bold="true" box="[420,436,1430,1456]" pageId="39" pageNumber="40">h</emphasis>
and
<emphasis id="B9363704FFD87D4FAF8CD1C7FE2A9D94" bold="true" box="[494,507,1430,1456]" pageId="39" pageNumber="40">z</emphasis>
of gonopod spiniform.
<collectingCountry id="F355AB86FFD87D4FAD71D1C4FCAA9D8B" box="[787,891,1429,1455]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
…………………………………………… …………………………………
<taxonomicName id="4C429095FFD87D4FAC0AD1E8FAA69DF7" authority="Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010" authorityName="Likhitrakarn, Golovatch, Prateepasen &amp; Panha" authorityYear="2010" box="[616,1399,1464,1491]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B9363704FFD87D4FAC0AD1E8FCD59DF7" bold="true" box="[616,772,1464,1491]" italics="true" pageId="39" pageNumber="40">T. bispinosus</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAD69D1E8FAA69DF7" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[779,1399,1465,1491]" pageId="39" pageNumber="40" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
Gonopod otherwise ………………………………………………………………………………12
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEDFD24CFAA79E5A" blockId="39.[189,1399,1565,1662]" pageId="39" pageNumber="40">
<keyStep id="30B6F3B3FFD87D4FAEDFD24CFAA79E5A" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD24CFAA79E5A" pageId="39" pageNumber="40">
12. Gonopod process
<emphasis id="B9363704FFD87D4FAFA0D24CFE039E13" bold="true" box="[450,466,1565,1591]" pageId="39" pageNumber="40">h</emphasis>
subflagelliform, process
<emphasis id="B9363704FFD87D4FAD62D24CFCC99E13" bold="true" box="[768,792,1565,1591]" pageId="39" pageNumber="40">m</emphasis>
extremely long and prominent.
<collectingCountry id="F355AB86FFD87D4FAAFBD24CFAD39E13" box="[1177,1282,1565,1591]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
………… …………………………………
<taxonomicName id="4C429095FFD87D4FAC0AD210FAA69E7F" ID-CoL="7CZMT" authority="Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010" authorityName="Likhitrakarn, Golovatch, Prateepasen &amp; Panha" authorityYear="2010" box="[616,1399,1600,1627]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="extremus">
<emphasis id="B9363704FFD87D4FAC0AD210FD269E7F" bold="true" box="[616,759,1601,1627]" italics="true" pageId="39" pageNumber="40">T. extremus</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAD62D211FAA69E7F" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[768,1399,1600,1627]" pageId="39" pageNumber="40" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
Gonopod otherwise ………………………………………………………………………………13
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEDFD2F4FAA69F22" blockId="39.[189,1399,1701,1798]" pageId="39" pageNumber="40">
<keyStep id="30B6F3B3FFD87D4FAEDFD2F4FAA69F22" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD2F4FAA69F22" pageId="39" pageNumber="40">
13. Gonopod process
<emphasis id="B9363704FFD87D4FAFAFD2F4FE0C9E9B" bold="true" box="[461,477,1701,1727]" pageId="39" pageNumber="40">h</emphasis>
prominent, spine
<emphasis id="B9363704FFD87D4FACA5D2F4FD039E9B" bold="true" box="[711,722,1701,1727]" pageId="39" pageNumber="40">s</emphasis>
high, rather slender, curved, nearly pointed.
<collectingCountry id="F355AB86FFD87D4FAB71D2F4FAA79E9B" box="[1299,1398,1701,1727]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
……………………………………………………………………………
<taxonomicName id="4C429095FFD87D4FAA5FD298FAA69EC6" ID-CoL="7CZPW" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[1085,1399,1736,1763]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sapaensis">
<emphasis id="B9363704FFD87D4FAA5FD298FB1C9EC6" bold="true" box="[1085,1229,1736,1763]" italics="true" pageId="39" pageNumber="40">T. sapaensis</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAAB0D298FAA69EC6" author="Nguyen A. D." box="[1234,1399,1736,1763]" pageId="39" pageNumber="40" pagination="289 - 311" refId="ref20147" refString="Nguyen A. D. 2012. Tylopus millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology 60 (2): 289 - 311." type="journal article" year="2012">Nguyen, 2012</bibRefCitation>
</taxonomicName>
Gonopod otherwise ………………………………………………………………………………14
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD87D4FAEDFD37CFAA69FF0" pageId="39" pageNumber="40">
<paragraph id="8BFDEB16FFD87D4FAEDFD37CFAA69FA9" blockId="39.[189,1399,1837,2004]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEDFD37CFAA69FA9" pageId="39" pageNumber="40">
14. Ƌ legs shorter, ca 1.21.3 times as long as midbody height. Gonopod lobe
<emphasis id="B9363704FFD87D4FAAEBD37CFB409F63" bold="true" box="[1161,1169,1837,1863]" pageId="39" pageNumber="40">l</emphasis>
velum-shaped and supplied with two denticles; spine
<emphasis id="B9363704FFD87D4FACEED301FD489F4E" bold="true" box="[652,665,1872,1898]" pageId="39" pageNumber="40">z</emphasis>
short and knife-shaped, while spine
<emphasis id="B9363704FFD87D4FAA34D301FBB79F4E" bold="true" box="[1110,1126,1872,1898]" pageId="39" pageNumber="40">h</emphasis>
rudimentary.
<collectingCountry id="F355AB86FFD87D4FAB6CD301FAA79F4E" box="[1294,1398,1872,1898]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
…………………………………
<taxonomicName id="4C429095FFD87D4FAC0AD325FAA69FA9" ID-CoL="7CZP4" authority="Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010" authorityName="Likhitrakarn, Golovatch, Prateepasen &amp; Panha" authorityYear="2010" box="[616,1399,1907,1934]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="B9363704FFD87D4FAC0AD325FD0C9FAA" bold="true" box="[616,733,1907,1934]" italics="true" pageId="39" pageNumber="40">T. veliger</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAC89D322FAA69FA9" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[747,1399,1907,1934]" pageId="39" pageNumber="40" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD87D4FAEA3D3C6FAA69FF0" blockId="39.[189,1399,1837,2004]" pageId="39" pageNumber="40">
<keyLead id="30B34823FFD87D4FAEA3D3C6FAA69FF0" pageId="39" pageNumber="40">
Ƌ legs longer, ca 1.61.7times as long as midbody height.Gonopod spine
<emphasis id="B9363704FFD87D4FAA53D3C6FBEF9F95" bold="true" box="[1073,1086,1943,1969]" pageId="39" pageNumber="40">z</emphasis>
small, placed closer to base of spine
<emphasis id="B9363704FFD87D4FAF30D3EBFEB39FF0" bold="true" box="[338,354,1978,2004]" pageId="39" pageNumber="40">h</emphasis>
.
<collectingCountry id="F355AB86FFD87D4FAF13D3EBFE089FF0" box="[369,473,1978,2004]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
…………
<taxonomicName id="4C429095FFD87D4FAC35D3EBFAA69FF0" ID-CoL="7CZPP" authority="Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010" authorityName="Likhitrakarn, Golovatch, Prateepasen &amp; Panha" authorityYear="2010" box="[599,1399,1977,2004]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="parajeekeli">
<emphasis id="B9363704FFD87D4FAC35D3EBFD2D9FF0" bold="true" box="[599,764,1977,2004]" italics="true" pageId="39" pageNumber="40">T. parajeekeli</emphasis>
<bibRefCitation id="EFD396E7FFD87D4FAD66D3EBFAA69FF0" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[772,1399,1978,2004]" pageId="39" pageNumber="40" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D40AEDFD55AFAA799AB" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD55AFAA7996D" blockId="40.[189,1399,267,399]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD55AFAA7996D" pageId="40" pageNumber="41">
15. Most of metaterga with a pattern of 2+2 and 2+2 setigerous tubercles in two rows, rear row somewhat less strongly developed than fore one.
<collectingCountry id="F355AB86FFD77D40AD41D57EFC44996D" box="[803,917,303,329]" name="Myanmar" pageId="40" pageNumber="41">Myanmar</collectingCountry>
and
<collectingCountry id="F355AB86FFD77D40ADA8D57FFBE3996C" box="[970,1074,302,328]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
<taxonomicName id="4C429095FFD77D40AA34D57EFAA7996D" ID-CoL="7CZMS" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[1110,1398,302,329]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B9363704FFD77D40AA34D57EFB6E996D" bold="true" box="[1110,1215,302,329]" italics="true" pageId="40" pageNumber="41">T. doriae</emphasis>
(
<bibRefCitation id="EFD396E7FFD77D40AAAFD57FFABF996D" author="Pocock R. I." box="[1229,1390,302,329]" pageId="40" pageNumber="41" pagination="346 - 368" refId="ref20188" refString="Pocock R. I. 1895. Report upon the Chilopoda and Diplopoda obtained by P. W. Bassett-Smith, Esq., Surgeon R. N., and J. J. Walker, Esq., R. N., during the cruise in the Chinese Seas of H. M. S. ' Penguin', Commander W. U. Moore commanding. Annals and Magazine of Natural History series 6, 15: 346 - 368. http: // biodiversitylibrary. org / page / 19265532" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D503FAA799AB" blockId="40.[189,1399,267,399]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D503FAA799AB" pageId="40" pageNumber="41"> Most of metaterga with rear row of setigerous tubercles or wrinkles more strongly developed than fore row, the latter (next to) wanting ………………………………………………………………16</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD77D40AEDFD5E7FAA79A33" blockId="40.[189,1398,438,536]" pageId="40" pageNumber="41">
<keyStep id="30B6F3B3FFD77D40AEDFD5E7FAA79A33" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD5E7FAA79A33" pageId="40" pageNumber="41">16. Transverse sulcus on metaterga starting with segment 4, either fully or almost fully developed there, always fully developed starting with segment 5 ……………………………………………………17 Transverse sulcus on metaterga starting only with segment 5 ……………………………………20</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD77D40AEDFD66FFAA69AC1" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD66FFAA79ABB" blockId="40.[189,1399,574,742]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD66FFAA79ABB" pageId="40" pageNumber="41">
17. Transverse sulcus fully developed and reaching base of paraterga already starting with segment 4. Gonopod tooth
<emphasis id="B9363704FFD77D40AFC2D633FE7C9A58" bold="true" box="[416,429,610,636]" pageId="40" pageNumber="41">z</emphasis>
at base of lobe
<emphasis id="B9363704FFD77D40AC3DD633FDB69A58" bold="true" box="[607,615,610,636]" pageId="40" pageNumber="41">l</emphasis>
coarsely serrate along proximal margin.
<collectingCountry id="F355AB86FFD77D40AA22D630FB729A5F" box="[1088,1187,609,635]" name="Vietnam" pageId="40" pageNumber="41">Vietnam</collectingCountry>
………………… ………………………………………………………………………………
<taxonomicName id="4C429095FFD77D40AA38D6D4FAA79ABB" ID-CoL="7CZMP" authority="(Attems, 1937)" baseAuthorityName="Attems" baseAuthorityYear="1937" box="[1114,1398,644,671]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hilaris">
<emphasis id="B9363704FFD77D40AA38D6D4FB129ABB" bold="true" box="[1114,1219,644,671]" italics="true" pageId="40" pageNumber="41">T. hilaris</emphasis>
(
<bibRefCitation id="EFD396E7FFD77D40AAB0D6D4FABF9ABB" author="Attems C." box="[1234,1390,645,671]" pageId="40" pageNumber="41" pagination="1 - 300" refId="ref19431" refString="Attems C. 1937. Myriopoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich 68: 1 - 300." type="journal article" year="1937">Attems, 1937</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D6F9FAA69AC1" blockId="40.[189,1399,574,742]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D6F9FAA69AC1" pageId="40" pageNumber="41">
Transverse sulcus fully developed only starting with segment 5. Gonopod tooth
<emphasis id="B9363704FFD77D40AAC6D6F9FB609AE6" bold="true" box="[1188,1201,680,706]" pageId="40" pageNumber="41">z</emphasis>
either devoid of serration or serrate along distal margin ……………………………………………………………18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D40AEDFD75CFAA79BB4" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD75CFAA69B49" blockId="40.[189,1399,780,913]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD75CFAA69B49" pageId="40" pageNumber="41">
18. Paraterga 2 rather broadly rounded caudolaterally. Gonopod relatively simple, process
<emphasis id="B9363704FFD77D40AB6CD75CFACF9B03" bold="true" box="[1294,1310,781,807]" pageId="40" pageNumber="41">h</emphasis>
poorly developed, no additional outgrowths near base.
