treatments-xml/data/03/F7/87/03F787A5BA04C811FF2620ADFC27FF43.xml

<document id="13375259A39A13611163ED9E18A27B9F" ID-CLB-Dataset="27583" ID-DOI="10.11646/zootaxa.4552.1.1" ID-GBIF-Dataset="537cc788-42c7-46ad-86e1-74e539b0ce74" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623150" ID-ZooBank="13CE299C-2085-4BBD-989D-75B417CE1CAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270626423" checkinUser="plazi" docAuthor="De, Lesley S., Taphorn, Donald C. &amp; Armbruster, Jonathan W." docDate="2019" docId="03F787A5BA04C811FF2620ADFC27FF43" docLanguage="en" docName="zootaxa.4552.1.1.pdf" docOrigin="Zootaxa 4552 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ancistrus patronus De &amp; Taphorn &amp; Armbruster 2019, new species" docType="treatment" docUuid="F4AC6A5C-BE37-4DDE-B3E1-39E34CEF5903" docUuidSource="ZooBank" docVersion="9" lastPageNumber="43" masterDocId="FFCEFFDDBA22C83BFFB1250AFFDEFFBB" masterDocTitle="Review of Ancistrus (Siluriformes: Loricariidae) from the northwestern Guiana Shield, Orinoco Andes, and adjacent basins with description of six new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="39" updateTime="1698716197891" updateUser="ExternalLinkService">
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<mods:title id="37586C6B1FB3856DBBE698966022BB39">Review of Ancistrus (Siluriformes: Loricariidae) from the northwestern Guiana Shield, Orinoco Andes, and adjacent basins with description of six new species</mods:title>
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<mods:namePart id="D14E92E7C45043902BAF52DD9BBB603B">De, Lesley S.</mods:namePart>
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(
<figureCitation id="13652A36BA04C81DFF2E20C0FEFBFA58" box="[159,293,1482,1507]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStartId-0="40.[151,250,1838,1860]" captionStartId-1="41.[151,250,695,717]" captionTargetBox-0="[151,1436,193,1817]" captionTargetBox-1="[151,1436,193,674]" captionTargetId-0="figure@40.[151,1436,193,1817]" captionTargetId-1="figure@41.[151,1436,193,674]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURE 19. Ancistrus patronus, holotype, in dorsal, lateral, and ventral views, AUM 39876, 83.6 mm SL. Scale = 1 cm. Photos by J.W. Armbruster." captionText-1="FIGURE 20. Live coloration of head of Ancistrus patronus in dorsal and lateral views, AUM 43868, 79.4 mm SL. Scale = 1 cm. Photos by N.K. Lujan." httpUri-0="https://zenodo.org/record/2623192/files/figure.png" httpUri-1="https://zenodo.org/record/2623194/files/figure.png" pageId="38" pageNumber="39">Figs. 19–20</figureCitation>
)
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. 
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(ex 
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39876) (1, 
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SL), male, 
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, 
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Orinoco 
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Basin
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, 
<location id="8E816068BA04C81DFF26233CFEE2F9F4" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFF26233CFEE2F9F4" box="[151,316,1590,1615]" country="Venezuela" county="Orinoco River Basin" municipality="Orinoco River" name="Ventuari River" pageId="38" pageNumber="39" stateProvince="Amazonas">Ventuari River</location>
at mouth of 
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, 
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NNE of Macaruco
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, 
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<quantity id="4CA69B56BA04C81DFC70233DFBC6F9F4" box="[961,1048,1590,1615]" metricMagnitude="5" metricUnit="m" metricValue="1.89" pageId="38" pageNumber="39" unit="km" value="189.0">189 km</quantity>
NE of San Fernando de Atabapo
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, 5.05588, -66.32742, 
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, 
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, 
<collectorName id="26AB5365BA04C81DFD462351FCA1F9C8" box="[759,895,1626,1651]" pageId="38" pageNumber="39">D. Werneke</collectorName>
, 
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, and 
<collectorName id="26AB5365BA04C81DFBFD2351FB71F9C8" box="[1100,1199,1627,1651]" pageId="38" pageNumber="39">O. León.</collectorName>
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.
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from 
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, 
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Basin: 
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<collectorName id="26AB5365BA04C81DFC192375FC3AF92C" box="[936,996,1663,1687]" pageId="38" pageNumber="39">State</collectorName>
: 
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<collectingDate id="EFA4E99BBA04C81DFB8C2374FB4FF92C" box="[1085,1169,1662,1687]" pageId="38" pageNumber="39" value="36-02-16">160236</collectingDate>
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(1, 
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SL), creek 
<locationDeviation id="9A532F91BA04C81DFF2623A8FE6FF900" box="[151,433,1698,1723]" location="Parguaza River" pageId="38" pageNumber="39">
<quantity id="4CA69B56BA04C81DFF2623A8FF3EF900" box="[151,224,1698,1723]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="km" value="15.0">15 km</quantity>
E Parguaza River
</locationDeviation>
ferry crossing on 
<location id="8E816068BA04C81DFD3723A8FC78F900" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFD3723A8FC78F900" box="[646,934,1698,1723]" country="Venezuela" name="Caicara-Puerto Ayacucho" pageId="38" pageNumber="39" stateProvince="Amazonas">Caicara-Puerto Ayacucho</location>
hwy, 6.35946, -67.05375, 
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<collectingDate id="EFA4E99BBA04C81DFB6A23A8FAABF900" box="[1243,1397,1698,1723]" pageId="38" pageNumber="39" value="1985-11-28">28-Nov-1985</collectingDate>
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, 
<collectorName id="26AB5365BA04C81DFA3323A9FF21F964" pageId="38" pageNumber="39">B. Chernoff</collectorName>
, 
<collectorName id="26AB5365BA04C81DFEBC23CDFEB5F964" box="[269,363,1734,1759]" pageId="38" pageNumber="39">W. Saul</collectorName>
, C. 
<collectorName id="26AB5365BA04C81DFE2F23CDFE23F965" box="[414,509,1735,1758]" pageId="38" pageNumber="39">Marrero</collectorName>
, R. 
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; 
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39271
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(6, 5, 50.8–86.0 mm SL), 
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, at raudales 
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, 
<locationDeviation id="9A532F91BA04C81DFEB923E1FD6EF8B8" box="[264,688,1770,1795]" location="San Juan de Manapiare" pageId="38" pageNumber="39">
<quantity id="4CA69B56BA04C81DFEB923E1FE89F8B8" box="[264,343,1770,1795]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="38" pageNumber="39" unit="km" value="2.7">2.7 km</quantity>
NE of San Juan de Manapiare
</locationDeviation>
, 
<location id="8E816068BA04C81DFD0A23E0FCC5F8B8" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFD0A23E0FCC5F8B8" box="[699,795,1770,1795]" country="Venezuela" county="Parucito River" municipality="Salomon" name="Ventuari" pageId="38" pageNumber="39" stateProvince="Amazonas">Ventuari</location>
/ 
<location id="8E816068BA04C81DFC9223E0FC47F8B8" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFC9223E0FC47F8B8" box="[803,921,1770,1795]" country="Venezuela" county="Parucito River" municipality="Salomon" name="Manapiare" pageId="38" pageNumber="39" stateProvince="Amazonas">Manapiare</location>
drainage, 5.34637, -66.03347, 
<collectingDate id="EFA4E99BBA04C81DFB4923E0FA46F8B8" box="[1272,1432,1770,1795]" pageId="38" pageNumber="39" value="2004-04-16">
<date id="FFE01073BA04C81DFB4923E0FA46F8B8" box="[1272,1432,1770,1795]" pageId="38" pageNumber="39" value="2004-04-16">16 April 2004</date>
</collectingDate>
, 
<collectorName id="26AB5365BA04C81DFF262205FEC0F89C" box="[151,286,1806,1831]" pageId="38" pageNumber="39">D. Werneke</collectorName>
, 
<collectorName id="26AB5365BA04C81DFE982205FE50F89C" box="[297,398,1806,1831]" pageId="38" pageNumber="39">N. Lujan</collectorName>
, 
<collectorName id="26AB5365BA04C81DFE282205FDEAF89C" box="[409,564,1807,1831]" pageId="38" pageNumber="39">O León-Mata</collectorName>
; 
<emphasis id="B92AEAA1BA04C81DFD8F2204FD59F89C" bold="true" box="[574,647,1806,1831]" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFD8F2204FD59F89C" box="[574,647,1806,1831]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38" pageNumber="39">ANSP</collectionCode>
</emphasis>
</materialsCitation>
205667 (4, 
<quantity id="4CA69B56BA04C81DFCBD2204FC71F89D" box="[780,943,1806,1831]" metricMagnitude="-2" metricUnit="m" metricValue="6.68" metricValueMax="7.57" metricValueMin="5.79" pageId="38" pageNumber="39" unit="mm" value="66.8" valueMax="75.7" valueMin="57.9">57.9–75.7 mm</quantity>
SL); 
<materialsCitation id="3B363CEEBA04C81DFC5C2204FD02F8D4" ID-GBIF-Occurrence="2234227011" collectingDate="2004-04-12" collectionCode="AUM" collectorName="N. Lujan &amp; Sabaj &amp; L. de Souza &amp; Werneke" country="Venezuela" location="Ventuari River" municipality="Manapiare River" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA04C81DFC5C2204FBA2F89C" bold="true" box="[1005,1148,1806,1831]" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFC5C2204FBEEF89C" box="[1005,1072,1806,1831]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="38" pageNumber="39" type="University or college">AUM</collectionCode>
39306
</emphasis>
(7, 5, 62.4–86.0 mm SL), 
<collectingMunicipality id="6B85ACC9BA04C81DFF262238FE87F8F0" box="[151,345,1842,1867]" pageId="38" pageNumber="39">Manapiare River</collectingMunicipality>
<locationDeviation id="9A532F91BA04C81DFEE82238FCF4F8F0" box="[345,810,1842,1867]" location="San Juan de Manapiare" pageId="38" pageNumber="39">
<quantity id="4CA69B56BA04C81DFEE82238FE68F8F0" box="[345,438,1842,1867]" metricMagnitude="4" metricUnit="m" metricValue="1.45" pageId="38" pageNumber="39" unit="km" value="14.5">14.5 km</quantity>
NW of San Juan de Manapiare
</locationDeviation>
, 
<location id="8E816068BA04C81DFC892238FC01F8F0" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFC892238FC01F8F0" box="[824,991,1842,1867]" country="Venezuela" municipality="Manapiare River" name="Ventuari River" pageId="38" pageNumber="39" stateProvince="Amazonas">Ventuari River</location>
drainage, 5.42863 -66.13616, 
<date id="FFE01073BA04C81DFAF22239FF0CF8D4" pageId="38" pageNumber="39" value="2004-04-12">
<collectingDate id="EFA4E99BBA04C81DFAF22239FF0CF8D4" pageId="38" pageNumber="39" value="2004-04-12">12-Apr- 2004</collectingDate>
</date>
, 
<collectorName id="26AB5365BA04C81DFF6D225DFE9CF8D4" box="[220,322,1878,1903]" pageId="38" pageNumber="39">N. Lujan</collectorName>
, M. 
