215 lines
20 KiB
XML
215 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5575697" ID-GBIF-Dataset="887f4c2c-1812-4aa5-b994-58388f6a45c5" ID-GBIF-Taxon="131977847" ID-Zenodo-Dep="5575697" checkinTime="1497949082728" checkinUser="jeremy" docAuthor="Deeleman-Reinhold, Christa" docDate="2001" docId="03B3FE72C96FFFB464FC8AE2DE8FFC14" docLanguage="en" docName="Deeleman-Reinhold_2001_LIOCRANIDAE_mat_cite_checked_IV.imf" docOrigin="Forest Spiders of South East Asia With a revision of the sac and ground spiders- Family Liocranidae, Leiden, Netherlands: Brill Leiden; Boston; Köln" docStyle="DocumentStyle{}" docTitle="Teutamus rhino Deeleman-Reinhold, 2001, sp. n" docType="treatment" docVersion="23" lastPageId="83" lastPageNumber="483" masterDocId="FF8A860AC93EFFE765528965DD50FFD6" masterDocTitle="Forest Spiders of South East Asia With a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae)." masterLastPageNumber="505" masterPageNumber="400" pageId="81" pageNumber="481" updateTime="1634566706600" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Forest Spiders of South East Asia With a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae).</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deeleman-Reinhold, Christa</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:originInfo>
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<mods:dateIssued>2001</mods:dateIssued>
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<mods:publisher>Brill Leiden; Boston; Köln</mods:publisher>
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<mods:place>
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<mods:placeTerm>Leiden, Netherlands</mods:placeTerm>
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</mods:place>
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</mods:originInfo>
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<mods:titleInfo>
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<mods:title>Forest Spiders of South East Asia With a revision of the sac and ground spiders- Family Liocranidae</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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||
<mods:extent unit="page">
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<mods:start>400</mods:start>
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<mods:end>505</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>book chapter</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.814704</mods:identifier>
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<mods:identifier type="GBIF-Dataset">887f4c2c-1812-4aa5-b994-58388f6a45c5</mods:identifier>
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<mods:identifier type="Zenodo-Dep">814704</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5575697" ID-GBIF-Taxon="131977847" ID-Zenodo-Dep="5575697" LSID="urn:lsid:plazi:treatment:03B3FE72C96FFFB464FC8AE2DE8FFC14" httpUri="http://treatment.plazi.org/id/03B3FE72C96FFFB464FC8AE2DE8FFC14" lastPageId="83" lastPageNumber="483" pageId="81" pageNumber="481">
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<subSubSection pageId="81" pageNumber="481" type="nomenclature">
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<paragraph blockId="81.[430,718,901,965]" pageId="81" pageNumber="481">
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<heading box="[430,718,903,931]" centered="true" fontSize="12" level="6" pageId="81" pageNumber="481" reason="1">
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<taxonomicName box="[430,636,903,931]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="481" phylum="Arthropoda" rank="species" species="rhino" status="sp. n">
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<emphasis box="[430,636,903,931]" italics="true" pageId="81" pageNumber="481">Teutamus rhino</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[648,710,903,931]" pageId="81" pageNumber="481" rank="species">sp. n</taxonomicNameLabel>
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,
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</heading>
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<heading box="[438,709,941,965]" centered="true" fontSize="12" level="6" pageId="81" pageNumber="481" reason="1">
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(
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<figureCitation box="[446,599,941,965]" captionStart="Figs 818 - 823" captionStartId="82.[147,191,1784,1803]" captionTargetBox="[158,1184,250,1661]" captionTargetId="figure@82.[158,1184,250,1661]" captionTargetPageId="82" captionText="Figs 818 - 823. Teutamus rhino sp. n. 818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal." httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="81" pageNumber="481">figs 818-823</figureCitation>
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,
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<figureCitation box="[613,701,941,965]" captionStart="Map 40" captionStartId="71.[40,84,1780,1799]" captionTargetBox="[56,1060,1020,1721]" captionTargetId="figure@71.[56,1060,1020,1721]" captionTargetPageId="71" captionText="Map 40: ● Teutamus politus ■ T. fertilis ▲ T. jambiensis ▼ T. rhino ⁎ T. rothorum ★ T. andrewdavisi ◙ T. vittatus Ө Unidentified specimen 1 - 7." httpUri="https://zenodo.org/record/814764/files/figure.png" pageId="81" pageNumber="481">map 40</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="81" pageNumber="481" type="materials_examined">
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<materialsCitation ID-GBIF-Occurrence="1569619176" collectingDate="1986-11-19" collectingDateMax="1986-11-25" collectingDateMin="1986-11-19" collectorName="S. Djojosudharmo" country="Indonesia" location="Ujung Kulon Reserve" municipality="Java" pageId="81" pageNumber="481" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<paragraph blockId="81.[44,601,1009,1035]" box="[44,601,1011,1035]" pageId="81" pageNumber="481">
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<emphasis box="[44,200,1011,1035]" italics="true" pageId="81" pageNumber="481">Type locality.</emphasis>
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— W
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<collectingMunicipality box="[270,323,1011,1035]" pageId="81" pageNumber="481">Java</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:03B3FE72C96FFFB464FC8AE2DE8FFC14:8EC519BFC96FFFB664008A96DF06FBDD" box="[338,598,1011,1035]" municipality="Java" name="Ujung Kulon Reserve" pageId="81" pageNumber="481">Ujung Kulon Reserve</location>
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.
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</paragraph>
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<paragraph blockId="81.[43,1104,1078,1175]" pageId="81" pageNumber="481">
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<emphasis box="[44,96,1081,1105]" italics="true" pageId="81" pageNumber="481">Type</emphasis>
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material.—
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<typeStatus box="[247,354,1081,1105]" pageId="81" pageNumber="481">Holotype</typeStatus>
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♂ from the type locality, swampy primary forest, from leaf litter,
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<collectingDate box="[93,254,1116,1140]" pageId="81" pageNumber="481" value="1986-11-19" valueMax="1986-11-25" valueMin="1986-11-19">
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<date box="[93,254,1116,1140]" pageId="81" pageNumber="481" value="1986-11-19" valueMax="1986-11-25" valueMin="1986-11-19">19-25.xi. 1986</date>
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</collectingDate>
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,
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<collectorName box="[269,482,1116,1140]" pageId="81" pageNumber="481">S. Djojosudharmo</collectorName>
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et. al.;
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<materialsCitation ID-GBIF-Occurrence="1569619168" collectingDate="1986-11-19" collectingDateMax="1986-11-25" collectingDateMin="1986-11-19" collectorName="S. Djojosudharmo" country="Indonesia" location="Ujung Kulon Reserve" municipality="Java" pageId="81" pageNumber="481" specimenCount="25" specimenCount-female="14" specimenCount-male="11" typeStatus="paratype">
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<typeStatus box="[572,687,1116,1140]" pageId="81" pageNumber="481">paratypes</typeStatus>
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: 11 ♂, 14 ♀ from the type locality, same data
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</materialsCitation>
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.