<collectingCountry id="F355AB86FFD77D40AD4CD761FC479B6E" box="[814,918,816,842]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
………………………………………… ………………………………………………………………
<taxonomicName id="4C429095FFD77D40ADC8D702FAA69B49" ID-CoL="7CZMD" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[938,1399,850,877]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B9363704FFD77D40ADC8D702FBC89B48" bold="true" box="[938,1049,850,877]" italics="true" pageId="40" pageNumber="41">T. affinis</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA41D702FAA69B49" author="Golovatch S. I. &amp; Enghoff H." box="[1059,1399,851,877]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D726FAA79BB4" blockId="40.[189,1399,780,913]" box="[193,1398,886,913]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D726FAA79BB4" box="[193,1398,886,913]" pageId="40" pageNumber="41"> Paraterga 2 pointed caudally. Gonopods more complex …………………………………………19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D40AEDFD7E9FAA69C7A" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD7E9FAA69C3C" blockId="40.[189,1399,951,1119]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD7E9FAA69C3C" pageId="40" pageNumber="41">
19. Coloration dark brown, without a cingulate pattern. Sternal lamina between Ƌ coxae 4 low and distinctly bimodal. Gonopod tooth
<emphasis id="B9363704FFD77D40ACB5D78AFD359BD1" bold="true" box="[727,740,987,1013]" pageId="40" pageNumber="41">z</emphasis>
prominent and serrate along distal margin.
<collectingCountry id="F355AB86FFD77D40AB6CD78AFAA79BD1" box="[1294,1398,987,1013]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
……………………………………………………………
<taxonomicName id="4C429095FFD77D40ADEFD7AFFAA69C3C" ID-CoL="7CZQ7" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[909,1399,1022,1048]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B9363704FFD77D40ADEFD7AFFBC19C3C" bold="true" box="[909,1040,1022,1048]" italics="true" pageId="40" pageNumber="41">T. rugosus</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA7FD7AFFAA69C3C" author="Golovatch S. I. &amp; Enghoff H." box="[1053,1399,1022,1048]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D073FAA69C7A" blockId="40.[189,1399,951,1119]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D073FAA69C7A" pageId="40" pageNumber="41">
Coloration pale, with a cingulate pattern. Sternal lamina between Ƌ coxae 4 high, subquadrate. Gonopod tooth
<emphasis id="B9363704FFD77D40AFFCD014FE7A9C7B" bold="true" box="[414,427,1093,1119]" pageId="40" pageNumber="41">z</emphasis>
smaller and spiniform.
<collectingCountry id="F355AB86FFD77D40ACDAD015FCF19C7A" box="[696,800,1092,1118]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
………
<taxonomicName id="4C429095FFD77D40AD1FD014FAA69C7A" ID-CoL="7CZPT" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[893,1399,1092,1119]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="semirugosus">
<emphasis id="B9363704FFD77D40AD1FD014FBE09C7A" bold="true" box="[893,1073,1092,1119]" italics="true" pageId="40" pageNumber="41">T. semirugosus</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA57D015FAA69C7A" author="Golovatch S. I. &amp; Enghoff H." box="[1077,1399,1092,1118]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D40AEDFD0D7FAA79D2D" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD0D7FAA69CC2" blockId="40.[189,1399,1158,1290]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD0D7FAA69CC2" pageId="40" pageNumber="41">
20. Paratergal corner protruding caudad behind rear margin only starting with segment 15, being obtuse-angled or subrectangular and lying more or less within the margin until segment 14.
<collectingCountry id="F355AB86FFD77D40AE88D09DFE9C9CC2" box="[234,333,1228,1254]" name="Vietnam" pageId="40" pageNumber="41">Vietnam</collectingCountry>
………………………………………………………………
<taxonomicName id="4C429095FFD77D40AA77D09CFAA69CC2" ID-CoL="7CZMY" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1045,1399,1228,1255]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hilaroides">
<emphasis id="B9363704FFD77D40AA77D09CFB7D9CC3" bold="true" box="[1045,1196,1228,1255]" italics="true" pageId="40" pageNumber="41">T. hilaroides</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AAD1D09DFAA69CC2" author="Golovatch S. I." box="[1203,1399,1228,1254]" pageId="40" pageNumber="41" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D0A1FAA79D2D" blockId="40.[189,1399,1158,1290]" box="[193,1398,1263,1290]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D0A1FAA79D2D" box="[193,1398,1263,1290]" pageId="40" pageNumber="41"> Paratergal corner protruding caudad behind rear margin before segment 14, mostly pointed …21</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD77D40AEDFD160FAA79D4B" blockId="40.[189,1399,1329,1391]" pageId="40" pageNumber="41">
<keyStep id="30B6F3B3FFD77D40AEDFD160FAA79D4B" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD160FAA79D4B" pageId="40" pageNumber="41">
21. Pattern of tergal setation on segments 18 and/or 19: 2+2 and 5+
<quantity id="4CBA46F3FFD77D40AA69D160FBEC9D6F" box="[1035,1085,1329,1356]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="40" pageNumber="41" unit="in" value="5.0">5 in</quantity>
two rows ……………22 Pattern of tergal setation at least on segments 519: 2+2 and 4+
<quantity id="4CBA46F3FFD77D40AA68D105FBEC9D4A" box="[1034,1085,1364,1390]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="40" pageNumber="41" unit="in" value="4.0">4 in</quantity>
two rows ……………25
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD77D40AEDFD1C7FAA79EA7" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD1C7FAA69E3D" blockId="40.[189,1399,1429,1667]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD1C7FAA69E3D" pageId="40" pageNumber="41">
22. Pattern of tergal setation 2+2 and 5+5 on both segments 18 and 19. Paraterga 2 pointed caudally. Epiproct with pre-apical incisions very close to apical knobs. Sternal lamina between Ƌ coxae 4 an unusually low and even ridge. Adenostyles on midbody Ƌ postfemora and, to a lesser extent, tibiae exceptionally prominent.
<collectingCountry id="F355AB86FFD77D40AC1AD1AEFD319E3D" box="[632,736,1535,1561]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
……
<taxonomicName id="4C429095FFD77D40AD49D1AEFAA69E3D" ID-CoL="7CZPL" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[811,1399,1535,1562]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="poolpermorum">
<emphasis id="B9363704FFD77D40AD49D1AEFBD59E3E" bold="true" box="[811,1028,1535,1562]" italics="true" pageId="40" pageNumber="41">T. poolpermorum</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA76D1AEFAA69E3D" author="Golovatch S. I. &amp; Enghoff H." box="[1044,1399,1535,1561]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D272FAA79EA7" blockId="40.[189,1399,1429,1667]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D272FAA79EA7" pageId="40" pageNumber="41"> Pattern of tergal setation 2+2 and 5+5 on segment 19. Paraterga 2 more or less narrowly rounded. Pre-apical incisions on epiproct more strongly removed from tip. Sternal lamina between Ƌ coxae 4 concave medially. Ventral adenostyles on Ƌ legs less prominent ……………………23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D40AEDFD2FBFAA79F75" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD2FBFAA69F2F" blockId="40.[189,1399,1706,1873]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD2FBFAA69F2F" pageId="40" pageNumber="41">
23. Body smaller: width ca 2.0 mm. Sternal lamina between Ƌ coxae 4 a pair of separated setigerous tubercles. Ventral adenostyles on Ƌ legs almost missing. Gonopods without any outgrowths near base of process
<emphasis id="B9363704FFD77D40AF04D2A0FEA79F2F" bold="true" box="[358,374,1777,1803]" pageId="40" pageNumber="41">h</emphasis>
.
<collectingCountry id="F355AB86FFD77D40AFE0D2A1FE3B9F2E" box="[386,490,1776,1802]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
…………………………………
<taxonomicName id="4C429095FFD77D40AD0FD2A0FAA69F2F" ID-CoL="7CZN3" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[877,1399,1776,1803]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="haplorugosus">
<emphasis id="B9363704FFD77D40AD0FD2A0FBFF9F2F" bold="true" box="[877,1070,1776,1803]" italics="true" pageId="40" pageNumber="41">T. haplorugosus</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA51D2A1FAA69F2F" author="Golovatch S. I. &amp; Enghoff H." box="[1075,1399,1776,1803]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD77D40AEA3D345FAA79F75" blockId="40.[189,1399,1706,1873]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEA3D345FAA79F75" pageId="40" pageNumber="41">
Body larger: width over 3.0 mm. Sternal lamina between Ƌ coxae 4 single. Ventral adenostyles on Ƌ legs more prominent. Gonopod with a spine near base of process
<emphasis id="B9363704FFD77D40AA5AD366FB999F75" bold="true" box="[1080,1096,1847,1873]" pageId="40" pageNumber="41">h</emphasis>
………………………24
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD77D41AEDFD329FAA699AB" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<paragraph id="8BFDEB16FFD77D40AEDFD329FAA69FFD" blockId="40.[189,1399,1912,2009]" pageId="40" pageNumber="41">
<keyLead id="30B34823FFD77D40AEDFD329FAA69FFD" pageId="40" pageNumber="41">
24. Sternal lamina between Ƌ coxae 4 high, emarginated. Adenostyles on Ƌ postfemora and tibiae well-developed. Gonopods rather simple, spine
<emphasis id="B9363704FFD77D40AD49D3CDFCE99F92" bold="true" box="[811,824,1948,1974]" pageId="40" pageNumber="41">z</emphasis>
inconspicuous.
<collectingCountry id="F355AB86FFD77D40AD9ED3CAFBB59F91" box="[1020,1124,1947,1973]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
……………………… ……………………………………………………………
<taxonomicName id="4C429095FFD77D40ADEFD3EEFAA69FFD" ID-CoL="7CZMB" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[909,1399,1982,2009]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="allorugosus">
<emphasis id="B9363704FFD77D40ADEFD3EEFBE49FFD" bold="true" box="[909,1077,1982,2009]" italics="true" pageId="40" pageNumber="41">T. allorugosus</emphasis>
<bibRefCitation id="EFD396E7FFD77D40AA5BD3EEFAA69FFD" author="Golovatch S. I. &amp; Enghoff H." box="[1081,1399,1983,2009]" pageId="40" pageNumber="41" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp;Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D57EFAA699AB" blockId="41.[193,1399,302,399]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D57EFAA699AB" pageId="41" pageNumber="42">
Sternal lamina between Ƌ coxae lower, slightly concave. Adenostyles on Ƌ postfemora and tibiae less strongly developed. Gonopods more complex, spine
<emphasis id="B9363704FFD67D41AA7CD503FBFA9948" bold="true" box="[1054,1067,338,364]" pageId="41" pageNumber="42">z</emphasis>
long and large.