<collectorName id="26AB5365BA04C81DFEC5225CFE6BF8D4" box="[372,437,1878,1903]" pageId="38" pageNumber="39">Sabaj</collectorName>
, 
<collectorName id="26AB5365BA04C81DFE71225DFD99F8D4" box="[448,583,1878,1903]" pageId="38" pageNumber="39">L. de Souza</collectorName>
, D. 
<collectorName id="26AB5365BA04C81DFDC4225CFD02F8D4" box="[629,732,1878,1903]" pageId="38" pageNumber="39">Werneke</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA04C81DFD57225CFA42F8D4" ID-GBIF-Occurrence="2234227002" box="[742,1436,1878,1903]" collectingDate="2004-04-08" collectionCode="AUM, ANSP" collectorName="M. Sabaj &amp; N. Lujan &amp; D. Werneke &amp; L. de Souza &amp; O. Leon." country="Venezuela" county="Orinoco River Basin" location="River Basin" municipality="Orinoco River" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA04C81DFD57225CFCABF8D4" bold="true" box="[742,885,1878,1903]" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFD57225CFCF7F8D4" box="[742,809,1878,1903]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="38" pageNumber="39" type="University or college">AUM</collectionCode>
39876
</emphasis>
(4, not measured), same data as holotype; 
<emphasis id="B92AEAA1BA04C81DFAE2225CFA42F8D4" bold="true" box="[1363,1436,1878,1903]" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFAE2225CFA42F8D4" box="[1363,1436,1878,1903]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38" pageNumber="39">ANSP</collectionCode>
</emphasis>
</materialsCitation>
205668 (3, 
<quantity id="4CA69B56BA04C81DFEAA2270FE1EF829" box="[283,448,1914,1939]" metricMagnitude="-2" metricUnit="m" metricValue="6.3549999999999995" metricValueMax="6.68" metricValueMin="6.029999999999999" pageId="38" pageNumber="39" unit="mm" value="63.55" valueMax="66.8" valueMin="60.3">60.3–66.8 mm</quantity>
SL); 
<materialsCitation id="3B363CEEBA04C81DFDB52270FA93F80C" ID-GBIF-Occurrence="2234227179" collectingDate="2004-04-01" collectionCode="AUM" collectorName="D. Werneke &amp; Lujan &amp; L. de Souza &amp; Leon Mata &amp; Sabaj" country="Venezuela" location="Orinoco River" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA04C81DFDB52270FD48F828" bold="true" box="[516,662,1914,1939]" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFDB52270FD99F828" box="[516,583,1914,1939]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="38" pageNumber="39" type="University or college">AUM</collectionCode>
40189
</emphasis>
(6, 1, 
<quantity id="4CA69B56BA04C81DFD552270FC94F829" box="[740,842,1914,1939]" metricMagnitude="-2" metricUnit="m" metricValue="6.01" pageId="38" pageNumber="39" unit="mm" value="60.1">60.1 mm</quantity>
SL), tributary to 
<location id="8E816068BA04C81DFBA72270FB65F828" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFBA72270FB65F828" box="[1046,1211,1914,1939]" country="Venezuela" name="Orinoco River" pageId="38" pageNumber="39" stateProvince="Amazonas">Orinoco River</location>
, 
<locationDeviation id="9A532F91BA04C81DFB792271FED7F80C" location="Puerto Ayacucho" pageId="38" pageNumber="39">
<quantity id="4CA69B56BA04C81DFB792271FACFF828" box="[1224,1297,1914,1939]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="39" unit="km" value="30.0">30 km</quantity>
S of Puerto Ayacucho
</locationDeviation>
, 5.38659, -67.61556, 
<date id="FFE01073BA04C81DFDB02295FD56F80C" box="[513,648,1951,1975]" pageId="38" pageNumber="39" value="2004-04-01">
<collectingDate id="EFA4E99BBA04C81DFDB02295FD56F80C" box="[513,648,1951,1975]" pageId="38" pageNumber="39" value="2004-04-01">1-Apr-2004</collectingDate>
</date>
, 
<collectorName id="26AB5365BA04C81DFD252295FCC3F80C" box="[660,797,1950,1975]" pageId="38" pageNumber="39">D. Werneke</collectorName>
, N. 
<collectorName id="26AB5365BA04C81DFCFC2294FC4FF80C" box="[845,913,1950,1975]" pageId="38" pageNumber="39">Lujan</collectorName>
, 
<collectorName id="26AB5365BA04C81DFC2F2295FBF6F80C" box="[926,1064,1950,1975]" pageId="38" pageNumber="39">L. de Souza</collectorName>
, O. 
<collectorName id="26AB5365BA04C81DFBE92295FB08F80D" box="[1112,1238,1951,1974]" pageId="38" pageNumber="39">Leon Mata</collectorName>
, M. 
<collectorName id="26AB5365BA04C81DFABA2294FA93F80C" box="[1291,1357,1950,1975]" pageId="38" pageNumber="39">Sabaj</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA04C81CFAE82294FE27FF0B" ID-GBIF-Occurrence="2234227099" collectingDate="2004-04-04" collectionCode="AUM" collectorName="M. Sabaj &amp; L. de Souza &amp; D. Werneke &amp; Lujan" country="Venezuela" lastPageId="39" lastPageNumber="40" location="Macuruco" municipality="Orinoco River" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA04C81DFAE82294FF03F860" bold="true" pageId="38" pageNumber="39">
<collectionCode id="ED4FAE76BA04C81DFAE82294FA42F80C" box="[1369,1436,1950,1975]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="38" pageNumber="39" type="University or college">AUM</collectionCode>
40578
</emphasis>
(1, 
<quantity id="4CA69B56BA04C81DFEBC22C9FEACF861" box="[269,370,1987,2011]" metricMagnitude="-2" metricUnit="m" metricValue="7.47" pageId="38" pageNumber="39" unit="mm" value="74.7">74.7 mm</quantity>
SL), 
<collectingMunicipality id="6B85ACC9BA04C81DFE0422C8FD89F860" box="[437,599,1986,2011]" pageId="38" pageNumber="39">Orinoco River</collectingMunicipality>
at 
<location id="8E816068BA04C81DFDCF22C9FD31F861" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA04C81DFDCF22C9FD31F861" box="[638,751,1987,2010]" country="Venezuela" municipality="Orinoco River" name="Macuruco" pageId="38" pageNumber="39" stateProvince="Amazonas">Macuruco</location>
landing, 3.95816, -067.03231, 
<date id="FFE01073BA04C81DFBE622C9FB03F860" box="[1111,1245,1987,2011]" pageId="38" pageNumber="39" value="2004-04-04">
<collectingDate id="EFA4E99BBA04C81DFBE622C9FB03F860" box="[1111,1245,1987,2011]" pageId="38" pageNumber="39" value="2004-04-04">4-Apr-2004</collectingDate>
</date>
, 
<collectorName id="26AB5365BA04C81DFB5B22C9FA8AF860" box="[1258,1364,1986,2011]" pageId="38" pageNumber="39">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA04C81CFAD022C9FF00FF0B" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">L. de Souza</collectorName>
, 
<collectorName id="26AB5365BA05C81CFF5F2592FEA2FF0B" box="[238,380,151,176]" pageId="39" pageNumber="40">D. Werneke</collectorName>
, N. 
<collectorName id="26AB5365BA05C81CFE05259DFE27FF0B" box="[436,505,151,176]" pageId="39" pageNumber="40">Lujan</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFDB8259DFB7AFF6F" ID-GBIF-Occurrence="2234226965" collectingDate="2004-04-13" collectionCode="AUM" collectorName="M. Sabaj &amp; N. Lujan &amp; Werneke &amp; L. de Souza" country="Venezuela" location="Manapiare River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFDB8259DFD40FF0B" bold="true" box="[521,670,151,176]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFDB8259DFD92FF0B" box="[521,588,151,176]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
41324
</emphasis>
(2, 1, 
<quantity id="4CA69B56BA05C81CFD462592FCBEFF14" box="[759,864,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="7.79" pageId="39" pageNumber="40" unit="mm" value="77.9">77.9 mm</quantity>
SL), 
<location id="8E816068BA05C81CFC18259DFBB5FF0B" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFC18259DFBB5FF0B" box="[937,1131,151,176]" country="Venezuela" name="Manapiare River" pageId="39" pageNumber="40" stateProvince="Amazonas">Manapiare River</location>
<locationDeviation id="9A532F91BA05C81CFBC92592FECCFF6E" location="San Juan de Manapiare" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFBC92592FB1AFF0B" box="[1144,1220,151,176]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="39" pageNumber="40" unit="km" value="14.0">14 km</quantity>
NW San Juan de Manapiare
</locationDeviation>
, 5.43704, -66.11408, 
<date id="FFE01073BA05C81CFDB425B6FD47FF6F" box="[517,665,188,212]" pageId="39" pageNumber="40" value="2004-04-13">
<collectingDate id="EFA4E99BBA05C81CFDB425B6FD47FF6F" box="[517,665,188,212]" pageId="39" pageNumber="40" value="2004-04-13">13-Apr-2004</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFD1525B7FCD2FF6E" box="[676,780,188,213]" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA05C81CFCA625B7FCA3FF6E" box="[791,893,188,213]" pageId="39" pageNumber="40">N. Lujan</collectorName>
, D. 
<collectorName id="26AB5365BA05C81CFC1A25B6FBCFFF6E" box="[939,1041,188,213]" pageId="39" pageNumber="40">Werneke</collectorName>
, 
<collectorName id="26AB5365BA05C81CFBAA25B7FB7AFF6F" box="[1051,1188,188,213]" pageId="39" pageNumber="40">L. de Souza</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFB1E25B6FB7DFEA6" ID-GBIF-Occurrence="2234227110" collectingDate="2004-04-11" collectionCode="AUM" collectorName="M. Sabaj &amp; L. de Souza &amp; D. Werneke &amp; Lujan" country="Venezuela" county="Ventuari River" location="Yutaje creek" municipality="Manapiare River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFB1E25B6FAE0FF6E" bold="true" box="[1199,1342,188,213]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFB1E25B6FB2CFF6E" box="[1199,1266,188,213]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
41494
</emphasis>
(1, 
<quantity id="4CA69B56BA05C81CFADA25B6FF1DFF4C" metricMagnitude="-2" metricUnit="m" metricValue="7.13" pageId="39" pageNumber="40" unit="mm" value="71.3">71.3 mm</quantity>
SL), 
<collectingCounty id="62804E3FBA05C81CFEBF25D5FE67FF43" box="[270,441,223,248]" pageId="39" pageNumber="40">Ventuari River</collectingCounty>
drainage, 
<collectingMunicipality id="6B85ACC9BA05C81CFD8A25D5FD20FF43" box="[571,766,223,248]" pageId="39" pageNumber="40">Manapiare River</collectingMunicipality>
at mouth of 
<location id="8E816068BA05C81CFC1625D5FBE2FF43" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFC1625D5FBE2FF43" box="[935,1084,223,248]" country="Venezuela" county="Ventuari River" municipality="Manapiare River" name="Yutaje creek" pageId="39" pageNumber="40" stateProvince="Amazonas">Yutaje creek</location>
, 
<locationDeviation id="9A532F91BA05C81CFBFF25EAFECCFEA6" location="San Juan de Manapiare" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFBFF25EAFB45FF43" box="[1102,1179,223,248]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="39" pageNumber="40" unit="km" value="14.0">14 km</quantity>
NW of San Juan de Manapiare
</locationDeviation>
, 5.43667, -66.11261, 
<date id="FFE01073BA05C81CFDB4240EFD48FEA7" box="[517,662,260,284]" pageId="39" pageNumber="40" value="2004-04-11">
<collectingDate id="EFA4E99BBA05C81CFDB4240EFD48FEA7" box="[517,662,260,284]" pageId="39" pageNumber="40" value="2004-04-11">11 Apr-2004</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFD13240FFCD4FEA6" box="[674,778,260,285]" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA05C81CFCA4240FFC43FEA7" box="[789,925,260,285]" pageId="39" pageNumber="40">L. de Souza</collectorName>
, 
<collectorName id="26AB5365BA05C81CFC18240FFBEFFEA6" box="[937,1073,260,285]" pageId="39" pageNumber="40">D. Werneke</collectorName>
, N. 
<collectorName id="26AB5365BA05C81CFBEF240EFB7DFEA6" box="[1118,1187,260,285]" pageId="39" pageNumber="40">Lujan</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFB1F240EFC15FEDE" ID-GBIF-Occurrence="2234227114" collectingDate="2005-03-11" collectionCode="AUM" collectorName="M. Sabaj &amp; L. de Souza &amp; Werneke &amp; Lujan" country="Venezuela" location="Siapa River" municipality="Casiquiare River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFB1F240EFAE0FEA6" bold="true" box="[1198,1342,260,285]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFB1F240EFB2FFEA6" box="[1198,1265,260,285]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
42176
</emphasis>
(1, 
<quantity id="4CA69B56BA05C81CFADA240EFF1DFE84" metricMagnitude="-2" metricUnit="m" metricValue="7.15" pageId="39" pageNumber="40" unit="mm" value="71.5">71.5 mm</quantity>
SL), 
<collectingMunicipality id="6B85ACC9BA05C81CFEB8242DFE17FEFB" box="[265,457,295,320]" pageId="39" pageNumber="40">Casiquiare River</collectingMunicipality>
drainage, 
<location id="8E816068BA05C81CFDF7242DFD13FEFB" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFDF7242DFD13FEFB" box="[582,717,295,320]" country="Venezuela" municipality="Casiquiare River" name="Siapa River" pageId="39" pageNumber="40" stateProvince="Amazonas">Siapa River</location>
at rapids, 
<locationDeviation id="9A532F91BA05C81CFCFC242DFAC6FEFB" box="[845,1304,295,320]" location="San Carlos de Rio Negro" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFCFC242DFC7BFEFB" box="[845,933,295,320]" metricMagnitude="5" metricUnit="m" metricValue="1.54" pageId="39" pageNumber="40" unit="km" value="154.0">154 km</quantity>
E of San Carlos de Rio Negro
</locationDeviation>
, 1.60339, - 65.71587, 
<date id="FFE01073BA05C81CFEA02446FE79FEDE" box="[273,423,332,357]" pageId="39" pageNumber="40" value="2005-03-11">
<collectingDate id="EFA4E99BBA05C81CFEA02446FE79FEDE" box="[273,423,332,357]" pageId="39" pageNumber="40" value="2005-03-11">11-Mar-2005</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFE042447FDC1FEDE" box="[437,543,332,357]" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA05C81CFD9F2447FD65FEDF" box="[558,699,332,357]" pageId="39" pageNumber="40">L. de Souza</collectorName>
, D. 
<collectorName id="26AB5365BA05C81CFD5F2446FC8AFEDE" box="[750,852,332,357]" pageId="39" pageNumber="40">Werneke</collectorName>
, N. 
<collectorName id="26AB5365BA05C81CFC372446FC15FEDE" box="[902,971,332,357]" pageId="39" pageNumber="40">Lujan</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFC682446FDACFE17" ID-GBIF-Occurrence="2234227049" collectingDate="2005-03-16" collectionCode="AUM" collectorName="N. Lujan &amp; D. Werneke &amp; M. Sabaj &amp; M. Arce-Hernandez &amp; R. Betancur" country="Venezuela" county="Casiquiare" location="Guama creek" municipality="Siapa River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFC682446FBB5FEDE" bold="true" box="[985,1131,332,357]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFC682446FBC2FEDE" box="[985,1052,332,357]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
43572
</emphasis>
(3, 2, 
<quantity id="4CA69B56BA05C81CFB0D2446FABFFEDF" box="[1212,1377,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.869999999999999" metricValueMax="6.51" metricValueMin="5.2299999999999995" pageId="39" pageNumber="40" unit="mm" value="58.699999999999996" valueMax="65.1" valueMin="52.3">52.3–65.1 mm</quantity>
SL), 
<collectingCounty id="62804E3FBA05C81CFF262465FED1FE33" box="[151,271,367,392]" pageId="39" pageNumber="40">Casiquiare</collectingCounty>
/ 
<collectingMunicipality id="6B85ACC9BA05C81CFEA62465FE45FE33" box="[279,411,367,392]" pageId="39" pageNumber="40">Siapa River</collectingMunicipality>
drainage, 
<location id="8E816068BA05C81CFDA5247AFD74FE33" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFDA5247AFD74FE33" box="[532,682,367,392]" country="Venezuela" county="Casiquiare" municipality="Siapa River" name="Guama creek" pageId="39" pageNumber="40" stateProvince="Amazonas">Guama creek</location>
draining 
<location id="8E816068BA05C81CFCA9247AFC1EFE33" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFCA9247AFC1EFE33" box="[792,960,367,392]" country="Venezuela" county="Casiquiare" municipality="Siapa River" name="Cerro Derecha" pageId="39" pageNumber="40" stateProvince="Amazonas">Cerro Derecha</location>
, 
<date id="FFE01073BA05C81CFC7C2465FBBBFE33" box="[973,1125,367,392]" pageId="39" pageNumber="40" value="2005-03-16">
<collectingDate id="EFA4E99BBA05C81CFC7C2465FBBBFE33" box="[973,1125,367,392]" pageId="39" pageNumber="40" value="2005-03-16">16-Mar-2005</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFBC0247AFB07FE33" box="[1137,1241,367,392]" pageId="39" pageNumber="40">N. Lujan</collectorName>
, 
<collectorName id="26AB5365BA05C81CFB57247AFAAEFE33" box="[1254,1392,367,392]" pageId="39" pageNumber="40">D. Werneke</collectorName>
, 
<collectorName id="26AB5365BA05C81CFACD247AFF06FE16" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA05C81CFF58249FFE0FFE16" box="[233,465,404,429]" pageId="39" pageNumber="40">M. Arce-Hernández</collectorName>
, 
<collectorName id="26AB5365BA05C81CFE52249FFDACFE17" box="[483,626,405,428]" pageId="39" pageNumber="40">R. Betancur</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFD32249EFA46FE6B" ID-GBIF-Occurrence="2234227071" collectingDate="2005-03-26" collectionCode="AUM" collectorName="N. Lujan &amp; M. Arce-Hernandez &amp; L. Richmond &amp; Grant &amp; Wesley" country="Venezuela" location="Cano Soromoni" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFD32249EFCC6FE16" bold="true" box="[643,792,404,429]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFD32249EFD18FE16" box="[643,710,404,429]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
43868
</emphasis>
(1, 
<quantity id="4CA69B56BA05C81CFCE1249EFC64FE17" box="[848,954,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="7.94" pageId="39" pageNumber="40" unit="mm" value="79.4">79.4 mm</quantity>
SL), 
<location id="8E816068BA05C81CFBB4249EFB60FE16" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFBB4249EFB60FE16" box="[1029,1214,404,429]" country="Venezuela" name="Cano Soromoni" pageId="39" pageNumber="40" stateProvince="Amazonas">Caño Soromoni</location>
, 
<locationDeviation id="9A532F91BA05C81CFB7E249EFECEFE6B" location="La Esmeralda" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFB7E249EFAEEFE16" box="[1231,1328,404,429]" metricMagnitude="4" metricUnit="m" metricValue="1.18" pageId="39" pageNumber="40" unit="km" value="11.8">11.8 km</quantity>
E of La Esmeralda
</locationDeviation>
, 3.19380, -65.65197, 
<date id="FFE01073BA05C81CFDB624BDFD41FE6B" box="[519,671,439,464]" pageId="39" pageNumber="40" value="2005-03-26">
<collectingDate id="EFA4E99BBA05C81CFDB624BDFD41FE6B" box="[519,671,439,464]" pageId="39" pageNumber="40" value="2005-03-26">26-Mar-2005</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFD1A24B2FCCCFE6B" box="[683,786,439,464]" pageId="39" pageNumber="40">N. Lujan</collectorName>
, 
<collectorName id="26AB5365BA05C81CFCAE24B2FBDCFE6B" box="[799,1026,439,464]" pageId="39" pageNumber="40">M. Arce-Hernández</collectorName>
, 
<collectorName id="26AB5365BA05C81CFBBF24B2FB79FE6B" box="[1038,1191,439,464]" pageId="39" pageNumber="40">L. Richmond</collectorName>
, B. 
<collectorName id="26AB5365BA05C81CFB6524B2FAC9FE6B" box="[1236,1303,440,464]" pageId="39" pageNumber="40">Grant</collectorName>
, E. 
<collectorName id="26AB5365BA05C81CFAF324BDFA46FE6B" box="[1346,1432,439,464]" pageId="39" pageNumber="40">Wesley</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFF2624D6FC1FFDA3" ID-GBIF-Occurrence="2234227127" collectingDate="2010-04-11" collectionCode="AUM" collectorName="V. Meza &amp; O. Santaella" country="Venezuela" county="Parucito River" location="Manapiare River" municipality="Solomon" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFF2624D6FEF7FE4E" bold="true" box="[151,297,476,501]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFF2624D6FF04FE4E" box="[151,218,476,501]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
53894
</emphasis>
(1, not measured), 
<collectingCounty id="62804E3FBA05C81CFDBC24D6FD6CFE4E" box="[525,690,476,501]" pageId="39" pageNumber="40">Parucito River</collectingCounty>
at raudales 
<collectingMunicipality id="6B85ACC9BA05C81CFCF124D6FC76FE4E" box="[832,936,476,501]" pageId="39" pageNumber="40">Solomon</collectingMunicipality>
, 
<location id="8E816068BA05C81CFC0424D6FBA9FE4E" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFC0424D6FBA9FE4E" box="[949,1143,476,501]" country="Venezuela" county="Parucito River" municipality="Solomon" name="Manapiare River" pageId="39" pageNumber="40" stateProvince="Amazonas">Manapiare River</location>
, 
<location id="8E816068BA05C81CFB3524D6FAF5FE4E" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFB3524D6FAF5FE4E" box="[1156,1323,476,501]" country="Venezuela" county="Parucito River" municipality="Solomon" name="Ventuari River" pageId="39" pageNumber="40" stateProvince="Amazonas">Ventuari River</location>
drainage, 5.34648, -66.03328, 
<collectorName id="26AB5365BA05C81CFE32270AFE3DFDAC" box="[387,483,512,535]" pageId="39" pageNumber="40">V. Meza</collectorName>
, 
<collectorName id="26AB5365BA05C81CFE5E270AFDA3FDA3" box="[495,637,511,536]" pageId="39" pageNumber="40">O. Santaella</collectorName>
, and N. 
<location id="8E816068BA05C81CFD6C24F5FCFFFDA3" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFD6C24F5FCFFFDA3" box="[733,801,511,536]" country="Venezuela" county="Parucito River" municipality="Solomon" name="Lujan" pageId="39" pageNumber="40" stateProvince="Amazonas">Lujan</location>
, 
<date id="FFE01073BA05C81CFC9C270AFC1FFDA3" box="[813,961,512,536]" pageId="39" pageNumber="40" value="2010-04-11">
<collectingDate id="EFA4E99BBA05C81CFC9C270AFC1FFDA3" box="[813,961,512,536]" pageId="39" pageNumber="40" value="2010-04-11">11-Apr-2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFC7F24F5FBFEFDDB" ID-GBIF-Occurrence="2234226976" collectingDate="2010-04-03" collectionCode="AUM" collectorName="N. Lujan &amp; D. Werneke &amp; M. Sabaj &amp; T. Carvalho" country="Venezuela" county="Ventuari River" location="Leon" municipality="Moriche" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFC7F24F5FB81FDA3" bold="true" box="[974,1119,511,536]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFC7F24F5FBCFFDA3" box="[974,1041,511,536]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
54088
</emphasis>
(1, not measured), 
<collectingCounty id="62804E3FBA05C81CFA8C24F5FF09FD86" pageId="39" pageNumber="40">Ventuari River</collectingCounty>
, cobble shoal ~ 
<quantity id="4CA69B56BA05C81CFE3A272EFE1DFD86" box="[395,451,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="km" value="1.0">1 km</quantity>
upstream of 
<collectingMunicipality id="6B85ACC9BA05C81CFDE8272EFD65FD86" box="[601,699,548,573]" pageId="39" pageNumber="40">Moriche</collectingMunicipality>
, 
<locationDeviation id="9A532F91BA05C81CFD76272EFB73FD86" box="[711,1197,548,573]" location="San Fernando de Atabapo" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFD76272EFCC0FD86" box="[711,798,548,573]" metricMagnitude="5" metricUnit="m" metricValue="1.67" pageId="39" pageNumber="40" unit="km" value="167.0">167 km</quantity>
ENE of San Fernando de Atabapo
</locationDeviation>
, 4.75346, -66.37260, 
<collectorName id="26AB5365BA05C81CFF262742FEDEFDDB" box="[151,256,583,608]" pageId="39" pageNumber="40">N. Lujan</collectorName>
, 
<collectorName id="26AB5365BA05C81CFEBF2742FE44FDDB" box="[270,410,583,608]" pageId="39" pageNumber="40">D. Werneke</collectorName>
, 
<collectorName id="26AB5365BA05C81CFE162742FDCCFDDB" box="[423,530,583,608]" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, 
<collectorName id="26AB5365BA05C81CFD912742FD77FDDB" box="[544,681,583,608]" pageId="39" pageNumber="40">T. Carvalho</collectorName>
, O. 
<location id="8E816068BA05C81CFD6C2742FCC7FDE4" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFD6C2742FCC7FDE4" box="[733,793,584,607]" country="Venezuela" county="Ventuari River" municipality="Moriche" name="Leon" pageId="39" pageNumber="40" stateProvince="Amazonas">Leon</location>
, V. 
<location id="8E816068BA05C81CFCF82742FC54FDE4" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFCF82742FC54FDE4" box="[841,906,584,607]" country="Venezuela" county="Ventuari River" municipality="Moriche" name="Meza" pageId="39" pageNumber="40" stateProvince="Amazonas">Meza</location>
, 
<date id="FFE01073BA05C81CFC292742FBFEFDDB" box="[920,1056,584,608]" pageId="39" pageNumber="40" value="2010-04-03">
<collectingDate id="EFA4E99BBA05C81CFC292742FBFEFDDB" box="[920,1056,584,608]" pageId="39" pageNumber="40" value="2010-04-03">3-Apr-2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFB9C274DFD0BFD3F" ID-GBIF-Occurrence="2234227046" collectionCode="AUM, KM" collectorName="Manapiare" country="Venezuela" location="Manapiare River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFB9C274DFB1EFDDB" bold="true" box="[1069,1216,583,608]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFB9C274DFBAEFDDB" box="[1069,1136,583,608]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
54420
</emphasis>
(1, not measured), 
<location id="8E816068BA05C81CFF262766FE8BFD3E" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFF262766FE8BFD3E" box="[151,341,620,645]" country="Venezuela" name="Manapiare River" pageId="39" pageNumber="40" stateProvince="Amazonas">Manapiare River</location>
at 
<collectorName id="26AB5365BA05C81CFECD2766FE2AFD3E" box="[380,500,620,645]" pageId="39" pageNumber="40">Manapiare</collectorName>
landing, 173.8 
<collectionCode id="ED4FAE76BA05C81CFD192767FD0BFD3F" box="[680,725,621,644]" country="Russia" name="Kotel'nich Museum" pageId="39" pageNumber="40">KM</collectionCode>
</materialsCitation>
<materialsCitation id="3B363CEEBA05C81CFD6F2766FD76FD13" ID-GBIF-Occurrence="2234227116" collectingDate="2010-04-13" collectorName="J. Birindelli" country="Venezuela" county="Puerto Ayacucho" location="Leon" municipality="Ventuari River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
ESE of 
<collectingCounty id="62804E3FBA05C81CFC892767FC22FD3E" box="[824,1020,620,645]" pageId="39" pageNumber="40">Puerto Ayacucho</collectingCounty>
, 
<collectingMunicipality id="6B85ACC9BA05C81CFBB82766FB71FD3E" box="[1033,1199,620,645]" pageId="39" pageNumber="40">Ventuari River</collectingMunicipality>
drainage, 5.32864, - 66.05067, 
<collectorName id="26AB5365BA05C81CFEBE279AFE46FD13" box="[271,408,655,680]" pageId="39" pageNumber="40">J. Birindelli</collectorName>
, O. 
<location id="8E816068BA05C81CFE78279AFDDBFD13" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFE78279AFDDBFD13" box="[457,517,656,680]" country="Venezuela" county="Puerto Ayacucho" municipality="Ventuari River" name="Leon" pageId="39" pageNumber="40" stateProvince="Amazonas">Léon</location>
, 
<date id="FFE01073BA05C81CFDA3279AFD76FD13" box="[530,680,656,680]" pageId="39" pageNumber="40" value="2010-04-13">
<collectingDate id="EFA4E99BBA05C81CFDA3279AFD76FD13" box="[530,680,656,680]" pageId="39" pageNumber="40" value="2010-04-13">13-Apr-2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B363CEEBA05C81CFD052785FF05FD4B" ID-GBIF-Occurrence="2234226964" collectingDate="2010-04-02" collectionCode="AUM, INHS" collectorName="N. Lujan &amp; M. Sabaj" country="Venezuela" county="Marueta creek" location="Carvalho" municipality="Werneke" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B92AEAA1BA05C81CFD052785FC98FD13" bold="true" box="[692,838,655,680]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFD052785FD29FD13" box="[692,759,655,680]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
54313
</emphasis>
(1, not measured), 
<collectingCounty id="62804E3FBA05C81CFB97279AFB15FD13" box="[1062,1227,655,680]" pageId="39" pageNumber="40">Marueta creek</collectingCounty>
, 
<locationDeviation id="9A532F91BA05C81CFB682785FE4EFD76" location="San Fernando de Arabapo" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFB682785FAF1FD13" box="[1241,1327,655,680]" metricMagnitude="5" metricUnit="m" metricValue="1.59" pageId="39" pageNumber="40" unit="km" value="159.0">159 km</quantity>
E of San Fernando de Arabapo
</locationDeviation>
, 4.29481, -66.28888, 
<collectorName id="26AB5365BA05C81CFD3B27BFFD2DFD76" box="[650,755,692,717]" pageId="39" pageNumber="40">N. Lujan</collectorName>
, 
<collectorName id="26AB5365BA05C81CFCB127BFFCB4FD76" box="[768,874,692,717]" pageId="39" pageNumber="40">M. Sabaj</collectorName>
, D. 
<collectingMunicipality id="6B85ACC9BA05C81CFC2A27BEFBDFFD76" box="[923,1025,692,717]" pageId="39" pageNumber="40">Werneke</collectingMunicipality>
, T. 
<location id="8E816068BA05C81CFB9D27BEFB48FD76" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFB9D27BEFB48FD76" box="[1068,1174,692,717]" country="Venezuela" county="Marueta creek" municipality="Werneke" name="Carvalho" pageId="39" pageNumber="40" stateProvince="Amazonas">Carvalho</location>
, V. 
<location id="8E816068BA05C81CFB7527BFFADBFD77" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFB7527BFFADBFD77" box="[1220,1285,693,716]" country="Venezuela" county="Marueta creek" municipality="Werneke" name="Meza" pageId="39" pageNumber="40" stateProvince="Amazonas">Meza</location>
, 
<date id="FFE01073BA05C81CFAA327BEFA47FD77" box="[1298,1433,692,716]" pageId="39" pageNumber="40" value="2010-04-02">
<collectingDate id="EFA4E99BBA05C81CFAA327BEFA47FD77" box="[1298,1433,692,716]" pageId="39" pageNumber="40" value="2010-04-02">2-Apr-2010</collectingDate>
</date>
. 
<emphasis id="B92AEAA1BA05C81CFF2627DDFF05FD4B" bold="true" box="[151,219,727,752]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFF2627DDFF05FD4B" box="[151,219,727,752]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="39" pageNumber="40" type="Other type of research institution/biorepository">INHS</collectionCode>
</emphasis>
</materialsCitation>
<materialsCitation id="3B363CEEBA05C81CFF5327DDFE1CFCAE" ID-GBIF-Occurrence="2234227094" collectingDate="1992-01-21" collectionCode="MCNG" collectorName="L. Page &amp; L. Nico &amp; P. Ceas &amp; Lyons" country="Venezuela" location="Pozo Azul" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
61554 (3, 
<quantity id="4CA69B56BA05C81CFEE527DDFE28FD54" box="[340,502,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="6.89" metricValueMax="7.959999999999999" metricValueMin="5.82" pageId="39" pageNumber="40" unit="mm" value="68.9" valueMax="79.6" valueMin="58.2">58.2–79.6 mm</quantity>
SL), stream near 
<location id="8E816068BA05C81CFD0D27D2FCEAFD4B" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFD0D27D2FCEAFD4B" box="[700,820,727,752]" country="Venezuela" name="Pozo Azul" pageId="39" pageNumber="40" stateProvince="Amazonas">Pozo Azul</location>
, 5.763219, -67.489034, 
<date id="FFE01073BA05C81CFBF227D2FB0CFD4B" box="[1091,1234,728,752]" pageId="39" pageNumber="40" value="1992-01-21">
<collectingDate id="EFA4E99BBA05C81CFBF227D2FB0CFD4B" box="[1091,1234,728,752]" pageId="39" pageNumber="40" value="1992-01-21">21-Jan-1992</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFB6C27D2FAEBFD4B" box="[1245,1333,728,752]" pageId="39" pageNumber="40">L. Page</collectorName>
, 
<collectorName id="26AB5365BA05C81CFAF127D2FA46FD4B" box="[1344,1432,727,752]" pageId="39" pageNumber="40">L. Nico</collectorName>
, 
<collectorName id="26AB5365BA05C81CFF2627F7FF32FCAF" box="[151,236,764,788]" pageId="39" pageNumber="40">P. Ceas</collectorName>
, J. 
<collectorName id="26AB5365BA05C81CFEA527F7FE82FCAE" box="[276,348,765,789]" pageId="39" pageNumber="40">Lyons</collectorName>
; 
<emphasis id="B92AEAA1BA05C81CFED827F6FE1CFCAE" bold="true" box="[361,450,764,789]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFED827F6FE1CFCAE" box="[361,450,764,789]" country="Venezuela" httpUri="http://grbio.org/cool/tyz9-grz8" name="Museo de Ciencias Naturales de la UNELLEZ en Guanare" pageId="39" pageNumber="40">MCNG</collectionCode>
</emphasis>
</materialsCitation>
<materialsCitation id="3B363CEEBA05C81CFE7B27F6FA42FC83" ID-GBIF-Occurrence="2234227141" collectingDate="1998-01-24" collectionCode="MCNG" country="Venezuela" location="Puerto Ayacucho" municipality="Agua Linda creek" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
38168 (1 of 3, 86.0 mm SL), 
<collectingMunicipality id="6B85ACC9BA05C81CFCAF27F6FC30FCAE" box="[798,1006,764,789]" pageId="39" pageNumber="40">Agua Linda creek</collectingMunicipality>
, 
<quantity id="4CA69B56BA05C81CFC4B27F6FB94FCAE" box="[1018,1098,764,789]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="40" unit="km" value="0.5">0.5 km</quantity>
upper waters 
<locationDeviation id="9A532F91BA05C81CFB5C27F6FE42FC83" firstLocation="Puerto Ayacucho" pageId="39" pageNumber="40" secondLocation="Caicara">between Puerto Ayacucho and Caicara</locationDeviation>
, approx. 
<quantity id="4CA69B56BA05C81CFDBA262AFD88FC83" box="[523,598,799,824]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="40" unit="km" value="30.0">30 km</quantity>
north of 
<location id="8E816068BA05C81CFD78262AFC4EFC83" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFD78262AFC4EFC83" box="[713,912,799,824]" country="Venezuela" municipality="Agua Linda creek" name="Puerto Ayacucho" pageId="39" pageNumber="40" stateProvince="Amazonas">Puerto Ayacucho</location>
, 5.645368, -67.456057 
<date id="FFE01073BA05C81CFB14262AFAEAFC83" box="[1189,1332,800,824]" pageId="39" pageNumber="40" value="1998-01-24">
<collectingDate id="EFA4E99BBA05C81CFB14262AFAEAFC83" box="[1189,1332,800,824]" pageId="39" pageNumber="40" value="1998-01-24">24-Jan-1998</collectingDate>
</date>
; 
<emphasis id="B92AEAA1BA05C81CFAF22615FA42FC83" bold="true" box="[1347,1436,799,824]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFAF22615FA42FC83" box="[1347,1436,799,824]" country="Venezuela" httpUri="http://grbio.org/cool/tyz9-grz8" name="Museo de Ciencias Naturales de la UNELLEZ en Guanare" pageId="39" pageNumber="40">MCNG</collectionCode>
</emphasis>
</materialsCitation>
<materialsCitation id="3B363CEEBA05C81CFF26264EFE8EFC3B" ID-GBIF-Occurrence="2234227016" collectingDate="2004-12-04" collectionCode="UF" country="Venezuela" location="Km" municipality="Manapiare River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
50038 (3, 64.8–73.0 mm SL), 
<collectingMunicipality id="6B85ACC9BA05C81CFE5B264EFD77FCE6" box="[490,681,836,861]" pageId="39" pageNumber="40">Manapiare River</collectingMunicipality>
, 14.5 
<location id="8E816068BA05C81CFD5D264FFCC8FCE7" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFD5D264FFCC8FCE7" box="[748,790,837,860]" country="Venezuela" municipality="Manapiare River" name="Km" pageId="39" pageNumber="40" stateProvince="Amazonas">Km</location>
al 
<locationDeviation id="9A532F91BA05C81CFC89264FFB48FCE6" box="[824,1174,836,861]" location="San Juan de Manapiare" pageId="39" pageNumber="40">NW de San Juan de Manapiare</locationDeviation>
, 5.428611, -66.136111, 
<date id="FFE01073BA05C81CFF262662FEFEFC3B" box="[151,288,872,896]" pageId="39" pageNumber="40" value="2004-12-04">
<collectingDate id="EFA4E99BBA05C81CFF262662FEFEFC3B" box="[151,288,872,896]" pageId="39" pageNumber="40" value="2004-12-04">4-Dec-2004</collectingDate>
</date>
; 
<emphasis id="B92AEAA1BA05C81CFE9A266DFE8EFC3B" bold="true" box="[299,336,871,896]" pageId="39" pageNumber="40">
<collectionCode id="ED4FAE76BA05C81CFE9A266DFE8EFC3B" box="[299,336,871,896]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="39" pageNumber="40" type="Museum">UF</collectionCode>
</emphasis>
</materialsCitation>
<materialsCitation id="3B363CEEBA05C81CFEE6266DFD77FC1E" ID-GBIF-Occurrence="2234227128" collectingDate="1981-03-20" collectorName="S. Reid &amp; C. Gilbert. &amp; State" country="Venezuela" county="Ventuari River" location="Tencua Falls" municipality="Tencua" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Apure" typeStatus="paratype">
77841 (1, 
<quantity id="4CA69B56BA05C81CFE78266DFDF3FCC4" box="[457,557,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="39" pageNumber="40" unit="mm" value="42.5">42.5 mm</quantity>
SL), 
<collectingCounty id="62804E3FBA05C81CFDDA266DFCD1FC3B" box="[619,783,871,896]" pageId="39" pageNumber="40">Ventuari River</collectingCounty>
at 
<collectingMunicipality id="6B85ACC9BA05C81CFC802662FC5AFCC4" box="[817,900,872,895]" pageId="39" pageNumber="40">Tencua</collectingMunicipality>
, just below 
<location id="8E816068BA05C81CFBB82662FB44FC3B" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFBB82662FB44FC3B" box="[1033,1178,871,896]" country="Venezuela" county="Ventuari River" municipality="Tencua" name="Tencua Falls" pageId="39" pageNumber="40" stateProvince="Apure">Tencua Falls</location>
, 5.050752, -65.625381 
<date id="FFE01073BA05C81CFF262686FEF0FC1F" box="[151,302,908,932]" pageId="39" pageNumber="40" value="1981-03-20">
<collectingDate id="EFA4E99BBA05C81CFF262686FEF0FC1F" box="[151,302,908,932]" pageId="39" pageNumber="40" value="1981-03-20">20-Mar-1981</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFE882686FE51FC1E" box="[313,399,908,933]" pageId="39" pageNumber="40">S. Reid</collectorName>
, 
<collectorName id="26AB5365BA05C81CFE282686FDCFFC1E" box="[409,529,908,933]" pageId="39" pageNumber="40">C. Gilbert.</collectorName>
<emphasis id="B92AEAA1BA05C81CFDA62686FD77FC1E" bold="true" box="[535,681,908,933]" pageId="39" pageNumber="40">
<collectingRegion id="499AF851BA05C81CFDA62686FDBDFC1E" box="[535,611,908,933]" country="Venezuela" name="Apure" pageId="39" pageNumber="40">Apure</collectingRegion>
<collectorName id="26AB5365BA05C81CFDDB2686FD77FC1E" box="[618,681,908,933]" pageId="39" pageNumber="40">State</collectorName>
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B363CEEBA05C81CFD042686FDF8FC56" ID-GBIF-Occurrence="2234226977" collectingDate="1999-12-26" collectionCode="AUM" collectorName="J. Armbruster &amp; M. Hardman &amp; J. Evans &amp; J. Thomas &amp; State" country="Venezuela" location="San Fernando" municipality="Potrerito creek" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<collectionCode id="ED4FAE76BA05C81CFD042686FD26FC1E" box="[693,760,908,933]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
22631 (10, 5, 
<quantity id="4CA69B56BA05C81CFC2B2686FB95FC1F" box="[922,1099,908,933]" metricMagnitude="-2" metricUnit="m" metricValue="7.1185" metricValueMax="8.787" metricValueMin="5.45" pageId="39" pageNumber="40" unit="mm" value="71.185" valueMax="87.87" valueMin="54.5">54.5–87.87 mm</quantity>
SL) 
<collectingMunicipality id="6B85ACC9BA05C81CFB332686FAF5FC1E" box="[1154,1323,908,933]" pageId="39" pageNumber="40">Potrerito creek</collectingMunicipality>
, ca. 
<locationDeviation id="9A532F91BA05C81CFAE72686FEE8FC7C" location="Puerto Paez" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFAE72686FA42FC1E" box="[1366,1436,908,933]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="km" value="15.0">15 km</quantity>
N Puerto Paez
</locationDeviation>
on road to 
<location id="8E816068BA05C81CFE0526BAFD8CFC73" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFE0526BAFD8CFC73" box="[436,594,943,968]" country="Venezuela" municipality="Potrerito creek" name="San Fernando" pageId="39" pageNumber="40" stateProvince="Bolivar">San Fernando</location>
, 6.41194, -67.53194, 
<date id="FFE01073BA05C81CFCF526A5FC05FC73" box="[836,987,943,968]" pageId="39" pageNumber="40" value="1999-12-26">
<collectingDate id="EFA4E99BBA05C81CFCF526A5FC05FC73" box="[836,987,943,968]" pageId="39" pageNumber="40" value="1999-12-26">26-Dec-1999</collectingDate>
</date>
, collectors: 
<collectorName id="26AB5365BA05C81CFBD126BAFB22FC73" box="[1120,1276,943,968]" pageId="39" pageNumber="40">J. Armbruster</collectorName>
, 
<collectorName id="26AB5365BA05C81CFAB726BAFA46FC73" box="[1286,1432,943,968]" pageId="39" pageNumber="40">M. Hardman</collectorName>
, 
<collectorName id="26AB5365BA05C81CFF2626DFFF29FC57" box="[151,247,981,1004]" pageId="39" pageNumber="40">J. Evans</collectorName>
, 
<collectorName id="26AB5365BA05C81CFEB226DFFEA5FC56" box="[259,379,980,1005]" pageId="39" pageNumber="40">J. Thomas</collectorName>
; 
<emphasis id="B92AEAA1BA05C81CFE3726DEFDF8FC56" bold="true" box="[390,550,980,1005]" pageId="39" pageNumber="40">
<collectingRegion id="499AF851BA05C81CFE3726DEFE01FC56" box="[390,479,980,1005]" country="Venezuela" name="Bolivar" pageId="39" pageNumber="40">Bolivar</collectingRegion>
<collectorName id="26AB5365BA05C81CFE5626DEFDF8FC56" box="[487,550,980,1005]" pageId="39" pageNumber="40">State</collectorName>
</emphasis>
</materialsCitation>
: 
<materialsCitation id="3B363CEEBA05C81CFD8226DEFAACFBAB" ID-GBIF-Occurrence="2234227121" collectingDate="1999-12-25" collectionCode="AUM" collectorName="J. Armbruster &amp; M. Hardman &amp; J. Evans &amp; Thomas" country="Venezuela" location="Caicara-Puerto Ayacucho" municipality="Ore creek" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<collectionCode id="ED4FAE76BA05C81CFD8226DEFDA8FC56" box="[563,630,980,1005]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="39" pageNumber="40" type="University or college">AUM</collectionCode>
22336 (1, 
<quantity id="4CA69B56BA05C81CFD4026DEFC8BFC57" box="[753,853,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="4.62" pageId="39" pageNumber="40" unit="mm" value="46.2">46.2 mm</quantity>
SL), 6.31583, -67.09611, 
<collectingMunicipality id="6B85ACC9BA05C81CFBCE26DEFB31FC56" box="[1151,1263,980,1005]" pageId="39" pageNumber="40">Ore creek</collectingMunicipality>
, ca. 
<locationDeviation id="9A532F91BA05C81CFA9226DEFEEDFBAB" location="Los Pijiguaos" pageId="39" pageNumber="40">
<quantity id="4CA69B56BA05C81CFA9226DEFAB5FC56" box="[1315,1387,980,1005]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="40" unit="km" value="50.0">50 km</quantity>
SW Los Pijiguaos
</locationDeviation>
, 
<location id="8E816068BA05C81CFE8F26FDFD82FBAB" LSID="urn:lsid:plazi:treatment:03F787A5BA04C811FF2620ADFC27FF43:8E816068BA05C81CFE8F26FDFD82FBAB" box="[318,604,1015,1040]" country="Venezuela" municipality="Ore creek" name="Caicara-Puerto Ayacucho" pageId="39" pageNumber="40" stateProvince="Bolivar">Caicara-Puerto Ayacucho</location>
road., 
<date id="FFE01073BA05C81CFD1B26FDFC9EFBAB" box="[682,832,1015,1040]" pageId="39" pageNumber="40" value="1999-12-25">
<collectingDate id="EFA4E99BBA05C81CFD1B26FDFC9EFBAB" box="[682,832,1015,1040]" pageId="39" pageNumber="40" value="1999-12-25">25-Dec-1999</collectingDate>
</date>
, 
<collectorName id="26AB5365BA05C81CFCFA26F2FC39FBAB" box="[843,999,1015,1040]" pageId="39" pageNumber="40">J. Armbruster</collectorName>
, 
<collectorName id="26AB5365BA05C81CFC4326F2FB5AFBAB" box="[1010,1156,1015,1040]" pageId="39" pageNumber="40">M. Hardman</collectorName>
, 
<collectorName id="26AB5365BA05C81CFB3E26F2FB31FBB4" box="[1167,1263,1016,1039]" pageId="39" pageNumber="40">J. Evans</collectorName>
, J. 
<collectorName id="26AB5365BA05C81CFAA526FDFAACFBAB" box="[1300,1394,1015,1040]" pageId="39" pageNumber="40">Thomas</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3446538BA05C81CFF762116FC37FAEF" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BE136B3BA05C81CFF762116FC37FAEF" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="40">
<emphasis id="B92AEAA1BA05C81CFF762116FE9FFB8E" bold="true" box="[199,321,1052,1077]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
<taxonomicName id="4C5E4D30BA05C81CFEE12116FDF3FB8F" authorityName="De &amp; Taphorn &amp; Armbruster" authorityYear="2019" box="[336,557,1052,1076]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="patronus">
<emphasis id="B92AEAA1BA05C81CFEE12116FDF3FB8F" box="[336,557,1052,1076]" italics="true" pageId="39" pageNumber="40">Ancistrus patronus</emphasis>
</taxonomicName>
differs from all other 
<taxonomicName id="4C5E4D30BA05C81CFCFF2116FC66FB88" authorityName="Kner" authorityYear="1854" box="[846,952,1052,1075]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA1BA05C81CFCFF2116FC66FB88" box="[846,952,1052,1075]" italics="true" pageId="39" pageNumber="40">Ancistrus</emphasis>
</taxonomicName>
in the region except 
<emphasis id="B92AEAA1BA05C81CFB7C2116FE36FBEC" italics="true" pageId="39" pageNumber="40">
<taxonomicName id="4C5E4D30BA05C81CFB7C2116FAAEFB8F" authorityName="Eigenmann" authorityYear="1912" box="[1229,1392,1052,1076]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="lithurgicus">A. lithurgicus</taxonomicName>
, 
<taxonomicName id="4C5E4D30BA05C81CFA352116FE8BFBE3" authorityName="Pellegrin" authorityYear="1912" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="macrophthalmus">A. macrophthalmus</taxonomicName>
, 
<taxonomicName id="4C5E4D30BA05C81CFED2214AFE36FBEC" box="[355,488,1088,1111]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="maximus">A. maximus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C5E4D30BA05C81CFD92214AFD77FBE3" baseAuthorityName="Muller &amp; Troschel" baseAuthorityYear="1849" box="[547,681,1088,1112]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="nudiceps">
<emphasis id="B92AEAA1BA05C81CFD92214AFD77FBE3" box="[547,681,1088,1112]" italics="true" pageId="39" pageNumber="40">A. nudiceps</emphasis>
</taxonomicName>
by having an entirely black body with tiny white to golden dots; from 
<taxonomicName id="4C5E4D30BA05C81CFF65216EFEF5FBC7" authorityName="De &amp; Taphorn &amp; Armbruster" authorityYear="2019" box="[212,299,1124,1148]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="leoni">
<emphasis id="B92AEAA1BA05C81CFF65216EFEF5FBC7" box="[212,299,1124,1148]" italics="true" pageId="39" pageNumber="40">A. leoni</emphasis>
</taxonomicName>
by having the abdomen almost as dark as the sides (vs. much lighter); from 
<taxonomicName id="4C5E4D30BA05C81CFBCC216EFADEFBC7" baseAuthorityName="Muller &amp; Troschel" baseAuthorityYear="1849" box="[1149,1280,1124,1148]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="nudiceps">
<emphasis id="B92AEAA1BA05C81CFBCC216EFADEFBC7" box="[1149,1280,1124,1148]" italics="true" pageId="39" pageNumber="40">A. nudiceps</emphasis>
</taxonomicName>
by reaching a much smaller maximum size (86 vs. 
<quantity id="4CA69B56BA05C81CFD822182FD4AFB1B" box="[563,660,1160,1184]" metricMagnitude="-1" metricUnit="m" metricValue="1.38" pageId="39" pageNumber="40" unit="mm" value="138.0">138 mm</quantity>
SL), by having the distance between the anteromedial plate and the snout greater than or equal to the distance between the anteromedial plate and a line formed by the anterior margins of the nasal apertures (vs. less than); and from 
<taxonomicName id="4C5E4D30BA05C81CFD0F21DAFCC6FB53" authorityName="De &amp; Taphorn &amp; Armbruster" authorityYear="2019" box="[702,792,1232,1256]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="leoni">
<emphasis id="B92AEAA1BA05C81CFD0F21DAFCC6FB53" box="[702,792,1232,1256]" italics="true" pageId="39" pageNumber="40">A. leoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5E4D30BA05C81CFC9B21DAFC1AFB53" authorityName="Eigenmann" authorityYear="1912" box="[810,964,1232,1256]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="lithurgicus">
<emphasis id="B92AEAA1BA05C81CFC9B21DAFC1AFB53" box="[810,964,1232,1256]" italics="true" pageId="39" pageNumber="40">A. lithurgicus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C5E4D30BA05C81CFBB921DAFB3BFB53" authorityName="Pellegrin" authorityYear="1912" box="[1032,1253,1232,1256]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="macrophthalmus">
<emphasis id="B92AEAA1BA05C81CFBB921DAFB3BFB53" box="[1032,1253,1232,1256]" italics="true" pageId="39" pageNumber="40">A. macrophthalmus</emphasis>
</taxonomicName>
by being deepbodied (vs. flattened), by having the median plate below the posterior edge of the adipose-fin membrane about twice as tall as wide (vs. just slightly taller than), and by the eyes being laterally placed (vs. dorsolaterally placed in 
<taxonomicName id="4C5E4D30BA05C81CFF262036FEF0FAEF" authorityName="Eigenmann" authorityYear="1912" box="[151,302,1340,1364]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="lithurgicus">
<emphasis id="B92AEAA1BA05C81CFF262036FEF0FAEF" box="[151,302,1340,1364]" italics="true" pageId="39" pageNumber="40">A. lithurgicus</emphasis>
</taxonomicName>
and almost completely dorsally placed in 
<taxonomicName id="4C5E4D30BA05C81CFCB92036FC3CFAEF" authorityName="Pellegrin" authorityYear="1912" box="[776,994,1340,1364]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="macrophthalmus">
<emphasis id="B92AEAA1BA05C81CFCB92036FC3CFAEF" box="[776,994,1340,1364]" italics="true" pageId="39" pageNumber="40">A. macrophthalmus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3446538BA05C812FF762055FB5FF80A" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40" type="description">
<paragraph id="8BE136B3BA05C81CFF762055FBBDF923" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="40">
<emphasis id="B92AEAA1BA05C81CFF762055FE87FAC3" bold="true" box="[199,345,1375,1400]" pageId="39" pageNumber="40">Description.</emphasis>
Morphometrics given in 
<tableCitation id="C6DC0308BA05C81CFDC62055FD10FAC3" box="[631,718,1375,1400]" captionStart="TABLE 5" captionStartId="37.[151,239,154,177]" captionText="TABLE 5. Morphometric features of Ancistrus nudiceps (N =60, except 1 N=51) and A. patronus n. sp. (N=47, except 2 N=44 and 3 N=46)." pageId="39" pageNumber="40">Table 5</tableCitation>
. Size range this study: 
<quantity id="4CA69B56BA05C81CFC622055FB92FAC3" box="[979,1100,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" metricValueMax="8.6" metricValueMin="4.3" pageId="39" pageNumber="40" unit="mm" value="64.5" valueMax="86.0" valueMin="43.0">43–86 mm</quantity>
SL. A relatively large, robust 
<taxonomicName id="4C5E4D30BA05C81CFF26208EFEDFFA20" authorityName="Kner" authorityYear="1854" box="[151,257,1412,1435]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="39" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA1BA05C81CFF26208EFEDFFA20" box="[151,257,1412,1435]" italics="true" pageId="39" pageNumber="40">Ancistrus</emphasis>
</taxonomicName>
, body broadest anteriorly, greatest body width near opercles, usually at base of pectoral spines, but sometimes body slightly wider near middle of pectoral-spine length in fish with plump abdomens, thence narrowing progressively to end of caudal peduncle. Head and body not very depressed, greatest body depth just anterior to dorsal-fin origin. Caudal peduncle deep, robust, compressed posteriorly. Dorsal profile of head steeply ascending through length of soft snout, then continuing in gentle concave arc to tip of supraoccipital process where it then descends to dorsal-fin origin and continues its decline to point below tip of adipose-fin spine; then angled up through dorsal procurrent caudal-fin spines. Ventral profile of mouth flat to beneath pectoral-fin insertions, then slightly concave to pelvic-fin insertions, then almost straight to lower caudal-fin spine.
</paragraph>
<paragraph id="8BE136B3BA05C81CFF7623AEFAC5F866" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="40">Head wide, interorbital width about equal to head depth, much more than half of head length. Snout rounded with large broad naked margin in males, less wide in females and juveniles. Snout length more than one-half head length. Eye large, interorbital area slightly convex. Oral disk ovate, wider than long. Lips covered with minute papillae, larger near mouth. Lower lip moderate in size, not reaching gill aperture, its border covered with very small papillae. Maxillary barbel very short, its length less than orbit diameter. Jaws short with premaxillary tooth rows forming strong arc and dentary tooth rows forming angle of&gt;135°. Dentary and premaxillary tooth rows strongly curved medially, lateralmost dentary tooth medial to lateralmost premaxillary tooth. Teeth small and numerous (58–120 per jaw ramus), asymmetrically bifid, medial cusp much larger and spatulate, lateral cusp minute and pointed, usually not reaching more than half length of medial cusp, but equal in worn teeth.</paragraph>
<caption id="DF21663BBA0AC813FF262224FE71F8D8" httpUri="https://zenodo.org/record/2623192/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1838,1860]" targetBox="[151,1436,193,1817]" targetPageId="40">
<paragraph id="8BE136B3BA0AC813FF262224FE71F8D8" blockId="40.[151,1436,1838,1891]" pageId="40" pageNumber="41">
<emphasis id="B92AEAA1BA0AC813FF262224FEFCF8F8" bold="true" box="[151,290,1838,1860]" pageId="40" pageNumber="41">FIGURE 19.</emphasis>
<taxonomicName id="4C5E4D30BA0AC813FE9A2225FE33F8FF" authorityName="De &amp; Taphorn &amp; Armbruster" authorityYear="2019" box="[299,493,1839,1860]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="patronus">
<emphasis id="B92AEAA1BA0AC813FE9A2225FE33F8FF" box="[299,493,1839,1860]" italics="true" pageId="40" pageNumber="41">Ancistrus patronus</emphasis>
</taxonomicName>
, holotype, in dorsal, lateral, and ventral views, AUM 39876, 83.6 mm SL. Scale = 1 cm. Photos by J.W. Armbruster.
</paragraph>
</caption>
<caption id="DF21663BBA0BC812FF2627BDFE79FD50" httpUri="https://zenodo.org/record/2623194/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,695,717]" targetBox="[151,1436,193,674]" targetPageId="41">
<paragraph id="8BE136B3BA0BC812FF2627BDFE79FD50" blockId="41.[151,1435,695,747]" pageId="41" pageNumber="42">
<emphasis id="B92AEAA1BA0BC812FF2627BDFEFFFD77" bold="true" box="[151,289,695,717]" pageId="41" pageNumber="42">FIGURE 20.</emphasis>
Live coloration of head of 
<taxonomicName id="4C5E4D30BA0BC812FD8A27B2FD25FD76" authorityName="De &amp; Taphorn &amp; Armbruster" authorityYear="2019" box="[571,763,696,717]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="patronus">
<emphasis id="B92AEAA1BA0BC812FD8A27B2FD25FD76" box="[571,763,696,717]" italics="true" pageId="41" pageNumber="42">Ancistrus patronus</emphasis>
</taxonomicName>
in dorsal and lateral views, AUM 43868, 79.4 mm SL. Scale = 1 cm. Photos by N.K. Lujan.
</paragraph>
</caption>
<paragraph id="8BE136B3BA0BC812FF762612FD1AFC26" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">Hypertrophied cheek odontodes strongly evertible, nine to 19, stout with tips hooked anteriorly, bases encased in thick fleshy sheaths. Exposed part of opercle roughly triangular, covered with short stout odontodes. Head smooth, bones on back of head not carinate; supraoccipital with margins between surrounding bones and plates usually clearly visible. Lateral plates not carinate.</paragraph>
<paragraph id="8BE136B3BA0BC812FF7626A2FE76FBCE" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">Ventral surface of head and abdomen naked, no exposed platelets anterior to anal-fin spine. Nuchal plate small and curved posterolaterally. Enlarged odontodes present along edges of lateral plates. Five series of lateral plates anteriorly, three series on caudal peduncle, mid-dorsal and mid-ventral plate series end on caudal peduncle beneath adipose fin. Last plate in median series much smaller than penultimate plate, and median plate below end of adipose fin about twice as high as wide. Base of caudal fin with about eight roughly triangular platelets covering bases of caudal-fin rays.</paragraph>
<paragraph id="8BE136B3BA0BC812FF76218AFAF4FACB" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">Dorsal-fin origin situated anterior to vertical through pelvic-fin insertion. First dorsal-fin ray not elongate, but longer than snout length; last dorsal-fin ray reaching adipose-fin spine when depressed. Dorsal-fin base length much longer than dorsal–adipose distance. Adipose fin stout, angled up from body, membrane visible beneath spine. Pectoral spine stout, moderately long, reaching just past pelvic-fin base but not to cloaca. Anal fin small but well developed; base of first anal-fin pterygiophore covered by skin, anal-fin origin posterior to vertical through base of last dorsal-fin ray. Pelvic fins reaching well past anal-fin origin but not past anal-fin tip, inserted at vertical about through second branched dorsal-fin ray. Caudal fin truncate, lower lobe slightly longer than upper.</paragraph>
<paragraph id="8BE136B3BA0BC812FF762076FB30FA02" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">Tiny odontodes present on body plates, largest on posterior margins of plates. All fin spines with small odontodes, more developed in pectoral-fin spine of males. All fin rays with tiny odontodes on rays.</paragraph>
<paragraph id="8BE136B3BA0BC812FF7620CFFC20F9F3" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">Meristics (N=47). Mid-dorsal plates 6–18, x= ̃13; median plates 22-24, x= ̃ 23; mid-ventral plates 16–19, x= ̃ 17–18; plates bordering dorsal-fin base 4–8, x̃ = 7; plates between dorsal and adipose fins 4–8 median 6–7; preadipose plates one. Fin-ray formulae invariate: dorsal I,7; pectoral I,6; pelvic i,5; anal i,4; caudal i,14,i. Caudal procurrent spines: dorsal: four to five, x= ̃five; ventral: two to four, x= ̃three.</paragraph>
<paragraph id="8BE136B3BA0BC812FF76235EFEDBF947" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">
<emphasis id="B92AEAA1BA0BC812FF76235EFE61F9D6" bold="true" box="[199,447,1620,1645]" pageId="41" pageNumber="42">Sexual dimorphism.</emphasis>
(
<figureCitation id="13652A36BA0BC812FE66235EFDEFF9D6" box="[471,561,1620,1645]" captionStart="FIGURE 5" captionStartId="9.[151,250,1210,1232]" captionTargetBox="[194,1392,337,1188]" captionTargetId="figure@9.[194,1392,337,1188]" captionTargetPageId="9" captionText="FIGURE 5. Heads, dorsal view, of A. Ancistrus nudiceps AUM 35623, 140.2 mm SL and B. A. patronus, holotype, AUM 39876, 83.6 mm SL demonstrating differences in head shape and tentacle pattern. Ancistrus nudiceps has a longer, more spatulate snout with the point from the edge of medial snout plates to the end of the snout (turquoise line) much shorter than the distance from the edge of the medial snout plates to a line through the anterior margins of the orbits (yellow line, dashed line is the line that demarks the anterior borders of the orbits) while the two lengths are approximately equal in A. patronus. In addition, the anterior margin of the snout plates form a rough V in A. nudiceps and a U in A. patronus (solid white line). Scale = 1 cm. Photos by J.W. Armbruster." httpUri="https://zenodo.org/record/2623161/files/figure.png" pageId="41" pageNumber="42">Fig. 5b</figureCitation>
) snout tentacles of nuptial males long, largest 1.5 times eye diameter. Posteromedial tentacles diverging in 
<emphasis id="B92AEAA1BA0BC812FD842373FD93F92B" box="[565,589,1657,1680]" italics="true" pageId="41" pageNumber="42">V-</emphasis>
shape along anteriorly triangular snout plates. Naked areas of snout without tentacles rugose, separated from naris by several, wide plates; naked area wide, distance from anteromedial plate to snout greater than or equal to distance from anteromedial plate to line formed between anterior edges of nasal apertures.
</paragraph>
<paragraph id="8BE136B3BA0BC812FF762202FC85F8D2" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">
<emphasis id="B92AEAA1BA0BC812FF762202FE4FF89A" bold="true" box="[199,401,1800,1825]" pageId="41" pageNumber="42">Color in alcohol.</emphasis>
(
<figureCitation id="13652A36BA0BC812FE122202FE28F89A" box="[419,502,1800,1825]" captionStart="FIGURE 19" captionStartId="40.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@40.[151,1436,193,1817]" captionTargetPageId="40" captionText="FIGURE 19. Ancistrus patronus, holotype, in dorsal, lateral, and ventral views, AUM 39876, 83.6 mm SL. Scale = 1 cm. Photos by J.W. Armbruster." httpUri="https://zenodo.org/record/2623192/files/figure.png" pageId="41" pageNumber="42">Fig. 19</figureCitation>
) Base color of all fins, dorsum, sides and ventrum of body black, brown or gray, with tiny white dots; dots often lost in preservative. First dorsal-fin membrane with dark pigment near base. Upper and lower tips of caudal fin sometimes white or cream colored.
</paragraph>
<paragraph id="8BE136B3BA0BC812FF76227EFB5FF80A" blockId="41.[151,1441,792,2005]" pageId="41" pageNumber="42">
<emphasis id="B92AEAA1BA0BC812FF76227EFE8FF836" bold="true" box="[199,337,1908,1933]" pageId="41" pageNumber="42">Life colors.</emphasis>
(
<figureCitation id="13652A36BA0BC812FED4227EFE62F836" box="[357,444,1908,1933]" captionStart="FIGURE 20" captionStartId="41.[151,250,695,717]" captionTargetBox="[151,1436,193,674]" captionTargetId="figure@41.[151,1436,193,674]" captionTargetPageId="41" captionText="FIGURE 20. Live coloration of head of Ancistrus patronus in dorsal and lateral views, AUM 43868, 79.4 mm SL. Scale = 1 cm. Photos by N.K. Lujan." httpUri="https://zenodo.org/record/2623194/files/figure.png" pageId="41" pageNumber="42">Fig. 20</figureCitation>
) Body and fins jet black with tiny white dots. First dorsal-fin membrane with dark pigment near base. Upper and lower tips of caudal fin sometimes white or cream colored.
</paragraph>
</subSubSection>
<subSubSection id="C3446538BA0BC812FF7622B6FA4BF86E" box="[199,1429,1980,2005]" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8BE136B3BA0BC812FF7622B6FA4BF86E" blockId="41.[151,1441,792,2005]" box="[199,1429,1980,2005]" pageId="41" pageNumber="42">
<emphasis id="B92AEAA1BA0BC812FF7622B6FEBEF86E" bold="true" box="[199,352,1980,2005]" pageId="41" pageNumber="42">Distribution.</emphasis>
Present in the upper río Orinoco from the río Cinaruco upstream and the río Casiquiare (
<figureCitation id="13652A36BA0BC812FAF322B6FA58F86E" box="[1346,1414,1980,2005]" captionStart="FIGURE 7" captionStartId="12.[151,250,1964,1986]" captionTargetBox="[151,1436,1165,1943]" captionTargetId="figure@12.[151,1436,1165,1943]" captionTargetPageId="12" captionText="FIGURE 7. Distribution of species of Ancistrus from the northwestern Guiana Shield and neighboring regions." httpUri="https://zenodo.org/record/2623165/files/figure.png" pageId="41" pageNumber="42">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3446538BA08C811FF76259DFC27FF43" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BE136B3BA08C811FF76259DFC27FF43" blockId="42.[151,1436,151,248]" pageId="42" pageNumber="43">
<emphasis id="B92AEAA1BA08C811FF76259DFE90FF0B" bold="true" box="[199,334,151,176]" pageId="42" pageNumber="43">Etymology.</emphasis>
Named for good fathers. 
<emphasis id="B92AEAA1BA08C811FDDE2593FD0BFF0B" box="[623,725,153,176]" italics="true" pageId="42" pageNumber="43">Patronus</emphasis>
is Latin for protector or defender and is derived from 
<emphasis id="B92AEAA1BA08C811FA882593FAAAFF0B" box="[1337,1396,153,176]" italics="true" pageId="42" pageNumber="43">pater</emphasis>
for father. Males of this genus guard their nests and protect their young until they are fairly large. With their cheek odontodes everted, males can block off nests and potentially injure predators.
</paragraph>
</subSubSection>
</treatment>
</document>