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<document id="CF6E8DEFC631365A3C939E32D1F30520" ID-ISSN="1175­5334" ID-ZooBank="505937B0-9F57-4068-82E6-8553826DD5AA" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629822157238" checkinUser="felipe" docAuthor="Judson, Sarah W. &amp; Nelson, C. Riley" docDate="2012" docId="03B387E7154D811DFF5AFEC8FC655470" docLanguage="en" docName="zt03541p118.pdf" docOrigin="Zootaxa 3541" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Megarcys ochracea Klapalek 1912" docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="FF8AFF9F15668136FFCDFFA7FFAF5015" masterDocTitle="3541" masterLastPageNumber="118" masterPageNumber="1" pageNumber="44" updateTime="1699117090469" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="9D6F8FE098FE72A229AE628D36C031FA">Judson, Sarah W.</mods:namePart>
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<treatment id="03B387E7154D811DFF5AFEC8FC655470" LSID="urn:lsid:plazi:treatment:03B387E7154D811DFF5AFEC8FC655470" httpUri="http://treatment.plazi.org/id/03B387E7154D811DFF5AFEC8FC655470" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="C300657A154D811DFF5AFEC8FE9351DA" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8BA536F1154D811DFF5AFEC8FDEB519C" blockId="43.[151,580,367,393]" box="[151,580,367,393]" pageId="43" pageNumber="44">
<heading id="D0ED819D154D811DFF5AFEC8FDEB519C" bold="true" box="[151,580,367,393]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<taxonomicName id="4C1A4D72154D811DFF5AFEC8FDEB519C" ID-CoL="3YWZY" authority="Klapalek, 1912" authorityName="Klapalek" authorityYear="1912" box="[151,580,367,393]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="B96EEAE3154D811DFF5AFEC8FDEB519C" bold="true" box="[151,580,367,393]" pageId="43" pageNumber="44">
<emphasis id="B96EEAE3154D811DFF5AFEC8FE2D519C" bold="true" box="[151,386,367,393]" italics="true" pageId="43" pageNumber="44">Megarcys ochracea</emphasis>
Klapálek, 1912
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA536F1154D811DFF5AFE11FE9351DA" blockId="43.[151,645,438,589]" box="[151,316,438,463]" pageId="43" pageNumber="44">
<heading id="D0ED819D154D811DFF5AFE11FE9351DA" bold="true" box="[151,316,438,463]" fontSize="10" level="2" pageId="43" pageNumber="44" reason="2">
<emphasis id="B96EEAE3154D811DFF5AFE11FE9351DA" bold="true" box="[151,316,438,463]" pageId="43" pageNumber="44">SYNONYMY</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300657A154D811DFF5AFE7DFE795259" pageId="43" pageNumber="44" type="description">
<paragraph id="8BA536F1154D811DFF5AFE7DFDC051E5" blockId="43.[151,645,438,589]" box="[151,623,474,496]" pageId="43" pageNumber="44">
<taxonomicName id="4C1A4D72154D811DFF5AFE7DFDC051E5" authority="Klapalek, 1912" authorityName="Klapalek" authorityYear="1912" box="[151,623,474,496]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea" subGenus="Megarcys">
<emphasis id="B96EEAE3154D811DFF5AFE7DFE6051E5" box="[151,463,474,496]" italics="true" pageId="43" pageNumber="44">Perlodes (Megarcys) ochracea</emphasis>
Klapálek, 1912
</taxonomicName>
</paragraph>
<paragraph id="8BA536F1154D811DFF5AFE5DFE72521A" blockId="43.[151,645,438,589]" box="[151,477,505,527]" pageId="43" pageNumber="44">
<taxonomicName id="4C1A4D72154D811DFF5AFE5DFE72521A" authority="Kasai, 1938" authorityName="Kasai" authorityYear="1938" box="[151,477,505,527]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="B96EEAE3154D811DFF5AFE5DFEF1521A" box="[151,350,505,527]" italics="true" pageId="43" pageNumber="44">Megarcys ochracea</emphasis>
Kasai, 1938
</taxonomicName>
</paragraph>
<paragraph id="8BA536F1154D811DFF5AFDBFFD2A523B" blockId="43.[151,645,438,589]" box="[151,645,536,558]" pageId="43" pageNumber="44">
<taxonomicName id="4C1A4D72154D811DFF5AFDBFFD2A523B" authority="Ricker, 1952" authorityName="Ricker" authorityYear="1952" box="[151,645,536,558]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea" subGenus="Megarcys">
<emphasis id="B96EEAE3154D811DFF5AFDBFFE52523B" box="[151,509,536,558]" italics="true" pageId="43" pageNumber="44">Arcynopteryx (Megarcys) ochracea</emphasis>
Ricker, 1952
</taxonomicName>
</paragraph>
<paragraph id="8BA536F1154D811DFF5AFD90FE795259" blockId="43.[151,645,438,589]" box="[151,470,566,589]" pageId="43" pageNumber="44">
<taxonomicName id="4C1A4D72154D811DFF5AFD90FE795259" authority="Illies, 1966" authorityName="Illies" authorityYear="1966" box="[151,470,566,589]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="B96EEAE3154D811DFF5AFD90FEF15258" box="[151,350,567,589]" italics="true" pageId="43" pageNumber="44">Megarcys ochracea</emphasis>
Illies, 1966
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C300657A154D811DFF5AFDDEFD7F5284" box="[151,720,633,658]" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BA536F1154D811DFF5AFDDEFD7F5284" blockId="43.[151,1437,633,1125]" box="[151,720,633,658]" pageId="43" pageNumber="44">
<materialsCitation id="3B723CAC154D811DFF5AFDDEFD635284" box="[151,716,633,658]" location="Hokkadate" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Sakhalin" typeStatus="holotype">
<emphasis id="B96EEAE3154D811DFF5AFDDEFE2B5287" bold="true" box="[151,388,633,658]" pageId="43" pageNumber="44">
<typeStatus id="54A18853154D811DFF5AFDDEFF4D5284" box="[151,226,633,657]" pageId="43" pageNumber="44">TYPE</typeStatus>
LOCALITY:
</emphasis>
USSR:
<collectingRegion id="49DEF813154D811DFE2DFDDEFDEC5284" box="[480,579,633,657]" country="Russia" name="Sakhalin" pageId="43" pageNumber="44">Sachalin</collectingRegion>
,
<location id="8EC5602A154D811DFD82FDDEFD635284" LSID="urn:lsid:plazi:treatment:03B387E7154D811DFF5AFEC8FC655470:8EC5602A154D811DFD82FDDEFD635284" box="[591,716,633,657]" name="Hokkadate" pageId="43" pageNumber="44" stateProvince="Sakhalin">Hokkadate</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C300657A154D811DFF0AFD3AFEBE53C0" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BA536F1154D811DFF0AFD3AFEBE53C0" blockId="43.[151,1437,633,1125]" pageId="43" pageNumber="44">
<emphasis id="B96EEAE3154D811DFF0AFD3AFEDF52A3" bold="true" box="[199,368,669,694]" pageId="43" pageNumber="44">DIAGNOSIS:</emphasis>
Adults retain the remnants of nymphal thoracic (
<figureCitation id="13212A74154D811DFC77FD3AFBB152A0" box="[954,1054,669,693]" captionStart="FIGURES 229234" captionStartId="80.[151,263,1210,1233]" captionTargetBox="[167,1421,202,1176]" captionTargetId="figure-61@80.[167,1421,202,1178]" captionTargetPageId="80" captionText="FIGURES 229234. Perlodidae genus key. 229: Mesosternum of Arcynopteryx polaris, ventral; 230: Right mandible of Arcynopteryx polaris; 231: Mesosternum of Diura bicaudata, ventral; 232: Right mandible of Diura bicaudata; 233: Thorax of Megarcys ochracea, ventral; 234: Thorax of Skwala pusilla, ventral." pageId="43" pageNumber="44">Fig. 233</figureCitation>
) and submental gills (
<figureCitation id="13212A74154D811DFAE5FD3AFA2052A0" box="[1320,1423,669,693]" captionStart="FIGURES 235244" captionStartId="81.[151,263,1838,1861]" captionTargetBox="[167,1420,193,1817]" captionTargetId="figure-108@81.[167,1421,193,1817]" captionTargetPageId="81" captionText="FIGURES 235244. Perlodidae genus key. 235: Basal cercal segments of Diura bicaudata; 236: Distal cercal segments of Diura sp.; 237: Basal cercal segments of Isoperla potanini; 238: Distal cercal segments of Isoperla potanini; 239: Lacinia of Filchneria mongolica. Illustration after Teslenko and Zhiltzova (2009); 240: Lacinia of Skwala brevis; 241: Maxillary palp, galea, and lacinia of right lacinia of Diura bicaudata, dorsal; 242: Head of Diura bicaudata, ventral; 243: Maxillary palp, galea, and lacinia of right lacinia of Megarcys ochracea, dorsal; 244: Head showing submental gills of Megarcys ochracea, ventral." pageId="43" pageNumber="44">Fig. 244</figureCitation>
). The male hemitergal processes are strongly curved (
<figureCitation id="13212A74154D811DFD22FD66FCFD52CC" box="[751,850,705,729]" captionStart="FIGURES 278282" captionStartId="86.[151,263,1893,1916]" captionTargetBox="[153,1435,1103,1872]" captionTargetId="figure-74@86.[151,1436,1103,1872]" captionTargetPageId="86" captionText="FIGURES 278282. Megarcys ochracea. 278: Habitus (preserved); 279: Male, dorsal; 280: Female terminalia, ventral; 281: Nymph head; 282: Predicted ENM map with documented occurrences and range estimate." pageId="43" pageNumber="44">Fig. 279</figureCitation>
) and point anteriorly. The epiproct of the male has lateral stylets which extend slightly over the hemitergal processes. The female subgenital plate has a narrow median notch guarded by two medially directed, subtriangular lobes (
<figureCitation id="13212A74154D811DFC63FCAEFBBC5334" box="[942,1043,777,801]" captionStart="FIGURES 278282" captionStartId="86.[151,263,1893,1916]" captionTargetBox="[153,1435,1103,1872]" captionTargetId="figure-74@86.[151,1436,1103,1872]" captionTargetPageId="86" captionText="FIGURES 278282. Megarcys ochracea. 278: Habitus (preserved); 279: Male, dorsal; 280: Female terminalia, ventral; 281: Nymph head; 282: Predicted ENM map with documented occurrences and range estimate." pageId="43" pageNumber="44">Fig. 280</figureCitation>
). Nymphal head pattern (
<figureCitation id="13212A74154D811DFAFFFCAEFA3C5334" box="[1330,1427,777,801]" captionStart="FIGURES 278282" captionStartId="86.[151,263,1893,1916]" captionTargetBox="[153,1435,1103,1872]" captionTargetId="figure-74@86.[151,1436,1103,1872]" captionTargetPageId="86" captionText="FIGURES 278282. Megarcys ochracea. 278: Habitus (preserved); 279: Male, dorsal; 280: Female terminalia, ventral; 281: Nymph head; 282: Predicted ENM map with documented occurrences and range estimate." pageId="43" pageNumber="44">Fig. 281</figureCitation>
) includes a distinct M-line on the clypeus and white patches posterior to the eyes and in the interocellar region, all coloration characters of which are shared with
<taxonomicName id="4C1A4D72154D811DFD2AFCF6FCD7537C" authorityName="Brinck" authorityYear="1949" box="[743,888,849,873]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="B96EEAE3154D811DFD2AFCF6FCD7537C" box="[743,888,849,873]" italics="true" pageId="43" pageNumber="44">A. compacta</emphasis>
</taxonomicName>
nymphs. Despite these superficial similarities,
<taxonomicName id="4C1A4D72154D811DFF5AFCD1FEA95398" authorityName="Klapalek" authorityYear="1912" box="[151,262,886,909]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAE3154D811DFF5AFCD1FEA95398" box="[151,262,886,909]" italics="true" pageId="43" pageNumber="44">Megarcys</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A4D72154D811DFE89FCD2FE725398" box="[324,477,885,909]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAE3154D811DFE89FCD2FE725398" box="[324,477,885,909]" italics="true" pageId="43" pageNumber="44">Arcynopteryx</emphasis>
</taxonomicName>
nymphs can be easily separated with genus-level characteristics, particularly the presence of gills. The nymph of
<taxonomicName id="4C1A4D72154D811DFDC0FC3DFDD353A4" authorityName="Klapalek" authorityYear="1912" box="[525,636,922,945]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAE3154D811DFDC0FC3DFDD353A4" box="[525,636,922,945]" italics="true" pageId="43" pageNumber="44">Megarcys</emphasis>
</taxonomicName>
has finger-like thoracic gills (
<figureCitation id="13212A74154D811DFC15FC3EFB9453A4" box="[984,1083,921,945]" captionStart="FIGURES 229234" captionStartId="80.[151,263,1210,1233]" captionTargetBox="[167,1421,202,1176]" captionTargetId="figure-61@80.[167,1421,202,1178]" captionTargetPageId="80" captionText="FIGURES 229234. Perlodidae genus key. 229: Mesosternum of Arcynopteryx polaris, ventral; 230: Right mandible of Arcynopteryx polaris; 231: Mesosternum of Diura bicaudata, ventral; 232: Right mandible of Diura bicaudata; 233: Thorax of Megarcys ochracea, ventral; 234: Thorax of Skwala pusilla, ventral." pageId="43" pageNumber="44">Fig. 233</figureCitation>
) in addition to submental gills (
<figureCitation id="13212A74154D811DFF52FC1AFEAB53C0" box="[159,260,957,981]" captionStart="FIGURES 235244" captionStartId="81.[151,263,1838,1861]" captionTargetBox="[167,1420,193,1817]" captionTargetId="figure-108@81.[167,1421,193,1817]" captionTargetPageId="81" captionText="FIGURES 235244. Perlodidae genus key. 235: Basal cercal segments of Diura bicaudata; 236: Distal cercal segments of Diura sp.; 237: Basal cercal segments of Isoperla potanini; 238: Distal cercal segments of Isoperla potanini; 239: Lacinia of Filchneria mongolica. Illustration after Teslenko and Zhiltzova (2009); 240: Lacinia of Skwala brevis; 241: Maxillary palp, galea, and lacinia of right lacinia of Diura bicaudata, dorsal; 242: Head of Diura bicaudata, ventral; 243: Maxillary palp, galea, and lacinia of right lacinia of Megarcys ochracea, dorsal; 244: Head showing submental gills of Megarcys ochracea, ventral." pageId="43" pageNumber="44">Fig. 244</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C300657A154D811DFF0AFC46FA9053EC" box="[199,1343,993,1018]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BA536F1154D811DFF0AFC46FA9053EC" blockId="43.[151,1437,633,1125]" box="[199,1343,993,1018]" pageId="43" pageNumber="44">
<emphasis id="B96EEAE3154D811DFF0AFC46FDA353EF" bold="true" box="[199,524,993,1018]" pageId="43" pageNumber="44">DISTRIBUTION—Global:</emphasis>
East Palearctic—
<emphasis id="B96EEAE3154D811DFD19FC46FCE753EF" bold="true" box="[724,840,993,1018]" pageId="43" pageNumber="44">Regional:</emphasis>
AOB, IDB^—
<emphasis id="B96EEAE3154D811DFC3CFC46FBE653EF" bold="true" box="[1009,1097,993,1018]" pageId="43" pageNumber="44">Aimag:</emphasis>
BU^, KhG^, SE, TO.
</paragraph>
</subSubSection>
<subSubSection id="C300657A154D811DFF0AFBA2FC655470" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BA536F1154D811DFF0AFBA2FC655470" blockId="43.[151,1437,633,1125]" pageId="43" pageNumber="44">
<emphasis id="B96EEAE3154D811DFF0AFBA2FED4540B" bold="true" box="[199,379,1029,1054]" pageId="43" pageNumber="44">DISCUSSION:</emphasis>
<materialsCitation id="3B723CAC154D811DFE4FFBA1FC695470" country="Mongolia" location="Selenge" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Selenge">
Most of the
<taxonomicName id="4C1A4D72154D811DFDC6FBA1FD375408" authorityName="Illies" authorityYear="1966" box="[523,664,1029,1053]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="B96EEAE3154D811DFDC6FBA1FD375408" box="[523,664,1029,1053]" italics="true" pageId="43" pageNumber="44">M. ochracea</emphasis>
</taxonomicName>
documented by our survey were from mid-range elevations (approx. 8001500 msl) and has only been documented in the
<collectingRegion id="49DEF813154D811DFCC8FB8EFCF25454" box="[773,861,1065,1089]" country="Mongolia" name="Selenge" pageId="43" pageNumber="44">Selenge</collectingRegion>
River basin, especially near Lake
<collectingRegion id="49DEF813154D811DFB3DFB8EFAF25454" box="[1264,1373,1065,1089]" country="Mongolia" name="Hovsgol" pageId="43" pageNumber="44">Khovsgol</collectingRegion>
(
<figureCitation id="13212A74154D811DFAA2FB8EFF6A5470" captionStart="FIGURES 278282" captionStartId="86.[151,263,1893,1916]" captionTargetBox="[153,1435,1103,1872]" captionTargetId="figure-74@86.[151,1436,1103,1872]" captionTargetPageId="86" captionText="FIGURES 278282. Megarcys ochracea. 278: Habitus (preserved); 279: Male, dorsal; 280: Female terminalia, ventral; 281: Nymph head; 282: Predicted ENM map with documented occurrences and range estimate." pageId="43" pageNumber="44">Fig. 282</figureCitation>
). The species was first recorded from
<collectingCountry id="F30D7661154D811DFDBDFBEAFD725470" box="[624,733,1101,1125]" name="Mongolia" pageId="43" pageNumber="44">Mongolia</collectingCountry>
by Zhiltzova (1972)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>