<collectingCountry id="F355AB86FFD67D41AB6CD503FAA79948" box="[1294,1398,338,364]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
………………………………………………………………………
<taxonomicName id="4C429095FFD67D41AA60D524FAA699AB" ID-CoL="7CZPQ" authority="Hoffman, 1973" authorityName="Hoffman" authorityYear="1973" box="[1026,1399,373,399]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="perarmatus">
<emphasis id="B9363704FFD67D41AA60D524FB6099AB" bold="true" box="[1026,1201,373,399]" italics="true" pageId="41" pageNumber="42">T. perarmatus</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AADCD524FAA699AB" author="Hoffman R. L." box="[1214,1399,373,399]" pageId="41" pageNumber="42" pagination="361 - 389" refId="ref19686" refString="Hoffman R. L. 1973. Descriptions and allocations of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida). Journal of Natural History 7 (4): 361 - 389. http: // dx. doi. org / 10.1080 / 00222937300770281" type="journal article" year="1973">Hoffman, 1973</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD67D41AEDFD5E7FAA79A79" pageId="41" pageNumber="42">
<paragraph id="8BFDEB16FFD67D41AEDFD5E7FAA69A33" blockId="41.[189,1399,438,606]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD5E7FAA69A33" pageId="41" pageNumber="42">
25. Paraterga 2 pointed caudally. Sternal lamina between Ƌ coxae 4 exceptionally densely setose, low and concave ventrally. Gonopods with a medium-sized process
<emphasis id="B9363704FFD67D41ADA8D58BFC0B99D0" bold="true" box="[970,986,474,500]" pageId="41" pageNumber="42">h</emphasis>
and a smaller lobular
<emphasis id="B9363704FFD67D41AABAD58BFB3499D0" bold="true" box="[1240,1253,474,500]" pageId="41" pageNumber="42">z</emphasis>
at base of
<emphasis id="B9363704FFD67D41AB3DD58BFABE99D0" bold="true" box="[1375,1391,474,500]" pageId="41" pageNumber="42">h</emphasis>
.
<collectingCountry id="F355AB86FFD67D41AE88D5ACFE839A33" box="[234,338,509,535]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
…………………………………………………….
<taxonomicName id="4C429095FFD67D41ADCCD5ACFAA69A33" ID-CoL="7CZN8" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[942,1399,508,535]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="jeekeli">
<emphasis id="B9363704FFD67D41ADCCD5ACFBCC9A33" bold="true" box="[942,1053,508,535]" italics="true" pageId="41" pageNumber="42">T. jeekeli</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AA45D5ACFAA69A33" author="Golovatch S. I. &amp; Enghoff H." box="[1063,1399,509,535]" pageId="41" pageNumber="42" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D671FAA79A79" blockId="41.[189,1399,438,606]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D671FAA79A79" pageId="41" pageNumber="42">
Paraterga 2 more or less narrowly rounded caudally. Sternal lamina between Ƌ coxae 4 higher and less strongly setose. Gonopod outgrowths
<emphasis id="B9363704FFD67D41AD42D612FCE19A79" bold="true" box="[800,816,579,605]" pageId="41" pageNumber="42">h</emphasis>
and
<emphasis id="B9363704FFD67D41AD0DD612FCAD9A79" bold="true" box="[879,892,579,605]" pageId="41" pageNumber="42">z</emphasis>
either almost wanting or very large …26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD67D41AEDFD6D5FAA69B2C" pageId="41" pageNumber="42">
<paragraph id="8BFDEB16FFD67D41AEDFD6D5FAA69AE6" blockId="41.[189,1399,644,777]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD6D5FAA69AE6" pageId="41" pageNumber="42">26. Sternal lamina between Ƌ coxae 4 with a straight ventral margin. Pleurosternal carinae poorly developed, in Ƌ slightly drawn behind rear margin only until segments 810 …………………27</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D69DFAA69B2C" blockId="41.[189,1399,644,777]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D69DFAA69B2C" pageId="41" pageNumber="42"> Sternal lamina between Ƌ coxae 4 slightly concave ventrally. Pleurosternal carinae better developed, in Ƌ slightly drawn behind rear margin at least until segment 15 …………………28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD67D41AEDFD761FAA69BDE" pageId="41" pageNumber="42">
<paragraph id="8BFDEB16FFD67D41AEDFD761FAA69BB4" blockId="41.[189,1406,816,1018]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD761FAA69BB4" pageId="41" pageNumber="42">
27. Body smaller: width up to
<quantity id="4CBA46F3FFD67D41AC56D761FD199B6E" box="[564,712,816,842]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="41" pageNumber="42" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.13.2 mm</quantity>
. Mid-dorsal line very clear on both halves of metaterga. Gonopods relatively simple, with both
<emphasis id="B9363704FFD67D41ACD8D702FD1B9B49" bold="true" box="[698,714,851,877]" pageId="41" pageNumber="42">h</emphasis>
and
<emphasis id="B9363704FFD67D41AD64D702FCC29B49" bold="true" box="[774,787,851,877]" pageId="41" pageNumber="42">z</emphasis>
almost wanting.
<collectingCountry id="F355AB86FFD67D41AD82D702FB999B49" box="[992,1096,851,877]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
………………………… ……………………………………………………………
<taxonomicName id="4C429095FFD67D41ADEFD727FAA69BB4" ID-CoL="7CZMZ" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[909,1399,886,913]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="B9363704FFD67D41ADEFD727FBCC9BB5" bold="true" box="[909,1053,886,913]" italics="true" pageId="41" pageNumber="42">T. hoffmani</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AA45D727FAA69BB4" author="Golovatch S. I. &amp; Enghoff H." box="[1063,1399,886,912]" pageId="41" pageNumber="42" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D7CBFAA69BDE" blockId="41.[189,1406,816,1018]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D7CBFAA69BDE" pageId="41" pageNumber="42">
Body larger: width 4.0
<quantity id="4CBA46F3FFD67D41AC71D7C8FDA29B90" box="[531,627,921,948]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="41" pageNumber="42" unit="mm" value="5.3">5.3 mm</quantity>
. Mid-dorsal line not so well-developed at least on rear halves of metaterga. Gonopods more complex, with both
<emphasis id="B9363704FFD67D41AD37D7ECFCB49BF3" bold="true" box="[853,869,957,983]" pageId="41" pageNumber="42">h</emphasis>
and
<emphasis id="B9363704FFD67D41ADC4D7ECFC629BF3" bold="true" box="[934,947,957,983]" pageId="41" pageNumber="42">z</emphasis>
very conspicuous (
<figureCitation id="1379F793FFD67D41AAC3D7EDFAE99BF3" box="[1185,1336,956,983]" captionStart="Fig" captionStartId="32.[189,232,1930,1956]" captionTargetBox="[369,1231,950,1895]" captionTargetId="figure@32.[359,1236,943,1899]" captionTargetPageId="32" captionText="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="41" pageNumber="42">
Figs 24A
<emphasis id="B9363704FFD67D41AB71D7ECFAF39BF3" bold="true" box="[1299,1314,957,983]" pageId="41" pageNumber="42"></emphasis>
B
</figureCitation>
,
<figureCitation id="1379F793FFD67D41AB2AD7EDFAB89BF3" box="[1352,1385,956,983]" captionStart="Fig" captionStartId="33.[187,230,1930,1956]" captionTargetBox="[198,1389,743,1904]" captionTargetId="figure@33.[198,1389,743,1904]" captionTargetPageId="33" captionText="Fig. 25. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ, right gonopod. AB. Mesal and lateral views, respectively. CF. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3837925" httpUri="https://zenodo.org/record/3837925/files/figure.png" pageId="41" pageNumber="42">25</figureCitation>
).
<collectingCountry id="F355AB86FFD67D41AE88D7B1FE839BDE" box="[234,338,992,1018]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
………………………………………………
<taxonomicName id="4C429095FFD67D41AD0DD7B1FAA69BDE" ID-CoL="7CZMK" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[879,1399,991,1018]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFD67D41AD0DD7B1FBC19BDE" bold="true" box="[879,1040,991,1018]" italics="true" pageId="41" pageNumber="42">T. baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AA7FD7B1FAA69BDE" author="Golovatch S. I. &amp; Enghoff H." box="[1053,1399,992,1018]" pageId="41" pageNumber="42" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD67D41AEDFD070FAA69CCF" pageId="41" pageNumber="42">
<paragraph id="8BFDEB16FFD67D41AEDFD070FAA69CA6" blockId="41.[189,1399,1057,1260]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD070FAA69CA6" pageId="41" pageNumber="42">
28. Metatergum 19 slightly rugulose posteriorly. Calluses on segment 2 with three, on following paraterga with two, incisions. Gonopods extremely complex, with numerous spiniform outgrowths.
<collectingCountry id="F355AB86FFD67D41AB6CD015FAA79C7A" box="[1294,1398,1092,1118]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
…………………………………………………………
<taxonomicName id="4C429095FFD67D41AD12D039FAA69CA6" ID-CoL="7CZQ2" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[880,1399,1127,1154]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="perplexus">
<emphasis id="B9363704FFD67D41AD12D039FBDB9CA6" bold="true" box="[880,1034,1127,1154]" italics="true" pageId="41" pageNumber="42">T. perplexus</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AA7AD036FAA69CA6" author="Golovatch S. I. &amp; Enghoff H." box="[1048,1399,1127,1154]" pageId="41" pageNumber="42" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D0DAFAA69CCF" blockId="41.[189,1399,1057,1260]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D0DAFAA69CCF" pageId="41" pageNumber="42">
Metatergum 19 entirely smooth. Calluses with two or three incisions on poreless and poriferous paraterga, respectively. Gonopod less strongly differentiated.
<collectingCountry id="F355AB86FFD67D41ADC8D0FFFBC39CEC" box="[938,1042,1198,1224]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
……………………………… ………………………………………………………………
<taxonomicName id="4C429095FFD67D41ADC8D083FAA69CCF" ID-CoL="7CZMM" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[938,1399,1233,1260]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="amicus">
<emphasis id="B9363704FFD67D41ADC8D083FBF19CCF" bold="true" box="[938,1056,1233,1260]" italics="true" pageId="41" pageNumber="42">T. amicus</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AA4BD080FAA69CCF" author="Golovatch S. I. &amp; Enghoff H." box="[1065,1399,1233,1259]" pageId="41" pageNumber="42" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD67D41AEDFD142FAA79D57" blockId="41.[189,1399,1298,1395]" pageId="41" pageNumber="42">
<keyStep id="30B6F3B3FFD67D41AEDFD142FAA79D57" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD142FAA79D57" pageId="41" pageNumber="42">29. Either most of Ƌ sterna with an oblique tubercle or spine, or only anterior sterna with a small cone, near coxae …………………………………………………………………………………………30 Neither spines nor tubercles on Ƌ sterna …………………………………………………………37</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEDFD1CBFAA69DFC" blockId="41.[189,1399,1434,1496]" pageId="41" pageNumber="42">
<keyStep id="30B6F3B3FFD67D41AEDFD1CBFAA69DFC" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD1CBFAA69DFC" pageId="41" pageNumber="42">30. Calluses virtually devoid of incisions………………………………………………………………31 Calluses always at least with one lateral incision …………………………………………………32</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD67D41AEDFD1AEFAA69EA7" pageId="41" pageNumber="42">
<paragraph id="8BFDEB16FFD67D41AEDFD1AEFAA69E19" blockId="41.[189,1399,1535,1667]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD1AEFAA69E19" pageId="41" pageNumber="42">
31. Ƌ coxae 4 with a pair of separated cones. Gonopod process
<emphasis id="B9363704FFD67D41ADC8D1AEFC6B9E3D" bold="true" box="[938,954,1535,1561]" pageId="41" pageNumber="42">h</emphasis>
small, while solenophore slender and short.
<collectingCountry id="F355AB86FFD67D41AF54D273FEAD9E18" box="[310,380,1570,1596]" name="China" pageId="41" pageNumber="42">China</collectingCountry>
……………………………………………………………
<taxonomicName id="4C429095FFD67D41AA4AD272FAA69E19" ID-CoL="7CZPS" authority="Golovatch, 1995" authorityName="Golovatch" authorityYear="1995" box="[1064,1399,1570,1597]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9363704FFD67D41AA4AD272FB769E18" bold="true" box="[1064,1191,1570,1597]" italics="true" pageId="41" pageNumber="42">T. sinensis</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AAD3D273FAA69E19" author="Golovatch S. I." box="[1201,1399,1570,1597]" pageId="41" pageNumber="42" pagination="89 - 97" refId="ref19545" refString="Golovatch S. I. 1995. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda Polydesmida), III. Arthropoda Selecta 4 (2): 89 - 97." type="journal article" year="1995">Golovatch, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEA3D217FAA69EA7" blockId="41.[189,1399,1535,1667]" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEA3D217FAA69EA7" pageId="41" pageNumber="42">
Ƌ coxae 4 with an entire, subtrapeziform, sternal lobe. Gonopod process
<emphasis id="B9363704FFD67D41AA5BD217FB989E44" bold="true" box="[1081,1097,1606,1632]" pageId="41" pageNumber="42">h</emphasis>
larger, while solenophore stout and long.
<collectingCountry id="F355AB86FFD67D41AFF9D238FE2F9EA7" box="[411,510,1641,1667]" name="Vietnam" pageId="41" pageNumber="42">Vietnam</collectingCountry>
…………………………………………………
<taxonomicName id="4C429095FFD67D41AA53D238FAA69EA7" ID-CoL="7CZMQ" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[1073,1399,1640,1667]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="B9363704FFD67D41AA53D238FB1A9EA7" bold="true" box="[1073,1227,1640,1667]" italics="true" pageId="41" pageNumber="42">T. golovatchi</emphasis>
<bibRefCitation id="EFD396E7FFD67D41AAB3D238FAA69EA7" author="Nguyen A. D." box="[1233,1399,1641,1667]" pageId="41" pageNumber="42" pagination="289 - 311" refId="ref20147" refString="Nguyen A. D. 2012. Tylopus millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology 60 (2): 289 - 311." type="journal article" year="2012">Nguyen, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD67D41AEDFD2FBFAA69F2F" blockId="41.[189,1399,1706,1803]" pageId="41" pageNumber="42">
<keyStep id="30B6F3B3FFD67D41AEDFD2FBFAA69F2F" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD2FBFAA69F2F" pageId="41" pageNumber="42">
32. A pair of small, fully separated, setose cones between Ƌ coxae 4 (
<figureCitation id="1379F793FFD67D41AA4FD2FBFB7C9EE1" box="[1069,1197,1706,1733]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="41" pageNumber="42">
Fig. 10I
<emphasis id="B9363704FFD67D41AAF3D2FBFB719EE0" bold="true" box="[1169,1184,1706,1732]" pageId="41" pageNumber="42"></emphasis>
J
</figureCitation>
). Gonopod with process
<emphasis id="B9363704FFD67D41AF2FD29CFE8C9EC3" bold="true" box="[333,349,1741,1767]" pageId="41" pageNumber="42">h</emphasis>
rather short, slightly flattened, blunt (
<figureCitation id="1379F793FFD67D41AD4DD29CFC529EC3" box="[815,899,1741,1767]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="41" pageNumber="42">Fig. 11</figureCitation>
AD).
<collectingCountry id="F355AB86FFD67D41ADB9D29FFBC59ECC" box="[987,1044,1742,1768]" name="Laos" pageId="41" pageNumber="42">Laos</collectingCountry>
…………
<taxonomicName id="4C429095FFD67D41AAF6D29FFADC9ECC" ID-CoL="7CZPY" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1172,1293,1742,1768]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFD67D41AAF6D29FFADC9ECC" bold="true" box="[1172,1293,1742,1768]" italics="true" pageId="41" pageNumber="42">T. retusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD67D41AB7BD29FFAA69ECC" box="[1305,1399,1742,1768]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
An entire lobe between Ƌ coxae 4. Gonopod otherwise …………………………………………33
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BFDEB16FFD67D41AEDFD363FAA69FB7" blockId="41.[189,1399,1842,1939]" pageId="41" pageNumber="42">
<keyStep id="30B6F3B3FFD67D41AEDFD363FAA69FB7" pageId="41" pageNumber="42">
<keyLead id="30B34823FFD67D41AEDFD363FAA69FB7" pageId="41" pageNumber="42">
33. Metaterga strongly rugose/tuberculate/granulate (
<figureCitation id="1379F793FFD67D41AD54D363FC589F68" box="[822,905,1842,1868]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="41" pageNumber="42">Fig. 18</figureCitation>
AF, H) ………………………………34 Metaterga entirely smooth and polished, devoid of evident tubercles, at best extremely faintly rugulose near transverse sulcus (
<figureCitation id="1379F793FFD67D41AC18D329FD6E9FB6" box="[634,703,1912,1938]" captionStart="Fig" captionStartId="3.[189,232,1894,1920]" captionTargetBox="[189,1398,906,1846]" captionTargetId="figure@3.[189,1398,906,1846]" captionTargetPageId="3" captionText="Fig. 1. Tylopus flavolineatus sp. nov., holotype, Ƌ (A, CI), paratype, Ƌ (B). AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 10 and 11, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, dorsal and subventral views, respectively. HI. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837856" httpUri="https://zenodo.org/record/3837856/files/figure.png" pageId="41" pageNumber="42">Fig. 1</figureCitation>
AF) ……………………………………………………35
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD57D42AEDFD55AFAA699AB" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD55AFAA6996D" blockId="42.[189,1399,267,399]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD55AFAA6996D" pageId="42" pageNumber="43">
34. Calluses mostly with one lateral, clear incision. Pleurosternal carinae missing after segment 17. Gonopod processes
<emphasis id="B9363704FFD57D42AFBCD57EFE3F996D" bold="true" box="[478,494,303,329]" pageId="42" pageNumber="43">h</emphasis>
,
<emphasis id="B9363704FFD57D42AC63D57EFDDF996D" bold="true" box="[513,526,303,329]" pageId="42" pageNumber="43">z</emphasis>
,
<emphasis id="B9363704FFD57D42AC43D57EFDE8996D" bold="true" box="[545,569,303,329]" pageId="42" pageNumber="43">m</emphasis>
and spine
<emphasis id="B9363704FFD57D42ACA1D57EFD03996D" bold="true" box="[707,722,303,329]" pageId="42" pageNumber="43">x</emphasis>
present (
<figureCitation id="1379F793FFD57D42AD24D57FFC09996D" box="[838,984,302,329]" captionStart="Fig" captionStartId="26.[189,232,1928,1954]" captionTargetBox="[335,1275,1069,1864]" captionTargetId="figure@26.[333,1278,1058,1870]" captionTargetPageId="26" captionText="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="42" pageNumber="43">
Fig. 19A
<emphasis id="B9363704FFD57D42ADD1D57EFC13996D" bold="true" box="[947,962,303,329]" pageId="42" pageNumber="43"></emphasis>
B
</figureCitation>
).
<collectingCountry id="F355AB86FFD57D42AD90D57EFBFA996D" box="[1010,1067,303,329]" name="Laos" pageId="42" pageNumber="43">Laos</collectingCountry>
<taxonomicName id="4C429095FFD57D42AA36D57EFADC996D" ID-CoL="7CZP9" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1108,1293,302,329]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFD57D42AA36D57EFADC996D" bold="true" box="[1108,1293,302,329]" italics="true" pageId="42" pageNumber="43">T. thunghaihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD57D42AB7AD57EFAA6996D" box="[1304,1399,303,329]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D503FAA699AB" blockId="42.[189,1399,267,399]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D503FAA699AB" pageId="42" pageNumber="43">
Calluses with two lateral setigerous incisions. Pleurosternal carinae missing after segment 12. Only gonopod process
<emphasis id="B9363704FFD57D42AF99D524FDDA99AB" bold="true" box="[507,523,373,399]" pageId="42" pageNumber="43">h</emphasis>
present.
<collectingCountry id="F355AB86FFD57D42AC14D524FD0899AB" box="[630,729,373,399]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
……………………………
<taxonomicName id="4C429095FFD57D42AA46D524FAA699AB" ID-CoL="7CZQ4" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[1060,1399,373,399]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="spinisterna">
<emphasis id="B9363704FFD57D42AA46D524FB1999AB" bold="true" box="[1060,1224,373,399]" italics="true" pageId="42" pageNumber="43">T. spinisterna</emphasis>
<bibRefCitation id="EFD396E7FFD57D42AAB2D524FAA699AB" author="Nguyen A. D." box="[1232,1399,373,399]" pageId="42" pageNumber="43" pagination="289 - 311" refId="ref20147" refString="Nguyen A. D. 2012. Tylopus millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology 60 (2): 289 - 311." type="journal article" year="2012">Nguyen, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD57D42AEDFD5E7FAA79A79" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD5E7FAA79A3C" blockId="42.[189,1399,438,606]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD5E7FAA79A3C" pageId="42" pageNumber="43">
35. Only rear sternum on most of Ƌ segments with a pair of small spines. Metaterga mostly with 2+2 and 3+3 setigerous tubercles in two transverse rows. Pleurosternal carinae missing after segment 9.
<collectingCountry id="F355AB86FFD57D42AF69D5ACFEAC9A33" box="[267,381,509,535]" name="Myanmar" pageId="42" pageNumber="43">Myanmar</collectingCountry>
……………………………………………………………
<taxonomicName id="4C429095FFD57D42AA49D5ACFAA79A3C" ID-CoL="7CZQ6" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[1067,1398,508,536]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="B9363704FFD57D42AA49D5ACFB659A33" bold="true" box="[1067,1204,508,535]" italics="true" pageId="42" pageNumber="43">T. silvestris</emphasis>
(
<bibRefCitation id="EFD396E7FFD57D42AAAAD5ACFABF9A3C" author="Pocock R. I." box="[1224,1390,509,536]" pageId="42" pageNumber="43" pagination="346 - 368" refId="ref20188" refString="Pocock R. I. 1895. Report upon the Chilopoda and Diplopoda obtained by P. W. Bassett-Smith, Esq., Surgeon R. N., and J. J. Walker, Esq., R. N., during the cruise in the Chinese Seas of H. M. S. ' Penguin', Commander W. U. Moore commanding. Annals and Magazine of Natural History series 6, 15: 346 - 368. http: // biodiversitylibrary. org / page / 19265532" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D671FAA79A79" blockId="42.[189,1399,438,606]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D671FAA79A79" pageId="42" pageNumber="43"> Fore and rear sterna of most of Ƌ segments with a pair of small tubercles and spines, respectively. Pleurosternal carinae present until segment 18 ……………………………………………………36</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD57D42AEDFD6D5FAA69B08" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD6D5FAA69AC2" blockId="42.[189,1399,644,812]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD6D5FAA69AC2" pageId="42" pageNumber="43">
36. Metaterga with four transverse rows of very small setigerous tubercles or setae. Ƌ tarsal brushes present until leg 6. Ƌ femora 5 with a very strong, distoventral, densely pilose adenostyle (
<figureCitation id="1379F793FFD57D42AB72D6F9FAB89AE6" box="[1296,1385,680,706]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[265,1316,539,1860]" captionTargetId="figure@4.[261,1319,533,1884]" captionTargetPageId="4" captionText="Fig. 2. Tylopus flavolineatus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, caudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837863" httpUri="https://zenodo.org/record/3837863/files/figure.png" pageId="42" pageNumber="43">Fig. 2F</figureCitation>
). Gonopod process
<emphasis id="B9363704FFD57D42AFD8D69AFE1B9AC1" bold="true" box="[442,458,715,741]" pageId="42" pageNumber="43">h</emphasis>
higher and pointed (
<figureCitation id="1379F793FFD57D42ACD4D69AFD2B9AC1" box="[694,762,715,741]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[265,1316,539,1860]" captionTargetId="figure@4.[261,1319,533,1884]" captionTargetPageId="4" captionText="Fig. 2. Tylopus flavolineatus sp. nov., holotype, Ƌ, left gonopod. AD. Mesal, lateral, caudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837863" httpUri="https://zenodo.org/record/3837863/files/figure.png" pageId="42" pageNumber="43">Fig. 2</figureCitation>
AD). (
<figureCitation id="1379F793FFD57D42AD30D69AFC799AC2" box="[850,936,715,742]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="42" pageNumber="43">Fig. 26</figureCitation>
).
<collectingCountry id="F355AB86FFD57D42ADD8D69AFBF39AC1" box="[954,1058,715,741]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
……
<taxonomicName id="4C429095FFD57D42AA00D69AFAC89AC2" ID-CoL="7CZN5" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1122,1305,715,742]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="flavolineatus" status="sp. nov.">
<emphasis id="B9363704FFD57D42AA00D69AFAC89AC2" bold="true" box="[1122,1305,715,742]" italics="true" pageId="42" pageNumber="43">T. flavolineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD57D42AB7CD69DFAA69AC2" box="[1310,1399,716,742]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D6BEFAA69B08" blockId="42.[189,1399,644,812]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D6BEFAA69B08" pageId="42" pageNumber="43">
Metaterga with two transverse rows of small tubercles or spines, respevtively Ƌ tarsal brushes missing. Ƌ legs normal. Gonopod process
<emphasis id="B9363704FFD57D42AC11D743FD529B08" bold="true" box="[627,643,786,812]" pageId="42" pageNumber="43">h</emphasis>
very small.
<collectingCountry id="F355AB86FFD57D42AD71D740FCA79B0F" box="[787,886,785,811]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
……………
<taxonomicName id="4C429095FFD57D42AA73D743FAA69B08" ID-CoL="7CZN9" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1041,1399,785,812]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B9363704FFD57D42AA73D743FB7D9B08" bold="true" box="[1041,1196,785,812]" italics="true" pageId="42" pageNumber="43">T. maculatus</emphasis>
<bibRefCitation id="EFD396E7FFD57D42AAD1D740FAA69B08" author="Golovatch S. I." box="[1203,1399,785,812]" pageId="42" pageNumber="43" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD57D42AEDFD702FAA69BB4" blockId="42.[189,1399,851,913]" pageId="42" pageNumber="43">
<keyStep id="30B6F3B3FFD57D42AEDFD702FAA69BB4" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD702FAA69BB4" pageId="42" pageNumber="43">
37. Paraterga virtually missing (
<figureCitation id="1379F793FFD57D42AC59D702FD599B49" box="[571,648,851,877]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="42" pageNumber="43">Figs 4</figureCitation>
AH, 7AH) …………………………………………………38 Paraterga well- or strongly developed (
<figureCitation id="1379F793FFD57D42ACD6D727FCE89BB4" box="[692,825,886,912]" captionStart="Fig" captionStartId="3.[189,232,1894,1920]" captionTargetBox="[189,1398,906,1846]" captionTargetId="figure@3.[189,1398,906,1846]" captionTargetPageId="3" captionText="Fig. 1. Tylopus flavolineatus sp. nov., holotype, Ƌ (A, CI), paratype, Ƌ (B). AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 10 and 11, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, dorsal and subventral views, respectively. HI. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837856" httpUri="https://zenodo.org/record/3837856/files/figure.png" pageId="42" pageNumber="43">Figs 1A, C</figureCitation>
,
<figureCitation id="1379F793FFD57D42AD27D727FC4C9BB5" box="[837,925,886,913]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="42" pageNumber="43">10B, D</figureCitation>
,
<figureCitation id="1379F793FFD57D42ADCBD727FBD09BB5" box="[937,1025,886,913]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="42" pageNumber="43">12B, D</figureCitation>
,
<figureCitation id="1379F793FFD57D42AA6FD727FBB49BB5" box="[1037,1125,886,913]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="42" pageNumber="43">16B, D</figureCitation>
,
<figureCitation id="1379F793FFD57D42AA13D727FB189BB5" box="[1137,1225,886,913]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="42" pageNumber="43">18B, D</figureCitation>
,
<figureCitation id="1379F793FFD57D42AAB7D727FAFA9BB5" box="[1237,1323,886,913]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="42" pageNumber="43">23B, D</figureCitation>
) …39
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD57D42AEDFD7E9FAA69CA6" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD7E9FAA69C3C" blockId="42.[189,1399,951,1154]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD7E9FAA69C3C" pageId="42" pageNumber="43">
38. Antennae shorter (
<figureCitation id="1379F793FFD57D42AFA5D7E6FDF29BF6" box="[455,547,951,978]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="42" pageNumber="43">Fig. 4A</figureCitation>
), drawn behind only body segment 2 when stretched dorsally. A small, rounded, densely setose, sternal cone between Ƌ coxae 4 (
<figureCitation id="1379F793FFD57D42ADECD78AFC289BD1" box="[910,1017,987,1013]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="42" pageNumber="43">
Fig. 4I
<emphasis id="B9363704FFD57D42ADBED78AFC3A9BD1" bold="true" box="[988,1003,987,1013]" pageId="42" pageNumber="43"></emphasis>
J
</figureCitation>
). Goponod process
<emphasis id="B9363704FFD57D42AA82D78AFB219BD1" bold="true" box="[1248,1264,987,1013]" pageId="42" pageNumber="43">h</emphasis>
prominent, while process
<emphasis id="B9363704FFD57D42AFF8D7AFFE769C3C" bold="true" box="[410,423,1022,1048]" pageId="42" pageNumber="43">z</emphasis>
long and rather slender (
<figureCitation id="1379F793FFD57D42ACBCD7AFFC9D9C3D" box="[734,844,1022,1049]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1964,1990]" captionStartId-1="8.[189,232,1930,1956]" captionTargetBox-0="[339,1250,1163,1911]" captionTargetBox-1="[189,1398,682,1888]" captionTargetId-0="figure@7.[317,1262,1158,1917]" captionTargetId-1="figure@8.[189,1398,682,1888]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 5. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, right gonopod. AC. Lateral, mesal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." captionText-1="Fig. 6. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, left gonopod. AB. Mesal and lateral views, respectively. CF. Distal part, sublateral, submesal, suboral and subcaudal views, respectively. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837881" figureDoi-1="http://doi.org/10.5281/zenodo.3837885" httpUri-0="https://zenodo.org/record/3837881/files/figure.png" httpUri-1="https://zenodo.org/record/3837885/files/figure.png" pageId="42" pageNumber="43">
Figs 5
<emphasis id="B9363704FFD57D42AD4ED7AFFCEA9C3C" bold="true" box="[812,827,1022,1048]" pageId="42" pageNumber="43"></emphasis>
6
</figureCitation>
).
<collectingCountry id="F355AB86FFD57D42AD07D7AFFC1C9C3C" box="[869,973,1022,1048]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
………
<taxonomicName id="4C429095FFD57D42AA52D7AFFADC9C3C" ID-CoL="7CZMW" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1072,1293,1021,1048]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFD57D42AA52D7AFFADC9C3C" bold="true" box="[1072,1293,1021,1048]" italics="true" pageId="42" pageNumber="43">T. hongkhraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD57D42AB7BD7AFFAA69C3C" box="[1305,1399,1022,1048]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D073FAA69CA6" blockId="42.[189,1399,951,1154]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D073FAA69CA6" pageId="42" pageNumber="43">
Antennae longer (
<figureCitation id="1379F793FFD57D42AFDBD070FDC39C1F" box="[441,530,1057,1084]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, subdorsal, ventral and sublateral views, respectively. IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="42" pageNumber="43">Fig. 7A</figureCitation>
), reaching behind body segment 3. An entire, high, inverted funnel-shaped, sternal lobe between each of Ƌ coxae 3 and 4 (
<figureCitation id="1379F793FFD57D42AC95D015FCB09C7B" box="[759,865,1092,1119]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (BJ), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, subdorsal, ventral and sublateral views, respectively. IJ. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="42" pageNumber="43">
Fig. 7I
<emphasis id="B9363704FFD57D42AD26D014FC829C7B" bold="true" box="[836,851,1093,1119]" pageId="42" pageNumber="43"></emphasis>
J
</figureCitation>
). Goponod without processes (
<figureCitation id="1379F793FFD57D42AAA0D015FAFB9C7A" box="[1218,1322,1092,1119]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="11.[187,230,1964,1990]" captionStartId-1="12.[189,232,1930,1956]" captionTargetBox-0="[299,1276,1063,1923]" captionTargetBox-1="[189,1398,683,1889]" captionTargetId-0="figure@11.[295,1278,1054,1923]" captionTargetId-1="figure@12.[189,1398,683,1889]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 8. Tylopus moniliformis sp. nov., paratype, Ƌ, left gonopod. AB. Mesal and lateral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." captionText-1="Fig. 9. Tylopus moniliformis sp. nov., holotype, Ƌ, left gonopod. AB. Lateral and mesal views, respectively. CF. Distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837889" figureDoi-1="http://doi.org/10.5281/zenodo.3837891" httpUri-0="https://zenodo.org/record/3837889/files/figure.png" httpUri-1="https://zenodo.org/record/3837891/files/figure.png" pageId="42" pageNumber="43">
Figs 8
<emphasis id="B9363704FFD57D42AB68D014FAC89C7B" bold="true" box="[1290,1305,1093,1119]" pageId="42" pageNumber="43"></emphasis>
9
</figureCitation>
).
<collectingCountry id="F355AB86FFD57D42AB5FD014FAA79C7B" box="[1341,1398,1093,1119]" name="Laos" pageId="42" pageNumber="43">Laos</collectingCountry>
………………………………………………………………………………
<taxonomicName id="4C429095FFD57D42AA38D039FAC59CA6" ID-CoL="7CZPF" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1114,1300,1127,1154]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFD57D42AA38D039FAC59CA6" bold="true" box="[1114,1300,1127,1154]" italics="true" pageId="42" pageNumber="43">T. moniliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD57D42AB7ED039FAA69CA6" box="[1308,1399,1128,1154]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD57D42AEDFD0F8FAA79D08" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD0F8FAA69CC2" blockId="42.[189,1399,1193,1324]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD0F8FAA69CC2" pageId="42" pageNumber="43">39. Metaterga entirely smooth and polished, devoid of evident tubercles, at best extremely faintly rugulose near transverse sulcus ……………………………………………………………………40</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D0A1FAA79D08" blockId="42.[189,1399,1193,1324]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D0A1FAA79D08" pageId="42" pageNumber="43"> Metaterga rather clearly rugose/tuberculate/granulate, posterior row of setae at least partly borne on tubercles……………………………………………………………………………………………46</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD57D42AEDFD105FAA69D90" blockId="42.[189,1399,1364,1461]" pageId="42" pageNumber="43">
<keyStep id="30B6F3B3FFD57D42AEDFD105FAA69D90" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD105FAA69D90" pageId="42" pageNumber="43">
40. Transverse sulcus on metaterga starting with segment 3 or 4, but fully developed and reaching bases of paraterga starting with segment 4 or 5. Gonopod process
<emphasis id="B9363704FFD57D42AD9ED126FBDD9DB5" bold="true" box="[1020,1036,1399,1425]" pageId="42" pageNumber="43">h</emphasis>
large (
<figureCitation id="1379F793FFD57D42AA02D126FB629DB5" box="[1120,1203,1399,1425]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. AD. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="42" pageNumber="43">Fig. 17</figureCitation>
AD) ………41 Transverse sulcus on metaterga starting only with segment 5. Gonopod process
<emphasis id="B9363704FFD57D42AA1AD1CAFB599D91" bold="true" box="[1144,1160,1435,1461]" pageId="42" pageNumber="43">h</emphasis>
small (
<figureCitation id="1379F793FFD57D42AAB9D1CBFAFE9D90" box="[1243,1327,1434,1460]" captionStart="Fig" captionStartId="30.[189,232,1966,1992]" captionTargetBox="[311,1244,1002,1916]" captionTargetId="figure@30.[304,1249,1000,1919]" captionTargetPageId="30" captionText="Fig. 22. Tylopus punctus sp. nov., holotype, Ƌ. AD. Left gonopod, lateral, mesal, subcaudal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837919" httpUri="https://zenodo.org/record/3837919/files/figure.png" pageId="42" pageNumber="43">Fig. 22</figureCitation>
) …44
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD57D42AEDFD18DFAA79E44" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD18DFAA69E3D" blockId="42.[189,1399,1500,1632]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD18DFAA69E3D" pageId="42" pageNumber="43">
41. Gonopod with a distinct sharp process on lobe
<emphasis id="B9363704FFD57D42AC9CD18DFCD79DD2" bold="true" box="[766,774,1500,1526]" pageId="42" pageNumber="43">l</emphasis>
. Ventral adenostyles on Ƌ legs absent. Southern
<collectingCountry id="F355AB86FFD57D42AB52D18DFAA79DD2" box="[1328,1398,1500,1526]" name="China" pageId="42" pageNumber="43">China</collectingCountry>
…………………………………………………………………………
<taxonomicName id="4C429095FFD57D42AA43D1AEFAA69E3D" ID-CoL="7CZQ3" authority="Golovatch, 2014" authorityName="Golovatch" authorityYear="2014" box="[1057,1399,1535,1562]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B9363704FFD57D42AA43D1AEFB469E3E" bold="true" box="[1057,1175,1535,1562]" italics="true" pageId="42" pageNumber="43">T. similis</emphasis>
<bibRefCitation id="EFD396E7FFD57D42AACBD1AEFAA69E3D" author="Golovatch S. I." box="[1193,1399,1535,1561]" pageId="42" pageNumber="43" pagination="1 - 19" refId="ref19610" refString="Golovatch S. I. 2014. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XV. Arthropoda Selecta 23 (1): 1 - 19." type="journal article" year="2014">Golovatch, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D272FAA79E44" blockId="42.[189,1399,1500,1632]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D272FAA79E44" pageId="42" pageNumber="43">
Gonopod without processes on lobe
<emphasis id="B9363704FFD57D42ACCAD273FD619E18" bold="true" box="[680,688,1570,1596]" pageId="42" pageNumber="43">l</emphasis>
. Ventral adenostyles on Ƌ legs: a distal knob on femur, a distomedial knob on postfemur, and a parabasal knob on both tibia and tarsus ……………42
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD57D42AEDFD2D6FAA79F0A" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD2D6FAA69EC3" blockId="42.[189,1399,1671,1838]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD2D6FAA69EC3" pageId="42" pageNumber="43">
42. Head wider than collum and subequal in width to segment 3. Gonopod process
<emphasis id="B9363704FFD57D42AA90D2D6FAD39E85" bold="true" box="[1266,1282,1671,1697]" pageId="42" pageNumber="43">h</emphasis>
at about midlength with a strong ventral outgrowth, while solenophore without processes.
<collectingCountry id="F355AB86FFD57D42AAD1D2FBFAC79EE0" box="[1203,1302,1706,1732]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
……… ………………………………………………………………………
<taxonomicName id="4C429095FFD57D42AA60D29FFAA69EC3" ID-CoL="7CZPM" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1026,1399,1741,1768]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="procurvus">
<emphasis id="B9363704FFD57D42AA60D29FFB719ECC" bold="true" box="[1026,1184,1742,1768]" italics="true" pageId="42" pageNumber="43">T. procurvus</emphasis>
<bibRefCitation id="EFD396E7FFD57D42AACFD29CFAA69EC3" author="Golovatch S. I." box="[1197,1399,1741,1767]" pageId="42" pageNumber="43" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD57D42AEA3D2A0FAA79F0A" blockId="42.[189,1399,1671,1838]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEA3D2A0FAA79F0A" pageId="42" pageNumber="43">
Head narrower than collum. Gonopod process
<emphasis id="B9363704FFD57D42AD42D2A0FCE19F2F" bold="true" box="[800,816,1777,1803]" pageId="42" pageNumber="43">h</emphasis>
without outgrowths, while process
<emphasis id="B9363704FFD57D42AA83D2A0FB219F2F" bold="true" box="[1249,1264,1777,1803]" pageId="42" pageNumber="43">x</emphasis>
present on solenophore ………………………………………………………………………………………43
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD57D43AEDFD304FAA6996D" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<paragraph id="8BFDEB16FFD57D42AEDFD304FAA69F92" blockId="42.[189,1399,1877,1974]" pageId="42" pageNumber="43">
<keyLead id="30B34823FFD57D42AEDFD304FAA69F92" pageId="42" pageNumber="43">
43. Antennae shorter (
<figureCitation id="1379F793FFD57D42AFADD304FDEF9F4B" box="[463,574,1877,1903]" captionStart="Fig" captionStartId="22.[189,232,1893,1919]" captionTargetBox="[189,1398,592,1841]" captionTargetId="figure@22.[189,1398,592,1841]" captionTargetPageId="22" captionText="Fig. 16. Tylopus dorsalis sp. nov., holotype, Ƌ. A. Habitus, live coloration. BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837907" httpUri="https://zenodo.org/record/3837907/files/figure.png" pageId="42" pageNumber="43">Fig. 16A</figureCitation>
), extending only behind body segment 2 when stretched dorsally. Ƌ tarsal brushes present until segment 6. Gonopod process
<emphasis id="B9363704FFD57D42ADBCD328FC3C9FB7" bold="true" box="[990,1005,1913,1939]" pageId="42" pageNumber="43">x</emphasis>
longer, the latters tip rounded dorsally (
<figureCitation id="1379F793FFD57D42AF39D3CAFE619F92" box="[347,432,1947,1974]" captionStart="Fig" captionStartId="23.[189,232,1964,1990]" captionTargetBox="[314,1249,559,1910]" captionTargetId="figure@23.[310,1255,559,1920]" captionTargetPageId="23" captionText="Fig. 17. Tylopus dorsalis sp. nov., holotype, Ƌ. AD. Right gonopod, lateral, mesal, caudal and oral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837909" httpUri="https://zenodo.org/record/3837909/files/figure.png" pageId="42" pageNumber="43">Fig. 17</figureCitation>
AD).
<collectingCountry id="F355AB86FFD57D42AC67D3CDFDEF9F92" box="[517,574,1948,1974]" name="Laos" pageId="42" pageNumber="43">Laos</collectingCountry>
……………………………………………………
<taxonomicName id="4C429095FFD57D42AAF0D3CDFAC09F92" ID-CoL="7CZN4" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1170,1297,1947,1974]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="dorsalis" status="sp. nov.">
<emphasis id="B9363704FFD57D42AAF0D3CDFAC09F92" bold="true" box="[1170,1297,1947,1974]" italics="true" pageId="42" pageNumber="43">T. dorsalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD57D42AB78D3CDFAA69F92" box="[1306,1399,1948,1974]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D55DFAA6996D" blockId="43.[193,1399,267,329]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D55DFAA6996D" pageId="43" pageNumber="44">
Antennae longer, reaching until body segment 5. Ƌ tarsal brushes present on all segments. Gonopod process
<emphasis id="B9363704FFD47D43AF28D57EFE88996D" bold="true" box="[330,345,303,329]" pageId="43" pageNumber="44">x</emphasis>
shorter, its tip pointed.
<collectingCountry id="F355AB86FFD47D43AC13D57FFD05996C" box="[625,724,302,328]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
……………………………
<taxonomicName id="4C429095FFD47D43AA7DD57EFAA6996D" ID-CoL="7CZN6" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1055,1399,302,329]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B9363704FFD47D43AA7DD57EFB7B996D" bold="true" box="[1055,1194,303,329]" italics="true" pageId="43" pageNumber="44">T. crassipes</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAD0D57FFAA6996D" author="Golovatch S. I." box="[1202,1399,302,329]" pageId="43" pageNumber="44" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD47D43AEDFD521FAA79A3C" pageId="43" pageNumber="44">
<paragraph id="8BFDEB16FFD47D43AEDFD521FAA7998A" blockId="43.[189,1399,368,536]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD521FAA7998A" pageId="43" pageNumber="44">
44. Small species: width
<quantity id="4CBA46F3FFD47D43AF8FD521FD9C99AE" box="[493,589,368,394]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="43" pageNumber="44" unit="mm" value="1.6">1.6 mm</quantity>
. Only a single row of 2+2 tergal setae. Metaterga entirely smooth. Adenostyles on Ƌ legs present.
<collectingCountry id="F355AB86FFD47D43AC3AD5C2FD6A9989" box="[600,699,403,429]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
……………………………………
<taxonomicName id="4C429095FFD47D43AA3ED5C5FAA7998A" ID-CoL="7CZQ5" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[1116,1398,403,430]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="sigma">
<emphasis id="B9363704FFD47D43AA3ED5C5FB119989" bold="true" box="[1116,1216,403,430]" italics="true" pageId="43" pageNumber="44">T. sigma</emphasis>
(
<bibRefCitation id="EFD396E7FFD47D43AAB2D5C2FABF998A" author="Attems C." box="[1232,1390,403,430]" pageId="43" pageNumber="44" pagination="133 - 230" refId="ref19454" refString="Attems C. 1953. Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, N. S. 5 A: 133 - 230." type="book chapter" year="1953">Attems, 1953</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D5E6FAA79A3C" blockId="43.[189,1399,368,536]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D5E6FAA79A3C" pageId="43" pageNumber="44"> Body wider than 2.0 mm. Metaterga with two transverse rows of setae or traceable insertion points. Metaterga very finely rugulose only near transverse sulcus. Adenostyles on Ƌ legs absent or represented by pads in their stead ……………………………………………………………45</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD47D43AEDFD66FFAA69AC2" pageId="43" pageNumber="44">
<paragraph id="8BFDEB16FFD47D43AEDFD66FFAA69ABB" blockId="43.[189,1399,574,742]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD66FFAA69ABB" pageId="43" pageNumber="44">
45. Paraterga broader:
<quantity id="4CBA46F3FFD47D43AFABD66FFDF29A7D" box="[457,547,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="43" pageNumber="44" unit="mm" value="3.1">3.1 mm</quantity>
wide. Pleurosternal carinae present until segment 13 or 14. Pads present on Ƌ femora, postfemora, tibiae (all distally) and tarsi (almost entirely).
<collectingCountry id="F355AB86FFD47D43AA36D630FB6D9A5F" box="[1108,1212,609,635]" name="Thailand" pageId="43" pageNumber="44">Thailand</collectingCountry>
……………… ……………………………………………………………
<taxonomicName id="4C429095FFD47D43ADEFD6D4FAA69ABB" ID-CoL="7CZPK" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[909,1399,644,671]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pulvinipes">
<emphasis id="B9363704FFD47D43ADEFD6D4FBF69ABB" bold="true" box="[909,1063,644,671]" italics="true" pageId="43" pageNumber="44">T. pulvinipes</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AA4CD6D4FAA69ABB" author="Golovatch S. I. &amp; Enghoff H." box="[1070,1399,645,671]" pageId="43" pageNumber="44" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D6F9FAA69AC2" blockId="43.[189,1399,574,742]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D6F9FAA69AC2" pageId="43" pageNumber="44">
Paraterga narrower:
<quantity id="4CBA46F3FFD47D43AF87D6F9FDA89AE6" box="[485,633,680,707]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="43" pageNumber="44" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
wide. Pleurosternal carinae present until segment 17. Ƌ legs without pads.
<collectingCountry id="F355AB86FFD47D43AFF1D69DFDD49AC2" box="[403,517,716,742]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
…………………………………………………………
<taxonomicName id="4C429095FFD47D43AAF1D69AFAC39AC1" ID-CoL="7CZPX" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1171,1298,715,741]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="punctus" status="sp. nov.">
<emphasis id="B9363704FFD47D43AAF1D69AFAC39AC1" bold="true" box="[1171,1298,715,741]" italics="true" pageId="43" pageNumber="44">T. punctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFD47D43AB79D69DFAA69AC2" box="[1307,1399,716,742]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD47D43AEDFD75DFAA69B6E" blockId="43.[189,1399,780,842]" pageId="43" pageNumber="44">
<keyStep id="30B6F3B3FFD47D43AEDFD75DFAA69B6E" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD75DFAA69B6E" pageId="43" pageNumber="44">46. Metatergal surface polished and smooth except for two rows of conspicuous tubercles ………47 Metaterga at least partly rugulose/rugose to granular; at most one row of tubercles ……………52</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEDFD720FAA79BF6" blockId="43.[189,1399,881,978]" pageId="43" pageNumber="44">
<keyStep id="30B6F3B3FFD47D43AEDFD720FAA79BF6" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD720FAA79BF6" pageId="43" pageNumber="44">
47. Ƌ legs without adenostyles (= tubercles). Gonopods with process
<emphasis id="B9363704FFD47D43AA4AD720FBE99BAF" bold="true" box="[1064,1080,881,907]" pageId="43" pageNumber="44">h</emphasis>
higher than solenophore, strongly twisted, pointed apically. Southern
<collectingCountry id="F355AB86FFD47D43AC8ED7C5FCE39B8A" box="[748,818,916,942]" name="China" pageId="43" pageNumber="44">China</collectingCountry>
…………………
<taxonomicName id="4C429095FFD47D43AA67D7C4FAA69B8A" ID-CoL="7CZQ8" authority="Golovatch, 2013" authorityName="Golovatch" authorityYear="2013" box="[1029,1399,916,943]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="schawalleri">
<emphasis id="B9363704FFD47D43AA67D7C4FB7F9B8B" bold="true" box="[1029,1198,916,943]" italics="true" pageId="43" pageNumber="44">T. schawalleri</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAD6D7C5FAA69B8A" author="Golovatch S. I." box="[1204,1399,916,942]" pageId="43" pageNumber="44" pagination="1 - 31" refId="ref19577" refString="Golovatch S. I. 2013. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIII. Arthropoda Selecta 22 (1): 1 - 31." type="journal article" year="2013">Golovatch, 2013</bibRefCitation>
</taxonomicName>
Ƌ legs with adenostyles (= tubercles) at least on segment 6. Gonopods otherwise.
<collectingCountry id="F355AB86FFD47D43AAACD7E6FAE09BF5" box="[1230,1329,951,977]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
…48
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD47D43AEDFD7A8FAA79D09" pageId="43" pageNumber="44">
<paragraph id="8BFDEB16FFD47D43AEDFD7A8FAA79CE7" blockId="43.[189,1399,1017,1325]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD7A8FAA79CE7" pageId="43" pageNumber="44">
48. Paraterga very poorly developed, rounded, low, projecting slightly behind rear margin like small knobs only on segments 18 and 19. Calluses virtually devoid of incisions. Transverse sulcus on metaterga poorly developed, starting already with segment 2, although fully developed only starting with segment 5. A paramedian pair of denticles between Ƌ coxae 5 behind a prominent, subquadrate lamina between Ƌ coxae 4. Gonopod process
<emphasis id="B9363704FFD47D43AA66D0D7FBC59C84" bold="true" box="[1028,1044,1158,1184]" pageId="43" pageNumber="44">h</emphasis>
entirely missing.
<collectingCountry id="F355AB86FFD47D43AB6CD0D7FAA79C84" box="[1294,1398,1158,1184]" name="Thailand" pageId="43" pageNumber="44">Thailand</collectingCountry>
………………………………………………
<taxonomicName id="4C429095FFD47D43AC98D0F8FAA79CE7" ID-CoL="7CZQB" authority="(Korsos &amp; Golovatch, 1989)" baseAuthorityName="Korsos &amp; Golovatch" baseAuthorityYear="1989" box="[762,1398,1193,1220]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="strongylosomoides">
<emphasis id="B9363704FFD47D43AC98D0F8FC2E9CE0" bold="true" box="[762,1023,1193,1220]" italics="true" pageId="43" pageNumber="44">T. strongylosomoides</emphasis>
(Korsós &amp;
<bibRefCitation id="EFD396E7FFD47D43AAC3D0F8FABF9CE7" author="Golovatch S. I." box="[1185,1390,1193,1219]" pageId="43" pageNumber="44" pagination="211 - 220" refId="ref19813" refString="Kors όs Z. &amp; Golovatch S. I. 1989. Addenda to the millipede fauna of Vietnam (Diplopoda). Acta Zoologica Hungarica 35: 211 - 220." type="journal article" year="1989">Golovatch, 1989</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D09CFAA79D09" blockId="43.[189,1399,1017,1325]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D09CFAA79D09" pageId="43" pageNumber="44">
Paraterga better developed, protruding behind rear margin at least starting with segment 5. Calluses always at least with one lateral incision. Transverse sulcus starting only with segments 35. Dentiform tubercles between Ƌ coxae 5 missing. Gonopod process
<emphasis id="B9363704FFD47D43AA6ED142FBCD9D09" bold="true" box="[1036,1052,1299,1325]" pageId="43" pageNumber="44">h</emphasis>
invariably present ………49
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFDEB16FFD47D43AEDFD105FAA79D91" blockId="43.[189,1398,1364,1461]" pageId="43" pageNumber="44">
<keyStep id="30B6F3B3FFD47D43AEDFD105FAA79D91" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD105FAA79D91" pageId="43" pageNumber="44">
49. Body larger:more than
<quantity id="4CBA46F3FFD47D43AF93D105FD919D4B" box="[497,576,1364,1391]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="mm" value="25.0">25 mm</quantity>
long.Ƌ tarsal brushes not present on all legs.Gonopod lobe
<emphasis id="B9363704FFD47D43AAB8D105FB339D4A" bold="true" box="[1242,1250,1364,1390]" pageId="43" pageNumber="44">l</emphasis>
normal…50 Body smaller: less than
<quantity id="4CBA46F3FFD47D43AC6FD126FDB39DB6" box="[525,610,1399,1426]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="43" pageNumber="44" unit="mm" value="18.0">18 mm</quantity>
long. Ƌ tarsal brushes present on all legs. Gonopod lobe
<emphasis id="B9363704FFD47D43AB78D126FAF39DB5" bold="true" box="[1306,1314,1399,1425]" pageId="43" pageNumber="44">l</emphasis>
with a spine apically ………………………………………………………………………………………51
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30B6F3B3FFD47D43AEDFD18DFAA69E44" pageId="43" pageNumber="44">
<paragraph id="8BFDEB16FFD47D43AEDFD18DFAA69E3D" blockId="43.[189,1399,1500,1632]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD18DFAA69E3D" pageId="43" pageNumber="44">
50. Ƌ tarsal brushes present on legs 115. Pleurosternal carinae present until segment 9. Gonopod process
<emphasis id="B9363704FFD47D43AF29D1AEFE8A9E3D" bold="true" box="[331,347,1535,1561]" pageId="43" pageNumber="44">h</emphasis>
larger, finger-shaped.
<collectingCountry id="F355AB86FFD47D43AC07D1AEFD199E3D" box="[613,712,1535,1561]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
…………………………
<taxonomicName id="4C429095FFD47D43AD95D1AEFAA69E3D" ID-CoL="7CZPV" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[1015,1399,1535,1562]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="roseiparaterga">
<emphasis id="B9363704FFD47D43AD95D1AEFB179E3D" bold="true" box="[1015,1222,1535,1561]" italics="true" pageId="43" pageNumber="44">T. roseiparaterga</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAADD251FAA69E3D" author="Nguyen A. D." box="[1231,1399,1535,1562]" pageId="43" pageNumber="44" pagination="289 - 311" refId="ref20147" refString="Nguyen A. D. 2012. Tylopus millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology 60 (2): 289 - 311." type="journal article" year="2012">Nguyen, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D272FAA69E44" blockId="43.[189,1399,1500,1632]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D272FAA69E44" pageId="43" pageNumber="44">
Ƌ tarsal brushes present on legs 530. Pleurosternal carinae present until segment 17. Gonopod process
<emphasis id="B9363704FFD47D43AF2AD217FE899E44" bold="true" box="[328,344,1606,1632]" pageId="43" pageNumber="44">h</emphasis>
small, slender and pointed. Southern
<collectingCountry id="F355AB86FFD47D43AD6FD214FC829E7B" box="[781,851,1605,1631]" name="China" pageId="43" pageNumber="44">China</collectingCountry>
………………
<taxonomicName id="4C429095FFD47D43AA6BD217FAA69E44" ID-CoL="7CZN7" authority="Liu &amp; Luo, 2013" authorityName="Liu &amp; Luo" authorityYear="2013" box="[1033,1399,1605,1632]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B9363704FFD47D43AA6BD217FB7C9E44" bold="true" box="[1033,1197,1605,1632]" italics="true" pageId="43" pageNumber="44">T. deharvengi</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAD1D214FAA69E44" author="Liu W. X. &amp; Luo X. Z." box="[1203,1399,1605,1632]" pageId="43" pageNumber="44" pagination="50 - 52" refId="ref20102" refString="Liu W. X. &amp; Luo X. Z. 2013. A new species of the millipede genus Tylopus Jeekel from southern China (Diplopoda, Polydesmida, Paradoxosomatidae). Acta Zootaxonomica Sinica 38 (1): 50 - 52." type="journal article" year="2013">Liu &amp; Luo, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD47D43AEDFD2D6FAA69FB3" pageId="43" pageNumber="44">
<paragraph id="8BFDEB16FFD47D43AEDFD2D6FAA69F2F" blockId="43.[189,1399,1671,1944]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEDFD2D6FAA69F2F" pageId="43" pageNumber="44">
51. Paraterga acutangular caudally and pointed beak-like already starting with collum. Tergal setigerous tubercles: 3+3 and 5+5 on segments 1619. Adenostyle pattern on Ƌ legs: a distal knob on femora and a parabasal knob on most of postfemora, tibiae and tarsi. Gonopod process
<emphasis id="B9363704FFD47D43AAE4D29CFB479EC3" bold="true" box="[1158,1174,1741,1767]" pageId="43" pageNumber="44">h</emphasis>
large, lamellar and sigmoid.
<collectingCountry id="F355AB86FFD47D43AF34D2A1FE689F2E" box="[342,441,1776,1802]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
……………………………………………………
<taxonomicName id="4C429095FFD47D43AA69D2A0FAA69F2F" ID-CoL="7CZMR" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1035,1399,1776,1803]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9363704FFD47D43AA69D2A0FB7C9F2F" bold="true" box="[1035,1197,1776,1803]" italics="true" pageId="43" pageNumber="44">T. granulatus</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAD1D2A1FAA69F2F" author="Golovatch S. I." box="[1203,1399,1776,1803]" pageId="43" pageNumber="44" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD47D43AEA3D345FAA69FB3" blockId="43.[189,1399,1671,1944]" pageId="43" pageNumber="44">
<keyLead id="30B34823FFD47D43AEA3D345FAA69FB3" pageId="43" pageNumber="44">
Paraterga acutangular caudally and pointed beak-like only starting with segment 4. 2+2 and 4+4 tergal setigerous tubercles on segments 1619. Adenostyle pattern on Ƌ legs: a proximal fingershaped tubercle crowned with a bunch of setae only on femora 6, 8 and 9. Gonopod process
<emphasis id="B9363704FFD47D43AB66D30BFAC59F50" bold="true" box="[1284,1300,1882,1908]" pageId="43" pageNumber="44">h</emphasis>
smaller, spiniform.
<collectingCountry id="F355AB86FFD47D43AF0FD32CFE019FB3" box="[365,464,1917,1943]" name="Vietnam" pageId="43" pageNumber="44">Vietnam</collectingCountry>
………………………………………………………
<taxonomicName id="4C429095FFD47D43AA20D32FFAA69FB3" ID-CoL="7CZP6" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[1090,1399,1917,1944]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="topali">
<emphasis id="B9363704FFD47D43AA20D32FFB779FBC" bold="true" box="[1090,1190,1917,1944]" italics="true" pageId="43" pageNumber="44">T. topali</emphasis>
<bibRefCitation id="EFD396E7FFD47D43AAD2D32CFAA69FB3" author="Golovatch S. I." box="[1200,1399,1917,1943]" pageId="43" pageNumber="44" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD55AFAA699B4" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD55AFAA6996D" blockId="44.[189,1399,267,400]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD55AFAA6996D" pageId="44" pageNumber="45">
52. Transverse sulcus on metaterga starting and fully developed only with segment 5. Ventral adenostyles present on all Ƌ podomeres except coxa.
<collectingCountry id="F355AB86FFD37D44ADCED57FFBDE996C" box="[940,1039,302,328]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
and/or
<collectingCountry id="F355AB86FFD37D44AA1ED57EFB64996D" box="[1148,1205,303,329]" name="Laos" pageId="44" pageNumber="45">Laos</collectingCountry>
……………53
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D503FAA699B4" blockId="44.[189,1399,267,400]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D503FAA699B4" pageId="44" pageNumber="45">
Transverse sulcus on metaterga starting with segment 4, but fully developed only starting with segment 5. Ventral tubercles only on some of Ƌ telopoditomeres.
<collectingCountry id="F355AB86FFD37D44AA50D524FB4B99AB" box="[1074,1178,373,399]" name="Thailand" pageId="44" pageNumber="45">Thailand</collectingCountry>
………………54
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD5E7FAA79AA5" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD5E7FAA69A33" blockId="44.[189,1399,438,641]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD5E7FAA69A33" pageId="44" pageNumber="45">
53. Metaterga rugulose also in front of transverse sulcus, at rear margin with several oblong tubercles. Sternal lamina between Ƌ coxae 4 a pair of setigerous knobs. Neither gonopod lobe
<emphasis id="B9363704FFD37D44AAC4D58BFB6F99D0" bold="true" box="[1190,1214,474,500]" pageId="44" pageNumber="45">m</emphasis>
nor lobe
<emphasis id="B9363704FFD37D44AB45D58BFAFE99D0" bold="true" box="[1319,1327,474,500]" pageId="44" pageNumber="45">l</emphasis>
spinigerous.
<collectingCountry id="F355AB86FFD37D44AF24D5ACFE789A33" box="[326,425,509,535]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
…………………………………………
<taxonomicName id="4C429095FFD37D44ADE6D5ACFAA69A33" ID-CoL="7CZP8" authority="Korsos &amp; Golovatch, 1989" authorityName="Korsos &amp; Golovatch" authorityYear="1989" box="[900,1399,508,535]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B9363704FFD37D44ADE6D5ACFBE69A33" bold="true" box="[900,1079,508,535]" italics="true" pageId="44" pageNumber="45">T. tamdaoensis</emphasis>
Korsós &amp;
<bibRefCitation id="EFD396E7FFD37D44AAD6D5ACFAA69A33" author="Golovatch S. I." box="[1204,1399,509,535]" pageId="44" pageNumber="45" pagination="211 - 220" refId="ref19813" refString="Kors όs Z. &amp; Golovatch S. I. 1989. Addenda to the millipede fauna of Vietnam (Diplopoda). Acta Zoologica Hungarica 35: 211 - 220." type="journal article" year="1989">Golovatch, 1989</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D671FAA79AA5" blockId="44.[189,1399,438,641]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D671FAA79AA5" pageId="44" pageNumber="45">
Metaterga rugose only behind transverse sulcus, without evident tubercles at rear margin. Sternal lamina between Ƌ coxae 4 single. Both gonopod lobe
<emphasis id="B9363704FFD37D44ADE0D612FC4B9A79" bold="true" box="[898,922,579,605]" pageId="44" pageNumber="45">m</emphasis>
and lobe
<emphasis id="B9363704FFD37D44AA74D612FBCF9A79" bold="true" box="[1046,1054,579,605]" pageId="44" pageNumber="45">l</emphasis>
crowned with a spine.
<collectingCountry id="F355AB86FFD37D44AB5FD615FAA79A7A" box="[1341,1398,580,606]" name="Laos" pageId="44" pageNumber="45">Laos</collectingCountry>
and
<collectingCountry id="F355AB86FFD37D44AF7ED637FEAE9AA4" box="[284,383,614,640]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
……………………………………………………………
<taxonomicName id="4C429095FFD37D44AA4BD636FAA79AA5" ID-CoL="7CZPD" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[1065,1398,614,641]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nodulipes">
<emphasis id="B9363704FFD37D44AA4BD636FB6D9AA5" bold="true" box="[1065,1212,614,641]" italics="true" pageId="44" pageNumber="45">T. nodulipes</emphasis>
(
<bibRefCitation id="EFD396E7FFD37D44AAACD636FABF9AA5" author="Attems C." box="[1230,1390,614,641]" pageId="44" pageNumber="45" pagination="133 - 230" refId="ref19454" refString="Attems C. 1953. Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, N. S. 5 A: 133 - 230." type="book chapter" year="1953">Attems, 1953</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD6F9FAA69B57" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD6F9FAA69B2D" blockId="44.[189,1399,680,883]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD6F9FAA69B2D" pageId="44" pageNumber="45">
54. Metaterga modestly rugulose only near transverse sulcus, posteriorly neither granular nor microtuberculate. Calluses broad. Gonopod process
<emphasis id="B9363704FFD37D44AD2DD69AFC8E9AC1" bold="true" box="[847,863,715,741]" pageId="44" pageNumber="45">h</emphasis>
simple, high, never particularly coiled; lobe
<emphasis id="B9363704FFD37D44AB0CD69AFAA79AC1" bold="true" box="[1390,1398,715,741]" pageId="44" pageNumber="45">l</emphasis>
very moderately serrate at apex ……………………………………………………………………55
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D743FAA69B57" blockId="44.[189,1399,680,883]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D743FAA69B57" pageId="44" pageNumber="45">
Metaterga distinctly rugose-granular/microtuberculate even on fore halves. Calluses narrow. Gonopod process
<emphasis id="B9363704FFD37D44AFA0D764FE039B6B" bold="true" box="[450,466,821,847]" pageId="44" pageNumber="45">h</emphasis>
better developed and more strongly coiled; lobe
<emphasis id="B9363704FFD37D44AA43D764FBF89B6B" bold="true" box="[1057,1065,821,847]" pageId="44" pageNumber="45">l</emphasis>
apically either bare or with a digitiform outgrowth ……………………………………………………………………………56
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD7C8FAA69C64" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD7C8FAA69BDE" blockId="44.[189,1399,921,1089]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD7C8FAA69BDE" pageId="44" pageNumber="45">
55. Caudal corner of paraterga pointed starting with segment 3. Pleurosternal carinae particularly well-developed, drawn behind rear margin until segment 16 or 17. Adenostyles often present on Ƌ prefemora …………………………………………
<taxonomicName id="4C429095FFD37D44AD06D7B1FAA69BDE" ID-CoL="7CZMV" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[868,1399,991,1018]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="degerboelae">
<emphasis id="B9363704FFD37D44AD06D7B1FBC69BDE" bold="true" box="[868,1047,991,1018]" italics="true" pageId="44" pageNumber="45">T. degerboelae</emphasis>
<bibRefCitation id="EFD396E7FFD37D44AA40D7B1FAA69BDE" author="Golovatch S. I. &amp; Enghoff H." box="[1058,1399,992,1018]" pageId="44" pageNumber="45" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D052FAA69C64" blockId="44.[189,1399,921,1089]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D052FAA69C64" pageId="44" pageNumber="45">
Caudal corners of paraterga mainly narrowly rounded, pointed only starting with segment 15. Pleurosternal carinae less strongly developed ………………………
<taxonomicName id="4C429095FFD37D44ADD1D077FAA69C64" ID-CoL="7CZPB" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[947,1399,1062,1089]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B9363704FFD37D44ADD1D077FBFE9C65" bold="true" box="[947,1071,1062,1089]" italics="true" pageId="44" pageNumber="45">T. pallidus</emphasis>
<bibRefCitation id="EFD396E7FFD37D44AA56D077FAA69C64" author="Golovatch S. I. &amp; Enghoff H." box="[1076,1399,1062,1088]" pageId="44" pageNumber="45" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD036FAA79D2B" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD036FAA69CEC" blockId="44.[189,1399,1127,1295]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD036FAA69CEC" pageId="44" pageNumber="45">
56. Coloration dark, brown. Sternal lamina between Ƌ coxae 4 a pair of separated setigerous tubercles preceded by another pair of very small tubercles between coxae 3. Gonopod lobe
<emphasis id="B9363704FFD37D44AAAED0DAFB059C81" bold="true" box="[1228,1236,1163,1189]" pageId="44" pageNumber="45">l</emphasis>
devoid of an apical process …………………………………………………
<taxonomicName id="4C429095FFD37D44ADA7D0FFFAA69CEC" ID-CoL="7CZMN" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[965,1399,1198,1224]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B9363704FFD37D44ADA7D0FFFBF59CEC" bold="true" box="[965,1060,1198,1224]" italics="true" pageId="44" pageNumber="45">T. asper</emphasis>
<bibRefCitation id="EFD396E7FFD37D44AA4ED0FFFAA69CEC" author="Golovatch S. I. &amp; Enghoff H." box="[1068,1399,1198,1224]" pageId="44" pageNumber="45" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D083FAA79D2B" blockId="44.[189,1399,1127,1295]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D083FAA79D2B" pageId="44" pageNumber="45">
Coloration uniformly pale. Sternal lamina between Ƌ coxae 4 single, not accompanied by additional tubercles in front or behind. Gonopod lobe
<emphasis id="B9363704FFD37D44AC98D0A4FCD39D2B" bold="true" box="[762,770,1269,1295]" pageId="44" pageNumber="45">l</emphasis>
with a strong, apical, finger-shaped process ……
<emphasis id="B9363704FFD37D44AB3BD0A4FAA79D2B" bold="true" box="[1369,1398,1269,1295]" pageId="44" pageNumber="45">57</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6F3B3FFD37D44AEDFD167FAA69E62" pageId="44" pageNumber="45">
<paragraph id="8BFDEB16FFD37D44AEDFD167FAA69D9D" blockId="44.[189,1406,1334,1607]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEDFD167FAA69D9D" pageId="44" pageNumber="45">
57. Mostly 3+3 tubercles at rear margin of metaterga. Sternal lamina between Ƌ coxae 4 distinctly emarginate. Larger adenostyles close to midlength on Ƌ postfemora and tibiae, femora with a distal knob. Gonopod process
<emphasis id="B9363704FFD37D44AC33D12DFDB09DB2" bold="true" box="[593,609,1404,1430]" pageId="44" pageNumber="45">h</emphasis>
slenderer and shorter, apex of lobule
<emphasis id="B9363704FFD37D44AA42D12DFBE99DB2" bold="true" box="[1056,1080,1404,1430]" pageId="44" pageNumber="45">m</emphasis>
not developed into a spine ………………………………………………………
<taxonomicName id="4C429095FFD37D44AD30D1F1FAA69D9D" ID-CoL="7CZPZ" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[850,1399,1439,1466]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subcoriaceus">
<emphasis id="B9363704FFD37D44AD30D1F1FBC29D9E" bold="true" box="[850,1043,1439,1466]" italics="true" pageId="44" pageNumber="45">T. subcoriaceus</emphasis>
<bibRefCitation id="EFD396E7FFD37D44AA7DD1CEFAA69D9D" author="Golovatch S. I. &amp; Enghoff H." box="[1055,1399,1439,1465]" pageId="44" pageNumber="45" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDEB16FFD37D44AEA3D192FAA69E62" blockId="44.[189,1406,1334,1607]" pageId="44" pageNumber="45">
<keyLead id="30B34823FFD37D44AEA3D192FAA69E62" pageId="44" pageNumber="45">
Mostly 4+4 tubercles at rear margin of metaterga. Sternal lamina between Ƌ coxae trapeziform. Larger adenostyles on both postfemora and tibiae more distal, femoral knob missing. Gonopod process
<emphasis id="B9363704FFD37D44AF29D258FE8A9E07" bold="true" box="[331,347,1545,1571]" pageId="44" pageNumber="45">h</emphasis>
unusually prominent, with a hook at base; apex of lobule
<emphasis id="B9363704FFD37D44AA6CD258FBF79E07" bold="true" box="[1038,1062,1545,1571]" pageId="44" pageNumber="45">m</emphasis>
spiniform ………………… ……………………………………………………………
<taxonomicName id="4C429095FFD37D44ADEFD27CFAA69E62" ID-CoL="7CZMG" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[909,1399,1580,1607]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="B9363704FFD37D44ADEFD27CFBF19E62" bold="true" box="[909,1056,1580,1607]" italics="true" pageId="44" pageNumber="45">T. coriaceus</emphasis>
<bibRefCitation id="EFD396E7FFD37D44AA4BD27DFAA69E62" author="Golovatch S. I. &amp; Enghoff H." box="[1065,1399,1580,1606]" pageId="44" pageNumber="45" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>