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</paragraph>
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</materialsCitation>
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<paragraph blockId="81.[44,329,1218,1245]" box="[44,329,1221,1245]" pageId="81" pageNumber="481">
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<emphasis box="[44,112,1221,1245]" italics="true" pageId="81" pageNumber="481">Other</emphasis>
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material.— None.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="81" pageNumber="481" type="diagnosis">
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<paragraph blockId="81.[43,1104,1288,1489]" pageId="81" pageNumber="481">
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<emphasis box="[44,169,1291,1315]" italics="true" pageId="81" pageNumber="481">Diagnosis.</emphasis>
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— Distinct from
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<taxonomicName box="[363,490,1291,1315]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="481" phylum="Arthropoda" rank="species" species="jambiensis">
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<emphasis box="[363,490,1291,1315]" italics="true" pageId="81" pageNumber="481">jambiensis</emphasis>
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</taxonomicName>
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by the pattern of the abdomen with oval white spots ventro-laterally in the pre-anal area, the less elongate carapace and in the male palp the tibial apophysis with more acute lower apophysis and the straight spinelike conductor. The present species can be separated from
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<taxonomicName box="[498,604,1396,1420]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="481" phylum="Arthropoda" rank="species" species="rothorum">
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<emphasis box="[498,604,1396,1420]" italics="true" pageId="81" pageNumber="481">rothorum</emphasis>
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</taxonomicName>
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in the male by the presence of an uptilted distal tibial apophysis. In the epigyne long sinuous ducts and round spermathecae are visible.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="83" lastPageNumber="483" pageId="81" pageNumber="481" type="description">
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<paragraph blockId="81.[43,1105,1532,1835]" lastBlockId="83.[43,1104,201,788]" lastPageId="83" lastPageNumber="483" pageId="81" pageNumber="481">
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<emphasis box="[43,189,1534,1558]" italics="true" pageId="81" pageNumber="481">Description.</emphasis>
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— MALE. Total length
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<quantity box="[473,581,1534,1558]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" pageId="81" pageNumber="481" unit="mm" value="4.45">4.45 mm</quantity>
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. Carapace length
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<quantity box="[796,902,1534,1558]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="81" pageNumber="481" unit="mm" value="2.25">2.25 mm</quantity>
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, width
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<quantity box="[996,1098,1534,1558]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="481" unit="mm" value="1.5">1.50 mm</quantity>
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, height
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<quantity box="[127,235,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="81" pageNumber="481" unit="mm" value="0.75">0.75 mm</quantity>
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, width head
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<quantity box="[394,503,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="81" pageNumber="481" unit="mm" value="0.9">0.90 mm</quantity>
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, width eye group
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<quantity box="[727,835,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="81" pageNumber="481" unit="mm" value="0.7">0.70 mm</quantity>
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; abdomen length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="81" pageNumber="481" unit="mm" value="2.2">2.20 mm</quantity>
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, width
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||
<quantity box="[188,295,1603,1627]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="81" pageNumber="481" unit="mm" value="1.3">1.30 mm</quantity>
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||
. Leg length: leg I
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||
<quantity box="[528,633,1603,1627]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" pageId="81" pageNumber="481" unit="mm" value="8.45">8.45 mm</quantity>
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||
(2.10-3.05-2.05-1.25); leg II
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||
<quantity box="[998,1102,1603,1627]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="81" pageNumber="481" unit="mm" value="6.6">6.60 mm</quantity>
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||
(1.75-2.40-1.45-1.00); leg III
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||
<quantity box="[396,498,1638,1662]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" pageId="81" pageNumber="481" unit="mm" value="5.65">5.65 mm</quantity>
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||
(1.50-1.75-1.50-0.90); leg IV
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||
<quantity box="[861,963,1638,1662]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="81" pageNumber="481" unit="mm" value="8.1">8.10 mm</quantity>
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||
(2.10-2.50- 2.25-1.25), palp 0.55-0.25-0.35-
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<quantity box="[427,540,1673,1697]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="81" pageNumber="481" unit="mm" value="0.75">0.75 mm</quantity>
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||
. Colour of carapace, mouthparts, sternum and legs as in
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<emphasis box="[179,343,1708,1732]" italics="true" pageId="81" pageNumber="481">
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<taxonomicName box="[179,338,1708,1732]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="481" phylum="Arthropoda" rank="species" species="jambiensis">T. jambiensis</taxonomicName>
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,
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</emphasis>
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legs with red-orange femora only, other leg segments paler. Abdomen with distinctive pattern (
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<figureCitation box="[453,548,1742,1766]" captionStart="Figs 818 - 823" captionStartId="82.[147,191,1784,1803]" captionTargetBox="[158,1184,250,1661]" captionTargetId="figure@82.[158,1184,250,1661]" captionTargetPageId="82" captionText="Figs 818 - 823. Teutamus rhino sp. n. 818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal." httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="81" pageNumber="481">fig. 820</figureCitation>
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): dorsally blackish with the exception of a long reddish median band in the anterior part, behind which in the middle a black transverse bar which is interrupted in the middle, running somewhat diagonally; dorsal white preanal spot laterally extended with “wings” which sometimes connect to a ventral white zone; this zone is separated from the spinnerets by a black semicirlce. Eyes as in
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<emphasis italics="true" pageId="83" pageNumber="483">jambi ensis.</emphasis>
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Femur 1 with 2pl, tibia I with 9pv, 8-9rv, tibia
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<date box="[666,705,274,298]" pageId="83" pageNumber="483">II 8</date>
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pv, 8rv, metatarsus
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<date box="[937,989,274,298]" pageId="83" pageNumber="483">I 5-6</date>
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pv, 5-6rv, metatarsus
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<date box="[178,219,309,333]" pageId="83" pageNumber="483">II 5</date>
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pv, 4rv. Dorsal scutum covering more than 3/4 of abdomen; ventral scutum as long as dorsal scutum, collar with 3 transverse grooves, rear margin irregular, frayed. Male palp
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<figureCitation box="[168,318,379,403]" captionStart="Figs 818 - 823" captionStartId="82.[147,191,1784,1803]" captionTargetBox="[158,1184,250,1661]" captionTargetId="figure@82.[158,1184,250,1661]" captionTargetPageId="82" captionText="Figs 818 - 823. Teutamus rhino sp. n. 818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal." httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="83" pageNumber="483">figs 818-819</figureCitation>
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, distal tibial apophysis as large as proximal one, uptilted, bulb apically with a chitinous spine-like conductor, no other apophysis apparent.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="82" pageNumber="482" targetBox="[158,1184,250,1661]" targetPageId="82">
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||
<paragraph blockId="82.[147,1198,1779,1835]" pageId="82" pageNumber="482">
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<emphasis box="[147,191,1784,1803]" italics="true" pageId="82" pageNumber="482">Figs</emphasis>
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818-823.
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<taxonomicName box="[315,413,1784,1803]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="482" phylum="Arthropoda" rank="genus">
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<emphasis box="[315,413,1784,1803]" italics="true" pageId="82" pageNumber="482">Teutamus</emphasis>
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</taxonomicName>
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<taxonomicName box="[428,481,1784,1803]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="482" phylum="Arthropoda" rank="species" species="rhino" status="sp. n.">rhino</taxonomicName>
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<taxonomicNameLabel box="[497,558,1784,1803]" pageId="82" pageNumber="482" rank="species">sp. n.</taxonomicNameLabel>
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818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal.
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</paragraph>
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</caption>
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<paragraph blockId="83.[43,1104,201,788]" pageId="83" pageNumber="483">
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FEMALE. Total length
|
||
<quantity box="[327,435,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="83" pageNumber="483" unit="mm" value="4.6">4.60 mm</quantity>
|
||
. Carapace length
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||
<quantity box="[652,759,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="83" pageNumber="483" unit="mm" value="2.15">2.15 mm</quantity>
|
||
, width
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||
<quantity box="[854,957,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="83" pageNumber="483" unit="mm" value="1.5">1.50 mm</quantity>
|
||
, head width
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<quantity box="[43,147,484,508]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="83" pageNumber="483" unit="mm" value="0.85">0.85 mm</quantity>
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||
, eye group width
|
||
<quantity box="[351,454,484,508]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="83" pageNumber="483" unit="mm" value="0.7">0.70 mm</quantity>
|
||
, abdomen length
|
||
<quantity box="[658,760,484,508]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="83" pageNumber="483" unit="mm" value="2.5">2.50 mm</quantity>
|
||
, width
|
||
<quantity box="[849,949,484,508]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="83" pageNumber="483" unit="mm" value="1.5">1.50 mm</quantity>
|
||
. Leg lengths: leg I
|
||
<quantity box="[102,202,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" pageId="83" pageNumber="483" unit="mm" value="8.45">8.45 mm</quantity>
|
||
(2.10-3.15-2.10-1.10), leg II
|
||
<quantity box="[546,647,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="83" pageNumber="483" unit="mm" value="6.7">6.70 mm</quantity>
|
||
(1.75-2.45-1.50-1.00), leg III
|
||
<quantity box="[1002,1102,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="83" pageNumber="483" unit="mm" value="5.8">5.80 mm</quantity>
|
||
(1.50-1.80-1.50-1.00), leg IV
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||
<quantity box="[410,515,554,578]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="83" pageNumber="483" unit="mm" value="8.4">8.40 mm</quantity>
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(2.25-2.65-2.50-1.00), palp 0.65-0.20-0.42-
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="83" pageNumber="483" unit="mm" value="0.65">0.65 mm</quantity>
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||
; epigyne
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||
<quantity box="[204,307,589,613]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="83" pageNumber="483" unit="mm" value="0.45">0.45 mm</quantity>
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||
wide,
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||
<quantity box="[388,490,589,613]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="83" pageNumber="483" unit="mm" value="0.4">0.40 mm</quantity>
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||
long. Abdominal pattern more diluted than in male, dorsal pattern white on a grey background, ventral surface of abdomen white except for the black pre-anal semicircle. Collar a little shorter than in male, with 1-2 grooves. Spine formula: tibiae I and II with 9 pairs, metatarsus I with 6, metatarsus II with 5 pairs. Epigyne
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<figureCitation captionStart="Figs 818 - 823" captionStartId="82.[147,191,1784,1803]" captionTargetBox="[158,1184,250,1661]" captionTargetId="figure@82.[158,1184,250,1661]" captionTargetPageId="82" captionText="Figs 818 - 823. Teutamus rhino sp. n. 818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal." httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="83" pageNumber="483">fig. 821</figureCitation>
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; vulva
|
||
<figureCitation box="[171,319,729,753]" captionStart="Figs 818 - 823" captionStartId="82.[147,191,1784,1803]" captionTargetBox="[158,1184,250,1661]" captionTargetId="figure@82.[158,1184,250,1661]" captionTargetPageId="82" captionText="Figs 818 - 823. Teutamus rhino sp. n. 818. Male palp, retrolateral. 819. Ventral. 820. Female, abdomen, lateral view. 821. Epigyne. 822. Vulva, in clove oil, ventral. 823. Dorsal." httpUri="https://zenodo.org/record/814774/files/figure.png" pageId="83" pageNumber="483">figs 822-823</figureCitation>
|
||
, spermathecae round, openings large and round, appearing plugged. The position and tilt of the ducts are variable so that each epigyne looks slightly different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[43,882,833,857]" pageId="83" pageNumber="483" type="distribution">
|
||
<paragraph blockId="83.[43,882,831,858]" box="[43,882,833,857]" pageId="83" pageNumber="483">
|
||
<emphasis box="[43,185,833,857]" italics="true" pageId="83" pageNumber="483">Distribution</emphasis>
|
||
.— Only known from the Ujung Kulon Reserve in W Java.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="83" pageNumber="483" type="etymology">
|
||
<paragraph blockId="83.[43,1102,900,962]" pageId="83" pageNumber="483">
|
||
<emphasis box="[43,166,903,927]" italics="true" pageId="83" pageNumber="483">Etymology</emphasis>
|
||
.— From the presence of the last surviving population in Indonesia of the impressive
|
||
<taxonomicName box="[178,434,938,962]" class="Mammalia" family="Rhinocerotidae" genus="Rhinoceros" kingdom="Animalia" order="Perissodactyla" pageId="83" pageNumber="483" phylum="Chordata" rank="species" species="sondaicus">
|
||
<emphasis box="[178,434,938,962]" italics="true" pageId="83" pageNumber="483">Rhinoceros sondaicus</emphasis>
|
||
</taxonomicName>
|
||
for which the Ujung Kulon Reserve is known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |