163 lines
21 KiB
XML
163 lines
21 KiB
XML
<document id="B803C7079E9012149B951F34A1E8950D" ID-DOI="10.5281/zenodo.191334" ID-GBIF-Dataset="e9864b86-35a3-42ac-a1b5-a8b02beaa811" ID-ISSN="1175-5326" ID-Zenodo-Dep="191334" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461126891429" checkinUser="plazi" docAuthor="Ahyong, Shane T. & Ng, Peter K. L." docDate="2009" docId="130A8784FFFDFFAFBCD3F958CD7D35B5" docLanguage="en" docName="zt02289p047.pdf" docOrigin="Zootaxa 2289" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aphanodactylidae" docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="EF33FFFCFFFCFFADBC44FF83CE113356" masterDocTitle="Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms" masterLastPageNumber="47" masterPageNumber="33" pageNumber="34" updateTime="1698596964965" updateUser="plazi">
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<mods:title id="4DEE3A8F4B1DCB291944EA3F0B70871F">Aphanodactylidae, a new family of thoracotreme crabs (Crustacea: Brachyura) symbiotic with polychaete worms</mods:title>
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<mods:namePart id="E3F219C10B839886F56B75377A28F088">Ahyong, Shane T.</mods:namePart>
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<mods:name id="46E739C6E806A3BDB7F8D8CDF2619793" type="personal">
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<mods:roleTerm id="050421EEBEC6841E4BD8ADB19046348F">Author</mods:roleTerm>
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<mods:namePart id="77231848216BAB9EAAD87AE4CB5827CB">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="E972F2A707FA95CB632735B02D935BB2">text</mods:typeOfResource>
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<mods:title id="BDC6CCE9A156F13676E2936533CA882F">Zootaxa</mods:title>
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<mods:part id="1F965A7413E8FD953262ADE3082599C6">
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<mods:date id="BA8BC960B46F007F292C2CB8ACDCCEF5">2009</mods:date>
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<mods:detail id="25DDA9CBFF1728AF97FFFE1B7275A783" type="volume">
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<mods:number id="908350322F502B735DE603F935A213F1">2289</mods:number>
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<mods:identifier id="DE5B47E79C32490419E16E5C9AFF822C" type="DOI">10.5281/zenodo.191334</mods:identifier>
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<mods:identifier id="76B5CC4D7261F0438E513ECCF5154639" type="ISSN">1175-5326</mods:identifier>
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<treatment id="130A8784FFFDFFAFBCD3F958CD7D35B5" ID-DOI="http://doi.org/10.5281/zenodo.6222425" ID-GBIF-Taxon="119626521" ID-Zenodo-Dep="6222425" LSID="urn:lsid:plazi:treatment:130A8784FFFDFFAFBCD3F958CD7D35B5" httpUri="http://treatment.plazi.org/id/130A8784FFFDFFAFBCD3F958CD7D35B5" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
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<subSubSection id="D3B96519FFFDFFACBCD3F958CFC835A3" box="[151,473,1755,1781]" pageId="1" pageNumber="34" type="nomenclature">
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<paragraph id="9B1C3692FFFDFFACBCD3F958CFC835A3" blockId="1.[151,473,1755,1781]" box="[151,473,1755,1781]" pageId="1" pageNumber="34">
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<heading id="C05481FEFFFDFFACBCD3F958CFC835A3" bold="true" box="[151,473,1755,1781]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
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<emphasis id="A9D7EA80FFFDFFACBCD3F958CFC835A3" bold="true" box="[151,473,1755,1781]" pageId="1" pageNumber="34">
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<taxonomicName id="5CA34D11FFFDFFACBCD3F958CF6A35A3" ID-CoL="7NHCB" box="[151,379,1755,1781]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="34" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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n. fam.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3B96519FFFDFFAFBCD3F8A6CC723162" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="diagnosis">
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<paragraph id="9B1C3692FFFDFFAFBCD3F8A6CDAE32B1" blockId="1.[151,1437,1829,2010]" lastBlockId="2.[151,1438,152,1763]" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
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<emphasis id="A9D7EA80FFFDFFACBCD3F8A6CF063469" bold="true" box="[151,279,1829,1855]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
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Carapace strongly sexually dimorphic (where both are known); female carapace transversely ovate to rectangular; male carapace narrower than that of female, ovate to subquadrate; surface glabrous or finely setose; smooth or punctate; regions poorly defined. Front deflected ventrally, faintly sinuous to bilobed in dorsal view. Orbital margins entire, well defined, infraorbital margin terminating mesially as angular tooth or rounded corner. Antennules folding transversely or obliquely. Antennae not excluded from orbit. Cornea pigmented. Maxilliped 3 ischium longer than merus, subquadrate or triangular; merus subquadrate; palp articulating at distolateral margin of merus, segments articulating end-to-end, decreasing in size distally; exopod with flagellum. Chelipeds equal; segments unarmed; surfaces smooth, finely to sparsely setose. Ambulatory legs relatively short, stout, P3 longest, P5 shortest, dorsal to other pereopods; P3–4 merus with row of spines or teeth on flexor margin; propodus distoflexor angle with 1 or more spines opposing dactylus. Dactyli short, claw-like, apices corneous. Female gonopore (vulva) on sternite 6, between suture 5/6 and 6/7. Male gonopore (genital papilla) emerging near anterior margin of sternite 8, distinctly mesial to coxa 8. Abdomen of female with all somites and telson distinct, mobile; males with somites and telson mobile or with somites 4–6 fused. Male abdomen narrowly triangular or linguiform.
