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<document id="C5176BCC6A3F40549E07A69465F14FFE" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629656873565" checkinUser="felipe" docAuthor="Chandler, Peter J." docDate="2009" docId="03EA87E1FFB6126311BBFBE0592E6F34" docLanguage="en" docName="zt02318p506.pdf" docOrigin="Zootaxa 2318" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Monocentrota matilei Bechev 1989" docType="treatment" docVersion="3" masterDocId="FFD3FF99FFB1126B112CFF9D58536D53" masterDocTitle="The fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) of Sardinia, with description of six new species *" updateTime="1699090310938" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DBDDCADFA95DDC7336939973CE85E9FA">The fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) of Sardinia, with description of six new species *</mods:title>
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<mods:namePart id="FC3D74AEF5F3BDE034DF6ACF6A649A10">Chandler, Peter J.</mods:namePart>
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<mods:date id="115549674E7659E2B41EA5E034F2C776">2009</mods:date>
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<mods:number id="FBC2CA8CFAA02E328B40B7ED938893B6">2009-12-22</mods:number>
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<treatment id="03EA87E1FFB6126311BBFBE0592E6F34" ID-DOI="http://doi.org/10.5281/zenodo.5316214" ID-Zenodo-Dep="5316214" LSID="urn:lsid:plazi:treatment:03EA87E1FFB6126311BBFBE0592E6F34" httpUri="http://treatment.plazi.org/id/03EA87E1FFB6126311BBFBE0592E6F34" lastPageId="8" pageId="7">
<subSubSection id="C359657CFFB6126C11BBFBE05A2C69C4" box="[151,639,1149,1175]" pageId="7" type="nomenclature">
<paragraph id="8BFC36F7FFB6126C11BBFBE05A2C69C4" blockId="7.[151,639,1149,1214]" box="[151,639,1149,1175]" pageId="7">
<heading id="D0B4819BFFB6126C11BBFBE05A2C69C4" bold="true" box="[151,639,1149,1175]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="B937EAE5FFB6126C11BBFBE05A2C69C4" bold="true" box="[151,639,1149,1175]" pageId="7">
6.
<taxonomicName id="4C434D74FFB6126C1198FBE05A3069C4" ID-CoL="4457C" authority="Bechev, 1989" authorityName="Bechev" authorityYear="1989" box="[180,611,1149,1175]" class="Insecta" family="Keroplatidae" genus="Monocentrota" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="matilei">
<emphasis id="B937EAE5FFB6126C1198FBE059EA69C4" bold="true" box="[180,441,1149,1175]" italics="true" pageId="7">Monocentrota matilei</emphasis>
<bibRefCitation id="EFD24B06FFB6126C10ECFBE05A3069C4" author="Bechev, D." box="[448,611,1149,1175]" pageId="7" pagination="173 - 174" refId="ref40919" refString="Bechev, D. (1989) Monocentrota matilei n. sp. from Bulgaria and Algeria (Insecta, Diptera: Keroplatidae). Reichenbachia, 26, 173 - 174." type="journal article" year="1989">Bechev, 1989</bibRefCitation>
</taxonomicName>
**
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359657CFFB6126C11BBFB39594369ED" box="[151,272,1188,1214]" pageId="7" type="description">
<paragraph id="8BFC36F7FFB6126C11BBFB39594369ED" blockId="7.[151,639,1149,1214]" box="[151,272,1188,1214]" pageId="7">
(
<figureCitation id="13782A72FFB6126C11B3FB39595469ED" box="[159,263,1188,1214]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="7">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C359657CFFB6126C11BBFB6C598D6A7A" pageId="7" type="materials_examined">
<paragraph id="8BFC36F7FFB6126C11BBFB6C5DDC685F" blockId="7.[151,1436,1265,2026]" box="[151,1423,1265,1292]" pageId="7">
<emphasis id="B937EAE5FFB6126C11BBFB6C59E16858" bold="true" box="[151,434,1265,1291]" pageId="7">
Material examined.
<geoCoordinate id="EE775030FFB6126C10BEFB6C59E16858" box="[402,434,1265,1291]" degrees="1" direction="south" minutes="4" orientation="latitude" pageId="7" precision="925" value="-1.0666667">S1</geoCoordinate>
</emphasis>
418.IV.06
<specimenCount id="9D45FD7EFFB6126C136DFB6F5A386859" box="[577,619,1266,1292]" pageId="7" type="male">1 ♂</specimenCount>
<specimenCount id="9D45FD7EFFB6126C135FFB6F5ACE6859" box="[627,669,1266,1292]" pageId="7" type="female">1 ♀</specimenCount>
.
<geoCoordinate id="EE775030FFB6126C1387FB6C5AB2685F" box="[683,737,1265,1292]" degrees="2" direction="south" minutes="2" orientation="latitude" pageId="7" precision="925" value="-2.0333333">
<emphasis id="B937EAE5FFB6126C1387FB6C5A996858" bold="true" box="[683,714,1265,1291]" pageId="7">S2</emphasis>
2
</geoCoordinate>
16.V.06
<specimenCount id="9D45FD7EFFB6126C127DFB6F5B286859" box="[849,891,1266,1292]" pageId="7" type="female">1 ♀</specimenCount>
, 1125.VII.06
<specimenCount id="9D45FD7EFFB6126C1502FB6F5C3F6859" box="[1070,1132,1266,1292]" pageId="7" type="male">4 ♂♂</specimenCount>
, 25.VII8.VIII.06
<specimenCount id="9D45FD7EFFB6126C1467FB6F5DDB6859" box="[1355,1416,1266,1292]" pageId="7" type="male">2 ♂♂</specimenCount>
.
</paragraph>
<paragraph id="8BFC36F7FFB6126C11EAFA855D7E6861" blockId="7.[151,1436,1265,2026]" box="[198,1325,1304,1330]" pageId="7">
<emphasis id="B937EAE5FFB6126C11EAFA8559016861" bold="true" box="[198,338,1304,1330]" pageId="7">Chorotype.</emphasis>
<materialsCitation id="3B2B3CAAFFB6126C1076FA855D7A6861" box="[346,1321,1304,1330]" location="Mediterranean" municipality="central European" pageId="7" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F88855FFB6126C1076FA8559C76861" box="[346,404,1304,1330]" pageId="7">Type</typeStatus>
6 [1.12 of
<bibRefCitation id="EFD24B06FFB6126C133AFA855B346861" author="Vigna Taglianti, A. &amp; Audisio, P. A. &amp; Biondi, M. &amp; Bologna, M. A. &amp; Carpaneto, G. M. &amp; De Biase, A. &amp; Fattorini, S. &amp; Piattella, E. &amp; Sindaco, R. &amp; Venchi, A. &amp; Zapparoli, M." box="[534,871,1304,1330]" pageId="7" pagination="31 - 59" refId="ref45744" refString="Vigna Taglianti, A., Audisio, P. A., Biondi, M., Bologna, M. A., Carpaneto, G. M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. &amp; Zapparoli, M. (1999) A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palaearctic region. Biogeographia, Lavori della Societa Italiana di Biogeografia (n. s.), 20, 31 - 59." type="journal article" year="1999">
Vigna Taglianti
<emphasis id="B937EAE5FFB6126C13FEFA845B5F6861" box="[722,780,1305,1330]" italics="true" pageId="7">et al.</emphasis>
(1999)
</bibRefCitation>
].
<location id="8E9C602CFFB6126C1256FA855C766861" LSID="urn:lsid:plazi:treatment:03EA87E1FFB6126311BBFBE0592E6F34:8E9C602CFFB6126C1256FA855C766861" box="[890,1061,1304,1330]" municipality="central European" name="Mediterranean" pageId="7">Mediterranean</location>
and
<collectingMunicipality id="6B98AC8DFFB6126C1572FA855D7A6861" box="[1118,1321,1304,1330]" pageId="7">central European</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFC36F7FFB6126C11EAFAA25A35680A" blockId="7.[151,1436,1265,2026]" box="[198,614,1343,1369]" pageId="7">
<emphasis id="B937EAE5FFB6126C11EAFAA259EF680A" bold="true" box="[198,444,1343,1369]" pageId="7">Italian distribution.</emphasis>
<collectingRegion id="4987F815FFB6126C10E8FAA25A76680A" box="[452,549,1343,1369]" country="Italy" name="Sardegna" pageId="7">Sardinia</collectingRegion>
only.
</paragraph>
<paragraph id="8BFC36F7FFB6126C11EAFAF8598D6A7A" blockId="7.[151,1436,1265,2026]" pageId="7">
<emphasis id="B937EAE5FFB6126C11EAFAF85940682C" bold="true" box="[198,275,1381,1407]" pageId="7">Notes.</emphasis>
<taxonomicName id="4C434D74FFB6126C1037FAFB5A4B682C" authorityName="Bechev" authorityYear="1989" box="[283,536,1382,1407]" class="Insecta" family="Keroplatidae" genus="Monocentrota" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="matilei">
<emphasis id="B937EAE5FFB6126C1037FAFB5A4B682C" box="[283,536,1382,1407]" italics="true" pageId="7">Monocentrota matilei</emphasis>
</taxonomicName>
may be misplaced generically as it differs from other species of the genus in having antennae with only 13 flagellomeres and simple slender apically tapered gonostyli. Other
<taxonomicName id="4C434D74FFB6126C11BBFA2E5A43689E" box="[151,528,1459,1485]" class="Insecta" family="Keroplatidae" genus="Monocentrota" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B937EAE5FFB6126C11BBFA2E5913689F" box="[151,320,1459,1484]" italics="true" pageId="7">Monocentrota</emphasis>
Edwards species
</taxonomicName>
have 14 flagellomeres as in most
<taxonomicName id="4C434D74FFB6126C129FFA2E5C1D689E" authorityName="Rondani" authorityYear="1856" box="[947,1102,1459,1485]" class="Insecta" family="Keroplatidae" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="family">Keroplatidae</taxonomicName>
and the gonostyli are blunt ended with internal apical teeth.
<bibRefCitation id="EFD24B06FFB6126C133FFA475AEC68A7" author="Bechev, D." box="[531,703,1498,1524]" pageId="7" pagination="173 - 174" refId="ref40919" refString="Bechev, D. (1989) Monocentrota matilei n. sp. from Bulgaria and Algeria (Insecta, Diptera: Keroplatidae). Reichenbachia, 26, 173 - 174." type="journal article" year="1989">Bechev (1989)</bibRefCitation>
described this species from
<collectingCountry id="F3547667FFB6126C1526FA475C3C68A7" box="[1034,1135,1498,1524]" name="Bulgaria" pageId="7">Bulgaria</collectingCountry>
and
<collectingCountry id="F3547667FFB6126C158AFA475D5268A7" box="[1190,1281,1498,1524]" name="Algeria" pageId="7">Algeria</collectingCountry>
.
<bibRefCitation id="EFD24B06FFB6126C1421FA475DCF68A7" author="Papp, L." box="[1293,1436,1498,1524]" pageId="7" pagination="309 - 339" refId="ref44787" refString="Papp, L. (2003) Further additions and corrections to the Hungarian checklist (Diptera). Folia Entomologica Hungarica, 64, 309 - 339." type="journal article" year="2003">Papp (2003)</bibRefCitation>
recorded it from
<collectingCountry id="F3547667FFB6126C1071F99D59906B49" box="[349,451,1536,1562]" name="Hungary" pageId="7">Hungary</collectingCountry>
and provided more detailed figures of the genitalia. The Sardinian specimens agree in most respects with the figures provided by Bechev and Papp but the genitalia are also figured here (
<figureCitation id="13782A72FFB6126C146FF9BA58F56B34" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="7">Figs 1 3</figureCitation>
) as they obviously differ from previous figures in the following respects: tergite 9 longer and more tapered apically and the cerci markedly smaller relative to this tergite (
<figureCitation id="13782A72FFB6126C12A4F9E95B836BDD" box="[904,976,1652,1678]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="7">Fig. 3</figureCitation>
) and teeth only apparent on the apical margin of the aedeagus (
<figureCitation id="13782A72FFB6126C1093F9065A5B6BE6" box="[447,520,1691,1717]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="7">Fig. 2</figureCitation>
) rather than more widely on its surface. The male Sardinian material is also described below to augment the description given by Bechev and to characterise the population. Further study may show that it is specifically distinct but study of further material from other areas would be desirable to determine if this is the case.
</paragraph>
</subSubSection>
<subSubSection id="C359657CFFB6126311EAF8A8592E6F34" lastPageId="8" pageId="7" type="description">
<paragraph id="8BFC36F7FFB6126C11EAF8A85D086A25" blockId="7.[151,1436,1265,2026]" pageId="7">
<emphasis id="B937EAE5FFB6126C11EAF8A8590D6A1C" bold="true" box="[198,350,1845,1871]" pageId="7">Description.</emphasis>
Male.
<emphasis id="B937EAE5FFB6126C109DF8AB59A26A1C" box="[433,497,1846,1871]" italics="true" pageId="7">Head</emphasis>
dark brown with short dark occipital bristling. Antenna about as long as thorax, dark brown, with 2 + 13 segments; flagellomeres not much longer than broad. Palpus brownish, slender.
</paragraph>
<paragraph id="8BFC36F7FFB6126311EAF81E5B1B6C9F" blockId="7.[151,1436,1265,2026]" lastBlockId="8.[151,1436,152,615]" lastPageId="8" pageId="7">
<emphasis id="B937EAE5FFB6126C11EAF81E59486ACF" box="[198,283,1923,1948]" italics="true" pageId="7">Thorax</emphasis>
dark brown with short black bristling on mesonotum in acrostichal and dorsocentral stripes, and irregular longer black setae laterally. Pleura bare except for scattered short setae on upper part of anepisternum. Laterotergite with dark bristling on upper half. Mediotergite bare.
<emphasis id="B937EAE5FFB6126C1568F84C5CD36AB9" box="[1092,1152,2001,2026]" italics="true" pageId="7">Wing</emphasis>
slightly brownish tinted on membrane, with costa and radial veins brown. Vein Sc short, ending in costa before level of base of Rs. Vein R
<subScript id="17C734B2FFB9126311CBFF5358BC6D8F" attach="left" box="[231,239,206,220]" fontSize="6" pageId="8">4</subScript>
short, ending in costa more than twice its length beyond tip of R
<subScript id="17C734B2FFB9126312C6FF535BA16D8F" attach="right" box="[1002,1010,206,220]" fontSize="6" pageId="8">1</subScript>
.
<collectingRegion id="4987F815FFB91263152DFF235C106D8B" box="[1025,1091,190,216]" country="New Zealand" name="West Coast" pageId="8">Costa</collectingRegion>
extends 0.5 distance from R
<subScript id="17C734B2FFB9126314B8FF535DCF6D8F" attach="right" box="[1428,1436,206,220]" fontSize="6" pageId="8">5</subScript>
to M
<subScript id="17C734B2FFB9126311FCFF65588B6C55" attach="left" box="[208,216,248,262]" fontSize="6" pageId="8">1</subScript>
. Basal part of M not developed; r-m fusion long but shorter than curved Rs and stem of median fork. Vein A
<subScript id="17C734B2FFB9126311C6FEBF58A16C63" attach="left" box="[234,242,290,304]" fontSize="6" pageId="8">1</subScript>
weak on apical half but faintly reaching costa. R
<subScript id="17C734B2FFB91263121BFEBF5B6C6C63" attach="left" box="[823,831,290,304]" fontSize="6" pageId="8">1</subScript>
and R
<subScript id="17C734B2FFB9126312A1FEBF5BC66C63" attach="left" box="[909,917,290,304]" fontSize="6" pageId="8">5</subScript>
setose, other veins including Rs, r-m fusion and R
<subScript id="17C734B2FFB9126311F0FED658B76C0A" attach="left" box="[220,228,331,345]" fontSize="6" pageId="8">4</subScript>
bare. Haltere yellow.
<emphasis id="B937EAE5FFB9126310C9FEA65A226C07" box="[485,625,315,340]" italics="true" pageId="8">Wing length</emphasis>
<quantity id="4CBB9B12FFB912631355FEA65B5B6C06" box="[633,776,315,341]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.6" metricValueMin="1.9" pageId="8" unit="mm" value="2.25" valueMax="2.6" valueMin="1.9">1.92.6 mm</quantity>
.
<emphasis id="B937EAE5FFB912631238FEA65B186C07" box="[788,843,315,340]" italics="true" pageId="8">Legs</emphasis>
brownish yellow, with short dark setulae, those on tibiae and tarsi in regular rows; no stronger setae developed. Fore leg with metatarsus half length of tibia. Tibial spurs brown, a single spur on each tibia, that on fore leg as long as width of tibial apex, on mid leg 1.5 times and on hind leg twice as long as width of tibial apex.
</paragraph>
<paragraph id="8BFC36F7FFB9126311EAFE44592E6F34" blockId="8.[151,1436,152,615]" pageId="8">
<emphasis id="B937EAE5FFB9126311EAFE4459666CA1" box="[198,309,473,498]" italics="true" pageId="8">Abdomen</emphasis>
including genitalia brown with short dark bristling.
<emphasis id="B937EAE5FFB9126312C5FE445C0B6CA1" box="[1001,1112,473,498]" italics="true" pageId="8">Genitalia</emphasis>
(
<figureCitation id="13782A72FFB91263155DFE445CB16CA0" box="[1137,1250,473,499]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="8">Figs 13</figureCitation>
) with broadly emarginate gonocoxites, simple slender apically tapered gonostylus and aedeagus short and broad with short marginal teeth. Tergite 9 (
<figureCitation id="13782A72FFB9126310F4FDBB5A726F13" box="[472,545,550,576]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="8">Fig. 3</figureCitation>
) simple, tapered and rounded apically, with small densely bristled cerci protruding apically.
</paragraph>
</subSubSection>
</treatment>
</document>