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</paragraph>
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<paragraph id="9B1C3692FFFEFFAFBC82FE70CC723162" blockId="2.[151,1438,152,1763]" pageId="2" pageNumber="35">
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<emphasis id="A9D7EA80FFFEFFAFBC82FE70CF89315B" bold="true" box="[198,408,499,525]" pageId="2" pageNumber="35">Included genera.</emphasis>
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<taxonomicName id="5CA34D11FFFEFFAFBDDBFE77CCE03158" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[415,753,500,526]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBDDBFE77CC4D315B" box="[415,604,500,525]" italics="true" pageId="2" pageNumber="35">Aphanodactylus</emphasis>
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<bibRefCitation id="FF324B63FFFEFFAFBE20FE77CCE03158" author="Tesch" box="[612,753,500,526]" pageId="2" pageNumber="47" refString="Tesch, J. (1918) The Decapoda Brachyura of the Siboga-Expedition. II. Goneplacidae and Pinnotheridae. Siboga- Expeditie, 39 c 1, 149 - 295, 12 pls." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
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</taxonomicName>
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[
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<typeStatus id="44188830FFFEFFAFBEBAFE77CD223158" box="[766,819,500,526]" pageId="2" pageNumber="35">type</typeStatus>
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genus],
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<taxonomicName id="5CA34D11FFFEFFAFBFDCFE77CAD43158" authority="Kammerer, 2006" authorityName="Kammerer" authorityYear="2006" box="[920,1221,500,526]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBFDCFE77CDE7315B" box="[920,1014,500,525]" italics="true" pageId="2" pageNumber="35">Gandoa</emphasis>
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<bibRefCitation id="FF324B63FFFEFFAFBFB9FE77CAD43158" author="Kammerer" box="[1021,1221,500,526]" pageId="2" pageNumber="46" refString="Kammerer, C. F. (2006) Notes on some preoccupied names in Arthropoda. Acta Zootaxonomica Sinica, 31 (2), 269 - 271." type="journal article" year="2006">Kammerer, 2006</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="5CA34D11FFFEFFAFB895FE77CB2F315B" box="[1233,1342,500,525]" class="Malacostraca" family="Aphanodactylidae" genus="Gustavus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="A9D7EA80FFFEFFAFB895FE77CB2F315B" box="[1233,1342,500,525]" italics="true" pageId="2" pageNumber="35">Gustavus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B2E457FBFFFEFFAFB902FE77CB863158" box="[1350,1431,500,526]" pageId="2" pageNumber="35" rank="genus">n. gen.</taxonomicNameLabel>
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, and
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<taxonomicName id="5CA34D11FFFEFFAFBC8CFD99CC4E3162" authority="Naruse, Fujita & Ng, 2009" authorityName="Naruse, Fujita & Ng" authorityYear="2009" box="[200,607,538,564]" class="Malacostraca" family="Pinnotheridae" genus="Uruma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBC8CFD99CF0A3165" box="[200,283,538,563]" italics="true" pageId="2" pageNumber="35">Uruma</emphasis>
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Naruse,
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<bibRefCitation id="FF324B63FFFEFFAFBDC0FD99CC4E3162" author="Fujita" box="[388,607,538,564]" pageId="2" pageNumber="47" refString="Naruse, T., Fujita, Y. & Ng, P. K. L. (2009) A new genus and new species of commensal crab (Crustacea: Brachyura: Pinnotheroidea) from Oura Bay, Okinawa Island, Japan. Zootaxa 2053, 59 - 68." type="journal article" year="2009">Fujita & Ng, 2009</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3B96519FFFEFFAFBC82FDC3CD7D35B5" pageId="2" pageNumber="35" type="discussion">
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<paragraph id="9B1C3692FFFEFFAFBC82FDC3CB83304A" blockId="2.[151,1438,152,1763]" pageId="2" pageNumber="35">
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<emphasis id="A9D7EA80FFFEFFAFBC82FDC3CF51310C" bold="true" box="[198,320,576,602]" pageId="2" pageNumber="35">Remarks.</emphasis>
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Members of the
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<taxonomicName id="5CA34D11FFFEFFAFBE4AFDC2CCF2310D" box="[526,739,577,603]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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n. fam. all have a rather uniform appearance: transversely rectangular or ovate carapace in females, narrower carapace in males; unornamented carapace surfaces with very little evidence of regionalisation; short, stout pereopods 2–4 with short, claw-like dactyli opposing spines on the distal propodal margins. These features of
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<taxonomicName id="5CA34D11FFFEFFAFBEAFFD36CDD53199" box="[747,964,693,719]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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are reminiscent of many
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<taxonomicName id="5CA34D11FFFEFFAFB8B3FD36CB8C3199" box="[1271,1437,693,719]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
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<emphasis id="A9D7EA80FFFEFFAFBCD3FD5FCF1F31A3" box="[151,270,732,757]" italics="true" pageId="2" pageNumber="35">sensu lato</emphasis>
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and are essentially the basis of earlier classifications placing aphanodactylids there. Some features remain to be confirmed for some genera; male
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<taxonomicName id="5CA34D11FFFEFFAFBEF9FC81CD0A304D" box="[701,795,770,795]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBEF9FC81CD0A304D" box="[701,795,770,795]" italics="true" pageId="2" pageNumber="35">Gandoa</emphasis>
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</taxonomicName>
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and female
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<taxonomicName id="5CA34D11FFFEFFAFBFEFFC81CDEF304D" box="[939,1022,770,795]" class="Malacostraca" family="Pinnotheridae" genus="Uruma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBFEFFC81CDEF304D" box="[939,1022,770,795]" italics="true" pageId="2" pageNumber="35">Uruma</emphasis>
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</taxonomicName>
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are, unfortunately, not yet known.
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</paragraph>
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<paragraph id="9B1C3692FFFEFFAFBC81FCAACA073574" blockId="2.[151,1438,152,1763]" pageId="2" pageNumber="35">
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Aphanodactylids, however, differ significantly from all known
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<taxonomicName id="5CA34D11FFFEFFAFBFA9FCAACA8C3015" box="[1005,1181,809,835]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
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in lacking the chief synapomorphy of pinnotherids, namely, the highly modified maxilliped 3. In pinnotherids the merus of maxilliped 3 is either considerably larger than, or fused with the ischium, forming a single unit (e.g.,
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<bibRefCitation id="FF324B63FFFEFFAFB979FCF5CF3230E1" author="Ahyong" pageId="2" pageNumber="46" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
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: fig. 1C); and the dactylus usually articulates proximally to the distal end of the propodus, and may be considerably enlarged or significantly reduced. Conversely, the maxilliped 3 of aphanodactylids is of the more typical, plesiomorphic thoracotreme form: the merus is smaller than the ischium and the palp is not enlarged. They resemble those found in a number of thoracotremes such as ocypodoids, and are what might be expected in stem-lineage Pinnotheroidea. Comparisons of the position of the male gonopores of aphanodactylids with specimens of other thoracotreme families revealed another noteworthy pattern. The position of the male gonopore in aphanodactylids, being distant from the base of coxa 8, closely resembles the condition of other pinnotherids (e.g.,
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<emphasis id="A9D7EA80FFFEFFAFBE09FB2FCD943793" box="[589,901,1196,1221]" italics="true" pageId="2" pageNumber="35">
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<taxonomicName id="5CA34D11FFFEFFAFBE09FB2FCCEB3793" box="[589,762,1196,1221]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Nepinnotheres</taxonomicName>
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,
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<taxonomicName id="5CA34D11FFFEFFAFBF42FB2FCD943793" box="[774,901,1196,1221]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Arcotheres</taxonomicName>
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</emphasis>
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and
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<taxonomicName id="5CA34D11FFFEFFAFBFF9FB2FCA5D3793" box="[957,1100,1196,1221]" class="Malacostraca" family="Pinnotheridae" genus="Viridotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="A9D7EA80FFFEFFAFBFF9FB2FCA5D3793" box="[957,1100,1196,1221]" italics="true" pageId="2" pageNumber="35">Viridotheres</emphasis>
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</taxonomicName>
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) as well as ocypodoids such as members of the
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<taxonomicName id="5CA34D11FFFEFFAFBDC9FB51CC6F37BA" box="[397,638,1234,1260]" class="Malacostraca" family="Macrophthalmidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Macrophthalmidae</taxonomicName>
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and
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<taxonomicName id="5CA34D11FFFEFFAFBEFAFB51CD4937BA" box="[702,856,1234,1260]" class="Malacostraca" family="Ocypodidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Ocypodidae</taxonomicName>
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(see also
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<bibRefCitation id="FF324B63FFFEFFAFBF98FB51CA9237BA" author="Guinot" box="[988,1155,1234,1260]" pageId="2" pageNumber="46" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum national d'Histoire naturelle, Paris, (A) Zoologie, 112, 1 - 354, pls. 1 - 27." type="journal article" year="1979">Guinot 1979</bibRefCitation>
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). As such, the overall pinnotherid-like habitus of aphanodactylids suggests that they are close to pinnotherids, though the plesiomorphic form of maxilliped 3 excludes them from placement within
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<taxonomicName id="5CA34D11FFFEFFAFBFB8FAA3CAB2366C" box="[1020,1187,1312,1338]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
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.
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<taxonomicName id="5CA34D11FFFEFFAFB8E9FAA3CB93366C" box="[1197,1410,1312,1338]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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is therefore tentatively assigned to Pinnotheroidea, alongside
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<taxonomicName id="5CA34D11FFFEFFAFBF8CFAC5CA933636" box="[968,1154,1350,1376]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
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. If our alignment of
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<taxonomicName id="5CA34D11FFFEFFAFBCD3FAEECF6336D1" box="[151,370,1389,1415]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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with
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<taxonomicName id="5CA34D11FFFEFFAFBDFDFAEECC7036D1" box="[441,609,1389,1415]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
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is correct, then the ocypodoid-like maxilliped 3 and male gonopore position might point towards some
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<typeStatus id="44188830FFFEFFAFBE79FA17CC6136F8" box="[573,624,1428,1454]" pageId="2" pageNumber="35">type</typeStatus>
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of ocypodoid ancestry for Pinnotheroidea. Molecular investigations of the phylogenetic position of
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<taxonomicName id="5CA34D11FFFEFFAFBE64FA39CD113682" box="[544,768,1466,1492]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
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and
|
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<taxonomicName id="5CA34D11FFFEFFAFBF7BFA39CDFA3682" box="[831,1003,1466,1492]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
|
||
are currently underway by several investigators, although a close relationship between pinnotherids and ocypodoids has already been suggested by
|
||
<bibRefCitation id="FF324B63FFFEFFAFBCF8F98BCFB43574" box="[188,421,1544,1570]" pageId="2" pageNumber="47" refString="Wetzer, R., Martin, J. W. & Boyce, S. (2009) Evolutionary origin of the gall crabs (Family Cryptochiridae) based on 16 S rDNA sequence data. In: Martin, J. W., Crandall, K. A. & Felder, D. L. (eds.), Decapod Crustacean Phylogenetics. Crustacean Issues, 18, 475 - 490." type="journal article">
|
||
Wetzer
|
||
<emphasis id="A9D7EA80FFFEFFAFBD51F98BCF5E3577" box="[277,335,1544,1569]" italics="true" pageId="2" pageNumber="35">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
and to a lesser extent by
|
||
<bibRefCitation id="FF324B63FFFEFFAFBE8AF98BCA033574" box="[718,1042,1544,1570]" pageId="2" pageNumber="47" refString="Palacios-Theil, E., Cuesta, J. A., Campos, E. & Felder, D. L. (2009) Molecular genetic re-examination of subfamilies and polyphyly in the family Pinnotheridae (Crustacea: Decapoda). In: Martin, J. W., Crandall, K. A. & Felder, D. L. (eds.), Decapod Crustacean Phylogenetics. Crustacean Issues 18, 457 - 474." type="journal article">
|
||
Palacios-Theil
|
||
<emphasis id="A9D7EA80FFFEFFAFBF3AF98BCDA83577" box="[894,953,1544,1569]" italics="true" pageId="2" pageNumber="35">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9B1C3692FFFEFFAFBC81F9ADCD7D35B5" blockId="2.[151,1438,152,1763]" pageId="2" pageNumber="35">
|
||
All known members of the
|
||
<taxonomicName id="5CA34D11FFFEFFAFBE53F9ADCCE9351E" box="[535,760,1582,1608]" class="Malacostraca" family="Aphanodactylidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Aphanodactylidae</taxonomicName>
|
||
, for which hosts are recorded, are associated with tube building polychaete worms (
|
||
<taxonomicName id="5CA34D11FFFEFFAFBDADF9D6CC933539" box="[489,642,1621,1647]" class="Polychaeta" family="Terebellidae" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="35" phylum="Annelida" rank="family">Terebellidae</taxonomicName>
|
||
). The unusual distal propodal spines of the ambulatory legs, which oppose the dactyli forming a subcheliform structure, and teeth present on the flexor margins of some of the legs, possibly assists the crabs in holding onto the tube walls or the surface of their polychaete hosts. The four aphanodactylid genera can be distinguished by the key below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |